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<document id="A82DCF26E65CE75D633156B8BA99A6A3" ID-DOI="10.11646/zootaxa.5297.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7993190" ID-ZooBank="454B72E3-2659-4F3D-9C9A-503DBABEE9CE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685602563737" checkinUser="plazi" docAuthor="Molineri, Carlos, Rodríguez, José Sebastián, Leiva, Marta &amp; Márquez, Javier Andrés" docDate="2023" docId="03E22F20A7672234FF1CF231FD78FBFA" docLanguage="en" docName="zootaxa.5297.2.4.pdf" docOrigin="Zootaxa 5297 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Progomphus complicatus Selys 1854" docType="treatment" docVersion="2" lastPageNumber="246" masterDocId="FFDB5758A7622233FF8BF37BFFDBFFEB" masterDocTitle="Diagnoses and key for the larvae of Progomphus Selys, 1854 from Argentina (Anisoptera: Gomphidae), with first larval descriptions for P. aberrans Belle, 1973 and P. kimminsi Belle, 1973" masterLastPageNumber="259" masterPageNumber="239" pageNumber="244" updateTime="1685603009843" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9A310CDACFA08379B93FC9CA0D4BE5CD">Diagnoses and key for the larvae of Progomphus Selys, 1854 from Argentina (Anisoptera: Gomphidae), with first larval descriptions for P. aberrans Belle, 1973 and P. kimminsi Belle, 1973</mods:title>
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<mods:namePart id="63998D1238FB5AB96230E789BDF23F4E">Molineri, Carlos</mods:namePart>
<mods:affiliation id="97F6449952FA267070B4834874EB9D64">Instituto de Biodiversidad Neotropical, CONICET-UNT (Consejo Nacional de Investigaciones Científicas y Técnicas-Universidad Nacional de Tucumán), Facultad de Ciencias Naturales e Instituto M. Lillo, Tucumán, Argentina.</mods:affiliation>
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<mods:namePart id="CFFC4FE6C74F01C010111EE19F6647BB">Rodríguez, José Sebastián</mods:namePart>
<mods:nameIdentifier id="168FFD62B5CCD906E9504008A9AA9AF9" type="ORCID">0000-0002-8917-2882</mods:nameIdentifier>
<mods:affiliation id="FDBE0E505CF092E4C83EE409AAFB87BC">Instituto de Biodiversidad Neotropical, CONICET-UNT (Consejo Nacional de Investigaciones Científicas y Técnicas-Universidad Nacional de Tucumán), Facultad de Ciencias Naturales e Instituto M. Lillo, Tucumán, Argentina. &amp; josephum @ hotmail. com; https: // orcid. org / 0000 - 0002 - 8917 - 2882</mods:affiliation>
<mods:nameIdentifier id="7A4894AEB71E494AAC5A397A671FF948" type="email">josephum@hotmail.com</mods:nameIdentifier>
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<mods:name id="2C778F5AF8C1C588455DEB69D260E5DC" type="personal">
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<mods:namePart id="D5AD53FDE0E610E2DB9E09A6204DFC3C">Leiva, Marta</mods:namePart>
<mods:affiliation id="13B0301EE87804ECA7C2960438ED070D">Facultad de Ciencias Forestales (FCF), Universidad Nacional de Santiago del Estero (UNSE), Avenida Belgrano (Sur) 1912, Santiago del Estero, Argentina.</mods:affiliation>
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<mods:namePart id="9DC29D6A4B2CD2F9915DCE432E77DED3">Márquez, Javier Andrés</mods:namePart>
<mods:affiliation id="D74BD59D224310D43A812DE4E21AC1A3">Departamento de Ciencias Naturales, Instituto de Ciencias de la Tierra, Biodiversidad y Ambiente (ICBIA), Universidad Nacional de Río Cuarto- CONICET, Córdoba, Argentina.</mods:affiliation>
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<mods:date id="50E3A4ECB70DDA4B001ACBEC54A3EC7D">2023</mods:date>
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<mods:number id="24BE0E6299E5EEC2F849FDF38078E982">2023-06-01</mods:number>
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<mods:identifier id="ECE5F2B4710F0CE56F0CFDC3E64FFFF7" type="DOI">10.11646/zootaxa.5297.2.4</mods:identifier>
<mods:identifier id="E2E74EAD90D86C7442272530E4F8AE46" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F99ACCA4712343FE45392C0EE9E0AB92" type="Zenodo-Dep">7993190</mods:identifier>
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<treatment id="03E22F20A7672234FF1CF231FD78FBFA" ID-DOI="http://doi.org/10.5281/zenodo.7993198" ID-Zenodo-Dep="7993198" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA" httpUri="http://treatment.plazi.org/id/03E22F20A7672234FF1CF231FD78FBFA" lastPageId="7" lastPageNumber="246" pageId="5" pageNumber="244">
<subSubSection id="C351CDBDA7672236FF1CF231FE30FDF2" pageId="5" pageNumber="244" type="nomenclature">
<paragraph id="8BF49E36A7672236FF1CF231FD3FFE8F" blockId="5.[151,740,330,357]" box="[151,740,330,357]" pageId="5" pageNumber="244">
<heading id="D0BC295AA7672236FF1CF231FD3FFE8F" bold="true" box="[151,740,330,357]" fontSize="11" level="1" pageId="5" pageNumber="244" reason="1">
<emphasis id="B93F4224A7672236FF1CF231FD3FFE8F" bold="true" box="[151,740,330,357]" pageId="5" pageNumber="244">
<taxonomicName id="4C4BE5B5A7672236FF1CF231FD80FE8F" ID-CoL="4MR44" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[151,603,330,357]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="244" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7672236FF1CF231FE11FE8E" bold="true" box="[151,458,330,357]" italics="true" pageId="5" pageNumber="244">Progomphus complicatus</emphasis>
<bibRefCitation id="EFDAE3C7A7672236FE59F231FD80FE8F" author="Selys, L. E. de" box="[466,603,330,356]" pageId="5" pageNumber="244" pagination="23 - 112" refId="ref11872" refString="Selys, L. E. de (1854) Synopsis des Gomphines. Bulletin de l'Academie royale des sciences de Belgique, 21 (2), 23 - 112." type="journal article" year="1854">Selys, 1854</bibRefCitation>
</taxonomicName>
(
<figureCitation id="137082B3A7672236FDE1F231FD07FE8F" box="[618,732,330,356]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[152,255,1945,1969]" captionStartId-1="6.[152,255,1943,1967]" captionTargetBox-0="[193,1401,924,1918]" captionTargetBox-1="[274,1313,181,1919]" captionTargetId-0="figure-282@5.[183,1404,921,1921]" captionTargetId-1="figure-18@6.[274,1313,181,1919]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." captionText-1="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi-0="http://doi.org/10.5281/zenodo.7993200" figureDoi-1="http://doi.org/10.5281/zenodo.7993202" httpUri-0="https://zenodo.org/record/7993200/files/figure.png" httpUri-1="https://zenodo.org/record/7993202/files/figure.png" pageId="5" pageNumber="244">Figs. 3–4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF49E36A7672236FF1CF2E8FE30FDF2" blockId="5.[151,1437,403,897]" pageId="5" pageNumber="244">
<materialsCitation id="3B23946BA7672236FF1CF2E8FAB1FE3A" collectingDate="2021-12-19" country="Argentina" county="Reared" elevation="668" latitude="-26.61046" location="rio Tapia" longLatPrecision="1" longitude="-65.26338" municipality="Tapia" pageId="5" pageNumber="244" specimenCount="3" specimenCount-female="3" stateProvince="Tucuman">
<collectingCounty id="6295E6BAA7672236FF1CF2E8FF3DFE46" box="[151,230,403,429]" pageId="5" pageNumber="244">Reared</collectingCounty>
material.
