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<document id="89B088A3E2960C4F795BA7152739E6B9" ID-DOI="10.5281/zenodo.283207" ID-GBIF-Dataset="cb513858-dd8a-4295-9b25-25ff3a02a9dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="283207" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460566474209" checkinUser="plazi" docAuthor="Delgado, Leonardo &amp; Mora-Aguilar, Eder F." docDate="2012" docId="03FF5D7DFF97FFC623CC10170BA7FEB6" docLanguage="en" docName="zt03597p052.pdf" docOrigin="Zootaxa 3597" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Strigoderma tenebrosa Delgado &amp; Mora-Aguilar, new species" docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FFC62505FF96FFC4235B12330845FF89" masterDocTitle="A new Mexican species and new taxonomic and distributional data of Strigoderma Burmeister (Coleoptera: Scarabaeidae: Rutelinae)" masterLastPageNumber="52" masterPageNumber="47" pageNumber="48" updateTime="1698321040446" updateUser="plazi">
<mods:mods id="F916114486C9DC41028ACD0644CCE457" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7B3A7C35D3179A836027391DD0F0F67">
<mods:title id="DA422BEEAA79C33C1AA967B3874698DA">A new Mexican species and new taxonomic and distributional data of Strigoderma Burmeister (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="7CFAB467BC173C798DF0A5E5477775F2" type="personal">
<mods:role id="B6A54C8B039878614943E85CC33BD144">
<mods:roleTerm id="A1C198706F3B47411686408828F6C348">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18F83A690DBCD0170C9FBA508BED232C">Delgado, Leonardo</mods:namePart>
</mods:name>
<mods:name id="559C2D7E1E887080122DA3AB7F09D706" type="personal">
<mods:role id="17ED68CEEAB9F84AB208F7A3675A2884">
<mods:roleTerm id="625207D2268A9815822C97D3569749BD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19D7E3E0262AA499FB0E73F03D2C63CC">Mora-Aguilar, Eder F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0F8C037DE4182280A29D7AFBEAA03F4E">text</mods:typeOfResource>
<mods:relatedItem id="0FA48450B274E8ECDB5F3619A25DD4E9" type="host">
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<mods:title id="A42DA2C56438236F9C312A13968FAF9D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="339BDCE0CB360DDB8161F9F4C207E28C">
<mods:date id="1CD9C412713996EDF24313AE881AE4A8">2012</mods:date>
<mods:detail id="600623D94E0EAACC4A64AA7768AE6C22" type="volume">
<mods:number id="E892F72EB7B79691D3740B68B3CB6E87">3597</mods:number>
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<mods:classification id="4A80997043F8211B3AB5697A5D858977">journal article</mods:classification>
<mods:identifier id="A5C3C217D898F04A5C10D7E442193D93" type="DOI">10.5281/zenodo.283207</mods:identifier>
<mods:identifier id="A163138B2D42757BB25A0BAB230C3A06" type="GBIF-Dataset">cb513858-dd8a-4295-9b25-25ff3a02a9dc</mods:identifier>
<mods:identifier id="6D99E5CAE2CC6C75B36AE268EECEFED8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FEEB5DA970F9CA35D07B0366C31E0164" type="Zenodo-Dep">283207</mods:identifier>
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<treatment id="03FF5D7DFF97FFC623CC10170BA7FEB6" ID-DOI="http://doi.org/10.5281/zenodo.6168986" ID-GBIF-Taxon="119557671" ID-Zenodo-Dep="6168986" LSID="urn:lsid:plazi:treatment:03FF5D7DFF97FFC623CC10170BA7FEB6" httpUri="http://treatment.plazi.org/id/03FF5D7DFF97FFC623CC10170BA7FEB6" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="48">
<subSubSection id="C34CBFE0FF97FFC523CC10170957FDD7" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="8BE9EC6BFF97FFC523CC10170BCDFDB4" blockId="1.[151,904,547,606]" box="[151,904,547,574]" pageId="1" pageNumber="48">
<heading id="D0A15B07FF97FFC523CC10170BCDFDB4" bold="true" box="[151,904,547,574]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="B9223079FF97FFC523CC10170BCDFDB4" bold="true" box="[151,904,547,574]" pageId="1" pageNumber="48">
<taxonomicName id="4C5697E8FF97FFC523CC10170AA9FDB4" authority="Delgado &amp; Mora-Aguilar" authorityName="Delgado &amp; Mora-Aguilar" box="[151,748,547,574]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp. nov.">
<emphasis id="B9223079FF97FFC523CC101709EDFDB7" bold="true" box="[151,424,548,574]" italics="true" pageId="1" pageNumber="48">Strigoderma tenebrosa</emphasis>
Delgado &amp; Mora-Aguilar
