treatments-xml/data/03/FA/87/03FA87D00006FF98FF734A860CCEFECB.xml

<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3857.2.8" ID-GBIF-Dataset="09b8963f-3aa7-4914-9242-09d5968cfe35" ID-ISSN="1175-5326" ID-Zenodo-Dep="229717" ID-ZooBank="6D902CD0-7A28-485F-A91E-E23EE74C8907" checkinTime="1460899676357" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel Ángel" docDate="2014" docId="03FA87D00006FF98FF734A860CCEFECB" docLanguage="en" docName="zt03857p300.pdf" docOrigin="Zootaxa 3857 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.10:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="10" docTitle="Macrodactylus megaphyllus Arce-Perez &amp; Moron, new species" docType="treatment" docVersion="8" lastPageNumber="295" masterDocId="FFC3FFA80007FF9BFFE44A110F43FFAE" masterDocTitle="New species and records of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Bolivia" masterLastPageNumber="300" masterPageNumber="292" pageNumber="293" updateTime="1665413707506" updateUser="tatiana">
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<mods:titleInfo>
<mods:title>New species and records of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Bolivia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arce-Pérez, Roberto</mods:namePart>
<mods:affiliation>Red Sistemática y Biodiversidad, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, Mexico.</mods:affiliation>
<mods:nameIdentifier type="email">roberto.arce@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morón, Miguel Ángel</mods:namePart>
<mods:affiliation>Red Sistemática y Biodiversidad, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, Mexico.</mods:affiliation>
<mods:nameIdentifier type="email">miguel.moron@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3857</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:extent unit="page">
<mods:start>292</mods:start>
<mods:end>300</mods:end>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3857.2.8</mods:identifier>
<mods:identifier type="GBIF-Dataset">09b8963f-3aa7-4914-9242-09d5968cfe35</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">229717</mods:identifier>
<mods:identifier type="ZooBank">6D902CD0-7A28-485F-A91E-E23EE74C8907</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6123030" ID-GBIF-Taxon="119587635" ID-Zenodo-Dep="6123030" LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB" httpUri="http://treatment.plazi.org/id/03FA87D00006FF98FF734A860CCEFECB" lastPageId="3" lastPageNumber="295" pageId="1" pageNumber="293">
<subSubSection box="[151,920,151,178]" pageId="1" pageNumber="293" type="nomenclature">
<paragraph blockId="1.[151,920,151,211]" box="[151,920,151,178]" pageId="1" pageNumber="293">
<heading bold="true" box="[151,920,151,178]" fontSize="11" level="1" pageId="1" pageNumber="293" reason="1">
<emphasis bold="true" box="[151,920,151,178]" pageId="1" pageNumber="293">
<taxonomicName authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" box="[151,764,151,178]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="megaphyllus" status="new species">
<emphasis bold="true" box="[151,490,151,177]" italics="true" pageId="1" pageNumber="293">Macrodactylus megaphyllus</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
, 
<taxonomicNameLabel box="[776,920,152,178]" pageId="1" pageNumber="293" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection box="[151,329,186,211]" pageId="1" pageNumber="293" type="description">
<paragraph blockId="1.[151,920,151,211]" box="[151,329,186,211]" pageId="1" pageNumber="293">
(
<figureCitation box="[159,266,186,211]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1620,1643]" captionTargetBox="[337,1249,407,1598]" captionTargetId="figure@1.[337,1250,407,1598]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Macrodactylus megaphyllus: 1) dorsal view, 2) lateral view. Macrodactylus yunganus: 3) dorsal view, 4) lateral view. Macrodactylus nobilis: 5) dorsal view, 6) lateral view." httpUri="https://zenodo.org/record/229718/files/figure.png" pageId="1" pageNumber="293">Figs. 1–2</figureCitation>
, 
