treatments-xml/data/03/FA/87/03FA8785FFF9657DC0E2F895B0A9FF51.xml

<document id="2B0A85780CE45577B0A47F9AC1C2506C" ID-DOI="10.11646/zootaxa.4034.2.4" ID-GBIF-Dataset="69aeb7a8-4960-41a0-a9ed-cbe23aab24e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="242957" ID-ZooBank="C09E1E16-5D8B-4F20-90A4-07050A1A560D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461300037478" checkinUser="plazi" docAuthor="Ribeiro, Siria, Santos-Jr, Alfredo P. &amp; Zaher, Hussam" docDate="2015" docId="03FA8785FFF9657DC0E2F895B0A9FF51" docLanguage="en" docName="zt04034p324.pdf" docOrigin="Zootaxa 4034 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leposternon bagual Ribeiro, Santos-Jr &amp; Zaher, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="319" masterDocId="FFC3FFFDFFF86576C075FF9EB40CFF85" masterDocTitle="A new species of Leposternon Wagler, 1824 (Squamata, Amphisbaenia) from northeastern Argentina" masterLastPageNumber="324" masterPageNumber="309" pageNumber="310" updateTime="1698617244783" updateUser="plazi">
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<mods:title id="228F4C5EF18989E4425188AD7AB0CA75">A new species of Leposternon Wagler, 1824 (Squamata, Amphisbaenia) from northeastern Argentina</mods:title>
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<mods:namePart id="1FA3074951D99C5EEA2A049A0523C960">Ribeiro, Siria</mods:namePart>
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<mods:namePart id="9DAE96A83E89774B7BEA89268DD41703">Santos-Jr, Alfredo P.</mods:namePart>
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<mods:namePart id="C7F2F84790410CF5130379A14AA594CA">Zaher, Hussam</mods:namePart>
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<mods:date id="34349E25216440CEAC0B2429E11F9D06">2015</mods:date>
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<treatment id="03FA8785FFF9657DC0E2F895B0A9FF51" ID-DOI="http://doi.org/10.5281/zenodo.6101445" ID-GBIF-Taxon="119643649" ID-Zenodo-Dep="6101445" LSID="urn:lsid:plazi:treatment:03FA8785FFF9657DC0E2F895B0A9FF51" httpUri="http://treatment.plazi.org/id/03FA8785FFF9657DC0E2F895B0A9FF51" lastPageId="11" lastPageNumber="319" pageId="1" pageNumber="310">
<subSubSection id="C3496518FFF96577C0E2F895B72EF828" pageId="1" pageNumber="310" type="nomenclature">
<paragraph id="8BEC3693FFF96577C0E2F895B5E1F8A3" blockId="1.[151,493,1803,1863]" box="[151,493,1803,1830]" pageId="1" pageNumber="310">
<heading id="D0A481FFFFF96577C0E2F895B5E1F8A3" bold="true" box="[151,493,1803,1830]" fontSize="11" level="1" pageId="1" pageNumber="310" reason="1">
<emphasis id="B927EA81FFF96577C0E2F895B5E1F8A3" bold="true" box="[151,493,1803,1830]" pageId="1" pageNumber="310">
<taxonomicName id="4C534D10FFF96577C0E2F895B584F8A0" ID-CoL="88J3G" box="[151,392,1803,1829]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="310" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFF96577C0E2F895B584F8A0" bold="true" box="[151,392,1803,1829]" italics="true" pageId="1" pageNumber="310">Leposternon bagual</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457FAFFF96577C1E5F892B5E1F8A3" box="[400,493,1804,1830]" pageId="1" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC3693FFF96577C0E2F8B0B51EF8C2" blockId="1.[151,493,1803,1863]" box="[151,274,1838,1863]" pageId="1" pageNumber="310">
(
<figureCitation id="13682A16FFF96577C0EAF8B0B505F8C2" box="[159,265,1838,1863]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1924,1946]" captionStartId-1="3.[151,250,1779,1801]" captionStartId-2="5.[151,250,1758,1780]" captionStartId-3="6.[151,250,697,719]" captionTargetBox-0="[166,1413,421,1888]" captionTargetBox-1="[150,1436,195,1756]" captionTargetBox-2="[151,1436,193,1736]" captionTargetBox-3="[169,1416,182,678]" captionTargetId-0="figure@2.[151,1436,409,1903]" captionTargetId-1="figure@3.[150,1437,195,1757]" captionTargetId-2="figure@5.[151,1436,193,1737]" captionTargetId-3="figure@6.[169,1423,177,683]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." captionText-1="FIGURE 2. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." captionText-2="FIGURE 3. Dorsal view of the skull in the genus Leposternon. (A) skull of Leposternon bagual sp. nov. (paratype, FML 11313) with rostral processes of the maxillae and nasals contacting each other on the facial portion of the skull; (B) skull of Leposternon aff. microcephalum (MRT 11984) with rostral processes of the maxillae and nasals not contacting each other on the facial portion of the skull; (C) skull of Leposternon polystegum (ZUFRJ 528) without rostral processes of the maxillae and nasals. Abbreviations: F, frontal; N, nasal; PF, prefrontal; MX, maxilla; NPPM, nasal process of premaxilla; RPM, rostral process of maxilla; RPPM, rostral process of premaxilla." captionText-3="FIGURE 4. Dorsal view of the heads of paratypes of Leposternon bagual sp. nov. (A) FML 11312, (B) FML 11313 and (C) FML 11318. Scale bar = 5 mm." httpUri-0="https://zenodo.org/record/242958/files/figure.png" httpUri-1="https://zenodo.org/record/242959/files/figure.png" httpUri-2="https://zenodo.org/record/242960/files/figure.png" httpUri-3="https://zenodo.org/record/242961/files/figure.png" pageId="1" pageNumber="310">Figs. 1–4</figureCitation>
)
</paragraph>
<paragraph id="8BEC3693FFF96577C0E2F8E7B72EF828" blockId="1.[151,847,1910,1966]" pageId="1" pageNumber="310">
<taxonomicName id="4C534D10FFF96577C0E2F8E7B5B4F808" box="[151,440,1911,1934]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="310" phylum="Chordata" rank="species" species="microcephalum">
<emphasis id="B927EA81FFF96577C0E2F8E7B5B4F808" box="[151,440,1911,1934]" italics="true" pageId="1" pageNumber="310">Leposternon microcephalum</emphasis>
</taxonomicName>
— 
<bibRefCitation id="EFC24B62FFF96577C1A6F8E9B743F808" author="Scrocchi" box="[467,847,1911,1933]" pageId="1" pageNumber="310" refString="Scrocchi, G. J. &amp; Giraudo, A. R. (2005) Reptiles de la Reserva El Bagual. In: Giacomo, A. G. &amp; Krapovickas, S. F. (Eds.), Historia natural y paisaje de la Reserva El Bagual, Provincia de Formosa, Argentina. Inventario de la fauna de vertebrados y de la flora vascular de un area protegida del Chaco Humedo. Temas de Naturaleza y Conservacion 4. Aves Argentinas / Asociacion Ornitologica del Plata, Buenos Aires, p. 155 - 198." type="journal article" year="2005">Scrocchi &amp; Giraudo (2005: 163, 181)</bibRefCitation>
<taxonomicName id="4C534D10FFF96577C0E2F807B5B5F828" box="[151,441,1943,1966]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="310" phylum="Chordata" rank="species" species="microcephalum">
<emphasis id="B927EA81FFF96577C0E2F807B5B5F828" box="[151,441,1943,1966]" italics="true" pageId="1" pageNumber="310">Leposternon microcephalum</emphasis>
</taxonomicName>
(
<emphasis id="B927EA81FFF96577C1B2F807B606F82B" box="[455,522,1945,1966]" italics="true" pageId="1" pageNumber="310">partim</emphasis>
)—Ribeiro 
<emphasis id="B927EA81FFF96577C2F5F807B6A0F828" box="[640,684,1943,1966]" italics="true" pageId="1" pageNumber="310">et al</emphasis>
. (2008: 35)
</paragraph>
</subSubSection>
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<paragraph id="8BEC3693FFF96574C0E2F844B63BFF51" blockId="1.[151,1436,2010,2035]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="311" pageId="1" pageNumber="310">
<emphasis id="B927EA81FFF96577C0E2F844B508F876" bold="true" box="[151,260,2010,2035]" pageId="1" pageNumber="310">
<typeStatus id="54E88831FFF96577C0E2F844B508F876" box="[151,260,2010,2035]" pageId="1" pageNumber="310" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<collectionCode id="ED42AE56FFF96577C16EF845B559F877" box="[283,341,2011,2034]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="1" pageNumber="310">FML</collectionCode>
11319 (field number REB 21242), an adult female from El 
<taxonomicName id="4C534D10FFF96577C42FF844B0A5F876" box="[1114,1193,2010,2035]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="bagual">Bagual</taxonomicName>
Ecological Reserve (
<geoCoordinate id="EE675054FFFA6574C0D5FF09B525FF35" box="[160,297,151,176]" direction="south" orientation="latitude" pageId="2" pageNumber="311" precision="15" value="-26.181389">26º10’53’’S</geoCoordinate>
; 
<geoCoordinate id="EE675054FFFA6574C14FFF09B5C2FF35" box="[314,462,151,176]" direction="west" orientation="longitude" pageId="2" pageNumber="311" precision="15" value="-58.944168">58º56’39’’W</geoCoordinate>
), Province of 
<collectingCountry id="F3447603FFFA6574C20FFF06B6D3FF2A" box="[634,735,152,175]" name="Taiwan" pageId="2" pageNumber="311">Formosa</collectingCountry>
, Department of Laishi, 
<collectingCountry id="F3447603FFFA6574C38FFF09B060FF35" box="[1018,1132,151,176]" name="Argentina" pageId="2" pageNumber="311">Argentina</collectingCountry>
, collected in 0 
<date id="FFED1053FFFA6574C56BFF09B4DEFF51" pageId="2" pageNumber="311" value="2002-02-05">5 February 2002</date>
, collector unknown (
<figureCitation id="13682A16FFFA6574C1C8FF22B607FF51" box="[445,523,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="2" pageNumber="311">Figs. 1</figureCitation>
, 
<figureCitation id="13682A16FFFA6574C26CFF22B625FF51" box="[537,553,188,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1779,1801]" captionTargetBox="[150,1436,195,1756]" captionTargetId="figure@3.[150,1437,195,1757]" captionTargetPageId="3" captionText="FIGURE 2. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242959/files/figure.png" pageId="2" pageNumber="311">2</figureCitation>
).
</paragraph>
<paragraph id="8BEC3693FFFA6574C0B2FF41B776FEE0" blockId="2.[151,1436,151,357]" pageId="2" pageNumber="311">
<emphasis id="B927EA81FFFA6574C0B2FF41B54AFF7D" bold="true" box="[199,326,223,248]" pageId="2" pageNumber="311">
<typeStatus id="54E88831FFFA6574C0B2FF41B54DFF7D" box="[199,321,223,248]" pageId="2" pageNumber="311" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED42AE56FFFA6574C13BFF7EB584FF72" box="[334,392,224,247]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="2" pageNumber="311">FML</collectionCode>
11312 (field number REB 20887), a male collected in 0 
<date id="FFED1053FFFA6574C464FF7EB0CEFF7D" box="[1041,1218,223,248]" pageId="2" pageNumber="311" value="1992-10-01">1 October 1992</date>
; 
<collectionCode id="ED42AE56FFFA6574C4A5FF7EB106FF72" box="[1232,1290,224,247]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="2" pageNumber="311">FML</collectionCode>
11313 (field number REB 20888), a female collected in 
<date id="FFED1053FFFA6574C20CFE9AB71DFE99" box="[633,785,260,284]" pageId="2" pageNumber="311" value="1992-06-14">14 June 1992</date>
; 
<collectionCode id="ED42AE56FFFA6574C369FE9BB75AFE99" box="[796,854,261,284]" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="2" pageNumber="311">FML</collectionCode>
11318 (field number REB 1227), a female collected in 0 
<date id="FFED1053FFFA6574C0B0FEB6B572FEC5" box="[197,382,295,320]" pageId="2" pageNumber="311" value="1993-02-01">1 February 1993</date>
; all three 
<typeStatus id="54E88831FFFA6574C199FEB6B654FEC5" box="[492,600,296,320]" pageId="2" pageNumber="311" type="paratype">paratypes</typeStatus>
from El 
<taxonomicName id="4C534D10FFFA6574C2CEFEB9B706FEC5" box="[699,778,295,320]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="321" phylum="Chordata" rank="species" species="bagual">Bagual</taxonomicName>
Ecological Reserve (
<geoCoordinate id="EE675054FFFA6574C38EFEB9B088FEC5" box="[1019,1156,295,320]" direction="south" orientation="latitude" pageId="2" pageNumber="311" precision="15" value="-26.181389">26º10’53’’S</geoCoordinate>
; 
<geoCoordinate id="EE675054FFFA6574C4E5FEB9B128FEC5" box="[1168,1316,295,320]" direction="west" orientation="longitude" pageId="2" pageNumber="311" precision="15" value="-58.944168">58º56’39’’W</geoCoordinate>
), Province of 
<collectingCountry id="F3447603FFFA6574C0C0FED3B516FEE1" box="[181,282,333,356]" name="Taiwan" pageId="2" pageNumber="311">Formosa</collectingCountry>
, Department of Laishi, 
<collectingCountry id="F3447603FFFA6574C256FED2B69AFEE0" box="[547,662,332,357]" name="Argentina" pageId="2" pageNumber="311">Argentina</collectingCountry>
; collector unknown.
</paragraph>
</subSubSection>
<caption id="DF2C661BFFFA6574C0E2F81AB693F83C" httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="2" pageNumber="311" targetBox="[166,1413,421,1888]" targetPageId="2">
<paragraph id="8BEC3693FFFA6574C0E2F81AB693F83C" blockId="2.[151,1436,1924,1977]" pageId="2" pageNumber="311">
<emphasis id="B927EA81FFFA6574C0E2F81AB519F81C" bold="true" box="[151,277,1924,1946]" pageId="2" pageNumber="311">FIGURE 1.</emphasis>
<taxonomicName id="4C534D10FFFA6574C169F81BB5E5F81F" box="[284,489,1924,1946]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="311" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFFA6574C169F81BB5E5F81F" box="[284,489,1924,1946]" italics="true" pageId="2" pageNumber="311">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFFA6574C187F81BB648F81F" bold="true" box="[498,580,1925,1946]" pageId="2" pageNumber="311">
<taxonomicNameLabel id="A21457FAFFFA6574C187F81BB648F81F" box="[498,580,1925,1946]" pageId="2" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="DF2C661BFFFB6575C0E2F96DB693F8A2" httpUri="https://zenodo.org/record/242959/files/figure.png" pageId="3" pageNumber="312" targetBox="[150,1436,195,1756]" targetPageId="3">
<paragraph id="8BEC3693FFFB6575C0E2F96DB693F8A2" blockId="3.[151,1436,1779,1831]" pageId="3" pageNumber="312">
<emphasis id="B927EA81FFFB6575C0E2F96DB518F88D" bold="true" box="[151,276,1779,1801]" pageId="3" pageNumber="312">FIGURE 2.</emphasis>
<taxonomicName id="4C534D10FFFB6575C169F96AB5E5F88C" box="[284,489,1779,1801]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="312" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFFB6575C169F96AB5E5F88C" box="[284,489,1779,1801]" italics="true" pageId="3" pageNumber="312">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFFB6575C187F96DB648F88D" bold="true" box="[498,580,1779,1800]" pageId="3" pageNumber="312">
<taxonomicNameLabel id="A21457FAFFFB6575C187F96DB648F88D" box="[498,580,1779,1800]" pageId="3" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm.
