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<document id="64E90DA5E7A6CAA1BAFA2B647B9D63A2" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. &amp; Colville, Jonathan F." docDate="2020" docId="03DFBE05E020FFCAFF64D10DFEB1FB2C" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia nigrosetosa Dombrow 2020, new species" docType="treatment" docUuid="2C34F785-38C8-4265-B59D-DE94C9023A05" docUuidSource="ZooBank" docVersion="9" lastPageNumber="64" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="61" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:title id="C1210C389399720B20E462B0C4BCF12B">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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<mods:namePart id="1DF322429048D5FB48670E4FE373B0E0">Dombrow, Holger E.</mods:namePart>
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<mods:namePart id="6849F2CF11DC712A4D35B70946E16404">Colville, Jonathan F.</mods:namePart>
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<mods:date id="02FDD48C2F5FEE5E167DB809EE777283">2020</mods:date>
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<heading id="D081B87FE020FFC9FF64D10DFD6EFABD" bold="true" box="[151,723,1302,1329]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="4">
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<taxonomicName id="4C767490E020FFC9FF64D10DFD8BFABC" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,566,1302,1329]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrosetosa" status="sp. nov.">
<emphasis id="B902D301E020FFC9FF64D10DFE0FFABC" bold="true" box="[151,434,1302,1329]" italics="true" pageId="60" pageNumber="61">Beckhoplia nigrosetosa</emphasis>
Dombrow
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,
<taxonomicNameLabel id="A2316E7AE020FFC9FDB0D10CFD6EFABD" box="[579,723,1302,1329]" pageId="60" pageNumber="61" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C36C5C98E020FFC9FF64D120FCE5FAF5" pageId="60" pageNumber="61" type="description">
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(
<figureCitation id="134D1396E020FFC9FF6CD120FF52FAD8" box="[159,239,1339,1365]" captionStart="FIGURES" captionStartId="10.[151,264,1152,1176]" captionTargetBox="[157,1431,969,1123]" captionTargetId="figure-102@10.[151,1436,964,1129]" captionTargetPageId="10" captionText="FIGURES. 67. General drawings of the reflexion of the anterior margin of the clypeus of Beckhoplia Dombrow, 2005 species, in lateral view. 6, distinctly reflexed, ♂♂; 7, less distinct reflexed, ♀♀. Scale bar 1 mm ." figureDoi="http://doi.org/10.5281/zenodo.4401922" httpUri="https://zenodo.org/record/4401922/files/figure.png" pageId="60" pageNumber="61">Figs. 7</figureCitation>
,
<figureCitation id="134D1396E020FFC9FF08D120FEA4FAD8" box="[251,281,1339,1365]" captionStart="FIGURES 1114" captionStartId="10.[151,264,1818,1842]" captionTargetBox="[169,1417,1580,1789]" captionTargetId="figure-185@10.[164,1423,1574,1794]" captionTargetPageId="10" captionText="FIGURES 1114. General drawings of the eye canthus of Hopliini species in lateral view ♂♂, otherwise indicated. 11, short (Nanniscus Burmeister, 1844) Scale bar 1 mm; 12, long (Anisochelus Burmeister, 1844); 13, very long (Beckhoplia Dombrow, 2005); 14, full (Beckhoplia Dombrow, 2005, ♀♀). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401926" httpUri="https://zenodo.org/record/4401926/files/figure.png" pageId="60" pageNumber="61">13</figureCitation>
,
<figureCitation id="134D1396E020FFC9FED6D120FED0FAD8" box="[293,365,1339,1365]" captionStart="FIGURES 2328" captionStartId="16.[151,264,1470,1494]" captionTargetBox="[193,1392,1182,1406]" captionTargetId="figure-130@16.[189,1398,1177,1410]" captionTargetPageId="16" captionText="FIGURES 2328. General drawings of the pronotum and scutellum of Beckhoplia Dombrow, 2005 species, in lateral view ♂♂. Pronotum: 23, posterior corner obtuse, angled; 24, rounded. Scutellum: 25, posterior corner obtuse, angled; 26, rounded; 27, tip pointed; 28, tip rounded. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401932" httpUri="https://zenodo.org/record/4401932/files/figure.png" pageId="60" pageNumber="61">26, 28</figureCitation>
,
<figureCitation id="134D1396E020FFC9FE8AD121FE2AFAD8" box="[377,407,1338,1365]" captionStart="FIGURES 5558" captionStartId="24.[151,264,1387,1411]" captionTargetBox="[155,1432,874,1323]" captionTargetId="figure-95@24.[151,1436,870,1327]" captionTargetPageId="24" captionText="FIGURES 5558. General drawing of the types of mesothoracic pretarsomere of Beckhoplia Dombrow, 2005 species, in lateral view showing the distinctly thickened, roundish cutting missing ventrally, ♂♂, otherwise indicated. 55, with a shortened smooth ridge, with roundish cutting; 56, with a shortened granulate ridge; 57, slender, with a full undulate ridge, ♀♀; 58, with a shortened smooth ridge. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401950" httpUri="https://zenodo.org/record/4401950/files/figure.png" pageId="60" pageNumber="61">56</figureCitation>
,
<figureCitation id="134D1396E020FFC9FE50D120FE56FAD8" box="[419,491,1339,1365]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="60" pageNumber="61">9091</figureCitation>
,
<figureCitation id="134D1396E020FFC9FE04D120FD9CFAD8" box="[503,545,1339,1365]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="60" pageNumber="61">114</figureCitation>
)
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http://zoobank.org/
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<treatmentCitationGroup id="AB66283DE020FFC9FF64D1BEFD23FA31" box="[151,670,1445,1469]" pageId="60" pageNumber="61">
<treatmentCitation id="0AD72902E020FFC9FF64D1BEFD82FA30" author="Dombrow, H." box="[151,575,1445,1469]" page="128" pageId="60" pageNumber="61" year="2005">
<taxonomicName id="4C767490E020FFC9FF64D1BEFD82FA30" authority="Dombrow, 2005: 128" authorityName="Dombrow" authorityPageNumber="128" authorityYear="2005" box="[151,575,1445,1469]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E020FFC9FF64D1BEFEDCFA30" box="[151,353,1445,1469]" italics="true" pageId="60" pageNumber="61">Beckhoplia colvillei</emphasis>
<bibRefCitation id="EFE772E2E020FFC9FE94D1BEFD82FA30" author="Dombrow, H." box="[359,575,1445,1469]" pageId="60" pageNumber="61" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005: 128</bibRefCitation>
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ex parte.
