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<document id="945BA45A7D03E841790E8377D45D9C97" ID-DOI="10.11646/zootaxa.5343.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324250" ID-ZooBank="20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069790509" checkinUser="plazi" docAuthor="Yamasaki, Takeshi, Hashimoto, Yoshiaki, Endo, Tomoji, Hyodo, Fujio, Itioka, Takao, Mohamed, Maryati &amp; Meleng, Paulus" docDate="2023" docId="03FA87E7FFBFFFD1FF7D3A58FD568F8F" docLanguage="en" docName="zootaxa.5343.1.3.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna chamaeleon Deeleman-Reinhold 2001" docType="treatment" docVersion="2" lastPageNumber="65" masterDocId="FFC3FF9FFFB7FFDBFFEA3E15FFEC8816" masterDocTitle="Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species" masterLastPageNumber="73" masterPageNumber="55" pageNumber="63" updateTime="1694071302809" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DFCB528E9FDC9A4FFA96F2969306E3EB">Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species</mods:title>
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<taxonomicName id="4C534D72FFBFFFD3FF7D3A58FCFD8C7E" ID-CoL="7F64K" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[151,785,1101,1128]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chamaeleon">
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<emphasis id="B927EAE3FFBFFFD3FF7D3A58FE258C7E" bold="true" box="[151,457,1101,1128]" italics="true" pageId="8" pageNumber="63">Utivarachna chamaeleon</emphasis>
<bibRefCitation id="EFC24B00FFBFFFD3FE3A3A5BFCFD8C7E" author="Deeleman-Reinhold, C. L." box="[464,785,1102,1128]" pageId="8" pageNumber="63" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
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<figureCitation id="13682A74FFBFFFD3FF7D3A67FEF88C9B" box="[151,276,1138,1165]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStartId-0="9.[151,250,1037,1061]" captionStartId-1="9.[151,250,1664,1688]" captionStartId-2="10.[151,250,879,903]" captionStartId-3="10.[151,250,1309,1333]" captionTargetBox-0="[161,1435,273,1013]" captionTargetBox-1="[152,1434,1157,1639]" captionTargetBox-2="[159,1436,181,855]" captionTargetBox-3="[151,1435,999,1285]" captionTargetId-0="figure-61@9.[151,1435,273,1013]" captionTargetId-1="figure-101@9.[151,1435,1157,1639]" captionTargetId-2="figure-17@10.[151,1436,181,855]" captionTargetId-3="figure-55@10.[151,1435,999,1285]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="10" captionText-0="FIGURE 12. Utivarachna chamaeleon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." captionText-1="FIGURE 13. Utivarachna chamaeleon, male, holotype (RNMH.ARA.15153). A right palp, retrolateral view; B same, ventral view; C same, prolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." captionText-2="FIGURE 14. Utivarachna chamaeleon Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." captionText-3="FIGURE 15. Utivarachna chamaeleon, female, paratype (RNMH.ARA.15153). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bar: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8324282" figureDoi-1="http://doi.org/10.5281/zenodo.8324284" figureDoi-2="http://doi.org/10.5281/zenodo.8324286" figureDoi-3="http://doi.org/10.5281/zenodo.8324290" httpUri-0="https://zenodo.org/record/8324282/files/figure.png" httpUri-1="https://zenodo.org/record/8324284/files/figure.png" httpUri-2="https://zenodo.org/record/8324286/files/figure.png" httpUri-3="https://zenodo.org/record/8324290/files/figure.png" pageId="8" pageNumber="63">Figs 1215</figureCitation>