</paragraph>
</caption>
<subSubSection id="C3496518FFFB6572C0B2F8CAB191FB95" lastPageId="4" lastPageNumber="313" pageId="3" pageNumber="312" type="diagnosis">
<paragraph id="8BEC3693FFFB6572C0B2F8CAB4D7FE71" blockId="3.[151,1436,1876,2009]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="313" pageId="3" pageNumber="312">
<emphasis id="B927EA81FFFB6575C0B2F8CAB54DF8E8" bold="true" box="[199,321,1876,1901]" pageId="3" pageNumber="312">Diagnosis.</emphasis>
<taxonomicName id="4C534D10FFFB6575C132F8CBB624F8E9" box="[327,552,1876,1900]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="312" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFFB6575C132F8CBB624F8E9" box="[327,552,1876,1900]" italics="true" pageId="3" pageNumber="312">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFFB6575C25BF8CAB68BF8E8" bold="true" box="[558,647,1876,1901]" pageId="3" pageNumber="312">
<taxonomicNameLabel id="A21457FAFFFB6575C25BF8CAB68BF8E8" box="[558,647,1876,1901]" pageId="3" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a medium-sized amphisbaenian (largest specimen total length 
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) and can be characterized as a 
<taxonomicName id="4C534D10FFFB6575C246F8E7B6B3F815" box="[563,703,1913,1936]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="312" phylum="Chordata" rank="genus">
<emphasis id="B927EA81FFFB6575C246F8E7B6B3F815" box="[563,703,1913,1936]" italics="true" pageId="3" pageNumber="312">Leposternon</emphasis>
</taxonomicName>
and distinguished from all South American amphisbaenians by having the nostril openings on the ventral surface of snout, rostral and nasals fused into a single shield, gular portion without segmental cover and prefrontal bones in facial view with nasal process long and ‘T’ shaped (details of skull in 
<bibRefCitation id="EFC24B62FFFC6572C164FF06B61DFF35" author="Gans" box="[273,529,152,176]" pageId="4" pageNumber="313" refString="Gans, C. &amp; Montero, R. (2009) An Atlas of Amphisbaenian Skull Anatomy. In: Adler, K. (Ed.), Biology of Reptilia. Society for the Study of Amphibians and Reptiles, New York, 21, pp. 621 - 738." type="journal article" year="2009">Gans &amp; Montero 2009</bibRefCitation>
) (Ribeiro 
<emphasis id="B927EA81FFFC6572C2F0FF07B6B2FF35" box="[645,702,152,176]" italics="true" pageId="4" pageNumber="313">et al.</emphasis>
2008). Additionally, the new species differs from all other South American amphisbaenians, except 
<taxonomicName id="4C534D10FFFC6572C25DFF23B6B8FF51" box="[552,692,189,212]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis id="B927EA81FFFC6572C25DFF23B6B8FF51" box="[552,692,189,212]" italics="true" pageId="4" pageNumber="313">Leposternon</emphasis>
</taxonomicName>
spp. and 
<taxonomicName id="4C534D10FFFC6572C35FFF22B0FFFF51" authority="Barbour, 1914" authorityName="Barbour" authorityYear="1914" box="[810,1267,188,213]" class="Reptilia" family="Amphisbaenidae" genus="Amphisbaena" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="anomala">
<emphasis id="B927EA81FFFC6572C35FFF22B023FF51" box="[810,1071,188,212]" italics="true" pageId="4" pageNumber="313">Amphisbaena anomala</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C437FF22B0E7FF51" author="Barbour" box="[1090,1259,188,213]" pageId="4" pageNumber="313" refString="Barbour, T. (1914) Some new reptiles. Proceedings of the New England Zoology Club, 4, 95 - 98." type="journal article" year="1914">Barbour, 1914</bibRefCitation>
)
</taxonomicName>
, in having the frontal portion of the head greatly depressed and pectoral shields irregular (not arranged in dermal regular annuli). It differs from the other species of the genus by the presence of rostral processes in the maxillae and nasals that contact each other in the facial portion of the skull (
<figureCitation id="13682A16FFFC6572C377FEB9B747FEC5" box="[770,843,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1758,1780]" captionTargetBox="[151,1436,193,1736]" captionTargetId="figure@5.[151,1436,193,1737]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of the skull in the genus Leposternon. (A) skull of Leposternon bagual sp. nov. (paratype, FML 11313) with rostral processes of the maxillae and nasals contacting each other on the facial portion of the skull; (B) skull of Leposternon aff. microcephalum (MRT 11984) with rostral processes of the maxillae and nasals not contacting each other on the facial portion of the skull; (C) skull of Leposternon polystegum (ZUFRJ 528) without rostral processes of the maxillae and nasals. Abbreviations: F, frontal; N, nasal; PF, prefrontal; MX, maxilla; NPPM, nasal process of premaxilla; RPM, rostral process of maxilla; RPPM, rostral process of premaxilla." httpUri="https://zenodo.org/record/242960/files/figure.png" pageId="4" pageNumber="313">Fig. 3</figureCitation>
, Table 1). The new species further differs from 
<taxonomicName id="4C534D10FFFC6572C5F0FEB7B6E0FEE1" authority="Ribeiro, Vaz-Silva" authorityName="Ribeiro, Vaz-Silva" authorityYear="2008" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="cerradensis">
<emphasis id="B927EA81FFFC6572C5F0FEB7B516FEE1" italics="true" pageId="4" pageNumber="313">L. cerradensis</emphasis>
Ribeiro, 
<bibRefCitation id="EFC24B62FFFC6572C1E6FED2B6E0FEE1" author="Vaz-Silva" box="[403,748,332,357]" pageId="4" pageNumber="313" refString="Ribeiro, S., Vaz-Silva, W. &amp; Santos-Jr., A. P. (2008) New pored Leposternon (Squamata, Amphisbaenia) from Brazilian Cerrado. Zootaxa, 1930, 18 - 38." type="journal article" year="2008">Vaz-Silva &amp; Santos-Jr, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C28AFED3B084FEE0" authority="Berthold, 1859" authorityName="Berthold" authorityYear="1859" box="[767,1160,332,357]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="infraorbitale">
<emphasis id="B927EA81FFFC6572C28AFED3B7B8FEE1" box="[767,948,332,356]" italics="true" pageId="4" pageNumber="313">L. infraorbitale</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C3BEFED2B08DFEE0" author="Berthold" box="[971,1153,332,357]" pageId="4" pageNumber="313" refString="Berthold, A. A. (1859) Einige neue Reptilien des akademisch zoologischen Museums in Gottingen. Nachrichten Georg August Universitaet Koniglichen Wissenschaftliche Gesellschaft, 17, 179 - 181." type="journal article" year="1859">Berthold, 1859</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C4E9FED3B163FEE1" box="[1180,1391,332,356]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="microcephalum">
<emphasis id="B927EA81FFFC6572C4E9FED3B163FEE1" box="[1180,1391,332,356]" italics="true" pageId="4" pageNumber="313">L. microcephalum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C5F0FED3B5DAFE0D" authority="Dumeril, 1851" authorityName="Dumeril" authorityYear="1851" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="octostegum">
<emphasis id="B927EA81FFFC6572C5F0FED3B51BFE0D" italics="true" pageId="4" pageNumber="313">L. octostegum</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C150FEF1B5C1FE0D" author="Dumeril" box="[293,461,367,392]" pageId="4" pageNumber="313" refString="Dumeril, A. H. A. (1851) Catalogue methodique de la collection des reptiles du Museum d'Histoire Naturelle de Paris. Gide et Baudry, Paris, 149 pp." type="book" year="1851">Duméril, 1851</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C194FEEFB747FE0D" authority="Hemprich, 1820" authorityName="Hemprich" authorityYear="1820" box="[481,843,367,392]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="scutigerum">
<emphasis id="B927EA81FFFC6572C194FEEFB670FE02" box="[481,636,368,392]" italics="true" pageId="4" pageNumber="313">L. scutigerum</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C2FFFEF1B74FFE0D" author="Hemprich" box="[650,835,367,392]" pageId="4" pageNumber="313" refString="Hemprich, W. F. G. 1820. Amphisbaenarum generis novas species duas descripsit. Verhandlungen der Gesellschaft Naturforschunden Freunde, 1 (2), 129 - 130." type="journal article" year="1820">Hemprich, 1820</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C534D10FFFC6572C3F4FEEFB0B8FE0D" authority="Peters, 1879" authorityName="Peters" authorityYear="1879" box="[897,1204,368,392]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B927EA81FFFC6572C3F4FEEFB01CFE0D" box="[897,1040,368,392]" italics="true" pageId="4" pageNumber="313">L. wuchereri</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C46BFEEEB0A7FE0D" author="Peters" box="[1054,1195,368,392]" pageId="4" pageNumber="313" refString="Peters, W. C. H. (1879) Qber die Amphisbaenen und eine zu denselben gehorige neue Art (Lepidosternon wuchereri). Monatsberichte der Berliner Akademie der Wissenschaften, 1879, 273 - 277." type="journal article" year="1879">Peters, 1879</bibRefCitation>
)
</taxonomicName>
by the presence of 3 infralabial shields; and from 
<taxonomicName id="4C534D10FFFC6572C199FE0BB69DFE29" box="[492,657,404,428]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="cerradensis">
<emphasis id="B927EA81FFFC6572C199FE0BB69DFE29" box="[492,657,404,428]" italics="true" pageId="4" pageNumber="313">L. cerradensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C2D1FE0BB10BFE29" authority="Porto, Soares &amp; Caramaschi, 2000" authorityName="Porto, Soares &amp; Caramaschi" authorityYear="2000" box="[676,1287,404,429]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="kisteumacheri">
<emphasis id="B927EA81FFFC6572C2D1FE0BB76FFE29" box="[676,867,404,428]" italics="true" pageId="4" pageNumber="313">L. kisteumacheri</emphasis>
<bibRefCitation id="EFC24B62FFFC6572C31BFE0BB10BFE29" author="Porto" box="[878,1287,404,429]" pageId="4" pageNumber="313" refString="Porto, M., Soares, M. &amp; Caramaschi, U. (2000) A new species of Leposternon (Amphisbaenia, Amphisbaenidae) from Minas Gerais, Brazil, with a key to the species of the genus (Amphisbaenia, Amphisbaenidae). Boletim do Museu Nacional, 412, 1 - 10." type="journal article" year="2000">Porto, Soares &amp; Caramaschi, 2000</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C534D10FFFC6572C563FE0BB6EFFE55" authority="Ribeiro, Nogueira, Cintra, Silva Jr. &amp; Zaher, 2011" authorityName="Ribeiro, Nogueira, Cintra, Silva Jr. &amp; Zaher" authorityYear="2011" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="maximus">
<emphasis id="B927EA81FFFC6572C563FE0BB190FE29" box="[1302,1436,405,428]" italics="true" pageId="4" pageNumber="313">L. maximus</emphasis>
Ribeiro, Nogueira, Cintra, Silva Jr. &amp; 
<bibRefCitation id="EFC24B62FFFC6572C22CFE29B6EFFE55" author="Zaher" box="[601,739,439,464]" pageId="4" pageNumber="313" refString="Ribeiro, S., Nogueira, C., Cintra, C. E., Silva, N. J. Jr. &amp; Zaher, H. (2011) Description of a new pored Leposternon (Squamata, Amphisbaenidae) from the Brazilian Cerrado. South American Journal of Herpetology, 6 (3), 177 - 188. http: // dx. doi. org / 10.2994 / 057.006.0303" type="journal article" year="2011">Zaher, 2011</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C534D10FFFC6572C35DFE27B082FE55" authority="Dumeril, 1851" authorityName="Dumeril" authorityYear="1851" box="[808,1166,439,464]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="polystegum">
<emphasis id="B927EA81FFFC6572C35DFE27B7C6FE55" box="[808,970,440,464]" italics="true" pageId="4" pageNumber="313">L. polystegum</emphasis>
(
<bibRefCitation id="EFC24B62FFFC6572C3A9FE29B08AFE55" author="Dumeril" box="[988,1158,439,464]" pageId="4" pageNumber="313" refString="Dumeril, A. H. A. (1851) Catalogue methodique de la collection des reptiles du Museum d'Histoire Naturelle de Paris. Gide et Baudry, Paris, 149 pp." type="book" year="1851">Duméril, 1851</bibRefCitation>
)
</taxonomicName>
by lacking precloacals pores.
</paragraph>
<paragraph id="8BEC3693FFFC6572C0B2FD9FB191FB95" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="313">
<taxonomicName id="4C534D10FFFC6572C0B2FD9FB5A4FD9D" box="[199,424,512,536]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="313" phylum="Chordata" rank="species" species="bagual">
<emphasis id="B927EA81FFFC6572C0B2FD9FB5A4FD9D" box="[199,424,512,536]" italics="true" pageId="4" pageNumber="313">Leposternon bagual</emphasis>
</taxonomicName>
is further characterized by the following combination of pholidotic, meristic, cranial, and morphometric characters (see also Table 1 for a synthesis of diagnostic features within the genus): 1) two supralabials; 2) supraocular present; 3) 240–267 dorsal postpectoral half-annuli; 4) 206–214 ventral postpectoral half-annuli; 5) 11–12 tail annuli; 6) 27–29 dorsal segments in midbody half-annulus; 7) 23–27 ventral segments in midbody half-annulus; 8) 96 precloacal vertebrae; 9) many small and irregularly shaped pectoral shields (
<figureCitation id="13682A16FFFC6572C541FD11B178FD2D" box="[1332,1396,655,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Fig. 1</figureCitation>
D); 10) adults specimens dark brown dorsally; 11) five premaxillary teeth; 12) four maxillary teeth; 13) six dentary teeth; 14) rostral process of nasal present; 15) rostral process of maxilla present; 16) premaxillary foramina present; 17) head length with 3.5–4.5% of snout-vent length; 18) rostronasal length with 22.6–23.5% of head length; 19) rostronasal width with 64.0–68.1% of the greatest width of head; 20) azygous length with 29.3–35.0% of head length; 21) azygous width representing 35.6–45.8% of the greatest width of head; 22) frontal suture length with 25.3–25.5% of head length; 23) parietal suture with 9.8–13.9% of head length; 24) temporal length with 18.9– 22.0% of head length; 25) temporal width with 13.2–17.8% of head width; 26) prefrontals length with 23.0–26.8% of head length; 27) prefrontals width with 38.3–41.7% of head width; 28) height posterior of head comprising 69.0–72.5% of head length; 30) comissure of the mouth length comprising 53.4–59.1% of head length; 29) midbody width with 3.2–4.4% of snout-vent length; and 30) tail length representing 4.9–5.6% of snout-vent length.
</paragraph>
</subSubSection>
<subSubSection id="C3496518FFFC6570C0B2FB82B63CF8B2" lastPageId="6" lastPageNumber="315" pageId="4" pageNumber="313" type="description">
<paragraph id="8BEC3693FFFC6572C0B2FB82B6FAFB25" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="313">
<emphasis id="B927EA81FFFC6572C0B2FB82B602FBB0" bold="true" box="[199,526,1052,1077]" pageId="4" pageNumber="313">
Description of the 
<typeStatus id="54E88831FFFC6572C1D2FB82B602FBB0" box="[423,526,1052,1077]" pageId="4" pageNumber="313" type="holotype">holotype</typeStatus>
</emphasis>
. The specimen is an adult female with 
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snout-vent length and 
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tail length (5.6% of total length); midbody diameter 
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; tail diameter of five annuli 
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; head length 9.0 mm (3.5% of snout-vent length); head anterior height 
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; head posterior height 
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; head anterior width 
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; and head posterior width 
<quantity id="4CAB9B76FFFC6572C26DFB19B662FB25" box="[536,622,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="313" unit="mm" value="6.8">6.8 mm</quantity>
(
<figureCitation id="13682A16FFFC6572C208FB19B6C6FB25" box="[637,714,1159,1184]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
, 
<figureCitation id="13682A16FFFC6572C2ADFB16B6E5FB25" box="[728,745,1160,1184]" captionStart="FIGURE 2" captionStartId="3.[151,250,1779,1801]" captionTargetBox="[150,1436,195,1756]" captionTargetId="figure@3.[150,1437,195,1757]" captionTargetPageId="3" captionText="FIGURE 2. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242959/files/figure.png" pageId="4" pageNumber="313">2</figureCitation>
).
</paragraph>
<paragraph id="8BEC3693FFFC6572C0B2FB32B0E1F938" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="313">
Dorsal portion of the head with four transversal row shields (
<figureCitation id="13682A16FFFC6572C309FB32B7CAFB41" box="[892,966,1196,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A). Rostronasal 1.1 times wider than long and 0.1 times wider than its heigth, in contact with the first supralabials laterally, and azygous and prefrontals posteriorly. Nostrils opening in the ventral portion of rostronasal scutes. Prefrontals, azygous and supraoculars slightly fused (
<figureCitation id="13682A16FFFC6572C131FA89B59CFAB5" box="[324,400,1303,1328]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A). Prefrontals with irregular shape, slightly fused with azygous, almost as long (
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) as wide (
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), in contact with first supralabials laterally, and azygous and supraocular posteriorly. Azygous with anterior portion project (
<figureCitation id="13682A16FFFC6572C225FAC1B696FAFD" box="[592,666,1375,1400]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A) and posterior portion straight, 0.2 times longer than wide, in contact with the supraoculars laterally, and frontals posteriorly (
<figureCitation id="13682A16FFFC6572C31EFA1AB7B9FA19" box="[875,949,1412,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A). Supraocular slightly quadrangular, 0.2 times wider than long, in contact with temporal posteriorly. Frontals paired (suture length 
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), anterior portion projected and posterior portion straight, almost as long (
<quantity id="4CAB9B76FFFC6572C3F6FA52B7E2FA61" box="[899,1006,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" pageId="4" pageNumber="313" unit="mm" value="2.28">2.28 mm</quantity>
) as wide (
<quantity id="4CAB9B76FFFC6572C41EFA52B0D4FA61" box="[1131,1240,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="313" unit="mm" value="2.31">2.31 mm</quantity>
), in contact with temporals laterally and parietals posteriorly. Temporals slightly rectangular, 0.5 times longer than wide, disposed on each side of frontals (
<figureCitation id="13682A16FFFC6572C1CEF98AB60BF9A9" box="[443,519,1556,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A), in contact with laterals parietals posteriorly. A pair of medial parietals (suture length 
<quantity id="4CAB9B76FFFC6572C134F9A6B590F9D5" box="[321,412,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="313" unit="mm" value="1.2">1.2 mm</quantity>
) (
<figureCitation id="13682A16FFFC6572C1C2F9A9B608F9D5" box="[439,516,1591,1616]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
A, 2A), roughly rectangular, 0.4 times wider than long, in contact with lateral parietals laterally and occipitals posteriorly. One small lateral parietal on each side of the pair of medial parietals, both with irregular shape, in contact with temporals anteriorly and body annuli posteriorly. A pair of occipitals, with irregular shape and size, in contact with the first row of dorsal anterior half-annuli posteriorly.