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<materialsCitation id="3B1E054EE020FFC9FF64D1F0FCCFF98B" ID-GBIF-Occurrence="3022772302" box="[151,882,1514,1542]" collectionCode="NCP" country="South Africa" location="Nieuwoudtville Wildflower Reserve" pageId="60" pageNumber="61" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CDB1B1E020FFC9FF64D1F0FF6FF988" box="[151,210,1515,1541]" pageId="60" pageNumber="61">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F3614F83E020FFC9FEB0D1F0FEC2F988" box="[323,383,1515,1541]" name="South Africa" pageId="60" pageNumber="61">RSA</collectingCountry>
,
<collectionCode id="ED6797D6E020FFC9FE79D1F1FE7AF988" box="[394,455,1514,1541]" country="United Kingdom" name="National Collection of Passiflora" pageId="60" pageNumber="61" type="Herbarium">NCP</collectionCode>
]:
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.
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<emphasis id="B902D301E020FFC9FF64D228FEF9F9C0" bold="true" box="[151,324,1587,1613]" pageId="60" pageNumber="61">Type material.</emphasis>
[269]
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<typeStatus id="54CDB1B1E020FFC9FE7FD228FE44F9C0" box="[396,505,1587,1613]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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: white label pr «
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,
<collectionCode id="ED6797D6E020FFC9FCE2D228FCFAF9C0" box="[785,839,1587,1613]" country="United Kingdom" name="National Collection of Passiflora" pageId="60" pageNumber="61" type="Herbarium">NCP</collectionCode>
, Nieuwoudt- | ville, Wildflower Res. |
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| 22/08/98, J. Colville» || yellow label hw «18» pr «.534 | coll. Dombrow» || red label pr «
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|
<taxonomicName id="4C767490E020FFC9FAE3D24CFA33F9FC" authorityName="Dombrow" authorityYear="2005" box="[1296,1422,1623,1649]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E020FFC9FAE3D24CFA33F9FC" box="[1296,1422,1623,1649]" italics="true" pageId="60" pageNumber="61">Beckhoplia</emphasis>
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|
<taxonomicName id="4C767490E020FFC9FF64D260FF4CF918" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[151,241,1659,1685]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
<emphasis id="B902D301E020FFC9FF64D260FF4CF918" box="[151,241,1659,1685]" italics="true" pageId="60" pageNumber="61">colvillei</emphasis>
</taxonomicName>
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| det.: Dombrow 2003» || green label pr «Host plant: |
<emphasis id="B902D301E020FFC9FC68D260FAB8F918" box="[923,1285,1659,1685]" italics="true" pageId="60" pageNumber="61">
<taxonomicName id="4C767490E020FFC9FC68D260FBFBF918" authorityName="Roessler" authorityYear="1959" baseAuthorityName="Burm. fil." box="[923,1094,1659,1685]" class="Magnoliopsida" family="Asteraceae" genus="Gazania" kingdom="Plantae" order="Asterales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="rigida">Gazania rigida</taxonomicName>
|
<taxonomicName id="4C767490E020FFC9FBADD260FAB8F918" box="[1118,1285,1659,1685]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="erosus">Senecio erosus</taxonomicName>
</emphasis>
» || blue label pr «Bioregion: | Shale Renosterveld | Vegetation Type: FRs2 | Nieuwoudtville Shale Renosterveld» || yellow label pr «SAMCOL | A071219» || red label pr «
<typeStatus id="54CDB1B1E020FFC9FD20D2D8FC80F950" box="[723,829,1731,1757]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
|
<taxonomicName id="4C767490E020FFC9FCA4D2D8FC68F950" authorityName="Dombrow" authorityYear="2005" box="[855,981,1731,1757]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E020FFC9FCA4D2D8FC68F950" box="[855,981,1731,1757]" italics="true" pageId="60" pageNumber="61">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E020FFC9FC1CD2DFFBCCF950" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1007,1137,1732,1757]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrosetosa" status="sp. nov.">
<emphasis id="B902D301E020FFC9FC1CD2DFFBCCF950" box="[1007,1137,1732,1757]" italics="true" pageId="60" pageNumber="61">nigrosetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE020FFC9FB88D2DFFB0DF953" box="[1147,1200,1732,1758]" pageId="60" pageNumber="61" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E020FFC9FF1ED2FCFEFEF88C" box="[237,323,1767,1793]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="60" pageNumber="61" type="Museum">SAMC</collectionCode>
).
<emphasis id="B902D301E020FFC9FE99D2FCFE5FF88C" bold="true" box="[362,482,1767,1793]" pageId="60" pageNumber="61">
<typeStatus id="54CDB1B1E020FFC9FE99D2FCFE5FF88C" box="[362,482,1767,1793]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE020FFC9FE1AD2FCFDFAF88F" box="[489,583,1767,1794]" pageId="60" pageNumber="61" type="male">179 ♂♂</specimenCount>
,
<specimenCount id="9D70C49AE020FFC9FDA0D2FCFD1FF88F" box="[595,674,1767,1794]" pageId="60" pageNumber="61" type="female">40 ♀♀</specimenCount>
: same labels as
<typeStatus id="54CDB1B1E020FFC9FCA4D2FCFC04F88C" box="[855,953,1767,1793]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
but || yellow label hw «18» pr «.535.713,.734.773 | coll. Dombrow» || red label pr «
<typeStatus id="54CDB1B1E020FFC9FD88D317FD62F8AB" box="[635,735,1804,1830]" pageId="60" pageNumber="61" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C767490E020FFC9FD00D310FCCCF8A8" authorityName="Dombrow" authorityYear="2005" box="[755,881,1803,1829]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E020FFC9FD00D310FCCCF8A8" box="[755,881,1803,1829]" italics="true" pageId="60" pageNumber="61">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E020FFC9FC77D310FC63F8A8" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[900,990,1803,1829]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
<emphasis id="B902D301E020FFC9FC77D310FC63F8A8" box="[900,990,1803,1829]" italics="true" pageId="60" pageNumber="61">colvillei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE020FFC9FC16D317FBA7F8AB" box="[997,1050,1804,1830]" pageId="60" pageNumber="61" rank="species">sp.n.</taxonomicNameLabel>
| det.: Dombrow 2003» || red label pr «