<figureCitation box="[278,322,187,211]" captionStart="FIGURES 7 – 17" captionStartId="2.[151,264,1584,1607]" captionTargetBox="[372,1211,631,1558]" captionTargetId="figure@2.[366,1220,625,1563]" captionTargetPageId="2" captionText="FIGURES 7 – 17. Diagnostic structures of Macrodactylus megaphyllus: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. Macrodactylus yunganus: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. Macrodactylus nobilis: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="1" pageNumber="293">7–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="293" type="diagnosis">
<paragraph blockId="1.[151,1436,258,355]" pageId="1" pageNumber="293">
<emphasis bold="true" box="[151,266,258,283]" pageId="1" pageNumber="293">Diagnosis</emphasis>
. Body integument black with golden reflections; clypeus, funicle, tibiae, and tarsi reddish yellow; elytra dark-red, dorsally with whitish-yellow setae. Antenna longer than head; club black, as long as funicle. Tarsi without rings of white setae. Parameres glabrous.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/229718/files/figure.png" pageId="1" pageNumber="293" targetBox="[337,1249,407,1598]" targetPageId="1">
<paragraph blockId="1.[151,1435,1620,1673]" pageId="1" pageNumber="293">
<emphasis bold="true" box="[151,313,1620,1643]" pageId="1" pageNumber="293">FIGURES 1–6.</emphasis>
<taxonomicName box="[319,604,1620,1641]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="megaphyllus">
<emphasis box="[319,604,1620,1641]" italics="true" pageId="1" pageNumber="293">Macrodactylus megaphyllus</emphasis>
</taxonomicName>
: 1) dorsal view, 2) lateral view. 
<taxonomicName box="[925,1179,1620,1642]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="yunganus">
<emphasis box="[925,1179,1620,1642]" italics="true" pageId="1" pageNumber="293">Macrodactylus yunganus</emphasis>
</taxonomicName>
: 3) dorsal view, 4) lateral view. 
<taxonomicName box="[211,437,1651,1672]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis box="[211,437,1651,1672]" italics="true" pageId="1" pageNumber="293">Macrodactylus nobilis</emphasis>
</taxonomicName>
: 5) dorsal view, 6) lateral view.
</paragraph>
</caption>
<subSubSection lastPageId="2" lastPageNumber="294" pageId="1" pageNumber="293" type="description">
<paragraph blockId="1.[151,1436,1712,2025]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="294" pageId="1" pageNumber="293">
<emphasis bold="true" box="[199,337,1712,1737]" pageId="1" pageNumber="293">Description</emphasis>
. 
<typeStatus box="[355,459,1712,1737]" pageId="1" pageNumber="293">Holotype</typeStatus>
male: Body length 
<quantity box="[694,778,1713,1737]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="293" unit="mm" value="14.0">14 mm</quantity>
, humeral width 
<quantity box="[969,1062,1713,1737]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="293" unit="mm" value="4.2">4.2 mm</quantity>
. Color: Body black with golden reflections; clypeus, funicle, tibiae, and tarsi reddish-yellow; coxae, trochanters, and femora black (femora reddish yellow in ventrobasal region); elytra reddish brown. Vestiture: dorsally with whitish-yellow, short setae that do not completely cover the surface; tarsi without rings of white setae. Head: Generally short and wide, with long setae on ocular canthus and keel in front of eyes. Clypeus trapezoidal, anterior border nearly straight. Funicle slightly longer than head, reddish yellow; antennal club black, nearly as long as funicle (
<number box="[996,1034,1892,1917]" pageId="1" pageNumber="293" value="2.6">2.6</number>
: 
<quantity box="[1037,1132,1892,1917]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="293" unit="mm" value="2.8">2.8 mm</quantity>
). Maxillary palpus reddish yellow; mentum black, wide, oval, longitudinally furrowed, with scattered setae on the surface. Thorax: Pronotum hexagonal, convex; length 
<quantity box="[453,555,1964,1989]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="293" unit="mm" value="3.55">3.55 mm</quantity>
, width 
<quantity box="[637,740,1964,1989]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="1" pageNumber="293" unit="mm" value="3.25">3.25 mm</quantity>