</paragraph>
<paragraph id="8BEC3693FFFC6572C0B2F959B032F811" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="313">
Ocular slightly quadrangular, 0.3 times longer than high, in contact with first infralabial, anteriorly, and inferior portion of the second supralabial and postocular posteriorly. Two supralabials. First supralabial larger than second supralabial, 0.7 times longer than high, 1.9 times longer and 0.4 times higher than second supralabial, in contact with the second supralabials posteriorly. Second supralabial slightly rectangular, 0.2 times its width, higher than long, and in contact with three postlabials posteriorly (
<figureCitation id="13682A16FFFC6572C331F8C9B79DF8F5" box="[836,913,1879,1904]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
B, 2B). Three postlabials of reduced size, the smallest one located at the inferior portion of the second supralabial (
<figureCitation id="13682A16FFFC6572C3EAF8E2B7E5F811" box="[927,1001,1916,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="4" pageNumber="313">Figs. 1</figureCitation>
B, 2B).
</paragraph>
<caption id="DF2C661BFFFD6573C0E2F940B6DAF808" httpUri="https://zenodo.org/record/242960/files/figure.png" pageId="5" pageNumber="314" targetBox="[151,1436,193,1736]" targetPageId="5">
<paragraph id="8BEC3693FFFD6573C0E2F940B6DAF808" blockId="5.[151,1436,1758,1933]" pageId="5" pageNumber="314">
<emphasis id="B927EA81FFFD6573C0E2F940B519F976" bold="true" box="[151,277,1758,1780]" pageId="5" pageNumber="314">FIGURE 3.</emphasis>
Dorsal view of the skull in the genus 
<taxonomicName id="4C534D10FFFD6573C2C4F941B723F971" box="[689,815,1759,1780]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="314" phylum="Chordata" rank="genus">
<emphasis id="B927EA81FFFD6573C2C4F941B723F971" box="[689,815,1759,1780]" italics="true" pageId="5" pageNumber="314">Leposternon</emphasis>
</taxonomicName>
. (A) skull of 
<taxonomicName id="4C534D10FFFD6573C3B1F941B09EF971" box="[964,1170,1758,1780]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="314" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFFD6573C3B1F941B09EF971" box="[964,1170,1758,1780]" italics="true" pageId="5" pageNumber="314">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFFD6573C4E9F941B0E3F971" bold="true" box="[1180,1263,1759,1780]" pageId="5" pageNumber="314">
<taxonomicNameLabel id="A21457FAFFFD6573C4E9F941B0E3F971" box="[1180,1263,1759,1780]" pageId="5" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype, FML 11313) with rostral processes of the maxillae and nasals contacting each other on the facial portion of the skull; (B) skull of 
<taxonomicName id="4C534D10FFFD6573C0E2F882B5EEF8B4" box="[151,482,1819,1841]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="314" phylum="Chordata" rank="species" species="microcephalum">
<emphasis id="B927EA81FFFD6573C0E2F882B51AF8B4" box="[151,278,1820,1841]" italics="true" pageId="5" pageNumber="314">Leposternon</emphasis>
aff. 
<emphasis id="B927EA81FFFD6573C133F885B5EEF8B4" box="[326,482,1819,1841]" italics="true" pageId="5" pageNumber="314">microcephalum</emphasis>
</taxonomicName>
(MRT 11984) with rostral processes of the maxillae and nasals not contacting each other on the facial portion of the skull; (C) skull of 
<taxonomicName id="4C534D10FFFD6573C232F8A5B733F8D5" box="[583,831,1850,1872]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="314" phylum="Chordata" rank="species" species="polystegum">
<emphasis id="B927EA81FFFD6573C232F8A5B733F8D5" box="[583,831,1850,1872]" italics="true" pageId="5" pageNumber="314">Leposternon polystegum</emphasis>
</taxonomicName>
(ZUFRJ 528) without rostral processes of the maxillae and nasals. Abbreviations: F, frontal; N, nasal; PF, prefrontal; MX, maxilla; NPPM, nasal process of premaxilla; RPM, rostral process of maxilla; RPPM, rostral process of premaxilla.
</paragraph>
</caption>
<caption id="DF2C661BFFFE6570C0E2FD27B5DAFD68" httpUri="https://zenodo.org/record/242961/files/figure.png" pageId="6" pageNumber="315" targetBox="[169,1416,182,678]" targetPageId="6">
<paragraph id="8BEC3693FFFE6570C0E2FD27B5DAFD68" blockId="6.[151,1436,697,749]" pageId="6" pageNumber="315">
<emphasis id="B927EA81FFFE6570C0E2FD27B518FD4B" bold="true" box="[151,276,697,719]" pageId="6" pageNumber="315">FIGURE 4.</emphasis>
Dorsal view of the heads of paratypes of 
<taxonomicName id="4C534D10FFFE6570C2B1FD24B79CFD4A" box="[708,912,697,719]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="315" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFFE6570C2B1FD24B79CFD4A" box="[708,912,697,719]" italics="true" pageId="6" pageNumber="315">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFFE6570C3EDFD27B7E6FD4B" bold="true" box="[920,1002,697,718]" pageId="6" pageNumber="315">
<taxonomicNameLabel id="A21457FAFFFE6570C3EDFD27B7E6FD4B" box="[920,1002,697,718]" pageId="6" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) FML 11312, (B) FML 11313 and (C) FML 11318. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BEC3693FFFE6570C0B2FC84B5F3FB8D" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">
Three infralabials, with the first one slightly triangular and in contact with the mental laterally and second infralabial posteriorly; the second one relatively narrow (3.3 times longer than the first infralabial), in contact with postmental and malars laterally and third infralabial posteriorly; the third one slightly rectangular, 0.2 times longer than first infralabial, in contact with postlabials and lateral malar laterally (
<figureCitation id="13682A16FFFE6570C38DFC1AB048FC18" box="[1016,1092,900,925]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="6" pageNumber="315">Figs. 1</figureCitation>
C, 2C). Mental with posterior margin 0.55 times wider than anterior margin, in contact with postmental posteriorly. Postmental 0.4 times longer than wide, with anterior portion straight and posterior projected, in contact with malar and posterior malar intern laterally and genials posteriorly.
</paragraph>
<paragraph id="8BEC3693FFFE6570C0B2FB8DB5C2FB58" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">Two small, irregularly shaped genials, disposed posteriorly to postmental. An elongated teardrop shaped malar, disposed on either sides of the post-mental, 1.4 times longer than wide, contacting posterior malars posteriorly. Two posterior malars located between the postmental and the lateral malar; the medial one in contact with the postmental and the outermost malar; and lateral one in contact with the lateral malar. Lateral malar 0.5 times longer than wide, tapering anterolaterally and rounded posteriorly, in contact with the postmalar row. Postmalar row formed by 10 small shields.</paragraph>
<paragraph id="8BEC3693FFFE6570C0B2FB79B58DFA7D" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">
Dorsal and ventral portions of the body divided by a lateral groove; dorsal region further divided by a vertebral groove; lateral grooves absent in the tail. Pectoral scales diamond-shaped, with central scales larger than lateral scales (
<figureCitation id="13682A16FFFE6570C09CFAB0B53FFAC2" box="[233,307,1326,1351]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="6" pageNumber="315">Figs. 1</figureCitation>
D, 2D). Eighteen dorsal anterior half-annuli, 253 dorsal postpectoral half annuli plus two incomplete half-annuli, 214 ventral postpectoral half-annuli, five lateral half-annuli, and 12 tail annuli. Twenty eigth dorsal segments per half-annulus at midbody, 23 ventral segments per half-annulus at midbody, and 35 segments forming the fifth tail annulus. Anal flaps semicircular, 1.0 times longer than half-annuli, with 11 narrow segments (
<figureCitation id="13682A16FFFE6570C542FA06B18DFA34" box="[1335,1409,1432,1457]" captionStart="FIGURE 1" captionStartId="2.[151,250,1924,1946]" captionTargetBox="[166,1413,421,1888]" captionTargetId="figure@2.[151,1436,409,1903]" captionTargetPageId="2" captionText="FIGURE 1. Leposternon bagual sp. nov. (holotype, FML 11319). (A) Dorsal, (B) lateral, (C) ventral view of the head; (D) pectoral and (E) cloacal regions. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242958/files/figure.png" pageId="6" pageNumber="315">Figs. 1</figureCitation>
E, 2E). Ninety six precloacal vertebrae, with dorsal and ventral half-annuli around the body being more than twice the number of vertebrae.
</paragraph>
<paragraph id="8BEC3693FFFE6570C0B2F99DB4FDF9BA" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">
<emphasis id="B927EA81FFFE6570C0B2F99DB65AF999" bold="true" box="[199,598,1539,1564]" pageId="6" pageNumber="315">
Color of 
<typeStatus id="54E88831FFFE6570C141F99DB596F999" box="[308,410,1539,1564]" pageId="6" pageNumber="315" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
. Head, trunk and tail are uniform dark brown, except for the belly that is whitish.
</paragraph>
<paragraph id="8BEC3693FFFE6570C0B2F9D4B63CF8B2" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">
<emphasis id="B927EA81FFFE6570C0B2F9D4B539F9E6" bold="true" box="[199,309,1610,1635]" pageId="6" pageNumber="315">Variation</emphasis>
. Variation on the meristic and morphometric characteristics of the 
<typeStatus id="54E88831FFFE6570C45EF9D5B09AF9E6" box="[1067,1174,1611,1635]" pageId="6" pageNumber="315" type="paratype">paratypes</typeStatus>
is presented in Table 2 and in the 
<figureCitation id="13682A16FFFE6570C163F9F3B577F903" box="[278,379,1645,1670]" captionStart="FIGURE 4" captionStartId="6.[151,250,697,719]" captionTargetBox="[169,1416,182,678]" captionTargetId="figure@6.[169,1423,177,683]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view of the heads of paratypes of Leposternon bagual sp. nov. (A) FML 11312, (B) FML 11313 and (C) FML 11318. Scale bar = 5 mm." httpUri="https://zenodo.org/record/242961/files/figure.png" pageId="6" pageNumber="315">Figure 4</figureCitation>
. Additionally, the 
<typeStatus id="54E88831FFFE6570C227F9F0B6B1F903" box="[594,701,1646,1670]" pageId="6" pageNumber="315" type="paratype">paratypes</typeStatus>
show some variation in the disposition and morphology of the cephalic shields in comparison to the 
<typeStatus id="54E88831FFFE6570C24AF90EB6A9F92C" box="[575,677,1680,1705]" pageId="6" pageNumber="315" type="holotype">holotype</typeStatus>
. FML 11312 has a poorly defined anterior border of the frontal; the posterior border of the left parietal is elongated, occupying the space where the occipital shield would be located on the right side. FML 11313 shows partially fused parietal and occipital shields. FML 11313 shows a pre-frontal portion with numerous diminute, irregularly shaped shields, a frontal portion with a rounded anterior border; and a tuberculate dorsal portion of the tail.
</paragraph>
</subSubSection>
<subSubSection id="C3496518FFFE657DC0B2F8DCB0A9FF51" lastPageId="11" lastPageNumber="320" pageId="6" pageNumber="315" type="etymology">
<paragraph id="8BEC3693FFFE6570C0B2F8DCB59DF86E" blockId="6.[151,1437,794,2027]" pageId="6" pageNumber="315">
<emphasis id="B927EA81FFFE6570C0B2F8DCB54AF8DE" bold="true" box="[199,326,1858,1883]" pageId="6" pageNumber="315">Etymology</emphasis>
. The specific epithet for the new species is used here as a noun in apposition, an allusion to the ecological reserve where the new species has been collected (El 
<taxonomicName id="4C534D10FFFE6570C3D3F8F8B7F9F8FA" box="[934,1013,1894,1919]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="321" phylum="Chordata" rank="species" species="bagual">Bagual</taxonomicName>
Reserve). The word 
<taxonomicName id="4C534D10FFFE6570C48CF8F8B14BF8FB" box="[1273,1351,1894,1918]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="315" phylum="Chordata" rank="species" species="bagual">
<emphasis id="B927EA81FFFE6570C48CF8F8B14BF8FB" box="[1273,1351,1894,1918]" italics="true" pageId="6" pageNumber="315">bagual</emphasis>
</taxonomicName>
comes originally from the indigenous word 
<emphasis id="B927EA81FFFE6570C246F814B673F827" box="[563,639,1930,1954]" italics="true" pageId="6" pageNumber="315">cahual</emphasis>
, which in Araucana culture (native inhabitants of 
<collectingCountry id="F3447603FFFE6570C4C5F814B0E0F826" box="[1200,1260,1930,1955]" name="Chile" pageId="6" pageNumber="315">Chile</collectingCountry>
and 
<collectingCountry id="F3447603FFFE6570C556F814B199F826" box="[1315,1429,1930,1955]" name="Argentina" pageId="6" pageNumber="315">Argentina</collectingCountry>
) is applied to cattle (
<bibRefCitation id="EFC24B62FFFE6570C107F830B6F9F842" author="Di-Giacomo" box="[370,757,1966,1991]" pageId="6" pageNumber="315" refString="Di-Giacomo, A. G. &amp; Krapovickas, S. F. (2005) Historia natural y paisaje de la Reserva El Bagual, Provincia de Formosa, Argentina. Inventario de la fauna de vertebrados y de la flora vascular de un area protegida del Chaco Humedo. Temas de Naturaleza y Conservacion 4. Aves Argentinas / Asociacion Ornitologica del Plata, Buenos Aires, 592 pp." type="book" year="2005">Di-Giacomo &amp; Krapovickas 2005</bibRefCitation>
). Among other denominations, 
<taxonomicName id="4C534D10FFFE6570C420F830B0AEF843" box="[1109,1186,1966,1990]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="315" phylum="Chordata" rank="species" species="bagual">
<emphasis id="B927EA81FFFE6570C420F830B0AEF843" box="[1109,1186,1966,1990]" italics="true" pageId="6" pageNumber="315">bagual</emphasis>
</taxonomicName>
is used by “Gauchos” to define a wild horse.
</paragraph>
<paragraph id="8BEC3693FFFF6571C008FDB9B37EFDEE" blockId="7.[125,1908,550,818]" pageId="7" pageNumber="316">
<emphasis id="B927EA81FFFF6571C008FDB9B4D2FDBF" bold="true" box="[125,222,551,570]" pageId="7" pageNumber="316">TABLE 1.</emphasis>
Pholidotic, cranial, and morphometric characters for the species of the genus 
<emphasis id="B927EA81FFFF6571C3E4FDB9B069FDBF" box="[913,1125,550,570]" italics="true" pageId="7" pageNumber="316">
<taxonomicName id="4C534D10FFFF6571C3E4FDB9B00CFDBC" box="[913,1024,551,569]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="genus">Leposternon</taxonomicName>
. Pholidosis
</emphasis>
: 
<collectingCountry id="F3447603FFFF6571C407FDB9B084FDBF" box="[1138,1160,551,570]" name="Iceland" pageId="7" pageNumber="316">IL</collectingCountry>
, Infralabial; PPO, precloacal pores; SL, Supralabial; DOA, Dorsal postpectoral halfannuli; VEA, Ventral postpectoral half-annuli; TA, Tail annuli; DS, Dorsal segments in midbody half-annulus; VS, Ventral segments in midbody half-annulus; PVE, Precloacal vertebrae; PEC, pectoral shields; COL, colorations specimens adults. 