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<treatmentCitation id="0AF210E0FFBFFFD3FF7D3AA3FD338CD9" author="Deeleman-Reinhold, C. L." box="[151,735,1206,1231]" page="378" pageId="8" pageNumber="63" year="2001">
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<emphasis id="B927EAE3FFBFFFD3FF7D3AA3FE7B8CD8" box="[151,407,1206,1230]" italics="true" pageId="8" pageNumber="63">Utivarachna chamaeleon</emphasis>
<bibRefCitation id="EFC24B00FFBFFFD3FE743AA3FD338CD9" author="Deeleman-Reinhold, C. L." box="[414,735,1206,1231]" pageId="8" pageNumber="63" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001: 378</bibRefCitation>
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, figs 586592;
<treatmentCitation id="0AF210E0FFBFFFD3FC963AA3FAB08CD9" author="Dankittipakul, P. &amp; Tavano, M. &amp; Singtripop, T." box="[892,1372,1206,1231]" page="137" pageId="8" pageNumber="63" year="2011">
<bibRefCitation id="EFC24B00FFBFFFD3FC963AA3FAB08CD9" author="Dankittipakul, P. &amp; Tavano, M. &amp; Singtripop, T." box="[892,1372,1206,1231]" pageId="8" pageNumber="63" pagination="133 - 146" refId="ref8829" refString="Dankittipakul, P., Tavano, M. &amp; Singtripop, T. (2011) Two new species of the spider genus Utivarachna Kishida, 1940 from Southwest Asia (Araneae, Corinnidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, 103, 133 - 146." type="journal article" year="2011">Dankittipakul, Tavano &amp; Singtripop, 2011: 137</bibRefCitation>
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, fig. 7 (only a figure of RTA without description).
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<emphasis id="B927EAE3FFBFFFD3FF7D3B0EFE528D23" bold="true" box="[151,446,1307,1333]" pageId="8" pageNumber="63">Type material examined.</emphasis>
<materialsCitation id="3B3B3CACFFBFFFD3FE2E3B0EFDAA8D4F" collectionCode="RMNH, R" collectorName="Bako National Park, W. &amp; Coll. Deeleman" country="Malaysia" location="Bako National Park" pageId="8" pageNumber="63" specimenCount="1" typeStatus="holotype">
<emphasis id="B927EAE3FFBFFFD3FE2E3B0EFDA08D2F" bold="true" box="[452,588,1307,1337]" pageId="8" pageNumber="63">
<typeStatus id="54E88853FFBFFFD3FE2E3B0EFDDD8D23" box="[452,561,1307,1333]" pageId="8" pageNumber="63">Holotype</typeStatus>
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(
<collectionCode id="ED42AE34FFBFFFD3FDB03B0EFD5F8D23" box="[602,691,1307,1333]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="63" type="Museum">RMNH</collectionCode>
.ARA. 15153),
<collectorName id="26A65327FFBFFFD3FCB63B0EFB888D20" box="[860,1124,1307,1334]" pageId="8" pageNumber="63">
<location id="8E8C602AFFBFFFD3FCB63B0EFBD68D23" LSID="urn:lsid:plazi:treatment:03FA87E7FFBFFFD1FF7D3A58FD568F8F:8E8C602AFFBFFFD3FCB63B0EFBD68D23" box="[860,1082,1307,1333]" country="Malaysia" name="Bako National Park" pageId="8" pageNumber="63">Bako National Park</location>
, W.
</collectorName>
<collectingCountry id="F3447661FFBFFFD3FB833B0EFB208D23" box="[1129,1228,1307,1333]" name="Malaysia" pageId="8" pageNumber="63">Sarawak</collectingCountry>
, 1.IV.[1985],
<collectorName id="26A65327FFBFFFD3FA8F3B0EFEE78D4F" pageId="8" pageNumber="63">Coll. Deeleman</collectorName>
, leg. C.L. &amp; P.
<collectionCode id="ED42AE34FFBFFFD3FE5B3B55FE2A8D4C" box="[433,454,1344,1370]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="63">R</collectionCode>
. Deeleman
</materialsCitation>
.