<typeStatus id="54CDB1B1E020FFC9FF30D32BFE9AF8C7" box="[195,295,1840,1866]" pageId="60" pageNumber="61" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C767490E020FFC9FECDD334FE01F8C4" authorityName="Dombrow" authorityYear="2005" box="[318,444,1839,1865]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E020FFC9FECDD334FE01F8C4" box="[318,444,1839,1865]" italics="true" pageId="60" pageNumber="61">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E020FFC9FE20D32BFDE8F8C4" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[467,597,1840,1865]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrosetosa" status="sp. nov.">
<emphasis id="B902D301E020FFC9FE20D32BFDE8F8C4" box="[467,597,1840,1865]" italics="true" pageId="60" pageNumber="61">nigrosetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE020FFC9FDAED32BFD2FF8C7" box="[605,658,1840,1866]" pageId="60" pageNumber="61" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E020FFC9FC2BD334FB93F8C4" box="[984,1070,1839,1865]" pageId="60" pageNumber="61">ESDW</collectionCode>
);
<specimenCount id="9D70C49AE020FFC9FBB1D334FB2CF8C7" box="[1090,1169,1839,1866]" pageId="60" pageNumber="61" type="male">39 ♂♂</specimenCount>
<specimenCount id="9D70C49AE020FFC9FB6AD334FB57F8C7" box="[1177,1258,1839,1866]" pageId="60" pageNumber="61" type="female">10 ♀♀</specimenCount>
: same labels as
<typeStatus id="54CDB1B1E020FFC9FF64D348FF44F8E0" box="[151,249,1875,1901]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
but || yellow label hw «55» pr «.708.756 | coll. Dombrow» || yellow label pr «SAMCOL | A071219» || red label pr «
<typeStatus id="54CDB1B1E020FFC9FEDAD363FE30F81F" box="[297,397,1912,1938]" pageId="60" pageNumber="61" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C767490E020FFC9FE52D36CFDA2F81C" authorityName="Dombrow" authorityYear="2005" box="[417,543,1911,1937]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E020FFC9FE52D36CFDA2F81C" box="[417,543,1911,1937]" italics="true" pageId="60" pageNumber="61">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E020FFC9FDC0D363FD08F81C" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[563,693,1912,1937]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrosetosa" status="sp. nov.">
<emphasis id="B902D301E020FFC9FDC0D363FD08F81C" box="[563,693,1912,1937]" italics="true" pageId="60" pageNumber="61">nigrosetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE020FFC9FD48D363FD4DF81F" box="[699,752,1912,1938]" pageId="60" pageNumber="61" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E020FFC9FBDFD36CFB3FF81C" box="[1068,1154,1911,1937]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="60" pageNumber="61" type="Museum">SAMC</collectionCode>
).
</paragraph>
<paragraph id="8BC90F13E020FFC9FF64D3A4FAAEF854" blockId="60.[151,1299,1983,2010]" box="[151,1299,1983,2010]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF64D3A4FE1FF854" bold="true" box="[151,418,1983,2009]" pageId="60" pageNumber="61">
Condition of
<typeStatus id="54CDB1B1E020FFC9FEC7D3A4FE20F854" box="[308,413,1983,2009]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
.
</emphasis>
Very good. Pinned. Dissected, parameres glued on separate card below beetle.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E021FFC8FF64D48DFCDCFE20" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="8BC90F13E021FFC8FF64D48DFCDCFE20" blockId="61.[151,1437,150,429]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF64D48DFE22FF3D" bold="true" box="[151,415,150,176]" pageId="61" pageNumber="62">Differential diagnosis.</emphasis>
Differs from
<taxonomicName id="4C767490E021FFC8FDCCD48CFCABFF3C" authority=": Elytra" authorityName="Elytra" box="[575,790,151,177]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FDCCD48CFD7FFF3C" box="[575,706,151,177]" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
: Elytra
</taxonomicName>
of
<taxonomicName id="4C767490E021FFC8FCCCD48CFC5FFF3D" authorityName="Dombrow" authorityYear="2020" box="[831,994,151,176]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E021FFC8FCCCD48CFC5FFF3D" box="[831,994,151,176]" italics="true" pageId="61" pageNumber="62">B. nigrosetosa</emphasis>
</taxonomicName>
widest medially. Elytra of
<taxonomicName id="4C767490E021FFC8FAEAD48CFA21FF3C" box="[1305,1436,151,177]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FAEAD48CFA21FF3C" box="[1305,1436,151,177]" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
</taxonomicName>
widest in anterior third. Lateral margin of elytra of
<taxonomicName id="4C767490E021FFC8FD23D4A0FCCCFF59" authorityName="Dombrow" authorityYear="2020" box="[720,881,187,212]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E021FFC8FD23D4A0FCCCFF59" box="[720,881,187,212]" italics="true" pageId="61" pageNumber="62">B. nigrosetosa</emphasis>
</taxonomicName>
narrowly darkened. Lateral margin of elytra of
<taxonomicName id="4C767490E021FFC8FA77D4A0FF44FF74" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FA77D4A0FF44FF74" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
</taxonomicName>
broadly blackened. Interno-posterior margin of parameres of
<taxonomicName id="4C767490E021FFC8FC59D4C4FBF7FF75" authorityName="Dombrow" authorityYear="2020" box="[938,1098,223,248]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E021FFC8FC59D4C4FBF7FF75" box="[938,1098,223,248]" italics="true" pageId="61" pageNumber="62">B. nigrosetosa</emphasis>
</taxonomicName>
parabola in dorsal view (
<figureCitation id="134D1396E021FFC8FA97D4C4FF0BFE90" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 90</figureCitation>
,
<figureCitation id="134D1396E021FFC8FF33D518FF50FE90" box="[192,237,259,285]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">114</figureCitation>
). Interno-posterior margin of parameres of
<taxonomicName id="4C767490E021FFC8FD3DD518FCF3FE90" box="[718,846,259,285]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FD3DD518FCF3FE90" box="[718,846,259,285]" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
</taxonomicName>
rounded in dorsal view (
<figureCitation id="134D1396E021FFC8FB90D518FB02FE90" box="[1123,1215,259,285]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 88</figureCitation>
,
<figureCitation id="134D1396E021FFC8FB39D518FB4AFE90" box="[1226,1271,259,285]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">113</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E021FFC8FE1AD53CFD37FECD" authorityName="Dombrow" authorityYear="2020" box="[489,650,295,320]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E021FFC8FE1AD53CFD37FECD" box="[489,650,295,320]" italics="true" pageId="61" pageNumber="62">B. nigrosetosa</emphasis>
</taxonomicName>
linear with a convex median extension in dorsal view (
<figureCitation id="134D1396E021FFC8FB0AD53CFAE5FECC" box="[1273,1368,295,321]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 90</figureCitation>