; with black setae on margins. Prosternal process long, reddish brown, curved, and flat in frontal view. Scutellum black, subtriangular with vestiture similar to pronotum. Elytra reddish brown with striae slightly covered by vestiture. Metasternum wide, black, slightly flattened; with rows of long, dense, short setae and rows of long, slender, yellow setae between mesocoxae and metacoxae. Abdomen: Color black, robust. Sternites 
<number box="[502,544,223,248]" pageId="2" pageNumber="294" value="3.5" valueMax="5.0" valueMin="2.0">2–5</number>
moderately concave at middle, sternites 
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with 
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spine-like, red, long, slender setae at each side of midline; length of sternite 
<number box="[761,775,260,285]" pageId="2" pageNumber="294" value="5.0">5</number>
twice length of sternite 
<number box="[1043,1059,260,284]" pageId="2" pageNumber="294" value="4.0">4</number>
, with numerous long setae, apical third glabrous, light yellow; sternite 
<number box="[582,596,295,320]" pageId="2" pageNumber="294" value="6.0">6</number>
ovate, with long setae. Pygidium elongate, ovate, with numerous setae concentrically distributed on disc, and few long setae in the apex. Legs: Profemur with a subtriangular, curved, and sharp projection near the base. Mesotibia widened toward the apex; with two straight, lightly curved toward the apical spurs; with spine-like setae around apex. Tarsi enlarged; without rings of setae but with long, spine-like setae around apex; all tarsal claws long and cleft. Genitalia: Total length 
<quantity box="[941,1048,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="294" unit="mm" value="2.95">2.95 mm</quantity>
. Parameres stout, wide, glabrous, completely fused dorsobasally, widely curved toward apex, forming round-oval shape in distal view; with length 1.00 mm and width 
<number box="[385,437,512,536]" pageId="2" pageNumber="294" value="1.04">1.04</number>
; apex long, widely lanceolate, slightly acute (
<figureCitation box="[964,1076,511,536]" captionStart="FIGURES 7 – 17" captionStartId="2.[151,264,1584,1607]" captionTargetBox="[372,1211,631,1558]" captionTargetId="figure@2.[366,1220,625,1563]" captionTargetPageId="2" captionText="FIGURES 7 – 17. Diagnostic structures of Macrodactylus megaphyllus: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. Macrodactylus yunganus: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. Macrodactylus nobilis: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="2" pageNumber="294">
Figs. 
<number box="[1030,1076,512,536]" pageId="2" pageNumber="294" value="8.0" valueMax="9.0" valueMin="7.0">7–9</number>
</figureCitation>
). Parameres with lifted internal margin forming a crest that terminates near the apex.
</paragraph>
<caption httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="2" pageNumber="294" targetBox="[372,1211,631,1558]" targetPageId="2">
<paragraph blockId="2.[151,1436,1584,1698]" pageId="2" pageNumber="294">
<emphasis bold="true" box="[151,327,1584,1607]" pageId="2" pageNumber="294">FIGURES 7–17.</emphasis>
Diagnostic structures of 
<taxonomicName box="[580,865,1585,1606]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="megaphyllus">
<emphasis box="[580,865,1585,1606]" italics="true" pageId="2" pageNumber="294">Macrodactylus megaphyllus</emphasis>
</taxonomicName>
: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. 
<taxonomicName box="[414,671,1615,1637]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="yunganus">
<emphasis box="[414,671,1615,1637]" italics="true" pageId="2" pageNumber="294">Macrodactylus yunganus</emphasis>
</taxonomicName>
: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. 
<taxonomicName box="[688,916,1646,1667]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis box="[688,916,1646,1667]" italics="true" pageId="2" pageNumber="294">Macrodactylus nobilis</emphasis>
</taxonomicName>
: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm.