<materialsCitation id="3B3B3CACFFBFFFD3FDBB3B2AFF178D6B" collectionCode="RMNH" collectorName="Bako National Park, W. &amp; Coll. Deeleman" country="Malaysia" location="Bako National Park" pageId="8" pageNumber="63" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B927EAE3FFBFFFD3FDBB3B2AFD258D4F" bold="true" box="[593,713,1343,1369]" pageId="8" pageNumber="63">
<typeStatus id="54E88853FFBFFFD3FDBB3B2AFD258D4F" box="[593,713,1343,1369]" pageId="8" pageNumber="63">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D55FD78FFBFFFD3FD3D3B2AFCED8D4C" box="[727,769,1343,1370]" pageId="8" pageNumber="63" type="female">2 ♀</specimenCount>
(
<collectionCode id="ED42AE34FFBFFFD3FCFA3B2AFC858D4F" box="[784,873,1343,1369]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="63" type="Museum">RMNH</collectionCode>
.ARA. 15153), same data. Label data are shown in
<figureCitation id="13682A74FFBFFFD3FF7D3B76FF178D6B" box="[151,251,1379,1405]" captionStart="FIGURE 12" captionStartId="9.[151,250,1037,1061]" captionTargetBox="[161,1435,273,1013]" captionTargetId="figure-61@9.[151,1435,273,1013]" captionTargetPageId="9" captionText="FIGURE 12. Utivarachna chamaeleon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324282" httpUri="https://zenodo.org/record/8324282/files/figure.png" pageId="8" pageNumber="63">Fig. 12E</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFBFFFD3FF2D3B92FEAA8E43" pageId="8" pageNumber="63" type="diagnosis">
<paragraph id="8BEC36F1FFBFFFD3FF2D3B92FEAA8E43" blockId="8.[151,1437,1307,2018]" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF2D3B92FEAE8DB7" bold="true" box="[199,322,1415,1441]" pageId="8" pageNumber="63">Diagnosis.</emphasis>
In the
<taxonomicName id="4C534D72FFBFFFD3FE4A3B92FDCA8DB7" authorityName="Kishida" authorityYear="1940" box="[416,550,1415,1441]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBFFFD3FE4A3B92FDCA8DB7" box="[416,550,1415,1441]" italics="true" pageId="8" pageNumber="63">fukasawana</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C534D72FFBFFFD3FD6E3B9DFCDF8DB7" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[644,819,1415,1441]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBFFFD3FD6E3B9DFCDF8DB7" box="[644,819,1415,1441]" italics="true" pageId="8" pageNumber="63">U. chamaeleon</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="4C534D72FFBFFFD3FB783B9DFAAC8DB7" authorityName="Kishida" authorityYear="1940" box="[1170,1344,1415,1441]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBFFFD3FB783B9DFAAC8DB7" box="[1170,1344,1415,1441]" italics="true" pageId="8" pageNumber="63">U. fukasawana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D72FFBFFFD3FA6B3B9DFEFE8DD3" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFBFFFD3FA6B3B9DFEFE8DD3" italics="true" pageId="8" pageNumber="63">U. ichneumon</emphasis>
</taxonomicName>
by short stalk of posterior carapace.
<taxonomicName id="4C534D72FFBFFFD3FD593BBEFC238DD3" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[691,975,1451,1477]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBFFFD3FD593BBEFC238DD3" box="[691,975,1451,1477]" italics="true" pageId="8" pageNumber="63">Utivarachna chamaeleon</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C534D72FFBFFFD3FB893BB9FE258DFF" authority="Dankittipakul, Tavano &amp; Singtripop, 2011" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFBFFFD3FB893BB9FB1F8DD3" box="[1123,1267,1451,1477]" italics="true" pageId="8" pageNumber="63">U. galyaniae</emphasis>
<bibRefCitation id="EFC24B00FFBFFFD3FB113BBEFE258DFF" author="Dankittipakul, P. &amp; Tavano, M. &amp; Singtripop, T." pageId="8" pageNumber="63" pagination="133 - 146" refId="ref8829" refString="Dankittipakul, P., Tavano, M. &amp; Singtripop, T. (2011) Two new species of the spider genus Utivarachna Kishida, 1940 from Southwest Asia (Araneae, Corinnidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, 103, 133 - 146." type="journal article" year="2011">