,
<figureCitation id="134D1396E021FFC8FA90D53CFA2DFECC" box="[1379,1424,295,321]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">114</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E021FFC8FD88D550FD43FEE8" box="[635,766,331,357]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FD88D550FD43FEE8" box="[635,766,331,357]" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
</taxonomicName>
convex in dorsal view (
<figureCitation id="134D1396E021FFC8FBE5D550FBCBFEE8" box="[1046,1142,331,357]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 88</figureCitation>
,
<figureCitation id="134D1396E021FFC8FB77D550FB0CFEE8" box="[1156,1201,331,357]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">113</figureCitation>
). Anterior margin of phallobase of
<taxonomicName id="4C767490E021FFC8FEC0D574FE6EFE05" authorityName="Dombrow" authorityYear="2020" box="[307,467,367,392]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E021FFC8FEC0D574FE6EFE05" box="[307,467,367,392]" italics="true" pageId="61" pageNumber="62">B. nigrosetosa</emphasis>
</taxonomicName>
broad parabola-shaped in dorsal view (
<figureCitation id="134D1396E021FFC8FC7AD574FC5AFE04" box="[905,999,367,393]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 90</figureCitation>
,
<figureCitation id="134D1396E021FFC8FC02D574FBA3FE04" box="[1009,1054,367,393]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">114</figureCitation>
). Anterior margin of phallobase of
<taxonomicName id="4C767490E021FFC8FF64D588FEA5FE20" box="[151,280,403,429]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E021FFC8FF64D588FEA5FE20" box="[151,280,403,429]" italics="true" pageId="61" pageNumber="62">B. suturalis</emphasis>
</taxonomicName>
curly bracket-shaped in dorsal view (
<figureCitation id="134D1396E021FFC8FD4DD588FCA1FE20" box="[702,796,403,429]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="61" pageNumber="62">Figs. 88</figureCitation>
,
<figureCitation id="134D1396E021FFC8FCDBD588FCE8FE20" box="[808,853,403,429]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="61" pageNumber="62">113</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E021FFCAFF64D5C1FAE8FCD1" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
<paragraph id="8BC90F13E021FFC8FF64D5C1FE96FE79" blockId="61.[151,1437,474,2013]" box="[151,299,474,500]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF64D5C1FE96FE79" bold="true" box="[151,299,474,500]" pageId="61" pageNumber="62">Description:</emphasis>
</paragraph>
<paragraph id="8BC90F13E021FFC8FF34D5E4FC83FD94" blockId="61.[151,1437,474,2013]" box="[199,830,510,537]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D5E4FE3DFD94" bold="true" box="[199,384,511,537]" pageId="61" pageNumber="62">Measurements:</emphasis>
length
<quantity id="4C8EA2F6E021FFC8FE27D5E4FDE3FD94" box="[468,606,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.4" metricValueMin="7.6" pageId="61" pageNumber="62" unit="mm" value="8.0" valueMax="8.4" valueMin="7.6">7.68.4 mm</quantity>
; width
<quantity id="4C8EA2F6E021FFC8FD43D5E5FC87FD94" box="[688,826,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="4.9" metricValueMin="4.5" pageId="61" pageNumber="62" unit="mm" value="4.7" valueMax="4.9" valueMin="4.5">4.54.9 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E021FFC8FF34D638FECCFDEC" blockId="61.[151,1437,474,2013]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D638FEE6FDB0" bold="true" box="[199,347,547,573]" pageId="61" pageNumber="62">Integument:</emphasis>
Black; funiculum pale-brown annulated; elytra transparently pale-brown, lateral and posterior margins blackened.
</paragraph>
<paragraph id="8BC90F13E021FFC8FF34D670FD42F96C" blockId="61.[151,1437,474,2013]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D670FEFAFD08" bold="true" box="[199,327,619,645]" pageId="61" pageNumber="62">Tegument:</emphasis>
<emphasis id="B902D301E021FFC8FEA3D670FE17FD08" box="[336,426,619,645]" italics="true" pageId="61" pageNumber="62">Clypeus</emphasis>
bare; loose setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins.
<emphasis id="B902D301E021FFC8FE89D694FE0AFD24" box="[378,439,655,681]" italics="true" pageId="61" pageNumber="62">Head</emphasis>
loose setose with long, erect, black setae.
<emphasis id="B902D301E021FFC8FC63D694FBA5FD24" box="[912,1048,655,681]" italics="true" pageId="61" pageNumber="62">Eye canthus</emphasis>
with a loose setose ciliary of long, black, setae.
<emphasis id="B902D301E021FFC8FEDBD6A8FECAFD40" box="[296,375,691,717]" italics="true" pageId="61" pageNumber="62">Scapus</emphasis>
loose setose with long, black setae, pale-brown ventrally, sparse and medium-long on
<emphasis id="B902D301E021FFC8FAB1D6A8FE86FD7C" italics="true" pageId="61" pageNumber="62">pedicellus; funiculum</emphasis>
with few very-short, erect, brown setae, shorter and semi-erect on
<emphasis id="B902D301E021FFC8FBB0D6CCFB36FD7C" box="[1091,1163,727,753]" italics="true" pageId="61" pageNumber="62">clavus</emphasis>
.
<emphasis id="B902D301E021FFC8FB68D6CCFAB7FD7D" box="[1179,1290,727,752]" italics="true" pageId="61" pageNumber="62">Pronotum</emphasis>
loose setose with long, erect, brown setae, shorter towards posterior margin; with a loose setose ciliary of long, brown setae on lateral margin.
<emphasis id="B902D301E021FFC8FEB3D704FE13FCB4" box="[320,430,799,825]" italics="true" pageId="61" pageNumber="62">Scutellum</emphasis>
loose setose with short and medium-long, semi-erect, brown setae.
<emphasis id="B902D301E021FFC8FB6CD704FB58FCB4" box="[1183,1253,799,825]" italics="true" pageId="61" pageNumber="62">Elytra</emphasis>
without pattern. Sparse setose with short, semi-erect, pale-brown and black setae, medium-long at humerus; few short, reclinate, whitish setae interspersed below apical torus; with two loose, longitudinal rows of medium-long, semi-erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating towards middle. Suture with a loose row of short and long, robust, semi-erect, black setae.
<emphasis id="B902D301E021FFC8FC53D7C8FC4CFC60" box="[928,1009,979,1005]" italics="true" pageId="61" pageNumber="62">Thorax</emphasis>
and
<emphasis id="B902D301E021FFC8FBDFD7C8FB33FC60" box="[1068,1166,979,1005]" italics="true" pageId="61" pageNumber="62">pleurites</emphasis>
loose pilose with long, black hairs.
<emphasis id="B902D301E021FFC8FEEED7ECFE1CFB9D" box="[285,417,1015,1040]" italics="true" pageId="61" pageNumber="62">Ventrite IV</emphasis>
loose setose with short, reclinate, brownish setae; long, semi-erect, black setae interspersed.