<emphasis id="B927EA81FFFF6571C18BFDC9B625FDEE" box="[510,553,599,619]" italics="true" pageId="7" pageNumber="316">Skull</emphasis>
: PMXT, premaxillary teeth; MXT, maxillary teeth; DT, dentary teeth; RPN, rostral process of nasal; RPMX, rostral process of maxilla; FPMX, foramina of
</paragraph>
<paragraph id="8BEC3693FFFF6571C0E8FDEFB37EFD01" blockId="7.[125,1908,550,818]" box="[157,1906,625,645]" pageId="7" pageNumber="316">
premaxilla. 
<emphasis id="B927EA81FFFF6571C171FDEFB59DFD00" box="[260,401,625,645]" italics="true" pageId="7" pageNumber="316">Morphometrics:</emphasis>
HL/SVL, head length/snout-vent length expressed as a percentage; SSLL/HL, second supralabial length/head length expressed as a percentage (analyzed only in species with
</paragraph>
<paragraph id="8BEC3693FFFF6571C0E7FD14B57EFCB7" blockId="7.[125,1908,550,818]" pageId="7" pageNumber="316">supralabials); RL/HL, rostronasal length/head length expressed as a percentage; RW/HW, rostronasal length/greatest width of head expressed as a percentage and analyze; RH/HH, rostronasal height/posterior height of head expressed as a percentage; AZL/HL, azygous length/head length expressed as a percentage; AZW/HW, azygous width/posterior height of head expressed as a percentage; AZL/FW, azygous length/azygous width expressed as a percentage; FSL/HL, frontal suture length/head length expressed as a percentage; PL/HL, parietal suture length/head length expressed as a percentage; TL/HL, temporal length/head length expressed as a percentage; TW/HW, temporal width/head head width expressed as a percentage; PFL/HL, prefrontal length/head length expressed as a percentage; PFW/HW, prefrontals width/head width expresses as a percentage; HH/HL, head posterior height/head length expressed as a percentage; CML/HL, commissure of the mouth length/head length expressed as a percentage; MB/SVL midbody width/snout-vent length expressed as a percentage; TL/SVL, tail length/snout-vent length expressed as a percentage; SD, standard deviation. Diagnostic characters are given in bold.</paragraph>
<paragraph id="8BEC3693FFFF6571C0F7FCC6B081FCED" blockId="7.[129,1902,853,1673]" box="[130,1165,853,875]" pageId="7" pageNumber="316">
Charact Species of 
<taxonomicName id="4C534D10FFFF6571C454FCC8B081FCED" box="[1057,1165,854,872]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="genus">
<emphasis id="B927EA81FFFF6571C454FCC8B081FCED" box="[1057,1165,854,872]" italics="true" pageId="7" pageNumber="316">Leposternon</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BEC3693FFFF6571C0E2FCEFB34AFC23" blockId="7.[129,1902,853,1673]" pageId="7" pageNumber="316">
ers 
<emphasis id="B927EA81FFFF6571C17EFCEBB35EFC0C" box="[267,1874,885,905]" italics="true" pageId="7" pageNumber="316">
<taxonomicName id="4C534D10FFFF6571C17EFCEBB54BFC0C" box="[267,327,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="bagual">bagual</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C1D4FCEBB60AFC0C" box="[417,518,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="cerradensis">cerradensis</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C233FCEBB6B9FC0C" box="[582,693,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="infraorbitale">infraorbitale</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C29EFCEBB768FC0C" box="[747,868,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="kisteumacheri">kisteumacheri</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C3D9FCE8B0B3FC0C" box="[940,1215,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="subSpecies" species="maximus" subSpecies="microcephalum">maximus microcephalum</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C483FCE8B154FC0D" box="[1270,1368,886,904]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="octostegum">octostegum</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C5D5FCEBB20EFC0C" box="[1440,1538,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="polystegum">polystegum</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C639FCE8B2A0FC0D" box="[1612,1708,886,904]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="scutigerum">scutigerum</taxonomicName>
<taxonomicName id="4C534D10FFFF6571C68FFCEBB35EFC0C" box="[1786,1874,885,905]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="316" phylum="Chordata" rank="species" species="wuchereri">wuchereri</taxonomicName>
</emphasis>
3/3 
<emphasis id="B927EA81FFFF6571C1B0FC0DB687FC23" bold="true" box="[453,651,915,934]" pageId="7" pageNumber="316">2/2 2/2</emphasis>
3/3 3/3 
<emphasis id="B927EA81FFFF6571C41AFC0DB139FC23" bold="true" box="[1135,1333,915,934]" pageId="7" pageNumber="316">2/2 1/1</emphasis>
3/3 
<emphasis id="B927EA81FFFF6571C61BFC0DB34AFC23" bold="true" box="[1646,1862,915,934]" pageId="7" pageNumber="316">1/1 1-2/1-2</emphasis>
</paragraph>
<paragraph id="8BEC3693FFFF6571C0F4FC29B34BFC6E" blockId="7.[129,1902,853,1673]" pageId="7" pageNumber="316">
PPO Absent 
<emphasis id="B927EA81FFFF6571C1B0FC29B5EDFC4F" bold="true" box="[453,481,951,970]" pageId="7" pageNumber="316">2-4</emphasis>
Absent 
<emphasis id="B927EA81FFFF6571C357FC29B7D4FC4F" bold="true" box="[802,984,951,970]" pageId="7" pageNumber="316">2 2</emphasis>
Absent Absent 
<emphasis id="B927EA81FFFF6571C5B6FC29B1D3FC4F" bold="true" box="[1475,1503,951,970]" pageId="7" pageNumber="316">2-4</emphasis>
Absent Absent 2/2 2/2 
<emphasis id="B927EA81FFFF6571C21AFC47B687FC69" bold="true" box="[623,651,985,1004]" pageId="7" pageNumber="316">3/3</emphasis>
2/2 2/2 2/2 
<emphasis id="B927EA81FFFF6571C56CFC47B1D3FC69" bold="true" box="[1305,1503,985,1004]" pageId="7" pageNumber="316">1/1 3/3</emphasis>
2/2 1-2/1-2
</paragraph>
<paragraph id="8BEC3693FFFF6571C0F4FC62B338FB8A" blockId="7.[129,1902,853,1673]" box="[129,1844,1020,1040]" pageId="7" pageNumber="316">
SOC 1/1 
<emphasis id="B927EA81FFFF6571C1C6FC63B5F8FB95" bold="true" box="[435,500,1021,1040]" pageId="7" pageNumber="316">Absent</emphasis>
1/1 0-1/0-1 1/1 1/1 
<emphasis id="B927EA81FFFF6571C572FC63B291FB95" bold="true" box="[1287,1693,1021,1040]" pageId="7" pageNumber="316">Absent Absent Absent</emphasis>
1/1
</paragraph>
<paragraph id="8BEC3693FFFF6571C0F4FBBEB347FBB1" blockId="7.[129,1902,853,1673]" box="[129,1867,1056,1076]" pageId="7" pageNumber="316">
DOA 240-267 
<emphasis id="B927EA81FFFF6571C1DAFBBFB5FBFBB1" bold="true" box="[431,503,1057,1076]" pageId="7" pageNumber="316">299-341</emphasis>
250-292 244-271 
<emphasis id="B927EA81FFFF6571C3DBFBBFB7FAFBB1" bold="true" box="[942,1014,1057,1076]" pageId="7" pageNumber="316">404-448</emphasis>
204-255 
<emphasis id="B927EA81FFFF6571C576FBBFB147FBB1" bold="true" box="[1283,1355,1057,1076]" pageId="7" pageNumber="316">353-398</emphasis>
253-363 259-295 231-263
</paragraph>
<paragraph id="8BEC3693FFFF6571C090FBA7B368FBC9" blockId="7.[129,1902,853,1673]" box="[229,1892,1081,1100]" pageId="7" pageNumber="316">(n = 4; Mean = (n = 6; Mean = (n = 20; Mean (n = 5; Mean = (n = 11; Mean (n = 119; Mean (n = 4; Mean = (n = 30; Mean (n = 5; Mean = (n = 5; Mean =</paragraph>
<paragraph id="8BEC3693FFFF6571C09EFBCCB330FBFB" blockId="7.[129,1902,853,1673]" pageId="7" pageNumber="316">256; SD ± 13) 317.2; SD ± = 270,0; SD ± 256.2; SD ± = 424.9; SD ± = 228.1; SD ± 372.3; SD ± = 302.8; SD ± 270.2; SD ± 247.0; SD ± 16.4) 11.7) 11.0) 14.1) 7.6) 20.5) 31.0) 14.2) 14.4)</paragraph>
<paragraph id="8BEC3693FFFF6571C0F4FB1AB346FB1D" blockId="7.[129,1902,853,1673]" box="[129,1866,1156,1176]" pageId="7" pageNumber="316">
VEA 206–214 
<emphasis id="B927EA81FFFF6571C1DAFB1AB7FAFB12" bold="true" box="[431,1014,1156,1175]" pageId="7" pageNumber="316">302-333 229-285 246-259 408-448</emphasis>
192-229 
<emphasis id="B927EA81FFFF6571C576FB1AB346FB1D" bold="true" box="[1283,1866,1156,1176]" pageId="7" pageNumber="316">342-390 252-349 250-278 226-254</emphasis>
</paragraph>
<paragraph id="8BEC3693FFFF6571C090FB03B368FB35" blockId="7.[129,1902,853,1673]" box="[229,1892,1181,1200]" pageId="7" pageNumber="316">(n = 4; Mean = (n = 6; Mean = (n = 20; Mean (n = 4; Mean = (n = 11; Mean (n = 120; Mean (n = 4; Mean = (n = 30 Mean = (n = 5; Mean = (n = 5; Mean =</paragraph>
<paragraph id="8BEC3693FFFF6571C084FB2BB1EFFAC0" blockId="7.[129,1902,853,1673]" pageId="7" pageNumber="316">
211; SD ± 3) 319.3; SD ± = 250.4; SD ± 253.5; SD ± = 426.9; SD ± = 207.6; SD ± 363.8; SD ± 294.2; SD ± 261.6; SD ± 239.0; SD ± 17.9) 15.0) 5.4) 11.3) 11.1) 19.8) 27.3) 11.5) 11.0) 11-12 
<emphasis id="B927EA81FFFF6571C1CFFB76B5E0FB7E" bold="true" box="[442,492,1256,1275]" pageId="7" pageNumber="316">
13- 
<date id="FFED1053FFFF6571C1A3FB76B5E0FB7E" box="[470,492,1256,1275]" pageId="7" pageNumber="316" value="1915-12-09" valueMax="1915-12-13" valueMin="1915-12-09">15</date>
</emphasis>
9-13 12-14 
<emphasis id="B927EA81FFFF6571C3CCFB76B7E7FB7E" bold="true" box="[953,1003,1256,1275]" pageId="7" pageNumber="316">
14- 
<date id="FFED1053FFFF6571C3A0FB76B7E7FB7E" box="[981,1003,1256,1275]" pageId="7" pageNumber="316" value="1916-11-08" valueMax="1916-11-14" valueMin="1916-11-08">16</date>
</emphasis>
8-14 11 9- 
<date id="FFED1053FFFF6571C5BAFB79B1E9FB7F" box="[1487,1509,1255,1274]" pageId="7" pageNumber="316" value="1918-11-12" valueMax="1918-11-14" valueMin="1918-11-12">18</date>
12-14 11-13 (n = 4) (n = 6; Mean = (n = 20; Mean (n = 5; Mean = (n = 11; Mean (n = 120; X (n = 4) (n = 30; Mean (n = 5; Mean = (n = 5; Mean = 13.7; SD ± 0.8) = 10.9; SD ± 13; SD ± 0.7) = 14.7; SD ± Mean = 10.7; = 13.4; SD ± 12.6; SD ± 0.9) 12.4; SD ± 0.9) 1.0) 0.5) SD ± 1.0) 1.9)
</paragraph>
<paragraph id="8BEC3693FFFF6571C17BFAD5B333FADB" blockId="7.[129,1902,853,1673]" box="[270,1855,1355,1374]" pageId="7" pageNumber="316">
27–29 
<emphasis id="B927EA81FFFF6571C1CFFAD5B5E0FADB" bold="true" box="[442,492,1355,1374]" pageId="7" pageNumber="316">32-37</emphasis>
22-28 20-33 24-33 17-31 27-34 22-41 21-27 
<emphasis id="B927EA81FFFF6571C778FAD5B333FADB" bold="true" box="[1805,1855,1355,1374]" pageId="7" pageNumber="316">16-19</emphasis>
</paragraph>
<paragraph id="8BEC3693FFFF6571C090FAFDB368FAF3" blockId="7.[129,1902,853,1673]" box="[229,1892,1379,1398]" pageId="7" pageNumber="316">(n = 4; Mean = (n = 6; Mean = (n = 20; Mean (n = 5; Mean = (n = 11; Mean (n = 117; Mean (n = 3; Mean = (n = 30; Mean (n = 5; Mean = (n = 4; Mean =</paragraph>
<paragraph id="8BEC3693FFFF6571C083FAE2B365FA0A" blockId="7.[129,1902,853,1673]" box="[246,1897,1404,1423]" pageId="7" pageNumber="316">28; SD ± 1) 34.8; SD ± 1.7) = 25.1; SD ± 26.8; SD ± 4.7) = 28.2; SD ± = 21.3; SD ± 30.7; SD ± 3.5) = 29.4; SD ± 23.2; SD ± 2.4) 17.5; SD ± 1.3)</paragraph>
<paragraph id="8BEC3693FFFF6571C219FA0BB1EFFA2D" blockId="7.[129,1902,853,1673]" box="[620,1507,1429,1448]" pageId="7" pageNumber="316">1.8) 2.6) 2.1) 3.6)</paragraph>
<paragraph id="8BEC3693FFFF6571C17BFA30B34CFA47" blockId="7.[129,1902,853,1673]" box="[270,1856,1454,1474]" pageId="7" pageNumber="316">
23–27 
<emphasis id="B927EA81FFFF6571C1CFFA31B5E0FA47" bold="true" box="[442,492,1455,1474]" pageId="7" pageNumber="316">30-36</emphasis>
25-35 22-32 23-32 17-31 27-28 20-33 25-29 
<emphasis id="B927EA81FFFF6571C778FA31B34CFA47" bold="true" box="[1805,1856,1455,1474]" pageId="7" pageNumber="316">16-21</emphasis>
</paragraph>
<paragraph id="8BEC3693FFFF6571C090FA59B368FA5F" blockId="7.[129,1902,853,1673]" box="[229,1892,1479,1498]" pageId="7" pageNumber="316">(n = 4; Mean = (n = 6; Mean = (n = 20; Mean (n = 5; Mean = (n = 11; Mean (n = 115; Mean (n = 4; Mean = (n = 30; Mean (n = 5; Mean = (n = 4; Mean =</paragraph>
<paragraph id="8BEC3693FFFF6571C083FA41B365FA76" blockId="7.[129,1902,853,1673]" box="[246,1897,1503,1523]" pageId="7" pageNumber="316">24; SD ± 2) 32.7; SD ± 2.5) = 30.1; SD ± 27.4; SD ± 3.6) = 28.5; SD ± = 22.6; SD ± 27.7; SD =0.6) = 25.0; SD ± 27.4; SD ± 1.7) 18.3; SD ± 2.1)</paragraph>
<paragraph id="8BEC3693FFFF6571C219FA66B1EFF98E" blockId="7.[129,1902,853,1673]" box="[620,1507,1528,1547]" pageId="7" pageNumber="316">3.1) 4.0) 2.5) 3.1)</paragraph>
<paragraph id="8BEC3693FFFF6571C0F4F98FB0B2F9B8" blockId="7.[129,1902,853,1673]" pageId="7" pageNumber="316">
PVE 96 
<emphasis id="B927EA81FFFF6571C1DAF98CB5F4F9A0" bold="true" box="[431,504,1554,1573]" pageId="7" pageNumber="316">118-121</emphasis>
92-98 
<emphasis id="B927EA81FFFF6571C282F98FB714F9A1" bold="true" box="[759,792,1553,1572]" pageId="7" pageNumber="316">118</emphasis>
(n = 1) not available 92-102 
<emphasis id="B927EA81FFFF6571C482F98FB114F9A1" bold="true" box="[1271,1304,1553,1572]" pageId="7" pageNumber="316">144</emphasis>
(n = 1) 
<emphasis id="B927EA81FFFF6571C5F8F98FB1D9F9A1" bold="true" box="[1421,1493,1553,1572]" pageId="7" pageNumber="316">117-131</emphasis>
(n = 2) 
<emphasis id="B927EA81FFFF6571C642F98FB273F9A1" bold="true" box="[1591,1663,1553,1572]" pageId="7" pageNumber="316">117-124</emphasis>
(n = 2) 
<emphasis id="B927EA81FFFF6571C68EF98FB31DF9A1" bold="true" box="[1787,1809,1553,1572]" pageId="7" pageNumber="316">91</emphasis>
(n = 1) (n = 1) (n= 4; Mean = (n = 4; Mean = (n = 43; means
</paragraph>
<paragraph id="8BEC3693FFFF6571C1E6F9DDB0B9F9D3" blockId="7.[129,1902,853,1673]" box="[403,1205,1603,1622]" pageId="7" pageNumber="316">119.2; SD 1.3) 95.0; SD ± 2.4) = 97.1; SD ±</paragraph>
<paragraph id="8BEC3693FFFF6571C413F9C2B09FF9EA" blockId="7.[129,1902,853,1673]" box="[1126,1171,1628,1647]" pageId="7" pageNumber="316">12.2)</paragraph>
<paragraph id="8BEC3693FFFF657FC62AF9EBB546FDA7" blockId="7.[129,1902,853,1673]" lastBlockId="9.[125,1896,527,1676]" lastPageId="9" lastPageNumber="318" pageId="7" pageNumber="316">
<emphasis id="B927EA81FFFF6571C62AF9EBB362F90D" box="[1631,1902,1653,1673]" italics="true" pageId="7" pageNumber="316">……continued on the next page</emphasis>
<emphasis id="B927EA81FFF0657EC0FCFDBAB4E4FDB2" bold="true" box="[137,232,548,567]" pageId="8" pageNumber="317">TABLE 1.</emphasis>
(Continued) 
<emphasis id="B927EA81FFF1657FC008FD91B4D0FDA7" bold="true" box="[125,220,527,546]" pageId="9" pageNumber="318">TABLE 1.</emphasis>
(Continued)
</paragraph>
<paragraph id="8BEC3693FFF0657EC00CFDCBB369F9EE" pageId="8" pageNumber="317">
<table id="F953C433FFF09A89C002FDCDB369F9E9" box="[119,1893,595,1644]" gridcols="9" gridrows="6" pageId="8" pageNumber="317">
<tr id="356334D1FFF09A89C002FDCDB369FD02" box="[119,1893,595,647]" gridrow="0" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FDCDB4B0FD02" box="[119,188,595,647]" gridcol="0" gridrow="0" pageId="8" pageNumber="317">Charact ers</th>