</paragraph>
</caption>
<paragraph blockId="2.[151,1437,1737,2015]" box="[199,411,1737,1762]" pageId="2" pageNumber="294">
<emphasis bold="true" box="[199,286,1737,1762]" pageId="2" pageNumber="294">Female</emphasis>
: Unknown
</paragraph>
<paragraph blockId="2.[151,1437,1737,2015]" pageId="2" pageNumber="294">
<emphasis bold="true" box="[199,309,1773,1798]" pageId="2" pageNumber="294">Variation</emphasis>
: One specimen with elytra clear yellow; femora black or reddish yellow in ventrobasal region; body length 
<quantity box="[228,352,1810,1834]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="2" pageNumber="294" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
; humeral width 3.0– 
<quantity box="[582,671,1810,1834]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="294" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294" type="materials_examined">
<paragraph blockId="2.[151,1437,1737,2015]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294">
<emphasis bold="true" box="[199,429,1845,1870]" pageId="2" pageNumber="294">Material examined</emphasis>
: 
<materialsCitation collectingDate="1999-01-28" collectionCode="CMNC" collectorName="R. Anderson" country="Bolivia" elevation="2700" latitude="-17.826946" location="Yungas de la Siberia" longLatPrecision="21" longitude="-64.7175" municipality="Santa Cruz" pageId="2" pageNumber="294" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[447,551,1845,1870]" pageId="2" pageNumber="294">Holotype</typeStatus>
<specimenCount box="[562,621,1845,1870]" pageId="2" pageNumber="294" type="male">male</specimenCount>
: 
<collectingCountry box="[635,753,1846,1870]" name="Bolivia" pageId="2" pageNumber="294">BOLIVIA</collectingCountry>
, 
<collectingMunicipality box="[768,898,1846,1870]" pageId="2" pageNumber="294">Santa Cruz</collectingMunicipality>
, 
<locationDeviation box="[913,1174,1845,1870]" pageId="2" pageNumber="294">
<quantity box="[913,988,1845,1870]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="294" unit="km" value="34.0">34 km</quantity>
NW Comarapa
</locationDeviation>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0005FF99FB414D260ADBF8E0" box="[1189,1432,1845,1871]" country="Bolivia" latitude="-17.826946" longLatPrecision="21" longitude="-64.7175" municipality="Santa Cruz" name="Yungas de la Siberia" pageId="2" pageNumber="294">Yungas de la Siberia</location>
, 
<geoCoordinate box="[151,277,1882,1906]" direction="south" orientation="latitude" pageId="2" pageNumber="294" precision="15" value="-17.826946">
<number box="[151,179,1882,1906]" pageId="2" pageNumber="294" value="17.0">17</number>
º 
<number box="[188,219,1882,1906]" pageId="2" pageNumber="294" value="49.0">49</number>
' 
<number box="[223,253,1882,1906]" pageId="2" pageNumber="294" value="37.0">37</number>
''S
</geoCoordinate>
; 
<geoCoordinate box="[292,427,1881,1906]" direction="west" orientation="longitude" pageId="2" pageNumber="294" precision="15" value="-64.7175">
<number box="[292,321,1881,1906]" pageId="2" pageNumber="294" value="64.0">64</number>
º 
<number box="[330,358,1881,1906]" pageId="2" pageNumber="294" value="43.0">43</number>
'03''W
</geoCoordinate>
, 
<date box="[441,556,1882,1906]" pageId="2" pageNumber="294" value="1999-01-28">
<collectingDate box="[441,556,1882,1906]" pageId="2" pageNumber="294" value="1999-01-28">
<number box="[441,470,1882,1906]" pageId="2" pageNumber="294" value="28.0">28</number>
-I- 
<number box="[497,556,1882,1906]" pageId="2" pageNumber="294" value="1999.0">1999</number>
</collectingDate>
</date>
, 
<quantity box="[571,661,1882,1906]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="294" unit="m" value="2700.0">
<elevation box="[571,661,1882,1906]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="294" unit="m" value="2700.0">2700 m</elevation>
</quantity>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0005FF99FD404D4B0CD5F8DC" box="[676,918,1881,1906]" country="Bolivia" latitude="-17.826946" longLatPrecision="21" longitude="-64.7175" municipality="Santa Cruz" name="wet elfin forest litter" pageId="2" pageNumber="294">wet elfin forest litter</location>
, 
<number box="[932,961,1881,1906]" pageId="2" pageNumber="294" value="99.0">99</number>
-005, 
<collectorName box="[1029,1174,1881,1906]" pageId="2" pageNumber="294">R. Anderson</collectorName>
(
<collectionCode box="[1192,1281,1882,1906]" pageId="2" pageNumber="294">CMNC</collectionCode>
).