<emphasis id="B902D301E021FFC8FF64D000FE95FBB8" box="[151,296,1051,1077]" italics="true" pageId="61" pageNumber="62">Propygidium</emphasis>
and
<emphasis id="B902D301E021FFC8FEACD000FE7AFBB8" box="[351,455,1051,1077]" italics="true" pageId="61" pageNumber="62">pygidium</emphasis>
loose setose with short, reclinate, pale-brown and whitish setae; long, semi-erect, black setae interspersed on propygidium, erect on pygidium; tip of pygidium with a loose setose ciliary of long, black setae.
<emphasis id="B902D301E021FFC8FF2CD078FE12FBF0" box="[223,431,1123,1149]" italics="true" pageId="61" pageNumber="62">Prothoracic femur</emphasis>
loose pilose with long, brown hairs; bare internally.
<emphasis id="B902D301E021FFC8FBFDD078FB6DFBF0" box="[1038,1232,1123,1149]" italics="true" pageId="61" pageNumber="62">Prothoracic tibia</emphasis>
loose setose with short, reclinate, pale-brown setae dorsally; erect, medium-long and black ventrally; medium-long, black, semi-erect setae interspersed dorsally; with a sparse setose longitudinal row of medium-long erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; whitish and very short at external ridge; a setaceous ciliary of short, brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth.
<emphasis id="B902D301E021FFC8FDFAD10CFD50FABC" box="[521,749,1303,1329]" italics="true" pageId="61" pageNumber="62">Mesothoracic femur</emphasis>
loose pilose with long, brown hairs; distinctly long, brown ventrally; long, erect, black setae interspersed dorsally and externally; bare internally.
<emphasis id="B902D301E021FFC8FB85D120FAF0FAD8" box="[1142,1357,1339,1365]" italics="true" pageId="61" pageNumber="62">Mesothoracic tibia</emphasis>
sparse setose with short, reclinate, white and pale-brown setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of medium-long, black bristles; missing internally.
<emphasis id="B902D301E021FFC8FE98D1BCFDF5FA4C" box="[363,584,1447,1473]" italics="true" pageId="61" pageNumber="62">Metathoracic femur</emphasis>
loose setose with medium-long and long, semi-erect, black setae; long, palebrown setae interspersed ventrally; bare internally.
<emphasis id="B902D301E021FFC8FD3CD1D0FC22FA68" box="[719,927,1483,1509]" italics="true" pageId="61" pageNumber="62">Metathoracic tibia</emphasis>
loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, black setae interspersed; denser, finer and long internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally.
<emphasis id="B902D301E021FFC8FB3ED208FAC0F9A1" box="[1229,1405,1555,1580]" italics="true" pageId="61" pageNumber="62">Tarsomere IIV</emphasis>
of all thoracic legs sparse setose with very short, reclinate, brown setae; with a setaceous, apical, marginal ciliary of short, dark red-brown setae lengthening towards
<emphasis id="B902D301E021FFC8FD4ED240FC6EF9F9" box="[701,979,1627,1653]" italics="true" pageId="61" pageNumber="62">metathoracic tarsomeres</emphasis>
, fine dorsally and more robust ventrally.
<emphasis id="B902D301E021FFC8FF64D264FE92F915" box="[151,303,1663,1688]" italics="true" pageId="61" pageNumber="62">Pretarsomere</emphasis>
of all thoracic legs with few, medium-long, semi-erect, red-brown setae at apical margin, shorter ventrally.
<emphasis id="B902D301E021FFC8FEF4D2B8FE1CF930" box="[263,417,1699,1725]" italics="true" pageId="61" pageNumber="62">Mesothoracic</emphasis>
and
<emphasis id="B902D301E021FFC8FE25D2B8FCBBF931" box="[470,774,1699,1725]" italics="true" pageId="61" pageNumber="62">metathoracic pretarsomere</emphasis>
additionally sparse setose with short, reclinate, pale-brown setae, lengthening towards
<emphasis id="B902D301E021FFC8FE3BD2DCFD45F96D" box="[456,760,1735,1761]" italics="true" pageId="61" pageNumber="62">metathoracic pretarsomere</emphasis>
.
</paragraph>
<paragraph id="8BC90F13E021FFC8FF34D2F0FAFDF8C0" blockId="61.[151,1437,474,2013]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D2F0FEADF888" bold="true" box="[199,272,1771,1797]" pageId="61" pageNumber="62">Head:</emphasis>
Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle III cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E021FFC8FF34D34CFA32F834" blockId="61.[151,1437,474,2013]" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D34CFEF4F8FC" bold="true" box="[199,329,1879,1905]" pageId="61" pageNumber="62">Pronotum:</emphasis>
Widest medially; loose punctate with small, shallow, roundish punctures, dense within medial furrow and posteriorly.Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly and posteriorly. With an indistinct, shallow, shortened medial furrow from posterior margin terminating on disk. Posterior corner rounded.