<collectingCountry id="F35CDEA6A7672236FED6F2E8FDDEFE46" box="[349,517,403,429]" name="Argentina" pageId="5" pageNumber="244">ARGENTINA</collectingCountry>
:
<specimenCount id="9D4D55BFA7672236FD98F2E8FDA3FE46" box="[531,632,403,429]" pageId="5" pageNumber="244" type="female">3 female</specimenCount>
adults with larval exuviae (IBN-O-1124, IBN-O-1300, IBN-O-1430) from
<collectingRegion id="498F50D4A7672236FF5CF2CCFE98FE3A" box="[215,323,439,465]" country="Argentina" name="Tucuman" pageId="5" pageNumber="244">Tucumán</collectingRegion>
,
<collectingMunicipality id="6B90044CA7672236FEDAF2CCFE49FE3A" box="[337,402,439,465]" pageId="5" pageNumber="244">Tapia</collectingMunicipality>
,
<location id="8E94C8EDA7672236FE2BF2CCFDD1FE3A" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FE2BF2CCFDD1FE3A" box="[416,522,439,465]" country="Argentina" county="Reared" latitude="-26.61046" longLatPrecision="1" longitude="-65.26338" municipality="Tapia" name="rio Tapia" pageId="5" pageNumber="244" stateProvince="Tucuman">río Tapia</location>
, puente RN9,
<geoCoordinate id="EE7FF8F1A7672236FD38F2CCFC99FE3A" box="[691,834,439,465]" degrees="26.61046" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-26.61046">26.61046° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FCD9F2CDFC30FE3A" box="[850,1003,438,465]" degrees="65.26338" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-65.26338">65.26338° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FC72F2CCFB9DFE3A" box="[1017,1094,439,465]" metricMagnitude="2" metricUnit="m" metricValue="6.68" pageId="5" pageNumber="244" unit="m" value="668.0">
<elevation id="00667905A7672236FC72F2CCFB9DFE3A" box="[1017,1094,439,465]" metricMagnitude="2" metricUnit="m" metricValue="6.68" pageId="5" pageNumber="244" unit="m" value="668.0">668 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FBDEF2CCFB3EFE3A" box="[1109,1253,439,465]" pageId="5" pageNumber="244" value="2021-12-19">
<collectingDate id="EFB1411EA7672236FBDEF2CCFB3EFE3A" box="[1109,1253,439,465]" pageId="5" pageNumber="244" value="2021-12-19">19/Dec/2021</collectingDate>
</date>
(collected)
</materialsCitation>
,
<materialsCitation id="3B23946BA7672236FAF3F2CCFEA2FE1E" collectingDate="2022-01-17" country="Argentina" county="Reared" elevation="668" isTailEnumeration="true" latitude="-26.61046" location="rio Tapia" longLatPrecision="1" longitude="-65.26338" municipality="Tapia" pageId="5" pageNumber="244" specimenCount="1" stateProvince="Tucuman">
<date id="FFF5B8F6A7672236FAF3F2CCFF26FE1E" pageId="5" pageNumber="244" value="2022-01-17">
<collectingDate id="EFB1411EA7672236FAF3F2CCFF26FE1E" pageId="5" pageNumber="244" value="2022-01-17">17/ Jan/2022</collectingDate>
</date>
(emerged)
</materialsCitation>
,
<materialsCitation id="3B23946BA7672236FE0FF2A0FE3DFDF2" collectingDate="2022-12-29" collectorName="C. Molineri &amp; G. Fontanarrosa" country="Argentina" county="Reared" elevation="668" isTailEnumeration="true" latitude="-26.61046" location="rio Tapia" longLatPrecision="1" longitude="-65.26338" municipality="Tapia" pageId="5" pageNumber="244" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman">
<collectorName id="26BEFBE0A7672236FE0FF2A0FDDDFE1E" box="[388,518,475,501]" pageId="5" pageNumber="244">C. Molineri</collectorName>
&amp;
<collectorName id="26BEFBE0A7672236FDA1F2A0FD05FE1E" box="[554,734,475,501]" pageId="5" pageNumber="244">G. Fontanarrosa</collectorName>
leg.;
<specimenCount id="9D4D55BFA7672236FC96F2A0FCB3FE1E" box="[797,872,475,501]" pageId="5" pageNumber="244" type="male">1 male</specimenCount>
adult with associated exuvia (IBN-O-1302), same data except date
<date id="FFF5B8F6A7672236FED8F284FE3DFDF2" box="[339,486,511,537]" pageId="5" pageNumber="244" value="2022-12-29">
<collectingDate id="EFB1411EA7672236FED8F284FE3DFDF2" box="[339,486,511,537]" pageId="5" pageNumber="244" value="2022-12-29">29/Dec/2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C351CDBDA7672234FF4CF158FD78FBFA" lastPageId="7" lastPageNumber="246" pageId="5" pageNumber="244" type="description">
<paragraph id="8BF49E36A7672236FF4CF158FBF7FC6A" blockId="5.[151,1437,403,897]" pageId="5" pageNumber="244">
Other material.