</materialsCitation>
<materialsCitation collectingDate="1999-01-27" collectionCode="CMNC" collectorName="F. Genier" country="Bolivia" elevation="2600" latitude="-17.826946" location="Kara Huasi" longLatPrecision="21" longitude="-64.7175" municipality="Santa-Cruz" pageId="2" pageNumber="294" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<typeStatus box="[1303,1412,1882,1906]" pageId="2" pageNumber="294">Paratypes</typeStatus>
<specimenCount pageId="2" pageNumber="294" type="male">
<number box="[1422,1436,1881,1906]" pageId="2" pageNumber="294" value="5.0">5</number>
males
</specimenCount>
: 
<collectingCountry box="[230,348,1918,1942]" name="Bolivia" pageId="2" pageNumber="294">BOLIVIA</collectingCountry>
, 
<collectingMunicipality box="[359,488,1918,1942]" pageId="2" pageNumber="294">Santa-Cruz</collectingMunicipality>
, 
<locationDeviation box="[499,768,1917,1942]" pageId="2" pageNumber="294">
<quantity box="[499,590,1917,1942]" metricMagnitude="4" metricUnit="m" metricValue="2.6399999999999997" pageId="2" pageNumber="294" unit="km" value="26.4">26.4 km</quantity>
NW Comarapa
</locationDeviation>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0005FF99FCEE4D6E0CC8F838" box="[778,907,1917,1942]" country="Bolivia" latitude="-17.826946" longLatPrecision="21" longitude="-64.7175" municipality="Santa-Cruz" name="Kara Huasi" pageId="2" pageNumber="294">Kara Huasi</location>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0005FF99FC724D6E0B3EF838" box="[918,1149,1917,1943]" country="Bolivia" latitude="-17.826946" longLatPrecision="21" longitude="-64.7175" municipality="Santa-Cruz" name="Yungas de la Siberia" pageId="2" pageNumber="294">Yungas de la Siberia</location>
, 
<geoCoordinate box="[1160,1286,1918,1942]" direction="south" orientation="latitude" pageId="2" pageNumber="294" precision="15" value="-17.825556">
<number box="[1160,1188,1918,1942]" pageId="2" pageNumber="294" value="17.0">17</number>
º 
<number box="[1197,1228,1918,1942]" pageId="2" pageNumber="294" value="49.0">49</number>
' 
<number box="[1231,1262,1918,1942]" pageId="2" pageNumber="294" value="32.0">32</number>
''S
</geoCoordinate>
; 
<geoCoordinate box="[1298,1432,1917,1942]" direction="west" orientation="longitude" pageId="2" pageNumber="294" precision="15" value="-64.68806">
<number box="[1298,1328,1917,1942]" pageId="2" pageNumber="294" value="64.0">64</number>
º 
<number box="[1337,1365,1917,1942]" pageId="2" pageNumber="294" value="41.0">41</number>
' 
<number box="[1367,1393,1917,1942]" pageId="2" pageNumber="294" value="17.0">17</number>
''W
</geoCoordinate>
, 
<date box="[151,266,1954,1978]" pageId="2" pageNumber="294" value="1999-01-27">
<collectingDate box="[151,266,1954,1978]" pageId="2" pageNumber="294" value="1999-01-27">
<number box="[151,180,1954,1978]" pageId="2" pageNumber="294" value="27.0">27</number>
-I- 
<number box="[207,266,1954,1978]" pageId="2" pageNumber="294" value="1999.0">1999</number>
</collectingDate>
</date>
, 
<quantity box="[278,366,1953,1978]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="294" unit="m" value="2600.0">
<elevation box="[278,366,1953,1978]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="294" unit="m" value="2600.0">2600 m</elevation>
</quantity>
, Yungas 
<emphasis box="[470,502,1954,1977]" italics="true" pageId="2" pageNumber="294">ex.</emphasis>
hand collecting, 
<number box="[699,729,1954,1978]" pageId="2" pageNumber="294" value="99.0">99</number>