</taxonomicName>
,
<taxonomicNameLabel id="A2118D02FF97FFC521A310100BCDFDB4" box="[760,904,547,573]" pageId="1" pageNumber="48" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9EC6BFF97FFC523CC10750957FDD7" blockId="1.[151,904,547,606]" box="[151,274,582,606]" pageId="1" pageNumber="48">
(
<figureCitation id="136DF0EEFF97FFC523C41075094CFDD7" box="[159,265,582,606]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34CBFE0FF97FFC523CC10BC0BDFFC9B" pageId="1" pageNumber="48" type="materials_examined">
<paragraph id="8BE9EC6BFF97FFC523CC10BC0BDFFC9B" blockId="1.[151,1436,654,2011]" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC523CC10BC090FFD2E" bold="true" box="[151,330,654,679]" pageId="1" pageNumber="48">
<typeStatus id="54ED52C9FF97FFC523CC10BC0897FD2E" box="[151,210,655,679]" pageId="1" pageNumber="48">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED52C9FF97FFC5220D10BD09FBFD2F" box="[342,446,654,678]" pageId="1" pageNumber="48" type="holotype">Holotype</typeStatus>
male, two males
<typeStatus id="54ED52C9FF97FFC521C210BC0B42FD2E" box="[665,775,655,679]" pageId="1" pageNumber="48" type="paratype">paratypes</typeStatus>
, and one female
<typeStatus id="54ED52C9FF97FFC5208110BC0C7FFD2E" box="[986,1082,655,679]" pageId="1" pageNumber="48" type="paratype">paratype</typeStatus>
labeled: “
<collectingCountry id="F341ACFBFF97FFC527ED10BD0D6EFD2F" box="[1206,1323,654,678]" name="Mexico" pageId="1" pageNumber="48">MEXICO</collectingCountry>
: Chiapas, Villaflores, Finca
<collectingCountry id="F341ACFBFF97FFC5223D108109B4FD43" box="[358,497,690,714]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="48">Santa Elena</collectingCountry>
,
<date id="FFE8CAABFF97FFC522A510810AC0FD43" box="[510,645,690,714]" pageId="1" pageNumber="48" value="2001-06-12">12-VI-2001</date>
, C. J. Morales col.” One male
<typeStatus id="54ED52C9FF97FFC520AB10800C14FD42" box="[1008,1105,691,715]" pageId="1" pageNumber="48" type="paratype">paratype</typeStatus>
with same data as
<typeStatus id="54ED52C9FF97FFC5266810810DDDFD43" box="[1331,1432,690,714]" pageId="1" pageNumber="48" type="holotype">holotype</typeStatus>
, except:
<date id="FFE8CAABFF97FFC523A810E5093EFD67" box="[243,379,726,750]" pageId="1" pageNumber="48" value="2001-06-22">22-VI-2001</date>
. The
<typeStatus id="54ED52C9FF97FFC5229B10E50A67FD67" box="[448,546,726,750]" pageId="1" pageNumber="48" type="holotype">holotype</typeStatus>
and one female
<typeStatus id="54ED52C9FF97FFC521B610E40B0BFD66" box="[749,846,727,751]" pageId="1" pageNumber="48" type="paratype">paratype</typeStatus>
are deposited in
<collectionCode id="ED4774AEFF97FFC5274410E40C23FD66" box="[1055,1126,727,751]" pageId="1" pageNumber="48">IEXA</collectionCode>
. Three male
<typeStatus id="54ED52C9FF97FFC5265810E40D2BFD66" box="[1283,1390,727,751]" pageId="1" pageNumber="48" type="paratype">paratypes</typeStatus>
are deposited in the following collections:
<collectionCode id="ED4774AEFF97FFC5211010C90AF5FC9B" box="[587,688,762,786]" pageId="1" pageNumber="48">CUACH</collectionCode>
,
<collectionCode id="ED4774AEFF97FFC521E010C90B4AFC9B" box="[699,783,762,786]" pageId="1" pageNumber="48">EMAC</collectionCode>
, and
<collectionCode id="ED4774AEFF97FFC5201210C90BD3FC9B" box="[841,918,762,786]" pageId="1" pageNumber="48">LLDC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CBFE0FF97FFC5239C112D0C1AF9C7" pageId="1" pageNumber="48" type="description">
<paragraph id="8BE9EC6BFF97FFC5239C112D0D6AFAFF" blockId="1.[151,1436,654,2011]" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC5239C112D091CFCBE" bold="true" box="[199,345,798,823]" pageId="1" pageNumber="48">Description.</emphasis>
<typeStatus id="54ED52C9FF97FFC5223A112D098CFCBF" box="[353,457,798,822]" pageId="1" pageNumber="48" type="holotype">Holotype</typeStatus>
male. Length:
<quantity id="4CAE418EFF97FFC52121112C0A93FCBF" box="[634,726,799,822]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="48" unit="mm" value="7.7">7.7 mm</quantity>
, width:
<quantity id="4CAE418EFF97FFC5206F112D0BD5FCBF" box="[820,912,798,822]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="48" unit="mm" value="4.1">4.1 mm</quantity>
. Form oval-elongate. Color black, with green and purple luster on head, pronotal disc, scutellum, legs, and venter.