<th id="76B25DADFFF09A89C0AFFDCDB569FD02" box="[218,357,595,647]" gridcol="1" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC174FDEDB531FD02" box="[257,317,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="bagual">
<emphasis id="B927EA81FFF0657EC174FDEDB531FD02" box="[257,317,627,647]" italics="true" pageId="8" pageNumber="317">bagual</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C1F6FDCDB601FD02" box="[387,525,595,647]" gridcol="2" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC1E2FDEDB5F0FD02" box="[407,508,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="cerradensis">
<emphasis id="B927EA81FFF0657EC1E2FDEDB5F0FD02" box="[407,508,627,647]" italics="true" pageId="8" pageNumber="317">cerradensis</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C258FDCDB6B6FD02" box="[557,698,595,647]" gridcol="3" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC248FDEDB6A0FD02" box="[573,684,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="infraorbitale">
<emphasis id="B927EA81FFF0657EC248FDEDB6A0FD02" box="[573,684,627,647]" italics="true" pageId="8" pageNumber="317">infraorbitale</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C2A2FDCDB769FD02" box="[727,869,595,647]" gridcol="4" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC297FDEDB757FD02" box="[738,859,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="kisteumacheri">
<emphasis id="B927EA81FFF0657EC297FDEDB757FD02" box="[738,859,627,647]" italics="true" pageId="8" pageNumber="317">kisteumacheri</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C3F7FDCDB0B5FD02" box="[898,1209,595,647]" gridcol="5" gridrow="0" pageId="8" pageNumber="317">
Species of 
<taxonomicName id="4C534D10FFF0657EC462FDD1B0BAFD02" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="subSpecies" species="maximus" subSpecies="microcephalum">
<emphasis id="B927EA81FFF0657EC462FDD1B0BAFD02" italics="true" pageId="8" pageNumber="317">Leposternon maximus microcephalum</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C4A3FDCDB169FD02" box="[1238,1381,595,647]" gridcol="6" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC499FDEAB143FD03" box="[1260,1359,628,646]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="octostegum">
<emphasis id="B927EA81FFF0657EC499FDEAB143FD03" box="[1260,1359,628,646]" italics="true" pageId="8" pageNumber="317">octostegum</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C5F4FDCDB203FD02" box="[1409,1551,595,647]" gridcol="7" gridrow="0" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC5E2FDEDB1F6FD02" box="[1431,1530,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="polystegum">
<emphasis id="B927EA81FFF0657EC5E2FDEDB1F6FD02" box="[1431,1530,627,647]" italics="true" pageId="8" pageNumber="317">polystegum</emphasis>
</taxonomicName>
</th>
<th id="76B25DADFFF09A89C65EFDCDB369FD02" box="[1579,1893,595,647]" gridcol="8" gridrow="0" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC636FDEAB345FD02" box="[1603,1865,627,647]" italics="true" pageId="8" pageNumber="317">
<taxonomicName id="4C534D10FFF0657EC636FDEAB2AFFD03" box="[1603,1699,628,646]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="scutigerum">scutigerum</taxonomicName>
<taxonomicName id="4C534D10FFF0657EC684FDEDB345FD02" box="[1777,1865,627,647]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="317" phylum="Chordata" rank="species" species="wuchereri">wuchereri</taxonomicName>
</emphasis>
</th>
</tr>
<tr id="356334D1FFF09A89C002FD0AB369FC02" box="[119,1893,660,903]" gridrow="1" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FD0AB4B0FC02" box="[119,188,660,903]" gridcol="0" gridrow="1" pageId="8" pageNumber="317">PEC COL PMXT</th>
<td id="76B25DADFFF09A89C0AFFD0AB569FC02" box="[218,357,660,903]" gridcol="1" gridrow="1" pageId="8" pageNumber="317">Many shields. All small and irregularly shape. Dark brown 5</td>
<td id="76B25DADFFF09A89C1F6FD0AB601FC02" box="[387,525,660,903]" gridcol="2" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC1FFFD0AB5E6FCB9" bold="true" pageId="8" pageNumber="317">Many shields. Mostly small and diamond- shaped. Absent</emphasis>
5
</td>
<td id="76B25DADFFF09A89C258FD0AB6B6FC02" box="[557,698,660,903]" gridcol="3" gridrow="1" pageId="8" pageNumber="317">
Many shields. All small and irregularly shape. 
<emphasis id="B927EA81FFF0657EC221FCB7B698FCB9" bold="true" box="[596,660,809,828]" pageId="8" pageNumber="317">Yellow</emphasis>
5–6
</td>
<td id="76B25DADFFF09A89C2A2FD0AB769FC02" box="[727,869,660,903]" gridcol="4" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC2AAFD0AB728FC02" bold="true" pageId="8" pageNumber="317">Many shields. Mostly small and diamond- shaped. Absent 7</emphasis>
</td>
<td id="76B25DADFFF09A89C3F7FD0AB0B5FC02" box="[898,1209,660,903]" gridcol="5" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC3FCFD0AB004FD22" bold="true" box="[905,1032,660,679]" pageId="8" pageNumber="317">Many shields.</emphasis>
Many shields. 
<emphasis id="B927EA81FFF0657EC3FBFD33B008FD45" bold="true" box="[910,1028,685,704]" pageId="8" pageNumber="317">Mostly small</emphasis>
All small and 
<emphasis id="B927EA81FFF0657EC3FDFD58B005FD5C" bold="true" box="[904,1033,710,729]" pageId="8" pageNumber="317">and diamond-</emphasis>
irregularly 
<emphasis id="B927EA81FFF0657EC3D3FD41B7E0FD77" bold="true" box="[934,1004,735,754]" pageId="8" pageNumber="317">shaped.</emphasis>
shape. 
<emphasis id="B927EA81FFF0657EC3D4FCB7B7FCFCB9" bold="true" box="[929,1008,809,828]" pageId="8" pageNumber="317">Uniform</emphasis>
Dark brown 
<emphasis id="B927EA81FFF0657EC3F8FCDCB7E7FCEB" bold="true" pageId="8" pageNumber="317">pinkish color pattern</emphasis>
Not available 
<emphasis id="B927EA81FFF0657EC41BFCEAB075FC02" bold="true" box="[1134,1145,884,903]" pageId="8" pageNumber="317">7</emphasis>
</td>
<td id="76B25DADFFF09A89C4A3FD0AB169FC02" box="[1238,1381,660,903]" gridcol="6" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC485FD0AB12FFC02" bold="true" pageId="8" pageNumber="317">Few large shields. Almost as long as wide. Absent 1</emphasis>
</td>
<td id="76B25DADFFF09A89C5F4FD0AB203FC02" box="[1409,1551,660,903]" gridcol="7" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC5FDFD0AB1C2FC02" bold="true" pageId="8" pageNumber="317">Many shields. Mostly small and diamond- shaped. Absent and yellow. 1</emphasis>
</td>
<td id="76B25DADFFF09A89C65EFD0AB369FC02" box="[1579,1893,660,903]" gridcol="8" gridrow="1" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC630FD0AB29FFCB8" bold="true" pageId="8" pageNumber="317">Few large Few large shields. Almost shields. All as long as wide. relatively long and two times longer than wide. Yellow</emphasis>
Dark brown 
<emphasis id="B927EA81FFF0657EC618FCEAB274FC02" bold="true" box="[1645,1656,884,903]" pageId="8" pageNumber="317">1</emphasis>
3, 5 and 7
</td>
</tr>
<tr id="356334D1FFF09A89C002FC06B369FC2E" box="[119,1893,920,939]" gridrow="2" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FC06B4B0FC2E" box="[119,188,920,939]" gridcol="0" gridrow="2" pageId="8" pageNumber="317">MXT</th>
<td id="76B25DADFFF09A89C0AFFC06B569FC2E" box="[218,357,920,939]" gridcol="1" gridrow="2" pageId="8" pageNumber="317">4/4</td>
<td id="76B25DADFFF09A89C1F6FC06B601FC2E" box="[387,525,920,939]" gridcol="2" gridrow="2" pageId="8" pageNumber="317">4/4</td>
<td id="76B25DADFFF09A89C258FC06B6B6FC2E" box="[557,698,920,939]" gridcol="3" gridrow="2" pageId="8" pageNumber="317">4/4</td>
<td id="76B25DADFFF09A89C2A2FC06B769FC2E" box="[727,869,920,939]" gridcol="4" gridrow="2" pageId="8" pageNumber="317">4/4</td>
<td id="76B25DADFFF09A89C3F7FC06B0B5FC2E" box="[898,1209,920,939]" gridcol="5" gridrow="2" pageId="8" pageNumber="317">Notavailable 4/4</td>
<td id="76B25DADFFF09A89C4A3FC06B169FC2E" box="[1238,1381,920,939]" gridcol="6" gridrow="2" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC565FC06B127FC2E" bold="true" box="[1296,1323,920,939]" pageId="8" pageNumber="317">3/3</emphasis>
</td>
<td id="76B25DADFFF09A89C5F4FC06B203FC2E" box="[1409,1551,920,939]" gridcol="7" gridrow="2" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC5CFFC06B1DAFC2E" bold="true" box="[1466,1494,920,939]" pageId="8" pageNumber="317">2/2</emphasis>
</td>
<td id="76B25DADFFF09A89C65EFC06B369FC2E" box="[1579,1893,920,939]" gridcol="8" gridrow="2" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC611FC06B326FC2E" bold="true" box="[1636,1834,920,939]" pageId="8" pageNumber="317">5/5 3/3</emphasis>
</td>
</tr>
<tr id="356334D1FFF09A89C002FC22B369FC4A" box="[119,1893,956,975]" gridrow="3" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FC22B4B0FC4A" box="[119,188,956,975]" gridcol="0" gridrow="3" pageId="8" pageNumber="317">DT</th>
<td id="76B25DADFFF09A89C0AFFC22B569FC4A" box="[218,357,956,975]" gridcol="1" gridrow="3" pageId="8" pageNumber="317">6/6</td>
<td id="76B25DADFFF09A89C1F6FC22B601FC4A" box="[387,525,956,975]" gridcol="2" gridrow="3" pageId="8" pageNumber="317">6/6</td>
<td id="76B25DADFFF09A89C258FC22B6B6FC4A" box="[557,698,956,975]" gridcol="3" gridrow="3" pageId="8" pageNumber="317">6/6</td>
<td id="76B25DADFFF09A89C2A2FC22B769FC4A" box="[727,869,956,975]" gridcol="4" gridrow="3" pageId="8" pageNumber="317">6/6</td>
<td id="76B25DADFFF09A89C3F7FC22B0B5FC4A" box="[898,1209,956,975]" gridcol="5" gridrow="3" pageId="8" pageNumber="317">Notavailable 6/6</td>
<td id="76B25DADFFF09A89C4A3FC22B169FC4A" box="[1238,1381,956,975]" gridcol="6" gridrow="3" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC565FC22B127FC4A" bold="true" box="[1296,1323,956,975]" pageId="8" pageNumber="317">5/5</emphasis>
</td>
<td id="76B25DADFFF09A89C5F4FC22B203FC4A" box="[1409,1551,956,975]" gridcol="7" gridrow="3" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC5CFFC22B1DAFC4A" bold="true" box="[1466,1494,956,975]" pageId="8" pageNumber="317">5/5</emphasis>
</td>
<td id="76B25DADFFF09A89C65EFC22B369FC4A" box="[1579,1893,956,975]" gridcol="8" gridrow="3" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC611FC22B28CFC4A" bold="true" box="[1636,1664,956,975]" pageId="8" pageNumber="317">5/5</emphasis>
4/4 and 6/6
</td>
</tr>
<tr id="356334D1FFF09A89C002FC7EB369FAB3" box="[119,1893,992,1334]" gridrow="4" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FC7EB4B0FAB3" box="[119,188,992,1334]" gridcol="0" gridrow="4" pageId="8" pageNumber="317">RPN RPMX FPMX</th>
<td id="76B25DADFFF09A89C0AFFC7EB569FAB3" box="[218,357,992,1334]" gridcol="1" gridrow="4" pageId="8" pageNumber="317">Present and contacting each other in the facial portion of skull. Present and contacting each other in the facial portion of skull. Present</td>
<td id="76B25DADFFF09A89C1F6FC7EB601FAB3" box="[387,525,992,1334]" gridcol="2" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC1DFFC7EB5E7FAB3" bold="true" pageId="8" pageNumber="317">Absent Absent Absent</emphasis>
</td>
<td id="76B25DADFFF09A89C258FC7EB6B6FAB3" box="[557,698,992,1334]" gridcol="3" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC249FC7EB681FA98" bold="true" pageId="8" pageNumber="317">Present, but relatively undeveloped and not contacting in the facial portion of skull. Present and not contacting in facial portion of skull.</emphasis>
Present
</td>
<td id="76B25DADFFF09A89C2A2FC7EB769FAB3" box="[727,869,992,1334]" gridcol="4" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC28BFC7EB733FAB3" bold="true" box="[766,831,992,1334]" pageId="8" pageNumber="317">Absent Absent Absent</emphasis>
</td>
<td id="76B25DADFFF09A89C3F7FC7EB0B5FAB3" box="[898,1209,992,1334]" gridcol="5" gridrow="4" pageId="8" pageNumber="317">
Not available 
<emphasis id="B927EA81FFF0657EC44EFC7EB080FB24" bold="true" pageId="8" pageNumber="317">Present, but relatively undeveloped and not contacting in the facial portion of skull.</emphasis>
Not available 
<emphasis id="B927EA81FFF0657EC449FB39B080FA98" bold="true" pageId="8" pageNumber="317">Present and not contacting in facial portion of skull.</emphasis>
Not available Present
</td>
<td id="76B25DADFFF09A89C4A3FC7EB169FAB3" box="[1238,1381,992,1334]" gridcol="6" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC488FC7EB132FAB3" bold="true" pageId="8" pageNumber="317">Absent Present and not contacting in facial portion of skull. Absent</emphasis>
</td>
<td id="76B25DADFFF09A89C5F4FC7EB203FAB3" box="[1409,1551,992,1334]" gridcol="7" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC5DDFC7EB1E5FAB3" bold="true" box="[1448,1513,992,1334]" pageId="8" pageNumber="317">Absent Absent Absent</emphasis>
</td>
<td id="76B25DADFFF09A89C65EFC7EB369FAB3" box="[1579,1893,992,1334]" gridcol="8" gridrow="4" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC627FC7EB29FFAB3" bold="true" pageId="8" pageNumber="317">Absent Present, but not contacting in the facial portion of skull. Absent Present and not contacting in facial portion of skull. Absent</emphasis>
Present
</td>
</tr>
<tr id="356334D1FFF09A89C002FAD8B369F9E9" box="[119,1893,1350,1644]" gridrow="5" pageId="8" pageNumber="317">
<th id="76B25DADFFF09A89C002FAD8B4B0F9E9" box="[119,188,1350,1644]" gridcol="0" gridrow="5" pageId="8" pageNumber="317">HL/SV L RL/HL RW/H W</th>
<td id="76B25DADFFF09A89C0AFFAD8B569F9E9" box="[218,357,1350,1644]" gridcol="1" gridrow="5" pageId="8" pageNumber="317">3.5–4.5 % (n = 3; Mean = 3.9; SD ± 0.5) 22.6–23.5% (n = 3; Mean = 23.3; SD ± 0.6) 64.0–68.1% (n = 3; Mean = 65.9; SD ± 2.1)</td>
<td id="76B25DADFFF09A89C1F6FAD8B601F9E9" box="[387,525,1350,1644]" gridcol="2" gridrow="5" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC1EAFAD9B604FA09" bold="true" pageId="8" pageNumber="317">2.6–3.3% (n = 6; Mean = 2.8; SD ± 0.2)</emphasis>
20.3–23.1% (n = 6;Mean = 21.6; SD ± 1.3) 
<emphasis id="B927EA81FFF0657EC1E1FA6BB601F9BC" bold="true" pageId="8" pageNumber="317">56.4–63.7% (n = 6; Mean= 60.2; SD ± 2.7)</emphasis>
</td>
<td id="76B25DADFFF09A89C258FAD8B6B6F9E9" box="[557,698,1350,1644]" gridcol="3" gridrow="5" pageId="8" pageNumber="317">3.3–5.1% (n = 17; Mean = 4.3; SD ± 0.5) 20.2–26.4% (n = 16; Mean = 23.9; SD ± 1.7) 49.3–74.3% (n = 17; Mean = 57.7; SD ± 7.4)</td>
<td id="76B25DADFFF09A89C2A2FAD8B769F9E9" box="[727,869,1350,1644]" gridcol="4" gridrow="5" pageId="8" pageNumber="317">
2. 