Dankittipakul, Tavano &amp; Singtripop,
<quantity id="4CAB9B14FFBFFFD3FE783BDAFE258DFF" box="[402,457,1487,1513]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="8" pageNumber="63" unit="in" value="2011.0">2011</quantity>
</bibRefCitation>
</taxonomicName>
in structures of palp of males and copulatory organs of females. However, in males,
<taxonomicName id="4C534D72FFBFFFD3FA6B3BC5FEF28E1B" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBFFFD3FA6B3BC5FEF28E1B" italics="true" pageId="8" pageNumber="63">U. chamaeleon</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C534D72FFBFFFD3FDA13BE1FD378E1B" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" box="[587,731,1523,1549]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFBFFFD3FDA13BE1FD378E1B" box="[587,731,1523,1549]" italics="true" pageId="8" pageNumber="63">U. galyaniae</emphasis>
</taxonomicName>
by the shape of its cymbium, which is flatter and apically more slender than that of
<taxonomicName id="4C534D72FFBFFFD3FE91380DFDE18E27" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" box="[379,525,1559,1585]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFBFFFD3FE91380DFDE18E27" box="[379,525,1559,1585]" italics="true" pageId="8" pageNumber="63">U. galyaniae</emphasis>
</taxonomicName>
. In females,
<taxonomicName id="4C534D72FFBFFFD3FD49380DFCA28E27" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[675,846,1559,1585]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBFFFD3FD49380DFCA28E27" box="[675,846,1559,1585]" italics="true" pageId="8" pageNumber="63">U. chamaeleon</emphasis>
</taxonomicName>
has slender bursae, whereas
<taxonomicName id="4C534D72FFBFFFD3FB70380DFAC08E27" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" box="[1178,1324,1559,1585]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFBFFFD3FB70380DFAC08E27" box="[1178,1324,1559,1585]" italics="true" pageId="8" pageNumber="63">U. galyaniae</emphasis>
</taxonomicName>
has more swollen bursae.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFBFFFD1FF2D384AFC678F63" lastPageId="10" lastPageNumber="65" pageId="8" pageNumber="63" type="description">
<paragraph id="8BEC36F1FFBFFFD3FF2D384AFC798F63" blockId="8.[151,1437,1307,2018]" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF2D384AFE9A8E6F" bold="true" box="[199,374,1631,1657]" pageId="8" pageNumber="63">Measurements</emphasis>
(
<typeStatus id="54E88853FFBFFFD3FE61384AFE038E6F" box="[395,495,1631,1657]" pageId="8" pageNumber="63">holotype</typeStatus>
♁/
<typeStatus id="54E88853FFBFFFD3FDF83874FD958E6A" box="[530,633,1633,1660]" pageId="8" pageNumber="63">paratype</typeStatus>
<specimenCount id="9D55FD78FFBFFFD3FD6C384AFD768E6F" box="[646,666,1631,1657]" pageId="8" pageNumber="63" type="female"></specimenCount>
). Carapace length, 2.95/2.80; cephalic width, 1.72/1.50; thoracic width, 2.10/1.96. Diameter of eyes: AME 0.21/0.20; ALE 0.18/0.16; PME 0.13/0.13; PLE 0.18/0.17. Interdistances between eyes: AMEAME 0.03/0.05; AMEALE 0.05/0.07; ALEALE 0.56/0.52; PMEPME 0.22/0.21; PME PLE 0.33/0.32; PLEPLE 1.17/1.15; ALEPLE 0.22/0.25; AMEPME 0.03/0.08. MOA: length 0.35/0.37; anterior width 0.45/0.40; posterior width 0.55/0.50. Clypeus height 0.51/0.46. Abdomen length 2.76/3.05; width 1.70/3.05. Legs: I 10.36 (2.96, 1.10, 2.70, 2.60, 1.00)/8.55 (2.43, 0.97, 2.20, 1.97, 0.98); II 8.63 (2.53, 0.93, 2.12, 2.15, 0.90)/7.88 (2.27, 0.81, 1.95, 1.90, 0.95); III 5.43 (1.57, 0.66, 1.20, 1.42, 0.58)/5.44 (1.55, 0.63, 1.23, 1.45, 0.58); IV 7.11 (2.06, 0.66, 1.60, 2.07, 0.72)/7.13 (2.00, 0.50, 1.72, 2.13, 0.78).