<emphasis id="B9223079FF97FFC520E811700BAAFCD2" box="[947,1007,835,859]" italics="true" pageId="1" pageNumber="48">Head</emphasis>
: Clypeus trapezoidal, lateral margins rounded, apex slightly emarginate and moderately reflexed; frons and clypeus with dense, deep and subrugose punctation; interocular distance equals 5.0 transverse eye diameters; antennae with 9 antennomeres, club subequal to funicle.
<emphasis id="B9223079FF97FFC5224B1183093AFC4E" box="[272,383,944,967]" italics="true" pageId="1" pageNumber="48">Pronotum</emphasis>
: hexagonal, moderately convex, almost as wide as long (1.0:1.2), sides widest at apical third, anterior angles acute, posterior angles obtuse, base sinuate to sides of scutellum; entire surface with punctation fine and confluent.
<emphasis id="B9223079FF97FFC5226411C509E8FB87" box="[319,429,1014,1038]" italics="true" pageId="1" pageNumber="48">Scutellum</emphasis>
: with fine and dense punctures.
<emphasis id="B9223079FF97FFC5204611C50B26FB87" box="[797,867,1014,1038]" italics="true" pageId="1" pageNumber="48">Elytra</emphasis>
: longer than wide (1.9:1.0), narrowed to the apex; surface granular, microreticulate, with some small punctures towards the apex; with four acute elevated ridges with vestigial ridges interspersed between.
<emphasis id="B9223079FF97FFC5211D160D0AF5FBDF" box="[582,688,1086,1110]" italics="true" pageId="1" pageNumber="48">Abdomen</emphasis>
: Apical region of propygidium with scarce, short setae; pygidium convex, wider than long (1.3:1.0), with punctation fine to moderate in size, moderately dense, with sparse, moderately setae; sternites concave, surface microreticulate with irregular punctation and each one with a transverse row of long, pale yellow setae.
<emphasis id="B9223079FF97FFC52121169F0A82FB4A" box="[634,711,1196,1219]" italics="true" pageId="1" pageNumber="48">Venter:</emphasis>
Prosternum with white setae in the middle; mestasternum with large, dense, white setae at the sides; disc glabrous; meso-metasternal process rounded, not elevated.
<emphasis id="B9223079FF97FFC5260416E30DD1FB6E" box="[1375,1428,1232,1255]" italics="true" pageId="1" pageNumber="48">Legs</emphasis>
: Protibiae bidentate; protarsi and mesotarsi short and thickened, apical protarsomere larger than the rest combined; inner proclaw elongate, as long as the apical protarsomere, cleft, inner ramus equals 0.3 the length of outer ramus (
<figureCitation id="136DF0EEFF97FFC523C4170908A3FADB" box="[159,230,1338,1362]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
); meso- and metatibia slightly widened at middle and with two diagonal carinae with accompanying spinules.
<emphasis id="B9223079FF97FFC5225A176D092EFAFF" box="[257,363,1374,1398]" italics="true" pageId="1" pageNumber="48">Genitalia</emphasis>
(
<figureCitation id="136DF0EEFF97FFC52221176D09A2FAFF" box="[378,487,1374,1398]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Figs. 2–3</figureCitation>
): Parameres elongate, convergent to the apex, scarcely bent in lateral view.