<emphasis id="B927EA81FFF0657EC377FAD8B76EFA53" bold="true" pageId="8" pageNumber="317">6–3.3% (n = 3; Mean = 2.9; SD ± 0.4) 24.0–27.4% (n = 4; Mean = 25.7; SD ± 1.4)</emphasis>
56.7–63.0% (n = 4; Mean = 59.9; SD ± 2.9)
</td>
<td id="76B25DADFFF09A89C3F7FAD8B0B5F9E9" box="[898,1209,1350,1644]" gridcol="5" gridrow="5" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC3EBFAD9B7F8FADF" bold="true" box="[926,1012,1351,1370]" pageId="8" pageNumber="317">1.9–2.8%</emphasis>
3.0–5.1% 
<emphasis id="B927EA81FFF0657EC3FDFAFEB006FAF6" bold="true" box="[904,1034,1376,1395]" pageId="8" pageNumber="317">(n = 11; Mean</emphasis>
(n = 113; Mean 
<emphasis id="B927EA81FFF0657EC3F7FAE6B01CFA0E" bold="true" box="[898,1040,1400,1419]" pageId="8" pageNumber="317">= 2.2; SD ± 0.2)</emphasis>
= 3.8; SD ± 0.4) 22.5–27.4% 21.9–33.7 % (n = 11; Mean (n = 115; Mean = 25.7; SD ± = 28.4; SD ± 1.7) 2.5) 57.9–75.4% 53.0–75.3% (n = 11; Mean (n = 115; Mean = 67.3; SD ± = 64.6; SD ± 5.0) 4.3)
</td>
<td id="76B25DADFFF09A89C4A3FAD8B169F9E9" box="[1238,1381,1350,1644]" gridcol="6" gridrow="5" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC486FAD9B150FA0E" bold="true" pageId="8" pageNumber="317">2.3–2.9% (n = 4; Mean = 2,7; SD ± 0.3)</emphasis>
22.1–23.8% (n = 4; Mean = 22.9; SD ± 0.9) 56.9–84.2% (n = 4; Mean = 64.3; SD ± 13.3)
</td>
<td id="76B25DADFFF09A89C5F4FAD8B203F9E9" box="[1409,1551,1350,1644]" gridcol="7" gridrow="5" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC5E8FAD9B1D5F9D7" bold="true" pageId="8" pageNumber="317">2.5–3.3% (n = 10; Mean = 2,7; SD ± 0.2) 28.9–34.3% (n = 14; Mean = 31.4; SD ± 1.6) 68.5–80.6% (n = 12; Mean = 71.9; SD ± 4.6)</emphasis>
</td>
<td id="76B25DADFFF09A89C65EFAD8B369F9E9" box="[1579,1893,1350,1644]" gridcol="8" gridrow="5" pageId="8" pageNumber="317">
<emphasis id="B927EA81FFF0657EC632FAD9B292FADF" bold="true" box="[1607,1694,1351,1370]" pageId="8" pageNumber="317">2.7–3.2%</emphasis>
2.6–4.1% 
<emphasis id="B927EA81FFF0657EC65EFAFEB2B8FAF6" bold="true" box="[1579,1716,1376,1395]" pageId="8" pageNumber="317">(n = 4; Mean =</emphasis>
(n = 4; Mean = 
<emphasis id="B927EA81FFF0657EC641FAE6B2BDFA0E" bold="true" box="[1588,1713,1400,1419]" pageId="8" pageNumber="317">3.0; SD ± 0.3)</emphasis>
3.1; SD ± 0.6) 
<emphasis id="B927EA81FFF0657EC648FA0CB2A4F98D" bold="true" pageId="8" pageNumber="317">28.3–29.3% 25.7–30.3% (n = 4; Mean = (n = 4; Mean = 28.9; SD ± 0.4) 27.6; SD ± 2.0) 78.6–84.3%</emphasis>
58.1–72.3% 
<emphasis id="B927EA81FFF0657EC65EF990B2B8F9A4" bold="true" box="[1579,1716,1550,1569]" pageId="8" pageNumber="317">(n = 4; Mean =</emphasis>
(n = 5; Mean = 
<emphasis id="B927EA81FFF0657EC65AF9B8B2BAF9BC" bold="true" box="[1583,1718,1574,1593]" pageId="8" pageNumber="317">81.4; SD ± 2.3)</emphasis>
67.4; SD ± 5.6) ……continued on the next page
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEC3693FFF1657FC00BFDA1B085FDC9" blockId="9.[125,1896,527,1676]" box="[126,1161,569,594]" pageId="9" pageNumber="318">
Charact Species of 
<taxonomicName id="4C534D10FFF1657FC468FDA4B085FDC9" box="[1053,1161,570,588]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="genus">
<emphasis id="B927EA81FFF1657FC468FDA4B085FDC9" box="[1053,1161,570,588]" italics="true" pageId="9" pageNumber="318">Leposternon</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BEC3693FFF1657FC0E6FDC6B342FDF7" blockId="9.[125,1896,527,1676]" box="[147,1870,600,626]" pageId="9" pageNumber="318">
ers 
<emphasis id="B927EA81FFF1657FC172FDC0B342FDF7" box="[263,1870,606,626]" italics="true" pageId="9" pageNumber="318">
<taxonomicName id="4C534D10FFF1657FC172FDC0B54FFDF7" box="[263,323,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="bagual">bagual</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC1E8FDC0B60EFDF7" box="[413,514,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="cerradensis">cerradensis</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC237FDC0B6BDFDF7" box="[578,689,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="infraorbitale">infraorbitale</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC292FDC0B76CFDF7" box="[743,864,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="kisteumacheri">kisteumacheri</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC3DDFDC1B0B0FDF7" box="[936,1212,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="subSpecies" species="maximus" subSpecies="microcephalum">maximus microcephalum</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC487FDC1B159FDF4" box="[1266,1365,607,625]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="octostegum">octostegum</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC5E9FDC0B1F3FDF7" box="[1436,1535,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="polystegum">polystegum</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC63DFDC1B2A4FDF4" box="[1608,1704,607,625]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="scutigerum">scutigerum</taxonomicName>
<taxonomicName id="4C534D10FFF1657FC683FDC0B342FDF7" box="[1782,1870,606,626]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="318" phylum="Chordata" rank="species" species="wuchereri">wuchereri</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC008FDE0B354FD17" blockId="9.[125,1896,527,1676]" box="[125,1880,638,658]" pageId="9" pageNumber="318">
AZL/H 29.3–35.0% 
<emphasis id="B927EA81FFF1657FC1ECFDE1B60AFD17" bold="true" box="[409,518,639,658]" pageId="9" pageNumber="318">40.1–51.4%</emphasis>
29.9–41.1% 
<emphasis id="B927EA81FFF1657FC29BFDE1B008FD17" bold="true" box="[750,1028,639,658]" pageId="9" pageNumber="318">35.4–40.5% 39.9–48.9%</emphasis>
22.7–43.2 % not applicable, 
<emphasis id="B927EA81FFF1657FC5EDFDE1B208FD17" bold="true" box="[1432,1540,639,658]" pageId="9" pageNumber="318">15.2–27.2%</emphasis>
Not applicable, 
<emphasis id="B927EA81FFF1657FC699FDE1B354FD17" bold="true" box="[1772,1880,639,658]" pageId="9" pageNumber="318">41.9–50.3%</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FD09B368FD2E" blockId="9.[125,1896,527,1676]" box="[225,1892,663,683]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFD06B61DFD2E" bold="true" box="[392,529,664,683]" pageId="9" pageNumber="318">(n = 6; Mean =</emphasis>
(n = 16; Mean 
<emphasis id="B927EA81FFF1657FC2A8FD06B01CFD2E" bold="true" box="[733,1040,664,683]" pageId="9" pageNumber="318">(n = 3; Mean = (n = 8; Mean =</emphasis>
(n = 96; Mean shield probably 
<emphasis id="B927EA81FFF1657FC5FFFD06B200FD2E" bold="true" box="[1418,1548,664,683]" pageId="9" pageNumber="318">(n = 14; Mean</emphasis>
shield absent. 
<emphasis id="B927EA81FFF1657FC6AEFD06B368FD2E" bold="true" box="[1755,1892,664,683]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FD2EB36AFD41" blockId="9.[125,1896,527,1676]" box="[226,1894,688,708]" pageId="9" pageNumber="318">
33.0; SD ± 3.3) 
<emphasis id="B927EA81FFF1657FC1F9FD2FB61FFD41" bold="true" box="[396,531,689,708]" pageId="9" pageNumber="318">40.7; SD ± 4.4)</emphasis>
= 36.2; SD ± 
<emphasis id="B927EA81FFF1657FC295FD2FB7BBFD46" bold="true" box="[736,951,688,708]" pageId="9" pageNumber="318">37.4; SD ± 2.8) 45.02</emphasis>
; 
<emphasis id="B927EA81FFF1657FC3B7FD2EB01BFD46" bold="true" box="[962,1047,688,707]" pageId="9" pageNumber="318">SD ± 3.1)</emphasis>
= 29.6; SD ± fused with 
<emphasis id="B927EA81FFF1657FC5E0FD2FB36AFD41" bold="true" box="[1429,1894,689,708]" pageId="9" pageNumber="318">= 18.8; SD ± 46.2; SD ± 3.8)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFD57B1D3FD59" blockId="9.[125,1896,527,1676]" box="[616,1503,713,732]" pageId="9" pageNumber="318">
3.1) 3.8) frontals 
<emphasis id="B927EA81FFF1657FC5C9FD57B1D3FD59" bold="true" box="[1468,1503,713,732]" pageId="9" pageNumber="318">3.2)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC008FD7CB354FD70" blockId="9.[125,1896,527,1676]" box="[125,1880,738,757]" pageId="9" pageNumber="318">
AZW/ 35.6–45.8% 35.5–48.6% 36.1–58.0% 
<emphasis id="B927EA81FFF1657FC29BFD7CB755FD70" bold="true" box="[750,857,738,757]" pageId="9" pageNumber="318">25.5–34.6%</emphasis>
29.6–69.2% 29.6–58.8 % 
<emphasis id="B927EA81FFF1657FC498FD7CB208FD70" bold="true" box="[1261,1540,738,757]" pageId="9" pageNumber="318">61.2–84.7% 13.7–20.4%</emphasis>
Not applicable, 
<emphasis id="B927EA81FFF1657FC699FD7CB354FD70" bold="true" box="[1772,1880,738,757]" pageId="9" pageNumber="318">59.0–67.4%</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FD65B368FC8B" blockId="9.[125,1896,527,1676]" box="[225,1892,763,782]" pageId="9" pageNumber="318">
(n = 3; Mean = (n = 6; Mean= (n = 16; Mean 
<emphasis id="B927EA81FFF1657FC2A8FD65B769FC8B" bold="true" box="[733,869,763,782]" pageId="9" pageNumber="318">(n = 3; Mean =</emphasis>
(n = 11; Mean (n = 105; Mean 
<emphasis id="B927EA81FFF1657FC4A9FD65B200FC8B" bold="true" box="[1244,1548,763,782]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 12; Mean</emphasis>
shield absent. 
<emphasis id="B927EA81FFF1657FC6AEFD65B368FC8B" bold="true" box="[1755,1892,763,782]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FC8DB36AFCA2" blockId="9.[125,1896,527,1676]" box="[226,1894,787,807]" pageId="9" pageNumber="318">
40.2; SD ± 5.2) 43.8; SD ± 4.4) = 44.9; SD ± 
<emphasis id="B927EA81FFF1657FC295FC8AB76BFCA2" bold="true" box="[736,871,788,807]" pageId="9" pageNumber="318">29.8; SD ± 4.6)</emphasis>
= 56.9; SD ± = 44.0; SD ± 
<emphasis id="B927EA81FFF1657FC486FC8AB36AFCA2" bold="true" box="[1267,1894,788,807]" pageId="9" pageNumber="318">68.9; SD ± = 16.9; SD ± 63.0; SD ± 4.3)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFCB2B1D3FCC5" blockId="9.[125,1896,527,1676]" box="[616,1503,812,832]" pageId="9" pageNumber="318">
5.6) 10.9) 5.7) 
<emphasis id="B927EA81FFF1657FC579FCB3B1D3FCC5" bold="true" box="[1292,1503,813,832]" pageId="9" pageNumber="318">10.7) 2.4)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC0D7FCDBB369FC26" blockId="9.[125,1896,527,1676]" pageId="9" pageNumber="318">
/H 25.3–25.5% 21.9–27.8% 21.0–29.4% 
<emphasis id="B927EA81FFF1657FC29BFCD8B008FCDC" bold="true" box="[750,1028,838,857]" pageId="9" pageNumber="318">28.9–36.6% 17.6–24.1%</emphasis>
15.2–30.6% Not applicable, 
<emphasis id="B927EA81FFF1657FC5EDFCD8B208FCDC" bold="true" box="[1432,1540,838,857]" pageId="9" pageNumber="318">41.1–47.9%</emphasis>
Not applicable, 
<emphasis id="B927EA81FFF1657FC770FCD8B333FCDC" bold="true" box="[1797,1855,838,857]" pageId="9" pageNumber="318">36.0%</emphasis>
(n = 2) (n = 4; Mean = (n = 16; Mean 
<emphasis id="B927EA81FFF1657FC29EFCC1B01CFCF7" bold="true" box="[747,1040,863,882]" pageId="9" pageNumber="318">(n = 2, other (n = 8; Mean =</emphasis>
(n = 90; Mean shield absent. 
<emphasis id="B927EA81FFF1657FC5F3FCC1B203FCF7" bold="true" box="[1414,1551,863,882]" pageId="9" pageNumber="318">(n = 5; Mean =</emphasis>
shield absent. 