</paragraph>
<paragraph id="8BEC36F1FFBFFFD2FF2D396AFB3A88EF" blockId="8.[151,1437,1307,2018]" lastBlockId="9.[151,1436,151,249]" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF2D396AFEE88F8F" bold="true" box="[199,260,1919,1945]" pageId="8" pageNumber="63">Male</emphasis>
(
<figureCitation id="13682A74FFBFFFD3FEF8396AFE708F8F" box="[274,412,1919,1945]" captionStart="FIGURE 12" captionStartId="9.[151,250,1037,1061]" captionTargetBox="[161,1435,273,1013]" captionTargetId="figure-61@9.[151,1435,273,1013]" captionTargetPageId="9" captionText="FIGURE 12. Utivarachna chamaeleon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324282" httpUri="https://zenodo.org/record/8324282/files/figure.png" pageId="8" pageNumber="63">Fig. 12AD</figureCitation>
). Carapace entirely granulated, dorsally oval, with short posterior stalk; AER almost straight in dorsal view; PER recurved in dorsal view; ocular tubercle accompanied with PLE; lateral margins of carapace strongly constricted between cephalic and thoracic parts; lateral margin of thoracic part undulating. Chelicerae granulated, with six prolateral and six retrolateral teeth. Clypeus with small wedge-shaped projection between chelicerae. Pedicel short. Abdomen oval; dorsum of anterior five-sixths and epigastric area weakly sclerotized. Legs granulated. Legs I and II elongated, with tooth-like cusps located ventrally from patella to tarsus.
</paragraph>
<caption id="DF2C6679FFBEFFD2FF7D3A18FF358C7B" ID-DOI="http://doi.org/10.5281/zenodo.8324282" ID-Zenodo-Dep="8324282" httpUri="https://zenodo.org/record/8324282/files/figure.png" pageId="9" pageNumber="64" startId="9.[151,250,1037,1061]" targetBox="[161,1435,273,1013]" targetPageId="9" targetType="figure">
<paragraph id="8BEC36F1FFBEFFD2FF7D3A18FF358C7B" blockId="9.[151,1437,1037,1133]" pageId="9" pageNumber="64">
<emphasis id="B927EAE3FFBEFFD2FF7D3A18FECF8C33" bold="true" box="[151,291,1037,1061]" pageId="9" pageNumber="64">FIGURE 12.</emphasis>
<taxonomicName id="4C534D72FFBEFFD2FEC63A18FCA78C33" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[300,843,1037,1061]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBEFFD2FEC63A18FDDC8C33" box="[300,560,1037,1061]" italics="true" pageId="9" pageNumber="64">Utivarachna chamaeleon</emphasis>
<bibRefCitation id="EFC24B00FFBEFFD2FDD03A18FCA78C33" author="Deeleman-Reinhold, C. L." box="[570,843,1037,1061]" pageId="9" pageNumber="64" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, male, holotype (RNMH.ARA.15153).