-003, 
<collectorName box="[794,898,1953,1978]" pageId="2" pageNumber="294">F. Génier</collectorName>
(
<specimenCount box="[918,992,1953,1978]" pageId="2" pageNumber="294" type="male">
<number box="[918,930,1954,1978]" pageId="2" pageNumber="294" value="1.0">1</number>
male
</specimenCount>
<collectionCode box="[1001,1089,1954,1978]" pageId="2" pageNumber="294">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="1999-01-30" collectionCode="CMNC, MXAL" collectorName="F. Genier" country="Bolivia" elevation="2700" lastPageId="3" lastPageNumber="295" latitude="-17.568611" location="Sehuenca" longLatPrecision="21" longitude="-65.27528" municipality="Cochabamba" pageId="2" pageNumber="294" specimenCount="2" specimenCount-male="2">
<collectingCountry box="[1109,1227,1954,1978]" name="Bolivia" pageId="2" pageNumber="294">BOLIVIA</collectingCountry>
, 
<collectingMunicipality box="[1240,1389,1953,1978]" pageId="2" pageNumber="294">Cochabamba</collectingMunicipality>
, 
<locationDeviation pageId="2" pageNumber="294">
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="294" unit="km" value="6.5">6.5 km</quantity>
NW Epizana
</locationDeviation>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0005FF99FE804DD40E91F870" box="[356,466,1989,2014]" country="Bolivia" latitude="-17.568611" longLatPrecision="21" longitude="-65.27528" municipality="Cochabamba" name="Sehuenca" pageId="2" pageNumber="294">Sehuenca</location>
, 
<geoCoordinate box="[479,607,1990,2014]" direction="south" orientation="latitude" pageId="2" pageNumber="294" precision="15" value="-17.568611">
<number box="[479,507,1990,2014]" pageId="2" pageNumber="294" value="17.0">17</number>
° 
<number box="[517,549,1990,2014]" pageId="2" pageNumber="294" value="34.0">34</number>
'07''S
</geoCoordinate>
; 
<geoCoordinate box="[621,757,1989,2014]" direction="west" orientation="longitude" pageId="2" pageNumber="294" precision="15" value="-65.27528">
<number box="[621,650,1989,2014]" pageId="2" pageNumber="294" value="65.0">65</number>
° 
<number box="[659,687,1989,2014]" pageId="2" pageNumber="294" value="16.0">16</number>
' 
<number box="[690,718,1989,2014]" pageId="2" pageNumber="294" value="31.0">31</number>
''W
</geoCoordinate>
, 
<date box="[770,885,1990,2014]" pageId="2" pageNumber="294" value="1999-01-30">
<collectingDate box="[770,885,1990,2014]" pageId="2" pageNumber="294" value="1999-01-30">
<number box="[770,800,1990,2014]" pageId="2" pageNumber="294" value="30.0">30</number>
-I- 
<number box="[827,885,1990,2014]" pageId="2" pageNumber="294" value="1999.0">1999</number>
</collectingDate>
</date>
, 
<quantity box="[898,986,1990,2014]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="294" unit="m" value="2700.0">
<elevation box="[898,986,1990,2014]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="294" unit="m" value="2700.0">2700 m</elevation>
</quantity>
, Yungas ex. hand collecting, 
<number box="[1321,1350,1990,2014]" pageId="2" pageNumber="294" value="99.0">99</number>
-019, 
<collectorName lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294">F. Génier</collectorName>
(
<specimenCount box="[250,329,151,176]" pageId="3" pageNumber="295" type="male">
<number box="[250,262,152,176]" pageId="3" pageNumber="295" value="1.0">1</number>
male
</specimenCount>
<collectionCode box="[342,427,152,176]" pageId="3" pageNumber="295">CMNC</collectionCode>
, 