<materialsCitation id="3B23946BA7672236FEF0F158FB6EFD8A" collectingDate="2022-12-17" collectorName="C. Molineri &amp; J. S. Rodriguez" country="Argentina" elevation="784" latitude="-26.18948" location="Pozo del Pescado" longLatPrecision="1" longitude="-65.28417" municipality="El Boyero" pageId="5" pageNumber="244" specimenCount="2" specimenCount-larva="2" stateProvince="Tucuman">
<collectingCountry id="F35CDEA6A7672236FEF0F158FDF8FDD6" box="[379,547,547,573]" name="Argentina" pageId="5" pageNumber="244">ARGENTINA</collectingCountry>
:
<specimenCount id="9D4D55BFA7672236FDBBF158FD50FDD6" box="[560,651,547,573]" pageId="5" pageNumber="244" type="larva">2 larvae</specimenCount>
(IBN-O-92, IBN-O-93) from
<collectingRegion id="498F50D4A7672236FC6CF158FB88FDD6" box="[999,1107,547,573]" country="Argentina" name="Tucuman" pageId="5" pageNumber="244">Tucumán</collectingRegion>
,
<collectingMunicipality id="6B90044CA7672236FBEAF158FB03FDD6" box="[1121,1240,547,573]" pageId="5" pageNumber="244">El Boyero</collectingMunicipality>
, arroyo
<location id="8E94C8EDA7672236FAB2F158FF2DFD8A" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FAB2F158FF2DFD8A" country="Argentina" latitude="-26.18948" longLatPrecision="1" longitude="-65.28417" municipality="El Boyero" name="Pozo del Pescado" pageId="5" pageNumber="244" stateProvince="Tucuman">Pozo del Pescado</location>
,
<geoCoordinate id="EE7FF8F1A7672236FE89F13CFE54FD8A" box="[258,399,583,609]" degrees="26.18948" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-26.18948">26.18948° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FE17F13DFDE9FD8A" box="[412,562,582,609]" degrees="65.28417" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-65.28417">65.28417° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FDB5F13CFD5CFD8A" box="[574,647,583,609]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="5" pageNumber="244" unit="m" value="784.0">
<elevation id="00667905A7672236FDB5F13CFD5CFD8A" box="[574,647,583,609]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="5" pageNumber="244" unit="m" value="784.0">784 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FD1FF13CFCFCFD8A" box="[660,807,583,609]" pageId="5" pageNumber="244" value="2022-12-17">
<collectingDate id="EFB1411EA7672236FD1FF13CFCFCFD8A" box="[660,807,583,609]" pageId="5" pageNumber="244" value="2022-12-17">17/Dec/2022</collectingDate>
</date>
,
<collectorName id="26BEFBE0A7672236FCB8F13CFC6CFD8A" box="[819,951,583,609]" pageId="5" pageNumber="244">C. Molineri</collectorName>
&amp;
<collectorName id="26BEFBE0A7672236FC57F13CFB5AFD8A" box="[988,1153,583,609]" pageId="5" pageNumber="244">J.S. Rodríguez</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FB4AF13CFD66FD42" collectingDate="2014-11-17" collectorName="E. Dominguez &amp; C. Molineri" country="Argentina" county="Caimancito" elevation="360" latitude="-23.71166" location="Aguas Calientes" longLatPrecision="1" longitude="-64.53444" municipality="rio San Francisco" pageId="5" pageNumber="244" specimenCount="1" specimenCount-larva="1" stateProvince="Jujuy">
<specimenCount id="9D4D55BFA7672236FB4AF13CFAD4FD8A" box="[1217,1295,583,609]" pageId="5" pageNumber="244" type="larva">1 larva</specimenCount>
(IBN-O-91) from
<collectingRegion id="498F50D4A7672236FF5CF110FECEFD6E" box="[215,277,619,645]" country="Argentina" name="Jujuy" pageId="5" pageNumber="244">Jujuy</collectingRegion>
,
<collectingCounty id="6295E6BAA7672236FEA9F110FE7DFD6E" box="[290,422,619,645]" pageId="5" pageNumber="244">Caimancito</collectingCounty>
,
<collectingMunicipality id="6B90044CA7672236FE38F110FDA7FD6E" box="[435,636,619,645]" pageId="5" pageNumber="244">río San Francisco</collectingMunicipality>
(bajo puente de
<location id="8E94C8EDA7672236FCB5F110FC20FD6E" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FCB5F110FC20FD6E" box="[830,1019,619,645]" country="Argentina" county="Caimancito" latitude="-23.71166" longLatPrecision="1" longitude="-64.53444" municipality="rio San Francisco" name="Aguas Calientes" pageId="5" pageNumber="244" stateProvince="Jujuy">Aguas Calientes</location>
),
<geoCoordinate id="EE7FF8F1A7672236FB9BF110FB46FD6E" box="[1040,1181,619,645]" degrees="23.71166" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-23.71166">23.71166° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FB27F111FAE4FD6E" box="[1196,1343,618,645]" degrees="64.53444" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-64.53444">64.53444° w</geoCoordinate>
,
<quantity id="4CB333D3A7672236FAC7F110FA4CFD6E" box="[1356,1431,619,645]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="244" unit="m" value="360.0">
<elevation id="00667905A7672236FAC7F110FA4CFD6E" box="[1356,1431,619,645]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="244" unit="m" value="360.0">360 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FF1CF1F4FEF5FD42" box="[151,302,655,681]" pageId="5" pageNumber="244" value="2014-11-17">