</paragraph>
<paragraph id="8BC90F13E021FFCBFF34D3D9FCE7FF59" blockId="61.[151,1437,474,2013]" lastBlockId="62.[151,1437,150,2013]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="B902D301E021FFC8FF34D3D9FEF5F851" bold="true" box="[199,328,1986,2012]" pageId="61" pageNumber="62">Scutellum:</emphasis>
Parabola-shaped; as broad as long; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in apical third. Lateral margin slightly convex. Tip pointed.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D4C5FDCDFE5D" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D4C5FEA0FF75" bold="true" box="[199,285,222,248]" pageId="62" pageNumber="63">Elytra:</emphasis>
Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; very slightly pulvinate laterally; humerus indistinct; with a broad roundish torus beside scutellum, expanding towards disk; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; with two loose, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating medially.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D5C1FBFBFDB1" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D5C1FEF8FE79" bold="true" box="[199,325,474,500]" pageId="62" pageNumber="63">Abdomen:</emphasis>
Ventrite I extremely shortened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linearly attenuate apically; tip linear.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D65DFA85FD7D" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D65DFE3BFDED" bold="true" box="[199,390,582,608]" pageId="62" pageNumber="63">Prothoracic leg:</emphasis>
Femur and tibia slender. Tibia with a medio-ventral ridge. Median tooth elongate. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere IIV combined extremely short; equal in length; slightly less than
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/
<quantity id="4C8EA2F6E022FFCBFCBFD6BAFCCDFD25" box="[844,880,654,687]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="62" pageNumber="63" unit="in" value="2.0">
<subScript id="17F20D56E022FFCBFCBFD6BAFCE9FD22" attach="left" box="[844,852,673,687]" fontSize="6" pageId="62" pageNumber="63">2</subScript>
in
</quantity>
length of tibia. Pretarsomere distinctly thickened; with a shortened granulate ridge ventrally; slightly more than
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<subScript id="17F20D56E022FFCBFCA5D6DEFCDBFD5E" attach="left" box="[854,870,709,723]" fontSize="6" pageId="62" pageNumber="63">10</subScript>
more in length of tarsomere IIV combined. Internal claw slightly longer than pretarsomere. External claw slightly more than
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<quantity id="4C8EA2F6E022FFCBFBF1D6F2FB92FD7D" box="[1026,1071,726,759]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="62" pageNumber="63" unit="in" value="10.0">
<subScript id="17F20D56E022FFCBFBF1D6F2FBAFFD7A" attach="left" box="[1026,1042,745,759]" fontSize="6" pageId="62" pageNumber="63">10</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D6E1FC23FC0D" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D6E1FE21FC99" bold="true" box="[199,412,762,788]" pageId="62" pageNumber="63">Mesothoracic leg:</emphasis>
Tibia slightly less than
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<quantity id="4C8EA2F6E022FFCBFD4DD716FD50FC99" box="[702,749,762,795]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="62" pageNumber="63" unit="in" value="10.0">
<subScript id="17F20D56E022FFCBFD4DD716FD73FC96" attach="left" box="[702,718,781,795]" fontSize="6" pageId="62" pageNumber="63">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined extremely short; slightly less than
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/
<quantity id="4C8EA2F6E022FFCBFE81D72BFE27FCB5" box="[370,410,798,830]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="62" pageNumber="63" unit="in" value="5.0">
<subScript id="17F20D56E022FFCBFE81D72BFEC7FCB3" attach="left" box="[370,378,816,830]" fontSize="6" pageId="62" pageNumber="63">5</subScript>
in
</quantity>
length of tibia. Pretarsomere with a shortened granulate ridge ventrally, roundish cutting missing; slightly longer than tarsomere IIV combined. External claw slightly less than
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<subScript id="17F20D56E022FFCBFB8FD74FFB39FCEF" attach="left" box="[1148,1156,852,866]" fontSize="6" pageId="62" pageNumber="63">5</subScript>
more in length of pretarsomere. Internal claw slightly less than
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<quantity id="4C8EA2F6E022FFCBFD90D762FD2DFC0D" box="[611,656,870,903]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="62" pageNumber="63" unit="in" value="10.0">
<subScript id="17F20D56E022FFCBFD90D762FDCEFC0A" attach="left" box="[611,627,889,903]" fontSize="6" pageId="62" pageNumber="63">10</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D791FD54FB9D" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D791FE21FC29" bold="true" box="[199,412,906,932]" pageId="62" pageNumber="63">Metathoracic leg:</emphasis>
Tibia slightly shorter than femur. Tarsomere IIV combined distinctly short; slightly less than
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<quantity id="4C8EA2F6E022FFCBFF2CD7DBFEB9FC45" box="[223,260,942,974]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="62" pageNumber="63" unit="in" value="5.0">
<subScript id="17F20D56E022FFCBFF2CD7DBFF5AFC43" attach="left" box="[223,231,960,974]" fontSize="6" pageId="62" pageNumber="63">5</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than
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<quantity id="4C8EA2F6E022FFCBFA84D7DAFA21FC45" box="[1399,1436,942,975]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="62" pageNumber="63" unit="in" value="2.0">
<subScript id="17F20D56E022FFCBFA84D7DAFAC2FC42" attach="left" box="[1399,1407,961,975]" fontSize="6" pageId="62" pageNumber="63">2</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slightly more than
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<subScript id="17F20D56E022FFCBFC4CD7FEFC72FC7E" attach="left" box="[959,975,997,1011]" fontSize="6" pageId="62" pageNumber="63">10</subScript>
more in length of pretarsomere. Internal claw slightly more than
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<quantity id="4C8EA2F6E022FFCBFE45D013FE66FB9D" box="[438,475,1014,1046]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="62" pageNumber="63" unit="in" value="5.0">
<subScript id="17F20D56E022FFCBFE45D013FE03FB9B" attach="left" box="[438,446,1032,1046]" fontSize="6" pageId="62" pageNumber="63">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D001FD90FBB9" blockId="62.[151,1437,150,2013]" box="[199,557,1050,1076]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D001FEECFBB9" bold="true" box="[199,337,1050,1076]" pageId="62" pageNumber="63">Parameres:</emphasis>
(
<figureCitation id="134D1396E022FFCBFE93D001FE55FBB9" box="[352,488,1050,1076]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="62" pageNumber="63">Figs. 9091</figureCitation>
,
<figureCitation id="134D1396E022FFCBFE07D001FD9CFBB9" box="[500,545,1050,1076]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="62" pageNumber="63">114</figureCitation>
).
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D05BFCCBFBD4" blockId="62.[151,1437,150,2013]" box="[199,886,1086,1115]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D05BFE05FBD5" bold="true" box="[199,440,1086,1115]" pageId="62" pageNumber="63">
<specimenCount id="9D70C49AE022FFCBFF34D05BFF4FFBD6" box="[199,242,1088,1115]" pageId="62" pageNumber="63" type="female">♀♀</specimenCount>
. Measurements:
</emphasis>
length
<quantity id="4C8EA2F6E022FFCBFDFFD025FD2BFBD4" box="[524,662,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.4" metricValueMin="8.4" pageId="62" pageNumber="63" unit="mm" value="8.9" valueMax="9.4" valueMin="8.4">8.49.4 mm</quantity>
; width
<quantity id="4C8EA2F6E022FFCBFD1BD025FCCFFBD4" box="[744,882,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="62" pageNumber="63" unit="mm" value="4.9" valueMax="5.3" valueMin="4.5">4.55.3 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D079FDA9FB2D" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D079FEE6FBF1" bold="true" box="[199,347,1122,1148]" pageId="62" pageNumber="63">Integument:</emphasis>
Black; elytra pale-brown, lateral margins darkened; thoracic legs dark red-brown; metathoracic tibia and thoracic tarsi red-brown.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D0B1FB97F8A5" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D0B1FEFAFB49" bold="true" box="[199,327,1194,1220]" pageId="62" pageNumber="63">Tegument:</emphasis>
<emphasis id="B902D301E022FFCBFEBCD0B1FE14FB49" box="[335,425,1194,1220]" italics="true" pageId="62" pageNumber="63">Clypeus</emphasis>
loose setose with short, erect, black setae, medium-long and brown on head.