<emphasis id="B927EA81FFF1657FC69CFCC1B350FCF7" bold="true" box="[1769,1884,863,882]" pageId="9" pageNumber="318">(n = 1; other</emphasis>
24.8; SD ± 2.5) = 25.0; SD ± 
<emphasis id="B927EA81FFF1657FC2ABFCE9B01DFC0F" bold="true" box="[734,1041,887,906]" pageId="9" pageNumber="318">specimens with 21.2; SD ± 2.1)</emphasis>
= 22.8; SD ± 
<emphasis id="B927EA81FFF1657FC5FFFCE9B364FC0F" bold="true" box="[1418,1896,887,906]" pageId="9" pageNumber="318">43.6; SD ± 2.6) specimens with</emphasis>
2.4) 
<emphasis id="B927EA81FFF1657FC2A8FC0EB767FC26" bold="true" box="[733,875,912,931]" pageId="9" pageNumber="318">fusion frontals)</emphasis>
3.3) 
<emphasis id="B927EA81FFF1657FC6AAFC0EB369FC26" bold="true" box="[1759,1893,912,931]" pageId="9" pageNumber="318">fused frontals)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC0E3FC36B345FC3E" blockId="9.[125,1896,527,1676]" box="[150,1865,936,956]" pageId="9" pageNumber="318">
/HL 9.8–13.9% 
<emphasis id="B927EA81FFF1657FC1D1FC37B5F6FC39" bold="true" box="[420,506,937,956]" pageId="9" pageNumber="318">7.5–8.3%</emphasis>
9.7–18.0% 
<emphasis id="B927EA81FFF1657FC28CFC37B743FC39" bold="true" box="[761,847,937,956]" pageId="9" pageNumber="318">4.8–9.4%</emphasis>
4.6–13.0% 7.6–17.8% Not applicable, 
<emphasis id="B927EA81FFF1657FC5D7FC37B1F5FC39" bold="true" box="[1442,1529,937,956]" pageId="9" pageNumber="318">4.3–5.3%</emphasis>
9.2–11.3% Fusioned
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FC5FB347FC51" blockId="9.[125,1896,527,1676]" box="[225,1867,961,981]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFC5CB61DFC50" bold="true" box="[392,529,962,981]" pageId="9" pageNumber="318">(n = 3; Mean =</emphasis>
(n = 17; Mean 
<emphasis id="B927EA81FFF1657FC2A8FC5CB769FC50" bold="true" box="[733,869,962,981]" pageId="9" pageNumber="318">(n = 3; Mean =</emphasis>
(n = 9; Mean = (n = 94; Mean shield absent. 
<emphasis id="B927EA81FFF1657FC5DAFC5CB1E0FC50" bold="true" box="[1455,1516,962,981]" pageId="9" pageNumber="318">(n = 2)</emphasis>
(n = 2) parietals.
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FC44B0BDFC68" blockId="9.[125,1896,527,1676]" box="[226,1201,986,1006]" pageId="9" pageNumber="318">
12.2; SD ± 2.1) 
<emphasis id="B927EA81FFF1657FC1E4FC45B601FC6B" bold="true" box="[401,525,987,1006]" pageId="9" pageNumber="318">7.9; SD ± 0.4)</emphasis>
= 13.6; SD ± 
<emphasis id="B927EA81FFF1657FC293FC45B76EFC6B" bold="true" box="[742,866,987,1006]" pageId="9" pageNumber="318">7.8; SD ± 2.6)</emphasis>
7.9; SD ± 2.5) = 12.4; SD ±
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFC6DB086FB83" blockId="9.[125,1896,527,1676]" box="[616,1162,1011,1030]" pageId="9" pageNumber="318">2.7) 2.3)</paragraph>
<paragraph id="8BEC3693FFF1657FC008FB92B354FB9A" blockId="9.[125,1896,527,1676]" box="[125,1880,1036,1055]" pageId="9" pageNumber="318">
TPL/H 18.9–22.0% 
<emphasis id="B927EA81FFF1657FC1ECFB92B609FB9A" bold="true" box="[409,517,1036,1055]" pageId="9" pageNumber="318">25.0–29.9%</emphasis>
15.8–27.4% 
<emphasis id="B927EA81FFF1657FC29BFB92B756FB9A" bold="true" box="[750,858,1036,1055]" pageId="9" pageNumber="318">27.1–31.4%</emphasis>
14.3–44.3% 14.0–23.8% 
<emphasis id="B927EA81FFF1657FC498FB92B20FFB9A" bold="true" box="[1261,1539,1036,1055]" pageId="9" pageNumber="318">27.5–31.8% 24.5–30.9%</emphasis>
Not applicable, 
<emphasis id="B927EA81FFF1657FC699FB92B354FB9A" bold="true" box="[1772,1880,1036,1055]" pageId="9" pageNumber="318">16.4–18.0%</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FBBBB368FBBD" blockId="9.[125,1896,527,1676]" box="[225,1892,1061,1080]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFBBBB61DFBBD" bold="true" box="[392,529,1061,1080]" pageId="9" pageNumber="318">(n = 5; Mean =</emphasis>
(n = 17; Mean 
<emphasis id="B927EA81FFF1657FC2A8FBBBB769FBBD" bold="true" box="[733,869,1061,1080]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 8; Mean = (n = 109; Mean 
<emphasis id="B927EA81FFF1657FC4A9FBBBB200FBBD" bold="true" box="[1244,1548,1061,1080]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 12; Mean</emphasis>
shield absent. 
<emphasis id="B927EA81FFF1657FC6AEFBBBB368FBBD" bold="true" box="[1755,1892,1061,1080]" pageId="9" pageNumber="318">(n = 3; Mean =</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FBA0B369FBD4" blockId="9.[125,1896,527,1676]" box="[226,1893,1086,1105]" pageId="9" pageNumber="318">
20.2; SD ± 1.6) 
<emphasis id="B927EA81FFF1657FC1FBFBA0B61CFBD4" bold="true" box="[398,528,1086,1105]" pageId="9" pageNumber="318">27.7; SD =1.7)</emphasis>
= 20.1; SD ± 
<emphasis id="B927EA81FFF1657FC295FBA0B76BFBD4" bold="true" box="[736,871,1086,1105]" pageId="9" pageNumber="318">28.7; SD ± 1.9)</emphasis>
29.3; SD ± 8.1) = 19.0; SD ± 
<emphasis id="B927EA81FFF1657FC495FBA0B369FBD4" bold="true" box="[1248,1893,1086,1105]" pageId="9" pageNumber="318">29.4; SD ± 2.0) = 27.2; SD ± 18.6; SD ± 1.9)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFBC8B1D3FBEF" blockId="9.[125,1896,527,1676]" box="[616,1503,1110,1130]" pageId="9" pageNumber="318">
3.0) 2.3) 
<emphasis id="B927EA81FFF1657FC5C9FBC9B1D3FBEF" bold="true" box="[1468,1503,1111,1130]" pageId="9" pageNumber="318">1.8)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC008FBF1B35AFB07" blockId="9.[125,1896,527,1676]" box="[125,1878,1135,1155]" pageId="9" pageNumber="318">
TPW/H 13.2–17.8% 
<emphasis id="B927EA81FFF1657FC1ECFBEEB60AFB06" bold="true" box="[409,518,1136,1155]" pageId="9" pageNumber="318">22.1–31.0%</emphasis>
7.1–24.5% 
<emphasis id="B927EA81FFF1657FC29BFBEEB009FB06" bold="true" box="[750,1029,1136,1155]" pageId="9" pageNumber="318">20.3–31.1% 19.2–28.6%</emphasis>
9.3–42.2% 
<emphasis id="B927EA81FFF1657FC498FBEEB20FFB06" bold="true" box="[1261,1539,1136,1155]" pageId="9" pageNumber="318">18.4–27.5% 29.9–39.5%</emphasis>
Not applicable, 15.8–18.0%
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FB16B36CFB1E" blockId="9.[125,1896,527,1676]" box="[225,1888,1160,1180]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFB17B61DFB19" bold="true" box="[392,529,1161,1180]" pageId="9" pageNumber="318">(n = 5; Mean =</emphasis>
(n = 18; Mean 
<emphasis id="B927EA81FFF1657FC2A8FB17B000FB19" bold="true" box="[733,1036,1161,1180]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 10; Mean</emphasis>
(n = 109; Mean 
<emphasis id="B927EA81FFF1657FC4A9FB17B200FB19" bold="true" box="[1244,1548,1161,1180]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 12; Mean</emphasis>
shield absent. (n = 3; Mean =
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FB3FB369FB31" blockId="9.[125,1896,527,1676]" box="[226,1893,1185,1204]" pageId="9" pageNumber="318">
15.6; SD ± 2.3) 
<emphasis id="B927EA81FFF1657FC1FBFB3FB61DFB31" bold="true" box="[398,529,1185,1204]" pageId="9" pageNumber="318">25.1; SD =3.2)</emphasis>
= 15.8; SD ± 
<emphasis id="B927EA81FFF1657FC295FB3FB00BFB31" bold="true" box="[736,1031,1185,1204]" pageId="9" pageNumber="318">26.3; SD ± 4.5) = 24.7; SD ±</emphasis>
= 19.7; SD ± 
<emphasis id="B927EA81FFF1657FC495FB3FB20AFB31" bold="true" box="[1248,1542,1185,1204]" pageId="9" pageNumber="318">21.1; SD ± 4.3) = 34.8; SD ±</emphasis>
16.8; SD ± 1.1)
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFB24B1D3FB48" blockId="9.[125,1896,527,1676]" box="[616,1503,1210,1229]" pageId="9" pageNumber="318">
4.2) 
<emphasis id="B927EA81FFF1657FC3C8FB24B7ECFB48" bold="true" box="[957,992,1210,1229]" pageId="9" pageNumber="318">3.0)</emphasis>
5.7) 
<emphasis id="B927EA81FFF1657FC5C9FB24B1D3FB48" bold="true" box="[1468,1503,1210,1229]" pageId="9" pageNumber="318">2.8)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657FC008FB4CB35BFB60" blockId="9.[125,1896,527,1676]" box="[125,1879,1234,1254]" pageId="9" pageNumber="318">
PFL/H 23.0–26.8% 
<emphasis id="B927EA81FFF1657FC1ECFB4DB609FB63" bold="true" box="[409,517,1235,1254]" pageId="9" pageNumber="318">37.0–44.3%</emphasis>
20.6–36.9% 
<emphasis id="B927EA81FFF1657FC29BFB4DB755FB63" bold="true" box="[750,857,1235,1254]" pageId="9" pageNumber="318">37.2–41.6%</emphasis>
22.2–27.2% 14.5–29.5% 
<emphasis id="B927EA81FFF1657FC498FB4DB154FB63" bold="true" box="[1261,1368,1235,1254]" pageId="9" pageNumber="318">36.4–41.3%</emphasis>
24.2–28.2% Not applicable, 20.9–26.1%
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FB75B36CFB7A" blockId="9.[125,1896,527,1676]" box="[225,1888,1259,1279]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFB72B61DFB7A" bold="true" box="[392,529,1260,1279]" pageId="9" pageNumber="318">(n = 5; Mean =</emphasis>
(n = 16; Mean 
<emphasis id="B927EA81FFF1657FC2A8FB72B769FB7A" bold="true" box="[733,869,1260,1279]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 2) (n = 71; Mean 
<emphasis id="B927EA81FFF1657FC4A9FB72B169FB7A" bold="true" box="[1244,1381,1260,1279]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 5; Mean = shield absent. (n = 3; Mean =
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FA9AB369FA92" blockId="9.[125,1896,527,1676]" box="[226,1893,1284,1304]" pageId="9" pageNumber="318">
24.8; SD ± 1.9) 
<emphasis id="B927EA81FFF1657FC1FBFA9BB61CFA9D" bold="true" box="[398,528,1285,1304]" pageId="9" pageNumber="318">39.6; SD =2.7)</emphasis>
= 27.5; SD ± 
<emphasis id="B927EA81FFF1657FC295FA9BB76BFA9D" bold="true" box="[736,871,1285,1304]" pageId="9" pageNumber="318">39.4; SD ± 2.2)</emphasis>
= 22.1; SD ± 
<emphasis id="B927EA81FFF1657FC495FA9BB16BFA9D" bold="true" box="[1248,1383,1285,1304]" pageId="9" pageNumber="318">37.8; SD ± 2.4)</emphasis>
26.0; SD ± 2.0) 23.0; SD ± 2.7)
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFA83B086FAB5" blockId="9.[125,1896,527,1676]" box="[616,1162,1309,1328]" pageId="9" pageNumber="318">4.3) 2.5)</paragraph>
<paragraph id="8BEC3693FFF1657FC008FAA8B35AFACC" blockId="9.[125,1896,527,1676]" box="[125,1878,1334,1353]" pageId="9" pageNumber="318">
PFW/H 38.3–41.7% 
<emphasis id="B927EA81FFF1657FC1ECFAA8B60AFACC" bold="true" box="[409,518,1334,1353]" pageId="9" pageNumber="318">18.9–27.3%</emphasis>
32.0–45.7% 
<emphasis id="B927EA81FFF1657FC29BFAA8B755FACC" bold="true" box="[750,857,1334,1353]" pageId="9" pageNumber="318">25.4–36.7%</emphasis>
Not available 28.4–52.0% 
<emphasis id="B927EA81FFF1657FC498FAA8B155FACC" bold="true" box="[1261,1369,1334,1353]" pageId="9" pageNumber="318">15.9–34.7%</emphasis>
not available Not applicable, 41.6–49.7%
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FAD1B36CFAE7" blockId="9.[125,1896,527,1676]" box="[225,1888,1359,1378]" pageId="9" pageNumber="318">
(n = 3; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFAD1B61DFAE7" bold="true" box="[392,529,1359,1378]" pageId="9" pageNumber="318">(n = 5; Mean =</emphasis>
(n = 16; Mean 
<emphasis id="B927EA81FFF1657FC2A8FAD1B769FAE7" bold="true" box="[733,869,1359,1378]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 90; Mean 
<emphasis id="B927EA81FFF1657FC4A9FAD1B169FAE7" bold="true" box="[1244,1381,1359,1378]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
shield absent. (n = 3; Mean =
</paragraph>
<paragraph id="8BEC3693FFF1657FC097FAF6B369FAFE" blockId="9.[125,1896,527,1676]" box="[226,1893,1384,1403]" pageId="9" pageNumber="318">
39.5; SD ± 1.9) 
<emphasis id="B927EA81FFF1657FC1FBFAF6B61DFAFE" bold="true" box="[398,529,1384,1403]" pageId="9" pageNumber="318">21.6; SD =3.0)</emphasis>
= 36.6; SD ± 
<emphasis id="B927EA81FFF1657FC295FAF6B76BFAFE" bold="true" box="[736,871,1384,1403]" pageId="9" pageNumber="318">28.8; SD ± 5.3)</emphasis>
= 43.9; SD ± 
<emphasis id="B927EA81FFF1657FC495FAF6B16BFAFE" bold="true" box="[1248,1383,1384,1403]" pageId="9" pageNumber="318">24.5; SD ± 7.8)</emphasis>
44.9; SD ± 4.3)
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFA1EB086FA16" blockId="9.[125,1896,527,1676]" box="[616,1162,1408,1427]" pageId="9" pageNumber="318">4.3) 4.1)</paragraph>
<paragraph id="8BEC3693FFF1657FC008FA07B35AFA29" blockId="9.[125,1896,527,1676]" box="[125,1878,1433,1453]" pageId="9" pageNumber="318">
CML/H 53.4–59.1% 
<emphasis id="B927EA81FFF1657FC1ECFA04B609FA28" bold="true" box="[409,517,1434,1453]" pageId="9" pageNumber="318">64.0–73.8%</emphasis>
49.2–69.6% 53.7–67.8% 56.7–64.5% 47.3–63.2% 
<emphasis id="B927EA81FFF1657FC498FA04B154FA28" bold="true" box="[1261,1368,1434,1453]" pageId="9" pageNumber="318">61.0–69.7%</emphasis>
54.0–57.4% 58.9–59.5% 54.0–57.1%
</paragraph>
<paragraph id="8BEC3693FFF1657FC094FA2CB353FA5B" blockId="9.[125,1896,527,1676]" pageId="9" pageNumber="318">
(n = 4; Mean = 
<emphasis id="B927EA81FFF1657FC1FDFA2DB61AFA43" bold="true" box="[392,534,1459,1478]" pageId="9" pageNumber="318">(n = 6; Mean =</emphasis>
(n = 17; Mean (n = 4; Mean = (n = 2) (n = 91; Mean 
<emphasis id="B927EA81FFF1657FC4A9FA2DB169FA43" bold="true" box="[1244,1381,1459,1478]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 3; Mean = (n = 2) (n = 3; Mean = 56.9; SD =2.8) 
<emphasis id="B927EA81FFF1657FC1FBFA52B61DFA5A" bold="true" box="[398,529,1484,1503]" pageId="9" pageNumber="318">68.1; SD =3.7)</emphasis>
= 59.9; SD ± 62.1; SD ± 6.0) = 55.4; SD ± 
<emphasis id="B927EA81FFF1657FC495FA52B16BFA5A" bold="true" box="[1248,1383,1484,1503]" pageId="9" pageNumber="318">65.5; SD ± 4.8)</emphasis>
55.7; SD=1.7) 0.7; SD ± 0.1)
</paragraph>
<paragraph id="8BEC3693FFF1657FC21DFA7AB086FA72" blockId="9.[125,1896,527,1676]" box="[616,1162,1508,1527]" pageId="9" pageNumber="318">5.6) 3.6)</paragraph>
<paragraph id="8BEC3693FFF1657FC0EBFA63B36CF9C7" blockId="9.[125,1896,527,1676]" pageId="9" pageNumber="318">
/SV 3.2–4.4% 2.4–3.2% 3.7–6.8% 
<emphasis id="B927EA81FFF1657FC28CFA63B7F5F995" bold="true" box="[761,1017,1533,1552]" pageId="9" pageNumber="318">2.6–3.0% 1.7–1.9%</emphasis>
2.5–5.5% 
<emphasis id="B927EA81FFF1657FC48DFA63B341F995" bold="true" box="[1272,1869,1533,1552]" pageId="9" pageNumber="318">2.0–2.9% 1.6–2.7% 2.4–2.9% 1.9–2.9%</emphasis>
(n =4; X = 3.7; (n = 6; Mean = (n = 17; Mean 
<emphasis id="B927EA81FFF1657FC2A8F988B000F9AC" bold="true" box="[733,1036,1558,1577]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 10; Mean</emphasis>
(n = 111; Mean 
<emphasis id="B927EA81FFF1657FC4A9F988B368F9AC" bold="true" box="[1244,1892,1558,1577]" pageId="9" pageNumber="318">(n = 4; Mean = (n = 10; Mean (n = 4; Mean = (n = 5; Mean =</emphasis>
SD =0.5) 2.7; SD =0.3) = 5.1; SD ± 0.9) 
<emphasis id="B927EA81FFF1657FC293F9B1B002F9C4" bold="true" box="[742,1038,1582,1602]" pageId="9" pageNumber="318">2.9; SD ± 0.2) = 1.8; SD ± 0.1</emphasis>
) = 3.8; SD ± 0.5) 
<emphasis id="B927EA81FFF1657FC490F9B1B36CF9C7" bold="true" box="[1253,1888,1583,1602]" pageId="9" pageNumber="318">2.4; SD ± 0.3) = 2.4; SD=0.3) 2.6; SD ± 0.2) 2.4; SD ± 0.3)</emphasis>
</paragraph>
<paragraph id="8BEC3693FFF1657CC0E2F9D9B104FEB4" blockId="9.[125,1896,527,1676]" lastBlockId="10.[151,1436,151,305]" lastPageId="10" lastPageNumber="319" pageId="9" pageNumber="318">
/SV 4.9–5.6% 4.9–5.8% 4.5–7.4% 5.3–6.7% 3.9–6.2% 3.8–7.5% 
<emphasis id="B927EA81FFF1657FC48DF9D6B142F9DE" bold="true" box="[1272,1358,1608,1627]" pageId="9" pageNumber="318">3.2–4.3%</emphasis>
4.2–5.0% 4.7–5.4% 4.6–5.8% (n =4; X = 5.3; (n = 6; Mean = (n = 17; Mean (n = 4; Mean = (n = 10; Mean (n = 114; Mean 
<emphasis id="B927EA81FFF1657FC4A9F9FEB169F9F6" bold="true" box="[1244,1381,1632,1651]" pageId="9" pageNumber="318">(n = 4; Mean =</emphasis>
(n = 9; Mean = (n = 4; Mean = (n = 5; Mean = SD ± 0.3) 5.3; SD =0.4) = 5.6; SD ± 0.7) 5.8; SD ± 0.7) = 4.8; SD ± 0.7) = 5.5; SD ± 0.6) 
<emphasis id="B927EA81FFF1657FC490F9E7B16DF909" bold="true" box="[1253,1377,1657,1676]" pageId="9" pageNumber="318">3.7; SD ± 0.5)</emphasis>
4.6; SD=0.3) 5.0; SD ± 0.3) 5.3; SD ± 0.5) 
<emphasis id="B927EA81FFF2657CC0E2FF06B506FF2A" bold="true" box="[151,266,152,175]" pageId="10" pageNumber="319">TABLE 2.</emphasis>
Meristic and morphometric measurements (in millimeters) of the 
<typeStatus id="54E88831FFF2657CC3A4FF06B7F3FF2A" box="[977,1023,152,175]" pageId="10" pageNumber="319">type</typeStatus>
series of 
<taxonomicName id="4C534D10FFF2657CC41DFF06B132FF2A" box="[1128,1342,151,175]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="319" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFF2657CC41DFF06B132FF2A" box="[1128,1342,151,175]" italics="true" pageId="10" pageNumber="319">Leposternon bagual</emphasis>
</taxonomicName>
<emphasis id="B927EA81FFF2657CC533FF06B190FF2A" bold="true" box="[1350,1436,152,175]" pageId="10" pageNumber="319">
<taxonomicNameLabel id="A21457FAFFF2657CC533FF06B190FF2A" box="[1350,1436,152,175]" pageId="10" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations of characters: CA, Caudal annuli; DAA, Dorsal anterior half-annuli; DPA, Dorsal postpectoral halfannuli; DS, Dorsal segments per half-annulus of midbody; F, female; HL, Head length; LA, Lateral half-annuli; M, male; MD, Midbody diameter; PV, Precloacal vertebrae; S, Sex; SC, Segments caudal (5th annulus); SVL, Snout-vent length; TL, Tail length; VPA, Ventral postpectoral half-annuli; VS, Ventral segments per half-annulus of midbody.