<collectingDate id="EFB1411EA7672236FF1CF1F4FEF5FD42" box="[151,302,655,681]" pageId="5" pageNumber="244" value="2014-11-17">17/Nov/2014</collectingDate>
</date>
,
<collectorName id="26BEFBE0A7672236FEB1F1F4FE05FD42" box="[314,478,655,681]" pageId="5" pageNumber="244">E. Dominguez</collectorName>
&amp;
<collectorName id="26BEFBE0A7672236FD8FF1F4FD53FD42" box="[516,648,655,681]" pageId="5" pageNumber="244">C. Molineri</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FD41F1F4FBD7FD26" collectingDate="2014-08-07" collectorName="J. S. Rodriguez" country="Argentina" elevation="385" latitude="-23.77222" location="Rio Zora" longLatPrecision="1" longitude="-64.6125" municipality="Caimancito" pageId="5" pageNumber="244" specimenCount="6" specimenCount-larva="6" stateProvince="Jujuy">
<specimenCount id="9D4D55BFA7672236FD41F1F4FCFFFD42" box="[714,804,655,681]" pageId="5" pageNumber="244" type="larva">6 larvae</specimenCount>
(IBN-O-95) from
<collectingRegion id="498F50D4A7672236FC71F1F4FBE3FD42" box="[1018,1080,655,681]" country="Argentina" name="Jujuy" pageId="5" pageNumber="244">Jujuy</collectingRegion>
,
<collectingMunicipality id="6B90044CA7672236FBCFF1F4FB13FD42" box="[1092,1224,655,681]" pageId="5" pageNumber="244">Caimancito</collectingMunicipality>
,
<location id="8E94C8EDA7672236FB5EF1F4FAE0FD42" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FB5EF1F4FAE0FD42" box="[1237,1339,655,681]" country="Argentina" latitude="-23.77222" longLatPrecision="1" longitude="-64.6125" municipality="Caimancito" name="Rio Zora" pageId="5" pageNumber="244" stateProvince="Jujuy">Río Zora</location>
por ruta interna,
<geoCoordinate id="EE7FF8F1A7672236FF7EF1C8FE58FD26" box="[245,387,691,717]" degrees="23.77222" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-23.77222">23.77222° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FE1AF1C9FDF2FD26" box="[401,553,690,717]" degrees="64.61250" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-64.6125">64.61250° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FDBEF1C9FD5BFD26" box="[565,640,690,717]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="5" pageNumber="244" unit="m" value="385.0">
<elevation id="00667905A7672236FDBEF1C9FD5BFD26" box="[565,640,690,717]" metricMagnitude="2" metricUnit="m" metricValue="3.85" pageId="5" pageNumber="244" unit="m" value="385.0">385 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FD05F1C8FCFFFD26" box="[654,804,691,717]" pageId="5" pageNumber="244" value="2014-08-07">
<collectingDate id="EFB1411EA7672236FD05F1C8FCFFFD26" box="[654,804,691,717]" pageId="5" pageNumber="244" value="2014-08-07">07/Aug/2014</collectingDate>
</date>
,
<collectorName id="26BEFBE0A7672236FCBAF1C8FC03FD26" box="[817,984,691,717]" pageId="5" pageNumber="244">J.S. Rodríguez</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FB91F1C8FA5AFD26" box="[1050,1409,691,717]" collectingDate="2014-08-07" collectorName="J. S. Rodriguez" country="Argentina" elevation="385" latitude="-23.77222" location="Rio Zora" longLatPrecision="1" longitude="-64.6125" municipality="Caimancito" pageId="5" pageNumber="244" specimenCount="9" specimenCount-larva="9" stateProvince="Jujuy">
<specimenCount id="9D4D55BFA7672236FB91F1C8FBAEFD26" box="[1050,1141,691,717]" pageId="5" pageNumber="244" type="larva">9 larvae</specimenCount>
(IBN-O-94) same data
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FA05F1C9FF20FCFE" collectingDate="2006-07-28" country="Argentina" county="Acheral" elevation="359" latitude="-27.11722" location="Arroyo Hollinado" longLatPrecision="1" longitude="-65.45661" municipality="Citricola" pageId="5" pageNumber="244" specimenCount="5" specimenCount-larva="5" stateProvince="Tucuman">
<specimenCount id="9D4D55BFA7672236FA05F1C9FF00FD1A" pageId="5" pageNumber="244" type="larva">5 larvae</specimenCount>
(IBN-O-99) from
<collectingRegion id="498F50D4A7672236FE38F1ACFDC4FD1A" box="[435,543,727,753]" country="Argentina" name="Tucuman" pageId="5" pageNumber="244">Tucumán</collectingRegion>
,
<collectingCounty id="6295E6BAA7672236FDA1F1ACFD5DFD1A" box="[554,646,727,753]" pageId="5" pageNumber="244">Acheral</collectingCounty>
,
<collectingMunicipality id="6B90044CA7672236FD18F1ACFD23FD1A" box="[659,760,727,753]" pageId="5" pageNumber="244">Citrícola</collectingMunicipality>
,
<location id="8E94C8EDA7672236FC8FF1ACFC0AFD1A" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FC8FF1ACFC0AFD1A" box="[772,977,727,753]" country="Argentina" county="Acheral" latitude="-27.11722" longLatPrecision="1" longitude="-65.45661" municipality="Citricola" name="Arroyo Hollinado" pageId="5" pageNumber="244" stateProvince="Tucuman">Arroyo Hollinado</location>
,
<geoCoordinate id="EE7FF8F1A7672236FC55F1ACFBB0FD1A" box="[990,1131,727,753]" degrees="27.11722" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-27.11722">27.11722° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FBF1F1ADFAC9FD1A" box="[1146,1298,726,753]" degrees="65.45661" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-65.45661">65.45661° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FA94F1ADFAB1FD1A" box="[1311,1386,726,753]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="244" unit="m" value="359.0">