</paragraph>
<paragraph id="8BE9EC6BFF97FFC5239C17B00C1AF9C7" blockId="1.[151,1436,654,2011]" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC5239C17B00A69FA12" bold="true" box="[199,556,1410,1435]" pageId="1" pageNumber="48">
<typeStatus id="54ED52C9FF97FFC5239C17B0097AFA12" box="[199,319,1411,1435]" pageId="1" pageNumber="48" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D5027E2FF97FFC5221517B109EFFA12" box="[334,426,1410,1435]" pageId="1" pageNumber="48" type="male">3 males</specimenCount>
,
<specimenCount id="9D5027E2FF97FFC522EC17B10A65FA12" box="[439,544,1410,1435]" pageId="1" pageNumber="48" type="female">1 female</specimenCount>
).
</emphasis>
Total length: 7.4–8.0 mm; width:
<quantity id="4CAE418EFF97FFC520F517B10C7CFA13" box="[942,1081,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="3.9" metricValueMin="3.8" pageId="1" pageNumber="48" unit="mm" value="3.8499999999999996" valueMax="3.9" valueMin="3.8">3.8–3.9 mm</quantity>
. Male
<typeStatus id="54ED52C9FF97FFC527DF17B00CB5FA12" box="[1156,1264,1411,1435]" pageId="1" pageNumber="48" type="paratype">paratypes</typeStatus>
differ from the male
<typeStatus id="54ED52C9FF97FFC523831795097FFA37" box="[216,314,1446,1470]" pageId="1" pageNumber="48" type="holotype">holotype</typeStatus>
in the following respects: head and pronotum with shiny reflections, and tarsi black to reddish brown. In the female
<typeStatus id="54ED52C9FF97FFC522DC17F809A2FA6A" box="[391,487,1483,1507]" pageId="1" pageNumber="48" type="paratype">paratype</typeStatus>
the elytral color black with irregular maculae dark reddish-brown, pronotal disc dull green and swollen, antennal club shorter than the funicle, elytra with the second broad interval without costae elevated, epipleura slightly wider, abdominal sternites convex in lateral view, teeth of protibia elongate and projected apically, tarsi not thickened and claws not enlarged, rami subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="C34CBFE0FF97FFC5239C1468093BF91F" pageId="1" pageNumber="48" type="etymology">
<paragraph id="8BE9EC6BFF97FFC5239C1468093BF91F" blockId="1.[151,1436,654,2011]" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC5239C1468090BF9FA" bold="true" box="[199,334,1627,1651]" pageId="1" pageNumber="48">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="B9223079FF97FFC5203A14680B9CF9FA" box="[865,985,1627,1651]" italics="true" pageId="1" pageNumber="48">tenebrosus</emphasis>
, meaning dark, in reference to the dark color of this species.
</paragraph>
</subSubSection>
<subSubSection id="C34CBFE0FF97FFC5239C14900C52F83F" pageId="1" pageNumber="48" type="discussion">
<paragraph id="8BE9EC6BFF97FFC5239C14900C52F83F" blockId="1.[151,1436,654,2011]" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC5239C1490097EF932" bold="true" box="[199,315,1699,1723]" pageId="1" pageNumber="48">Remarks.</emphasis>
<taxonomicName id="4C5697E8FF97FFC5221E14900A0CF932" box="[325,585,1699,1723]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B9223079FF97FFC5221E14900A0CF932" box="[325,585,1699,1723]" italics="true" pageId="1" pageNumber="48">Strigoderma tenebrosa</emphasis>
</taxonomicName>
possesses characters states that place it in the
<taxonomicName id="4C5697E8FF97FFC5272914900DD9F932" box="[1138,1436,1699,1723]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="B9223079FF97FFC5272914900DD9F932" box="[1138,1436,1699,1723]" italics="true" pageId="1" pageNumber="48">Strigoderma costulipennis</emphasis>
</taxonomicName>
species group: surface of elytra granular and with acute or shallow elevated ridges.