</paragraph>
<paragraph id="8BEC3693FFF2657CC0EAFED3B114FEE6" blockId="10.[159,1304,333,468]" box="[159,1304,333,355]" pageId="10" pageNumber="319">
Characters 
<typeStatus id="54E88831FFF2657CC1FEFED3B5E5FEE6" box="[395,489,333,355]" pageId="10" pageNumber="319" type="holotype">Holotype</typeStatus>
<typeStatus id="54E88831FFF2657CC21AFED0B6DEFEE6" box="[623,722,334,355]" pageId="10" pageNumber="319" type="paratype">Paratypes</typeStatus>
Mean ±SD
</paragraph>
<paragraph id="8BEC3693FFF2657CC1FEFEE9B7D7FE09" blockId="10.[159,1304,333,468]" box="[395,987,375,396]" pageId="10" pageNumber="319">FML FML FML FML</paragraph>
<paragraph id="8BEC3693FFF2657CC1FEFE0BB7EAFE2E" blockId="10.[159,1304,333,468]" box="[395,998,405,427]" pageId="10" pageNumber="319">11319 11312 11313 11318</paragraph>
<paragraph id="8BEC3693FFF2657CC0EAFE20B7B1FE51" blockId="10.[159,1304,333,468]" box="[159,957,446,468]" pageId="10" pageNumber="319">Sex F M F M</paragraph>
<paragraph id="8BEC3693FFF2657CC0EAFE76B11BFC41" pageId="10" pageNumber="319">
<table id="F953C433FFF29A89C0EAFE76B128FC41" box="[159,1316,488,964]" gridcols="7" gridrows="4" pageId="10" pageNumber="319">
<tr id="356334D1FFF29A89C0EAFE76B128FDD5" box="[159,1316,488,592]" gridrow="0" pageId="10" pageNumber="319">
<th id="76B25DADFFF29A89C0EAFE76B4DAFDD5" box="[159,214,488,592]" gridcol="0" gridrow="0" pageId="10" pageNumber="319">DAA DPA VPA</th>
<th id="76B25DADFFF29A89C1FEFE76B5BBFDD5" box="[395,439,488,592]" gridcol="1" gridrow="0" pageId="10" pageNumber="319">18 253 214</th>
<th id="76B25DADFFF29A89C21AFE76B697FDD5" box="[623,667,488,592]" gridcol="2" gridrow="0" pageId="10" pageNumber="319">16 266 211</th>
<th id="76B25DADFFF29A89C36CFE76B733FDD5" box="[793,831,488,592]" gridcol="3" gridrow="0" pageId="10" pageNumber="319">17 240 206</th>
<th id="76B25DADFFF29A89C3D2FE76B00DFDD5" box="[935,1025,488,592]" gridcol="4" gridrow="0" pageId="10" pageNumber="319">15 232 + n* 205 + n*</th>
<th id="76B25DADFFF29A89C420FE76B082FDD5" box="[1109,1166,488,592]" gridcol="5" gridrow="0" pageId="10" pageNumber="319">16.5 253.0 210.3</th>
<th id="76B25DADFFF29A89C491FE76B128FDD5" box="[1252,1316,488,592]" gridcol="6" gridrow="0" pageId="10" pageNumber="319">± 1.3 ± 13.0 ± 4.0</th>
</tr>
<tr id="356334D1FFF29A89C0EAFDFAB128FD49" box="[159,1316,612,716]" gridrow="1" pageId="10" pageNumber="319">
<th id="76B25DADFFF29A89C0EAFDFAB4DAFD49" box="[159,214,612,716]" gridcol="0" gridrow="1" pageId="10" pageNumber="319">CA LA DS</th>
<td id="76B25DADFFF29A89C1FEFDFAB5BBFD49" box="[395,439,612,716]" gridcol="1" gridrow="1" pageId="10" pageNumber="319">12 4/5 28</td>
<td id="76B25DADFFF29A89C21AFDFAB697FD49" box="[623,667,612,716]" gridcol="2" gridrow="1" pageId="10" pageNumber="319">11 5/5 28</td>
<td id="76B25DADFFF29A89C36CFDFAB733FD49" box="[793,831,612,716]" gridcol="3" gridrow="1" pageId="10" pageNumber="319">11 5/5 29</td>
<td id="76B25DADFFF29A89C3D2FDFAB00DFD49" box="[935,1025,612,716]" gridcol="4" gridrow="1" pageId="10" pageNumber="319">10 4/4 29</td>
<td id="76B25DADFFF29A89C420FDFAB082FD49" box="[1109,1166,612,716]" gridcol="5" gridrow="1" pageId="10" pageNumber="319">16.5 28.5</td>
<td id="76B25DADFFF29A89C491FDFAB128FD49" box="[1252,1316,612,716]" gridcol="6" gridrow="1" pageId="10" pageNumber="319">± 1.3 ± 0.6</td>
</tr>
<tr id="356334D1FFF29A89C0EAFD7EB128FCCD" box="[159,1316,736,840]" gridrow="2" pageId="10" pageNumber="319">
<th id="76B25DADFFF29A89C0EAFD7EB4DAFCCD" box="[159,214,736,840]" gridcol="0" gridrow="2" pageId="10" pageNumber="319">VS CS HL</th>
<td id="76B25DADFFF29A89C1FEFD7EB5BBFCCD" box="[395,439,736,840]" gridcol="1" gridrow="2" pageId="10" pageNumber="319">23 35 9.0</td>
<td id="76B25DADFFF29A89C21AFD7EB697FCCD" box="[623,667,736,840]" gridcol="2" gridrow="2" pageId="10" pageNumber="319">24 37 10.6</td>
<td id="76B25DADFFF29A89C36CFD7EB733FCCD" box="[793,831,736,840]" gridcol="3" gridrow="2" pageId="10" pageNumber="319">27 36 8.0</td>
<td id="76B25DADFFF29A89C3D2FD7EB00DFCCD" box="[935,1025,736,840]" gridcol="4" gridrow="2" pageId="10" pageNumber="319">26 30 7.3</td>
<td id="76B25DADFFF29A89C420FD7EB082FCCD" box="[1109,1166,736,840]" gridcol="5" gridrow="2" pageId="10" pageNumber="319">25.0 34.5 8.7</td>
<td id="76B25DADFFF29A89C491FD7EB128FCCD" box="[1252,1316,736,840]" gridcol="6" gridrow="2" pageId="10" pageNumber="319">± 1.8 ±3.1 ± 1.4</td>
</tr>
<tr id="356334D1FFF29A89C0EAFCC2B128FC41" box="[159,1316,860,964]" gridrow="3" pageId="10" pageNumber="319">
<th id="76B25DADFFF29A89C0EAFCC2B4DAFC41" box="[159,214,860,964]" gridcol="0" gridrow="3" pageId="10" pageNumber="319">SVL MD TL</th>
<td id="76B25DADFFF29A89C1FEFCC2B5BBFC41" box="[395,439,860,964]" gridcol="1" gridrow="3" pageId="10" pageNumber="319">257 9.2 14.5</td>
<td id="76B25DADFFF29A89C21AFCC2B697FC41" box="[623,667,860,964]" gridcol="2" gridrow="3" pageId="10" pageNumber="319">294 9.4 15.5</td>
<td id="76B25DADFFF29A89C36CFCC2B733FC41" box="[793,831,860,964]" gridcol="3" gridrow="3" pageId="10" pageNumber="319">178 7.9 8.8</td>
<td id="76B25DADFFF29A89C3D2FCC2B00DFC41" box="[935,1025,860,964]" gridcol="4" gridrow="3" pageId="10" pageNumber="319">ca. 174* 6.3 11.4</td>
<td id="76B25DADFFF29A89C420FCC2B082FC41" box="[1109,1166,860,964]" gridcol="5" gridrow="3" pageId="10" pageNumber="319">243 8.2 12.5</td>
<td id="76B25DADFFF29A89C491FCC2B128FC41" box="[1252,1316,860,964]" gridcol="6" gridrow="3" pageId="10" pageNumber="319">± 59.3 ±1.5 ± 3.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEC3693FFF2657CC0E2FC7BB5FBFC79" blockId="10.[151,503,997,1020]" box="[151,503,997,1020]" pageId="10" pageNumber="319">* Specimen with mutilated body.</paragraph>
<caption id="DF2C661BFFF2657CC0E2F8D1B742F8E1" box="[151,846,1871,1893]" httpUri="https://zenodo.org/record/242962/files/figure.png" pageId="10" pageNumber="319" targetBox="[168,1431,1074,1842]" targetPageId="10">
<paragraph id="8BEC3693FFF2657CC0E2F8D1B742F8E1" blockId="10.[151,846,1871,1893]" box="[151,846,1871,1893]" pageId="10" pageNumber="319">
<emphasis id="B927EA81FFF2657CC0E2F8D1B51EF8E1" bold="true" box="[151,274,1871,1893]" pageId="10" pageNumber="319">FIGURE 5.</emphasis>
Geographic distribution of 
<taxonomicName id="4C534D10FFF2657CC259F8CEB6FAF8E0" box="[556,758,1871,1893]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="319" phylum="Chordata" rank="species" species="bagual" status="sp. nov.">
<emphasis id="B927EA81FFF2657CC259F8CEB6FAF8E0" box="[556,758,1871,1893]" italics="true" pageId="10" pageNumber="319">Leposternon bagual</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457FAFFF2657CC288F8D1B742F8E1" box="[765,846,1871,1892]" pageId="10" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BEC3693FFF2657DC0B2F828B0A9FF51" blockId="10.[151,1437,1974,2035]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="320" pageId="10" pageNumber="319">
<emphasis id="B927EA81FFF2657CC0B2F828B68EF84A" bold="true" box="[199,642,1974,1999]" pageId="10" pageNumber="319">Geographic distribution and habitat.</emphasis>
So far, the new species is only known to occur in the 
<typeStatus id="54E88831FFF2657CC49FF829B116F84A" box="[1258,1306,1975,1999]" pageId="10" pageNumber="319">type</typeStatus>
locality, El 
<taxonomicName id="4C534D10FFF2657CC0E2F844B4EAF876" box="[151,230,2010,2035]" class="Squamata" family="Amphisbaenidae" genus="Leposternon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="321" phylum="Chordata" rank="species" species="bagual">Bagual</taxonomicName>
Ecological Reserve, located in the Province of 
<collectingCountry id="F3447603FFF2657CC35DF845B781F877" box="[808,909,2011,2034]" name="Taiwan" pageId="10" pageNumber="319">Formosa</collectingCountry>
, Laishi Department, 
<collectingCountry id="F3447603FFF2657CC4F2F844B0FBF876" box="[1159,1271,2010,2035]" name="Argentina" pageId="10" pageNumber="319">Argentina</collectingCountry>
(
<figureCitation id="13682A16FFF2657CC579F844B15AF876" box="[1292,1366,2010,2035]" captionStart="FIGURE 5" captionStartId="10.[151,250,1871,1893]" captionTargetBox="[168,1431,1074,1842]" captionTargetId="figure@10.[151,1436,1070,1850]" captionTargetPageId="10" captionText="FIGURE 5. Geographic distribution of Leposternon bagual sp. nov." httpUri="https://zenodo.org/record/242962/files/figure.png" pageId="10" pageNumber="319">Fig. 5</figureCitation>
). The Ecological Reserve has an extension of 3.336 ha within the fitogeographic region of the Humid Chaco, mainly dominated by floodplain pastures interspersed by forested areas (
<bibRefCitation id="EFC24B62FFF3657DC31BFF22B094FF51" author="Mercolli" box="[878,1176,188,213]" pageId="11" pageNumber="320" refString="Mercolli, C. &amp; Yanosky, A. A. (2001) Greater Rhea predation in the Eastern Chaco Argentino. Ararajuba, 9, 139 - 141." type="journal article" year="2001">Mercolli &amp; Yanosky 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>