<elevation id="00667905A7672236FA94F1ADFAB1FD1A" box="[1311,1386,726,753]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="244" unit="m" value="359.0">359 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FAF3F1ACFF20FCFE" pageId="5" pageNumber="244" value="2006-07-28">
<collectingDate id="EFB1411EA7672236FAF3F1ACFF20FCFE" pageId="5" pageNumber="244" value="2006-07-28">28/ Jul/2006</collectingDate>
</date>
</materialsCitation>
, C. Molineri
<emphasis id="B93F4224A7672236FE07F180FE18FCFE" box="[396,451,763,789]" italics="true" pageId="5" pageNumber="244">et al.</emphasis>
leg.;
<materialsCitation id="3B23946BA7672236FD8BF181FC87FCFE" box="[512,860,762,789]" collectingDate="2006-07-28" country="Argentina" county="Acheral" elevation="359" latitude="-27.11722" location="Arroyo Hollinado" longLatPrecision="1" longitude="-65.45661" municipality="Citricola" pageId="5" pageNumber="244" specimenCount="5" specimenCount-larva="5" stateProvince="Tucuman">
<specimenCount id="9D4D55BFA7672236FD8BF181FD83FCFE" box="[512,600,762,789]" pageId="5" pageNumber="244" type="larva">5 larvae</specimenCount>
(IBN-O-97) same data
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FCEDF180FA4CFCFE" box="[870,1431,762,789]" collectingDate="2008-09-15" country="Argentina" county="Acheral" elevation="359" latitude="-27.11722" location="Arroyo Hollinado" longLatPrecision="1" longitude="-65.45661" municipality="Citricola" pageId="5" pageNumber="244" specimenCount="1" specimenCount-larva="1" stateProvince="Tucuman">
<specimenCount id="9D4D55BFA7672236FCEDF180FC69FCFE" box="[870,946,763,789]" pageId="5" pageNumber="244" type="larva">1 larva</specimenCount>
(IBN-O-96) same data except
<date id="FFF5B8F6A7672236FA8CF181FA4CFCFE" box="[1287,1431,762,789]" pageId="5" pageNumber="244" value="2008-09-15">
<collectingDate id="EFB1411EA7672236FA8CF181FA4CFCFE" box="[1287,1431,762,789]" pageId="5" pageNumber="244" value="2008-09-15">15/Sep/2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B23946BA7672236FF1CF064FA43FCD2" box="[151,1432,798,825]" collectingDate="2006-07-28" country="Argentina" elevation="359" latitude="-27.11643" location="rio Aranillas" longLatPrecision="1" longitude="-65.45904" municipality="Acheral" pageId="5" pageNumber="244" specimenCount="1" specimenCount-larva="1" stateProvince="Tucuman">
<specimenCount id="9D4D55BFA7672236FF1CF064FF3EFCD2" box="[151,229,799,825]" pageId="5" pageNumber="244" type="larva">1 larva</specimenCount>
(IBN-O-98) from
<collectingRegion id="498F50D4A7672236FE3CF064FDF8FCD2" box="[439,547,799,825]" country="Argentina" name="Tucuman" pageId="5" pageNumber="244">Tucumán</collectingRegion>
,
<collectingMunicipality id="6B90044CA7672236FDA7F064FD53FCD2" box="[556,648,799,825]" pageId="5" pageNumber="244">Acheral</collectingMunicipality>
, RP 38,
<location id="8E94C8EDA7672236FD6CF064FCAEFCD2" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FD6CF064FCAEFCD2" box="[743,885,799,825]" country="Argentina" latitude="-27.11643" longLatPrecision="1" longitude="-65.45904" municipality="Acheral" name="rio Aranillas" pageId="5" pageNumber="244" stateProvince="Tucuman">río Aranillas</location>
,
<geoCoordinate id="EE7FF8F1A7672236FC0AF064FBD6FCD2" box="[897,1037,799,825]" degrees="27.11643" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-27.11643">27.11643° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FB91F065FB6BFCD2" box="[1050,1200,798,825]" degrees="65.45904" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-65.45904">65.45904° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FB30F065FADEFCD2" box="[1211,1285,798,825]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="244" unit="m" value="359.0">
<elevation id="00667905A7672236FB30F065FADEFCD2" box="[1211,1285,798,825]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="244" unit="m" value="359.0">359 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FA9AF064FA43FCD2" box="[1297,1432,799,825]" pageId="5" pageNumber="244" value="2006-07-28">
<collectingDate id="EFB1411EA7672236FA9AF064FA43FCD2" box="[1297,1432,799,825]" pageId="5" pageNumber="244" value="2006-07-28">28/Jul/2006</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B23946BA7672236FF1CF038FEAFFCB6" box="[151,372,835,861]" country="Argentina" location="Molineri" pageId="5" pageNumber="244" specimenCount="1">
<location id="8E94C8EDA7672236FF1CF038FF22FCB6" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FF1CF038FF22FCB6" box="[151,249,835,861]" country="Argentina" name="Molineri" pageId="5" pageNumber="244">Molineri</location>
<emphasis id="B93F4224A7672236FE8FF038FE9AFCB6" box="[260,321,835,861]" italics="true" pageId="5" pageNumber="244">et al.</emphasis>
leg
</materialsCitation>
.;