<taxonomicName id="4C5697E8FF97FFC5272114F50D38F957" box="[1146,1405,1734,1758]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B9223079FF97FFC5272114F50D38F957" box="[1146,1405,1734,1758]" italics="true" pageId="1" pageNumber="48">Strigoderma tenebrosa</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C5697E8FF97FFC5222F14D80A40F88A" box="[372,517,1771,1795]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tomentosa">
<emphasis id="B9223079FF97FFC5222F14D80A40F88A" box="[372,517,1771,1795]" italics="true" pageId="1" pageNumber="48">S. tomentosa</emphasis>
</taxonomicName>
by the pronotal punctation fine and dense, and the acute, elevated elytral ridges (pronotum coarsely punctuate with shallow, elytral ridges in
<taxonomicName id="4C5697E8FF97FFC52001153C0BA8F8AE" box="[858,1005,1807,1831]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tomentosa">
<emphasis id="B9223079FF97FFC52001153C0BA8F8AE" box="[858,1005,1807,1831]" italics="true" pageId="1" pageNumber="48">S. tomentosa</emphasis>
</taxonomicName>
).
<taxonomicName id="4C5697E8FF97FFC52753153D0D4FF8AF" box="[1032,1290,1806,1830]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B9223079FF97FFC52753153D0D4FF8AF" box="[1032,1290,1806,1830]" italics="true" pageId="1" pageNumber="48">Strigoderma tenebrosa</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="4C5697E8FF97FFC5238C150009CFF8C2" box="[215,394,1843,1867]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="B9223079FF97FFC5238C150009CFF8C2" box="[215,394,1843,1867]" italics="true" pageId="1" pageNumber="48">S. costulipennis</emphasis>
</taxonomicName>
by the total length greater than
<quantity id="4CAE418EFF97FFC5205915010B1FF8C3" box="[770,858,1842,1866]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="48" unit="mm" value="7.4">7.4 mm</quantity>
(7.0 mm or less in
<taxonomicName id="4C5697E8FF97FFC5271F15000CB2F8C2" box="[1092,1271,1843,1867]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="B9223079FF97FFC5271F15000CB2F8C2" box="[1092,1271,1843,1867]" italics="true" pageId="1" pageNumber="48">S. costulipennis</emphasis>
</taxonomicName>
), by the inner proclaw in the male as long as the apical tarsomere with the inner ramus 0.3 the length of the outer ramus (
<figureCitation id="136DF0EEFF97FFC5260B15650DD4F8E7" box="[1360,1425,1878,1902]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
) (male inner proclaw in
<taxonomicName id="4C5697E8FF97FFC522C615480A0BF81A" box="[413,590,1915,1939]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="B9223079FF97FFC522C6154809EEF81B" box="[413,427,1915,1938]" italics="true" pageId="1" pageNumber="48">S</emphasis>
.
<emphasis id="B9223079FF97FFC522E115480A0BF81A" box="[442,590,1915,1939]" italics="true" pageId="1" pageNumber="48">costulipennis</emphasis>
</taxonomicName>
shorter than the apical tarsomere with the inner ramus 0.7 the length of the outer ramus as in
<figureCitation id="136DF0EEFF97FFC5220515AD09E4F83F" box="[350,417,1950,1974]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Fig. 4</figureCitation>
), and by the male genitalia (
<figureCitation id="136DF0EEFF97FFC5218415AD0B0DF83F" box="[735,840,1950,1974]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Figs. 2–3</figureCitation>
versus
<figureCitation id="136DF0EEFF97FFC520C515AD0C4FF83F" box="[926,1034,1950,1974]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1917,1940]" captionTargetBox="[215,1359,380,1875]" captionTargetId="figure@2.[209,1364,375,1897]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Diagnostic structures of Strigoderma species. 1 – 3) S. tenebrosa: 1) Protarsal claws; 2) Parameres in frontal view; 3) Parameres in lateral view. 4 – 6) S. costulipennis: 4) Protarsal claws; 5) Parameres in frontal view; 6) Parameres in lateral view. 7 – 9) S. longicollis: 7) Parameres in frontal view; 8) Parameres in lateral view; 9) Tips of parameres in ventral view. Scale lines = 1 mm." httpUri="https://zenodo.org/record/283208/files/figure.png" pageId="1" pageNumber="48">Figs. 5–6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34CBFE0FF97FFC6239C15F10BA7FEB6" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48" type="distribution">