<emphasis id="B927EAE3FFBEFFD2FB003A18FB108C33" bold="true" box="[1258,1276,1037,1061]" pageId="9" pageNumber="64">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFBEFFD2FF3C3A24FF0B8C5F" bold="true" box="[214,231,1073,1097]" pageId="9" pageNumber="64">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBEFFD2FE523A24FE268C5F" bold="true" box="[440,458,1073,1097]" pageId="9" pageNumber="64">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFBEFFD2FC943A24FC7C8C5F" bold="true" box="[894,912,1073,1097]" pageId="9" pageNumber="64">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFBEFFD2FB4E3A24FB598C5F" bold="true" box="[1188,1205,1073,1097]" pageId="9" pageNumber="64">E</emphasis>
label. Scale bars AD: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFBEFFD2FF7D3895FA9F8EAA" ID-DOI="http://doi.org/10.5281/zenodo.8324284" ID-Zenodo-Dep="8324284" httpUri="https://zenodo.org/record/8324284/files/figure.png" pageId="9" pageNumber="64" startId="9.[151,250,1664,1688]" targetBox="[152,1434,1157,1639]" targetPageId="9" targetType="figure">
<paragraph id="8BEC36F1FFBEFFD2FF7D3895FA9F8EAA" blockId="9.[151,1436,1664,1724]" pageId="9" pageNumber="64">
<emphasis id="B927EAE3FFBEFFD2FF7D3895FECC8E8E" bold="true" box="[151,288,1664,1688]" pageId="9" pageNumber="64">FIGURE 13.</emphasis>
<taxonomicName id="4C534D72FFBEFFD2FECD3895FDC48E8E" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[295,552,1664,1688]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBEFFD2FECD3895FDC48E8E" box="[295,552,1664,1688]" italics="true" pageId="9" pageNumber="64">Utivarachna chamaeleon</emphasis>
</taxonomicName>
, male, holotype (RNMH.ARA.15153).
<emphasis id="B927EAE3FFBEFFD2FC553895FC3D8E8E" bold="true" box="[959,977,1664,1688]" pageId="9" pageNumber="64">A</emphasis>
right palp, retrolateral view;
<emphasis id="B927EAE3FFBEFFD2FB173895FAE28E8E" bold="true" box="[1277,1294,1664,1688]" pageId="9" pageNumber="64">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBEFFD2FF3C38B1FF048EAA" bold="true" box="[214,232,1700,1724]" pageId="9" pageNumber="64">C</emphasis>
same, prolateral view;
<emphasis id="B927EAE3FFBEFFD2FE3F38B1FE0B8EAA" bold="true" box="[469,487,1700,1724]" pageId="9" pageNumber="64">D</emphasis>
same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFBEFFD2FF2D391BFE7E8FAE" blockId="9.[151,1437,1806,1976]" pageId="9" pageNumber="64">
Palp (
<figureCitation id="13682A74FFBEFFD2FEE1391BFE768F3E" box="[267,410,1806,1832]" captionStart="FIGURE 13" captionStartId="9.[151,250,1664,1688]" captionTargetBox="[152,1434,1157,1639]" captionTargetId="figure-101@9.[151,1435,1157,1639]" captionTargetPageId="9" captionText="FIGURE 13. Utivarachna chamaeleon, male, holotype (RNMH.ARA.15153). A right palp, retrolateral view; B same, ventral view; C same, prolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.8324284" httpUri="https://zenodo.org/record/8324284/files/figure.png" pageId="9" pageNumber="64">Fig. 13AD</figureCitation>
). Cymbium pear-shaped, with slender apex; cymbium length about twice its width. Bulb inverted ovoid, prolateral half and subtegulum sclerotized, and retrolateral half partly membranous. Embolus coils inclining dorsally, and covered by cymbium. Sperm duct U-curved in posterior part of bulb, running retrolaterally with weak s-curve from posterior part to embolus. RTA strongly extended, apically developed into semi-transparent plate with apical claw.
</paragraph>
<caption id="DF2C6679FFBDFFD1FF7D3D7AFA7F8BBC" ID-DOI="http://doi.org/10.5281/zenodo.8324286" ID-Zenodo-Dep="8324286" httpUri="https://zenodo.org/record/8324286/files/figure.png" pageId="10" pageNumber="65" startId="10.[151,250,879,903]" targetBox="[159,1436,181,855]" targetPageId="10" targetType="figure">
<paragraph id="8BEC36F1FFBDFFD1FF7D3D7AFA7F8BBC" blockId="10.[151,1436,879,939]" pageId="10" pageNumber="65">
<emphasis id="B927EAE3FFBDFFD1FF7D3D7AFECD8B91" bold="true" box="[151,289,879,903]" pageId="10" pageNumber="65">FIGURE 14.</emphasis>
<taxonomicName id="4C534D72FFBDFFD1FEC33D7AFCAE8B91" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[297,834,879,903]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBDFFD1FEC33D7AFDC78B91" box="[297,555,879,903]" italics="true" pageId="10" pageNumber="65">Utivarachna chamaeleon</emphasis>
<bibRefCitation id="EFC24B00FFBDFFD1FDD93D7AFCAE8B91" author="Deeleman-Reinhold, C. L." box="[563,834,879,903]" pageId="10" pageNumber="65" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, female, paratype (RNMH.ARA.15153).