<materialsCitation id="3B23946BA7672236FE03F038FBF7FC6A" collectingDate="2017-10-27" collectorName="C. Molineri &amp; J. Marquez &amp; J. S. Rodriguez" country="Argentina" elevation="784" latitude="-26.18948" location="Pozo del Pescado" longLatPrecision="1" longitude="-65.28417" municipality="El Boyero" pageId="5" pageNumber="244" specimenCount="2" specimenCount-larva="2" stateProvince="Tucuman">
<specimenCount id="9D4D55BFA7672236FE03F038FE3EFCB6" box="[392,485,835,861]" pageId="5" pageNumber="244" type="larva">2 larvae</specimenCount>
(IBN-O-100) from
<collectingRegion id="498F50D4A7672236FD5BF038FCE7FCB6" box="[720,828,835,861]" country="Argentina" name="Tucuman" pageId="5" pageNumber="244">Tucumán</collectingRegion>
,
<collectingMunicipality id="6B90044CA7672236FCC0F038FC1FFCB6" box="[843,964,835,861]" pageId="5" pageNumber="244">El Boyero</collectingMunicipality>
, arroyo
<location id="8E94C8EDA7672236FBA3F038FB2CFCB6" LSID="urn:lsid:plazi:treatment:03E22F20A7672234FF1CF231FD78FBFA:8E94C8EDA7672236FBA3F038FB2CFCB6" box="[1064,1271,835,861]" country="Argentina" latitude="-26.18948" longLatPrecision="1" longitude="-65.28417" municipality="El Boyero" name="Pozo del Pescado" pageId="5" pageNumber="244" stateProvince="Tucuman">Pozo del Pescado</location>
,
<geoCoordinate id="EE7FF8F1A7672236FA8DF038FA4CFCB6" box="[1286,1431,835,861]" degrees="26.18948" direction="south" orientation="latitude" pageId="5" pageNumber="244" precision="1" value="-26.18948">26.18948° S</geoCoordinate>
,
<geoCoordinate id="EE7FF8F1A7672236FF1CF01DFEF6FC6A" box="[151,301,870,897]" degrees="65.28417" direction="west" orientation="longitude" pageId="5" pageNumber="244" precision="1" value="-65.28417">65.28417° W</geoCoordinate>
,
<quantity id="4CB333D3A7672236FEB3F01CFE5AFC6A" box="[312,385,871,897]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="5" pageNumber="244" unit="m" value="784.0">
<elevation id="00667905A7672236FEB3F01CFE5AFC6A" box="[312,385,871,897]" metricMagnitude="2" metricUnit="m" metricValue="7.84" pageId="5" pageNumber="244" unit="m" value="784.0">784 m</elevation>
</quantity>
,
<date id="FFF5B8F6A7672236FE06F01CFDC7FC6A" box="[397,540,871,897]" pageId="5" pageNumber="244" value="2017-10-27">
<collectingDate id="EFB1411EA7672236FE06F01CFDC7FC6A" box="[397,540,871,897]" pageId="5" pageNumber="244" value="2017-10-27">27/Oct/2017</collectingDate>
</date>
,
<collectorName id="26BEFBE0A7672236FDACF01CFD77FC6A" box="[551,684,871,897]" pageId="5" pageNumber="244">C. Molineri</collectorName>
,
<collectorName id="26BEFBE0A7672236FD3CF01CFCEFFC6A" box="[695,820,871,897]" pageId="5" pageNumber="244">J. Márquez</collectorName>
&amp;
<collectorName id="26BEFBE0A7672236FCDCF01CFC27FC6A" box="[855,1020,871,897]" pageId="5" pageNumber="244">J.S. Rodríguez</collectorName>
leg.
</materialsCitation>
</paragraph>
<caption id="DF34CEBEA7672236FF13F4E2FE40F812" ID-DOI="http://doi.org/10.5281/zenodo.7993200" ID-Zenodo-Dep="7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="5" pageNumber="244" startId="5.[152,255,1945,1969]" targetBox="[193,1401,924,1918]" targetPageId="5" targetType="figure">
<paragraph id="8BF49E36A7672236FF13F4E2FE40F812" blockId="5.[151,1436,1945,2041]" pageId="5" pageNumber="244">
<emphasis id="B93F4224A7672236FF13F4E2FEC2F85A" bold="true" box="[152,281,1945,1969]" pageId="5" pageNumber="244">FIGURE 3.</emphasis>
<taxonomicName id="4C4BE5B5A7672236FEABF4E1FE2FF85A" authority="Selys" authorityName="Selys" authorityYear="1854" box="[288,500,1945,1969]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="244" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7672236FEABF4E1FE6FF85A" box="[288,436,1945,1969]" italics="true" pageId="5" pageNumber="244">P. complicatus</emphasis>
Selys
</taxonomicName>
, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view.
</paragraph>
</caption>
<caption id="DF34CEBEA7642235FF13F4ECFB91F838" ID-DOI="http://doi.org/10.5281/zenodo.7993202" ID-Zenodo-Dep="7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="6" pageNumber="245" startId="6.[152,255,1943,1967]" targetBox="[274,1313,181,1919]" targetPageId="6" targetType="figure">
<paragraph id="8BF49E36A7642235FF13F4ECFB91F838" blockId="6.[151,1436,1943,2003]" pageId="6" pageNumber="245">
<emphasis id="B93F4224A7642235FF13F4ECFEC1F844" bold="true" box="[152,282,1943,1967]" pageId="6" pageNumber="245">FIGURE 4.</emphasis>
<taxonomicName id="4C4BE5B5A7642235FEA9F4E3FE2CF844" authority="Selys" authorityName="Selys" authorityYear="1854" box="[290,503,1943,1967]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="245" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7642235FEA9F4E3FE6DF844" box="[290,438,1943,1967]" italics="true" pageId="6" pageNumber="245">P. complicatus</emphasis>
Selys
</taxonomicName>
, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4
<superScript id="7C3E337EA7642235FA1AF4ECFA47F84E" attach="none" box="[1425,1436,1943,1957]" fontSize="6" pageId="6" pageNumber="245">th</superScript>
antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral.
</paragraph>
</caption>
<paragraph id="8BF49E36A7652234FF4CF3ECFE60FD6E" blockId="7.[151,1437,151,1041]" pageId="7" pageNumber="246">
Larval diagnosis.