<specimenCount box="[444,525,151,176]" pageId="3" pageNumber="295" type="male">
<number box="[444,458,152,176]" pageId="3" pageNumber="295" value="1.0">1</number>
male
</specimenCount>
<collectionCode box="[538,626,152,176]" pageId="3" pageNumber="295">MXAL</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="1999-12-14" collectionCode="MXAL" collectorName="G. Goemans" country="Bolivia" elevation="3250" location="Camino entre Cochabamba y Santa Cruz" pageId="3" pageNumber="295" specimenCount="2" specimenCount-male="2">
<collectingCountry box="[650,768,152,176]" name="Bolivia" pageId="3" pageNumber="295">BOLIVIA</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:03FA87D00006FF98FF734A860CCEFECB:8E8C601D0004FF98FCF44A860BB4FF1E" box="[784,1271,151,176]" country="Bolivia" name="Camino entre Cochabamba y Santa Cruz" pageId="3" pageNumber="295">Camino entre Cochabamba y Santa Cruz</location>
, 
<date box="[1288,1432,152,176]" pageId="3" pageNumber="295" value="1999-12-14">
<collectingDate box="[1288,1432,152,176]" pageId="3" pageNumber="295" value="1999-12-14">
<number box="[1288,1315,152,176]" pageId="3" pageNumber="295" value="14.0">14</number>
-XII- 
<number box="[1374,1432,152,176]" pageId="3" pageNumber="295" value="1999.0">1999</number>
</collectingDate>
</date>
, 
<quantity box="[151,308,188,213]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="3" pageNumber="295" unit="m" value="3250.0" valueMax="3500.0" valueMin="3000.0">
<elevation box="[151,308,188,213]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="3" pageNumber="295" unit="m" value="3250.0" valueMax="3500.0" valueMin="3000.0">3000–3500 m</elevation>
</quantity>
, Volando, 
<collectorName box="[426,563,188,212]" pageId="3" pageNumber="295">G. Goemans</collectorName>
(
<specimenCount box="[579,665,188,213]" pageId="3" pageNumber="295" type="male">
<number box="[579,594,188,212]" pageId="3" pageNumber="295" value="2.0">2</number>
males
</specimenCount>
<collectionCode box="[673,761,188,212]" pageId="3" pageNumber="295">MXAL</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="295" type="biology_ecology">
<paragraph blockId="3.[151,1437,151,357]" pageId="3" pageNumber="295">
<emphasis bold="true" box="[199,382,223,248]" pageId="3" pageNumber="295">Natural history</emphasis>
. Specimens were collected during daylight hours flying or on the leaves of unidentified small trees in wet elfin forest in December (
<number box="[576,591,260,284]" pageId="3" pageNumber="295" value="2.0">2</number>
) and January (
<number box="[757,772,260,284]" pageId="3" pageNumber="295" value="4.0">4</number>
) at elevations of 
<quantity box="[966,1124,260,285]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="3" pageNumber="295" unit="m" value="3050.0" valueMax="3500.0" valueMin="2600.0">2600–3500 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="295" type="discussion">
<paragraph blockId="3.[151,1437,151,357]" pageId="3" pageNumber="295">
<emphasis bold="true" box="[199,308,295,320]" pageId="3" pageNumber="295">Remarks</emphasis>
. Males of the new species appear distinct from other 
<taxonomicName box="[908,1075,296,320]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis box="[908,1075,296,320]" italics="true" pageId="3" pageNumber="295">Macrodactylus</emphasis>
</taxonomicName>
because of the notably enlarged antennomeres, whose combined lengths exceeds the length of head.
</paragraph>
</subSubSection>
</treatment>
</document>