<emphasis id="B927EAE3FFBDFFD1FB073D7AFB138B91" bold="true" box="[1261,1279,879,903]" pageId="10" pageNumber="65">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFBDFFD1FF3C3D86FF0B8BBD" bold="true" box="[214,231,915,939]" pageId="10" pageNumber="65">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBDFFD1FE533D86FE278BBD" bold="true" box="[441,459,915,939]" pageId="10" pageNumber="65">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFBDFFD1FC6B3D86FC7F8BBD" bold="true" box="[897,915,915,939]" pageId="10" pageNumber="65">D</emphasis>
leg I, left, prolateral view. Scale bars AD: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFBDFFD1FF7D3B08FB378D4F" ID-DOI="http://doi.org/10.5281/zenodo.8324290" ID-Zenodo-Dep="8324290" httpUri="https://zenodo.org/record/8324290/files/figure.png" pageId="10" pageNumber="65" startId="10.[151,250,1309,1333]" targetBox="[151,1435,999,1285]" targetPageId="10" targetType="figure">
<paragraph id="8BEC36F1FFBDFFD1FF7D3B08FB378D4F" blockId="10.[151,1436,1309,1369]" pageId="10" pageNumber="65">
<emphasis id="B927EAE3FFBDFFD1FF7D3B08FECE8D23" bold="true" box="[151,290,1309,1333]" pageId="10" pageNumber="65">FIGURE 15.</emphasis>
<taxonomicName id="4C534D72FFBDFFD1FEC03B08FDC08D23" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[298,556,1309,1333]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="chamaeleon">
<emphasis id="B927EAE3FFBDFFD1FEC03B08FDC08D23" box="[298,556,1309,1333]" italics="true" pageId="10" pageNumber="65">Utivarachna chamaeleon</emphasis>
</taxonomicName>
, female, paratype (RNMH.ARA.15153).
<emphasis id="B927EAE3FFBDFFD1FC313B08FC018D23" bold="true" box="[987,1005,1309,1333]" pageId="10" pageNumber="65">A</emphasis>
epigyne, ventral view;
<emphasis id="B927EAE3FFBDFFD1FB0E3B08FB198D23" bold="true" box="[1252,1269,1309,1333]" pageId="10" pageNumber="65">B</emphasis>
internal organs, ventral view;
<emphasis id="B927EAE3FFBDFFD1FEC83B54FED88D4F" bold="true" box="[290,308,1345,1369]" pageId="10" pageNumber="65">C</emphasis>
same, dorsal view. Scale bar: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFBDFFD1FF2D3BBEFCFD8E6F" blockId="10.[151,1437,1451,1945]" pageId="10" pageNumber="65">
Coloration and setation (
<figureCitation id="13682A74FFBDFFD1FE093BBEFD9D8DD3" box="[483,625,1451,1477]" captionStart="FIGURE 12" captionStartId="9.[151,250,1037,1061]" captionTargetBox="[161,1435,273,1013]" captionTargetId="figure-61@9.[151,1435,273,1013]" captionTargetPageId="9" captionText="FIGURE 12. Utivarachna chamaeleon Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324282" httpUri="https://zenodo.org/record/8324282/files/figure.png" pageId="10" pageNumber="65">Fig. 12AD</figureCitation>
). Carapace brown, covered with fine setae. Chelicerae brown, covered with fine setae; prolateral margin of fang furrow with long setae. Abdomen cream, covered with fine setae; in dorsal view, anterior lateral sides and posterior end tinged with black; sclerotized dorsum and epigastric area amber-colored. Legs covered with fine setae; legs I and II brown to light brown in coxa, trochanter, femur, and patella but cream in tibia, metatarsus, and tarsus; legs III and IV light brown. Distal part of femur, patella, tibia, metatarsus, and tarsus tinged with black or dark-colored rings in all legs.