<taxonomicName id="4C4BE5B5A7652234FE19F3ECFD6BFF5A" authorityName="Selys" authorityYear="1854" box="[402,688,151,177]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7652234FE19F3ECFD6BFF5A" box="[402,688,151,177]" italics="true" pageId="7" pageNumber="246">Progomphus complicatus</emphasis>
</taxonomicName>
can be separated from other larvae in the genus by the following combination of characters: fourth antennomere relatively long and slender (
<figureCitation id="137082B3A7652234FB8AF3C0FB83FF3E" box="[1025,1112,187,213]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="7" pageNumber="246">Fig. 4b</figureCitation>
); mandibular formula (
<figureCitation id="137082B3A7652234FAEFF3C0FF14FF12" captionStart="FIGURE 3" captionStartId="5.[152,255,1945,1969]" captionTargetBox="[193,1401,924,1918]" captionTargetId="figure-282@5.[183,1404,921,1921]" captionTargetPageId="5" captionText="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="7" pageNumber="246">Figs. 3a–b</figureCitation>
) L 1 2 3 4 0 a (m 1,2,3, 4) b, R 1 2 3 4 y a (m 1,2) b; galeolacinia with 4 ventral teeth, the first one very reduced, shorter than ½ length of the second (
<figureCitation id="137082B3A7652234FD20F278FD25FEF6" box="[683,766,259,285]" captionStart="FIGURE 3" captionStartId="5.[152,255,1945,1969]" captionTargetBox="[193,1401,924,1918]" captionTargetId="figure-282@5.[183,1404,921,1921]" captionTargetPageId="5" captionText="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="7" pageNumber="246">Fig. 3c</figureCitation>
), palp acute apically; prementum subrectangular (
<figureCitation id="137082B3A7652234FAB5F278FA54FEF6" box="[1342,1423,259,285]" captionStart="FIGURE 3" captionStartId="5.[152,255,1945,1969]" captionTargetBox="[193,1401,924,1918]" captionTargetId="figure-282@5.[183,1404,921,1921]" captionTargetPageId="5" captionText="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="7" pageNumber="246">Fig. 3f</figureCitation>
), ligula without submedian tubercles, margin with two rows of flat setae, short and blunt on ventral row, much longer and acute with irregular margin in the dorsal row (
<figureCitation id="137082B3A7652234FD5FF230FCFCFE8E" box="[724,807,331,357]" captionStart="FIGURE 3" captionStartId="5.[152,255,1945,1969]" captionTargetBox="[193,1401,924,1918]" captionTargetId="figure-282@5.[183,1404,921,1921]" captionTargetPageId="5" captionText="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="7" pageNumber="246">Fig. 3e</figureCitation>
); lateral margins of prementum with 4 short and 1 long spines submedially, and 1 short spine apically (
<figureCitation id="137082B3A7652234FD3FF214FCDEFE62" box="[692,773,367,393]" captionStart="FIGURE 3" captionStartId="5.[152,255,1945,1969]" captionTargetBox="[193,1401,924,1918]" captionTargetId="figure-282@5.[183,1404,921,1921]" captionTargetPageId="5" captionText="FIGURE 3. P. complicatus Selys, female last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, detail of flat marginal setae on ligula; f, prementum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7993200" httpUri="https://zenodo.org/record/7993200/files/figure.png" pageId="7" pageNumber="246">Fig. 3f</figureCitation>
); labial palp with smooth margins, inner apical projection subtriangular reaching basal 1/3 of movable hook, movable hook stout slightly shorter to outer palpal margin; fore claws cheliform, meso- and hindtarsal claws normal (long and acutely pointed, flattened), hind claw with a subapical seta; thorax and legs with blackish setae; without tibial distal spurs; hind tarsi darker than the rest of the leg; posterolateral spines present on S5–9 (
<figureCitation id="137082B3A7652234FDF0F284FD0BFDF2" box="[635,720,511,537]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="7" pageNumber="246">Fig. 4d</figureCitation>
); relatively large acute dorsal tubercles on terga S2–9 (
<figureCitation id="137082B3A7652234FAB7F284FA54FDF2" box="[1340,1423,511,537]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="7" pageNumber="246">Fig. 4e</figureCitation>
); sterna S2–8 formed by three plates (a large central one, and two smaller laterally), sterna S1 and S9–10 formed by one plate (
<figureCitation id="137082B3A7652234FE87F13CFEBAFD8A" box="[268,353,583,609]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="7" pageNumber="246">Fig. 4d</figureCitation>
); apical abdominal segments and anal pyramid elongated, relative length of sternite S8: S9: S10: epiproct = 1.1: 1.1: 1: 1.1.
</paragraph>
<paragraph id="8BF49E36A7652234FF4CF1F4FE6BFC22" blockId="7.[151,1437,151,1041]" pageId="7" pageNumber="246">
<bibRefCitation id="EFDAE3C7A7652234FF4CF1F4FEBEFD42" author="Santos, N. D." box="[199,357,655,681]" pageId="7" pageNumber="246" pagination="171 - 174" refId="ref11810" refString="Santos, N. D. (1968) Contribucao ao conhecimento da fauna do Estado da Guanabara. 61. Notas sobre a ninfa de Progomphus complicatus (?) Selys, 1854 e seu imago (Gomphidae, Odonata). Atas da Sociedade de Biologia de Rio de Janeiro, 11 (5), 171 - 174." type="journal article" year="1968">Santos (1968)</bibRefCitation>
described the larval stage of
<taxonomicName id="4C4BE5B5A7652234FD3CF1F4FC80FD42" authorityName="Selys" authorityYear="1854" box="[695,859,655,681]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7652234FD3CF1F4FC80FD42" box="[695,859,655,681]" italics="true" pageId="7" pageNumber="246">P. complicatus</emphasis>
</taxonomicName>
, but association of adults was only by supposition (
<emphasis id="B93F4224A7652234FF2AF1C8FF67FD27" box="[161,188,691,716]" italics="true" pageId="7" pageNumber="246">i.e</emphasis>
., shared locality). We have reared
<specimenCount id="9D4D55BFA7652234FDD6F1C8FD15FD26" box="[605,718,691,717]" pageId="7" pageNumber="246" type="female">3 females</specimenCount>
and
<specimenCount id="9D4D55BFA7652234FC84F1C8FC84FD26" box="[783,863,691,717]" pageId="7" pageNumber="246" type="male">1 male</specimenCount>
adults from larva that confirm the larval—adult association. We provide above some data and illustrations, not mentioned in the original description that now have proven to be useful to distinguish the species (
<emphasis id="B93F4224A7652234FD2BF180FD1AFCFF" box="[672,705,763,788]" italics="true" pageId="7" pageNumber="246">e.g</emphasis>
., mandibular formula, shape and arrangement of maxillary teeth, and other details of abdomen). Among the Argentinean species,
<taxonomicName id="4C4BE5B5A7652234FC09F064FBFCFCD2" authorityName="Selys" authorityYear="1854" box="[898,1063,799,825]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7652234FC09F064FBFCFCD2" box="[898,1063,799,825]" italics="true" pageId="7" pageNumber="246">P. complicatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C4BE5B5A7652234FB4CF064FA90FCD2" authorityName="Belle" authorityYear="1973" box="[1223,1355,799,825]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B93F4224A7652234FB4CF064FA90FCD2" box="[1223,1355,799,825]" italics="true" pageId="7" pageNumber="246">P. aberrans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4BE5B5A7652234FA03F064FF27FCB6" authorityName="Belle" authorityYear="1973" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="kimminsi">
<emphasis id="B93F4224A7652234FA03F064FF27FCB6" italics="true" pageId="7" pageNumber="246">P. kimminsi</emphasis>
</taxonomicName>
, because all of them present a slender, subcylindrical distal antennomere, absence of tibial spurs, and S5 with posterolateral spine. Nevertheless, the larva of
<taxonomicName id="4C4BE5B5A7652234FD6DF01CFC52FC6A" authorityName="Selys" authorityYear="1854" box="[742,905,871,897]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="246" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7652234FD6DF01CFC52FC6A" box="[742,905,871,897]" italics="true" pageId="7" pageNumber="246">P. complicatus</emphasis>
</taxonomicName>
can be easily separated from them by its short anal pyramid (vs long and slender in the other two species), and by the absence of posterolateral spine on S4 (present in the other two species).