<emphasis id="B902D301E022FFCBFAE7D0B0FA21FB49" box="[1300,1436,1194,1220]" italics="true" pageId="62" pageNumber="63">Eye canthus</emphasis>
with a loose setose ciliary of medium-long, brown setae. Setae on
<emphasis id="B902D301E022FFCBFC8ED0D4FC74FB65" box="[893,969,1231,1256]" italics="true" pageId="62" pageNumber="63">scapus</emphasis>
and
<emphasis id="B902D301E022FFCBFC0CD0D5FBCCFB65" box="[1023,1137,1230,1256]" italics="true" pageId="62" pageNumber="63">pedicellus</emphasis>
red-brown, pale-brown on
<emphasis id="B902D301E022FFCBFF64D0E9FEBAFA81" box="[151,263,1266,1292]" italics="true" pageId="62" pageNumber="63">funiculum</emphasis>
and
<emphasis id="B902D301E022FFCBFECDD0E9FE3BFA81" box="[318,390,1266,1292]" italics="true" pageId="62" pageNumber="63">clavus</emphasis>
.
<emphasis id="B902D301E022FFCBFE60D0E8FDBFFA81" box="[403,514,1267,1292]" italics="true" pageId="62" pageNumber="63">Pronotum</emphasis>
loose setose with short and long, semi-erect, brown setae; short, reclinate, whitish setae sometimes interspersed on disk, in posterior half of medial furrow and at posterior margin; with a loose setose ciliary of long, brown setae on lateral margin.
<emphasis id="B902D301E022FFCBFD42D121FCA2FAD9" box="[689,799,1338,1364]" italics="true" pageId="62" pageNumber="63">Scutellum</emphasis>
loose setose with short, semi-erect, brown setae; short, reclinate, white setae interspersed.
<emphasis id="B902D301E022FFCBFDD3D145FDDBFAF5" box="[544,614,1374,1400]" italics="true" pageId="62" pageNumber="63">Elytra</emphasis>
loose setose with short, reclinate, pale-brown setae; long, semi-erect and erect, black setae interspersed, distinctly long anteriorly; sometimes few, short, reclinate, whitish setae below apical torus.
<emphasis id="B902D301E022FFCBFF2ED1BDFE93FA4D" box="[221,302,1446,1472]" italics="true" pageId="62" pageNumber="63">Thorax</emphasis>
and
<emphasis id="B902D301E022FFCBFE97D1BDFE7BFA4D" box="[356,454,1446,1472]" italics="true" pageId="62" pageNumber="63">pleurites</emphasis>
loose pilose with long, pale-brown hairs.
<emphasis id="B902D301E022FFCBFC69D1BCFC4CFA4D" box="[922,1009,1447,1472]" italics="true" pageId="62" pageNumber="63">Ventrite</emphasis>
loose setose with short, reclinate, yellowish-white setae; medium-long and long, semi-erect, brown setae interspersed.
<emphasis id="B902D301E022FFCBFBD9D1D1FB06FA69" box="[1066,1211,1482,1508]" italics="true" pageId="62" pageNumber="63">Propygidium</emphasis>
and
<emphasis id="B902D301E022FFCBFB01D1D1FAE7FA69" box="[1266,1370,1482,1508]" italics="true" pageId="62" pageNumber="63">pygidium</emphasis>
loose setose with short, reclinate, yellowish-white setae; medium-long, semi-erect, pale-brown setae interspersed. Tip of pygidium with a loose setose ciliary of long, brown, setae.
<emphasis id="B902D301E022FFCBFCC1D209FBBCF9A1" box="[818,1025,1554,1580]" italics="true" pageId="62" pageNumber="63">Prothoracic femur</emphasis>
loose pilose with long, brown hairs, paler towards tibia.
<emphasis id="B902D301E022FFCBFE85D22DFD8BF9DD" box="[374,566,1590,1616]" italics="true" pageId="62" pageNumber="63">Prothoracic tibia</emphasis>
loose setose with short, semi-erect, pale-brown setae, erect and medium-long ventrally; a loose setose ciliary of short, pale-brown setae on internal ridge and external ridge; a setaceous ciliary of short, pale-brown setae on anterior margin.
<emphasis id="B902D301E022FFCBFD68D265FCD3F915" box="[667,878,1662,1688]" italics="true" pageId="62" pageNumber="63">Mesothoracic tibia</emphasis>
sparse setose with short, reclinate white setae; robust, medium-long, semi-erect, pale-brown setae interspersed. Ciliary of posterior margin as in male but red-brown.
<emphasis id="B902D301E022FFCBFF64D2DDFECBF96D" box="[151,374,1734,1760]" italics="true" pageId="62" pageNumber="63">Metathoracic femur</emphasis>
loose setose with long, semi-erect, pale-brown setae.
<emphasis id="B902D301E022FFCBFC21D2DDFB1EF96D" box="[978,1187,1734,1760]" italics="true" pageId="62" pageNumber="63">Metathoracic tibia</emphasis>
loose setose with medium-long and long, semi-erect, red-brown setae; more robust dorsally; denser, fine and long internally. Fringe of posterior margin as in male but red-brown. Setae of thoracic tarsi pale red-brown.
</paragraph>
<paragraph id="8BC90F13E022FFCBFF34D329FCD7F835" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D329FEADF8C1" bold="true" box="[199,272,1842,1868]" pageId="62" pageNumber="63">Head:</emphasis>
Clypeus very slightly depressed laterally; dense punctate with large, deep, roundish punctures, micropunctate within.Anterior margin of clypeus slightly vertically reflexed. Lateral margin very slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with very large, deep, roundish, not merging punctures. Eye below head level. Funicle I cylindrical; clavus short elliptic.