</paragraph>
<paragraph id="8BEC36F1FFBDFFD1FF2D3896FEAD8ED4" blockId="10.[151,1437,1451,1945]" pageId="10" pageNumber="65">
<emphasis id="B927EAE3FFBDFFD1FF2D3896FEF28E8B" bold="true" box="[199,286,1667,1693]" pageId="10" pageNumber="65">Female</emphasis>
(
<figureCitation id="13682A74FFBDFFD1FED83896FE2E8E8B" box="[306,450,1667,1693]" captionStart="FIGURE 14" captionStartId="10.[151,250,879,903]" captionTargetBox="[159,1436,181,855]" captionTargetId="figure-17@10.[151,1436,181,855]" captionTargetPageId="10" captionText="FIGURE 14. Utivarachna chamaeleon Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324286" httpUri="https://zenodo.org/record/8324286/files/figure.png" pageId="10" pageNumber="65">Fig. 14AD</figureCitation>
). Carapace and chelicerae almost same as in male. Abdomen oval, with sclerotized epigastric area.
</paragraph>
<paragraph id="8BEC36F1FFBDFFD1FF2D38DEFD858F47" blockId="10.[151,1437,1451,1945]" pageId="10" pageNumber="65">
Copulatory organs (
<figureCitation id="13682A74FFBDFFD1FE4038DEFDDB8EF0" box="[426,567,1739,1766]" captionStart="FIGURE 15" captionStartId="10.[151,250,1309,1333]" captionTargetBox="[151,1435,999,1285]" captionTargetId="figure-55@10.[151,1435,999,1285]" captionTargetPageId="10" captionText="FIGURE 15. Utivarachna chamaeleon, female, paratype (RNMH.ARA.15153). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bar: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8324290" httpUri="https://zenodo.org/record/8324290/files/figure.png" pageId="10" pageNumber="65">Fig. 15AC</figureCitation>
). Copulatory atrium nearly as long as wide. Copulatory openings located at anterior margin of atrium. Copulatory ducts extending anteriorly with S-curving, and then turning posteriorly with helical coils. Bursae long, elongate oval, extending posteriorly. Spermathecae oval. Fertilization ducts straight, extending from spermathecae posteriorly.
</paragraph>
<paragraph id="8BEC36F1FFBDFFD1FF2D394EFC678F63" blockId="10.[151,1437,1451,1945]" box="[199,907,1883,1910]" pageId="10" pageNumber="65">
Coloration and setation (
<figureCitation id="13682A74FFBDFFD1FE36394EFD8B8F63" box="[476,615,1883,1909]" captionStart="FIGURE 14" captionStartId="10.[151,250,879,903]" captionTargetBox="[159,1436,181,855]" captionTargetId="figure-17@10.[151,1436,181,855]" captionTargetPageId="10" captionText="FIGURE 14. Utivarachna chamaeleon Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15153). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324286" httpUri="https://zenodo.org/record/8324286/files/figure.png" pageId="10" pageNumber="65">Fig. 14AD</figureCitation>
). Almost same as in male.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFBDFFD1FF2D396AFD568F8F" box="[199,698,1919,1945]" pageId="10" pageNumber="65" type="distribution">
<paragraph id="8BEC36F1FFBDFFD1FF2D396AFD568F8F" blockId="10.[151,1437,1451,1945]" box="[199,698,1919,1945]" pageId="10" pageNumber="65">
<emphasis id="B927EAE3FFBDFFD1FF2D396AFE8D8F8F" bold="true" box="[199,353,1919,1945]" pageId="10" pageNumber="65">Distribution.</emphasis>
Central Kalimantan,
<collectingCountry id="F3447661FFBDFFD1FDB8396AFD5A8F8F" box="[594,694,1919,1945]" name="Malaysia" pageId="10" pageNumber="65">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>