<paragraph id="8BE9EC6BFF97FFC6239C15F10BA7FEB6" blockId="1.[151,1436,654,2011]" lastBlockId="2.[151,1436,151,319]" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<emphasis id="B9223079FF97FFC5239C15F10925F852" bold="true" box="[199,352,1986,2011]" pageId="1" pageNumber="48">Distribution.</emphasis>
<taxonomicName id="4C5697E8FF97FFC5223715F00A35F852" box="[364,624,1987,2011]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B9223079FF97FFC5223715F00A35F852" box="[364,624,1987,2011]" italics="true" pageId="1" pageNumber="48">Strigoderma tenebrosa</emphasis>
</taxonomicName>
is known only from a locality situated at the Central Depression of Chiapas,
<collectingCountry id="F341ACFBFF94FFC6225812A40919FF26" box="[259,348,151,175]" name="Mexico" pageId="2" pageNumber="49">Mexico</collectingCountry>
. This species was collected in areas with tropical deciduous forest at
<quantity id="4CAE418EFF94FFC627D612A40C92FF26" box="[1165,1239,151,175]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="49" unit="m" value="600.0">600 m</quantity>
of elevation. By contrast,
<taxonomicName id="4C5697E8FF94FFC6225A128E09F0FF5D" box="[257,437,188,212]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="B9223079FF94FFC6225A128E09F0FF5D" box="[257,437,188,212]" italics="true" pageId="2" pageNumber="49">S. costulipennis</emphasis>
</taxonomicName>
is distributed west of the Isthmus of Tehuantepec, in the states of Colima, Guerrero, Jalisco, Michoacán, Morelos, Nayarit, Oaxaca, and Puebla (
<bibRefCitation id="EFC7919AFF94FFC6206A12EC0B84FF7E" author="Moron" box="[817,961,223,247]" pageId="2" pageNumber="49" refString="Moron, M. A. (1997) Rutelinae. In: M. A. Moron, B. C. Ratcliffe, &amp; C. Deloya (eds.), Atlas de los escarabajos de Mexico. Coleoptera Lamelicornia. Vol. I. Familia Melolonthidae. CONABIO y Sociedad Mexicana de Entomologia, A. C. Mexico, pp. 9 - 49." type="book chapter" year="1997">Morón 1997</bibRefCitation>
), although the distant record of the state of Coahuila mapped by
<bibRefCitation id="EFC7919AFF94FFC622D413370A62FE95" author="Bader" box="[399,551,260,284]" pageId="2" pageNumber="49" refString="Bader, A. M. (1992) A review of the North and Central American Strigoderma (Coleoptera: Scarabaeidae). Transactions of the American Entomological Society, 118, 269 - 330." type="journal article" year="1992">Bader (1992)</bibRefCitation>
needs confirmation.
<taxonomicName id="4C5697E8FF94FFC6204713370C60FE95" box="[796,1061,260,284]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="tomentosa">
<emphasis id="B9223079FF94FFC6204713370C60FE95" box="[796,1061,260,284]" italics="true" pageId="2" pageNumber="49">Strigoderma tomentosa</emphasis>
</taxonomicName>
was described from the state of Oaxaca,
<collectingCountry id="F341ACFBFF94FFC623A213140914FEB6" box="[249,337,295,319]" name="Mexico" pageId="2" pageNumber="49">Mexico</collectingCountry>
, and after cited by
<bibRefCitation id="EFC7919AFF94FFC6217D13140AFFFEB6" author="Bader" box="[550,698,295,319]" pageId="2" pageNumber="49" refString="Bader, A. M. (1992) A review of the North and Central American Strigoderma (Coleoptera: Scarabaeidae). Transactions of the American Entomological Society, 118, 269 - 330." type="journal article" year="1992">Bader (1992)</bibRefCitation>
from
<collectingCountry id="F341ACFBFF94FFC621A413140B37FEB6" box="[767,882,295,319]" name="Nicaragua" pageId="2" pageNumber="49">Nicaragua</collectingCountry>
(
<figureCitation id="136DF0EEFF94FFC620DA13140B90FEB6" box="[897,981,295,319]" captionStart="FIGURE 10" captionStartId="3.[151,250,1230,1253]" captionTargetBox="[202,1404,187,1168]" captionTargetId="figure@3.[151,1436,161,1209]" captionTargetPageId="3" captionText="FIGURE 10. Geographical distribution of S. costulipennis (dots), S. tomentosa (square), and S. tenebrosa (triangle)." httpUri="https://zenodo.org/record/283209/files/figure.png" pageId="2" pageNumber="49">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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