</paragraph>
<paragraph id="8BF49E36A7652234FF4CF0A8FD78FBFA" blockId="7.[151,1437,151,1041]" pageId="7" pageNumber="246">
Distribution:
<collectingCountry id="F35CDEA6A7652234FED5F0A8FE16FC06" box="[350,461,979,1005]" name="Argentina" pageId="7" pageNumber="246">Argentina</collectingCountry>
(
<collectingRegion id="498F50D4A7652234FE54F0A8FD80FC06" box="[479,603,979,1005]" country="Argentina" name="Catamarca" pageId="7" pageNumber="246">Catamarca</collectingRegion>
,
<collectingRegion id="498F50D4A7652234FDE2F0A8FD7CFC06" box="[617,679,979,1005]" country="Argentina" name="Jujuy" pageId="7" pageNumber="246">Jujuy</collectingRegion>
, Misiones,
<collectingRegion id="498F50D4A7652234FCA0F0A8FCBDFC06" box="[811,870,979,1005]" country="Argentina" name="Salta" pageId="7" pageNumber="246">Salta</collectingRegion>
,
<collectingRegion id="498F50D4A7652234FCF8F0A8FC04FC06" box="[883,991,979,1005]" country="Argentina" name="Tucuman" pageId="7" pageNumber="246">Tucumán</collectingRegion>
,
<bibRefCitation id="EFDAE3C7A7652234FC67F0A8FB1CFC06" author="Lozano, F. &amp; Palacio, A. &amp; Ramos, L. &amp; Muzon, J." box="[1004,1223,979,1005]" pageId="7" pageNumber="246" pagination="113 - 153" refId="ref11512" refString="Lozano, F., del Palacio, A., Ramos, L. &amp; Muzon, J. (2020) The Odonata of Argentina: state of knowledge and updated checklist. International Journal of Odonatology, 23 (2), 113 - 153. https: // doi. org / 10.1080 / 13887890.2020.1737585" type="journal article" year="2020">
Lozano
<emphasis id="B93F4224A7652234FBC2F0A8FB5FFC06" box="[1097,1156,979,1005]" italics="true" pageId="7" pageNumber="246">et al.</emphasis>
2020
</bibRefCitation>
) and
<collectingCountry id="F35CDEA6A7652234FA81F0A8FA95FC06" box="[1290,1358,979,1005]" name="Brazil" pageId="7" pageNumber="246">Brazil</collectingCountry>
(
<bibRefCitation id="EFDAE3C7A7652234FAEBF0A8FF09FBFA" author="Belle, J." pageId="7" pageNumber="246" pagination="191 - 308" refId="ref11040" refString="Belle, J. (1973) A revision of the new world genus Progomphus Selys, 1854 (Anisoptera: Gomphidae). Odonatologica, 2 (4), 191 - 308." type="journal article" year="1973">Belle 1973</bibRefCitation>
;
<bibRefCitation id="EFDAE3C7A7652234FF57F08CFE65FBFA" author="Almeida, M. V. &amp; Pinto, A. P. &amp; Carvalho, A. D. &amp; Takiya, D. M." box="[220,446,1015,1041]" pageId="7" pageNumber="246" pagination="417 - 423" refId="ref10949" refString="Almeida, M. V., Pinto, A. P., Carvalho, A. D. &amp; Takiya, D. M. (2013) When rare is just a matter of sampling: unexpected dominance of clubtail dragonflies (Odonata, Gomphidae) through different collecting methods at Parque Nacional da Serra do Cipo, Minas Gerais State, Brazil. Revista brasileira de Entomologia, 57, 417 - 423. https: // doi. org / 10.1590 / S 0085 - 56262013005000042" type="journal article" year="2013">
Almeida
<emphasis id="B93F4224A7652234FECFF08CFEA7FBFA" box="[324,380,1015,1041]" italics="true" pageId="7" pageNumber="246">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFDAE3C7A7652234FE41F08CFD4CFBFA" author="Garcia Junior, M. D. &amp; Rakes, M. &amp; Pazini, J. D. &amp; Pasini, R. A. &amp; Garcia, F. R. &amp; Grutzmacher, A. D." box="[458,663,1015,1041]" pageId="7" pageNumber="246" pagination="107 - 117" refId="ref11197" refString="Garcia Junior, M. D., Rakes, M., Pazini, J. D., Pasini, R. A., Garcia, F. R. &amp; Grutzmacher, A. D. (2019) The diversity of Odonata adults's at Pampa Biome from Brazil. Revista de Biologia Tropical, 67 (1), 107 - 117. https: // doi. org / 10.15517 / rbt. v 67 i 1.33285" type="journal article" year="2019">
García
<emphasis id="B93F4224A7652234FD90F08CFD8FFBFA" box="[539,596,1015,1041]" italics="true" pageId="7" pageNumber="246">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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