</paragraph>
<paragraph id="8BC90F13E022FFCAFF34D3D9FDF0FF59" blockId="62.[151,1437,150,2013]" lastBlockId="63.[151,1436,150,866]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<emphasis id="B902D301E022FFCBFF34D3D9FEF4F851" bold="true" box="[199,329,1986,2012]" pageId="62" pageNumber="63">Pronotum:</emphasis>
Widest in anterior third; moderate pulvinate laterally; punctures additionally with a micropunctuation within. Lateral margin linearly attenuate in anterior third, slightly convex attenuate posteriorly. With an indistinct, shallow, complete medial furrow.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D4C5FBB4FE91" blockId="63.[151,1436,150,866]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D4C5FEF5FF75" bold="true" box="[199,328,222,248]" pageId="63" pageNumber="64">Scutellum:</emphasis>
Dense punctate with small, shallow, roundish punctures; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating medially. Tip rounded.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D53DFBF6FECD" blockId="63.[151,1436,150,866]" box="[199,1099,294,321]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D53DFEA0FECD" bold="true" box="[199,285,294,320]" pageId="63" pageNumber="64">Elytra:</emphasis>
Covering propygidium. Scutellar torus missing; lateral trench indistinct.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D551FC85FE21" blockId="63.[151,1436,150,866]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D551FEF8FEE9" bold="true" box="[199,325,330,356]" pageId="63" pageNumber="64">Abdomen:</emphasis>
Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite IIV broadened medially; ventrite V broadest. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper fifth, linearly attenuate apically.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D5ADFA81FDED" blockId="63.[151,1436,150,866]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D5ADFE3AFE5D" bold="true" box="[199,391,438,464]" pageId="63" pageNumber="64">Prothoracic leg:</emphasis>
Femur and tibia robust. Tibia smooth ventrally. Median tooth broad triangular-shaped. Basal tooth very small. Tarsomere IIV combined distinctly short; tarsomere I petiolate chalice-shaped, longest, tarsomere IIIV equal in length; slightly less than
<superScript id="7C03A25BE023FFCAFD96D5E5FDD0FD81" attach="right" box="[613,621,510,524]" fontSize="6" pageId="63" pageNumber="64">7</superScript>
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<quantity id="4C8EA2F6E023FFCAFD86D60AFD19FD95" box="[629,676,510,543]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="63" pageNumber="64" unit="in" value="10.0">
<subScript id="17F20D56E023FFCAFD86D60AFD38FD92" attach="left" box="[629,645,529,543]" fontSize="6" pageId="63" pageNumber="64">10</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; slightly more than
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<quantity id="4C8EA2F6E023FFCAFD5BD62FFD72FDB1" box="[680,719,546,578]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="63" pageNumber="64" unit="in" value="5.0">
<subScript id="17F20D56E023FFCAFD5BD62FFD0DFDCF" attach="left" box="[680,688,564,578]" fontSize="6" pageId="63" pageNumber="64">5</subScript>
in
</quantity>
length of tarsomere IIV combined. Internal claw slender; not broadened medially; as long as pretarsomere. External claw slightly less than
<superScript id="7C03A25BE023FFCAFC0DD65DFBBBFDD9" attach="right" box="[1022,1030,582,596]" fontSize="6" pageId="63" pageNumber="64">4</superScript>
/
<quantity id="4C8EA2F6E023FFCAFBFDD643FB8EFDED" box="[1038,1075,582,614]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="63" pageNumber="64" unit="in" value="5.0">
<subScript id="17F20D56E023FFCAFBFDD643FBABFDEB" attach="left" box="[1038,1046,600,614]" fontSize="6" pageId="63" pageNumber="64">5</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D671FBFEFD7D" blockId="63.[151,1436,150,866]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D671FE26FD09" bold="true" box="[199,411,618,644]" pageId="63" pageNumber="64">Mesothoracic leg:</emphasis>
Tibia slightly less than
<superScript id="7C03A25BE023FFCAFD55D671FD13FDF5" attach="right" box="[678,686,618,632]" fontSize="6" pageId="63" pageNumber="64">4</superScript>
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<quantity id="4C8EA2F6E023FFCAFD45D667FD66FD09" box="[694,731,618,650]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="63" pageNumber="64" unit="in" value="5.0">
<subScript id="17F20D56E023FFCAFD45D667FD03FD07" attach="left" box="[694,702,636,650]" fontSize="6" pageId="63" pageNumber="64">5</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined very short; slightly less than
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/
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length of tibia. Pretarsomere slender; with a shortened smooth ridge ventrally; slightly less than
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in
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length of tarsomere IIV combined. External claw slender; not broadened medially; slightly less than
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more in length of pretarsomere. Internal claw slightly less than
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in
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length of external claw.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF34D6E1FAE8FCD1" blockId="63.[151,1436,150,866]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF34D6E1FE26FC99" bold="true" box="[199,411,762,788]" pageId="63" pageNumber="64">Metathoracic leg:</emphasis>
Femur slightly thickened. Tibia distinctly extended dorso-apically; slightly less than
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in
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length of femur. Tarsomere IIV combined short. Pretarsomere slender; slightly more than
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in
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length of tarsomere IIV combined. External claw slightly shorter than pretarsomere. Internal claw
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in
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length of external claw.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E023FFCAFF64D791FCE8FC60" pageId="63" pageNumber="64" type="distribution">
<paragraph id="8BC90F13E023FFCAFF64D791FBEDFC29" blockId="63.[151,1104,906,933]" box="[151,1104,906,933]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF64D791FE8CFC29" bold="true" box="[151,305,906,932]" pageId="63" pageNumber="64">Distribution.</emphasis>
<collectingRegion id="49B2C1F1E023FFCAFEC4D791FDF5FC29" box="[311,584,906,932]" country="South Africa" name="Northern Cape" pageId="63" pageNumber="64">Northern Cape Province</collectingRegion>
(
<figureCitation id="134D1396E023FFCAFDA4D791FD05FC29" box="[599,696,906,932]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="63" pageNumber="64">Fig. 140</figureCitation>
), only known from the
<typeStatus id="54CDB1B1E023FFCAFC4CD790FC52FC28" box="[959,1007,907,933]" pageId="63" pageNumber="64">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC90F13E023FFCAFF64D7C9FCE8FC60" blockId="63.[151,853,978,1005]" box="[151,853,978,1005]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF64D7C9FEE3FC61" bold="true" box="[151,350,978,1004]" pageId="63" pageNumber="64">
Vegetation
<typeStatus id="54CDB1B1E023FFCAFEEFD7C9FEE4FC61" box="[284,345,978,1004]" pageId="63" pageNumber="64">Type</typeStatus>
.
</emphasis>
Nieuwoudtville Shale Renosterveld [FRs 2].
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E023FFCAFF64D001FB90FBB9" box="[151,1069,1050,1077]" pageId="63" pageNumber="64" type="biology_ecology">
<paragraph id="8BC90F13E023FFCAFF64D001FB90FBB9" blockId="63.[151,1069,1050,1077]" box="[151,1069,1050,1077]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF64D001FE95FBB9" bold="true" box="[151,296,1050,1076]" pageId="63" pageNumber="64">Host Plants.</emphasis>
<taxonomicName id="4C767490E023FFCAFEDED001FE11FBB9" box="[301,428,1050,1076]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
:
<taxonomicName id="4C767490E023FFCAFE44D000FDDCFBB9" authorityName="Roessler" authorityYear="1959" baseAuthorityName="Burm. fil." box="[439,609,1050,1077]" class="Magnoliopsida" family="Asteraceae" genus="Gazania" kingdom="Plantae" order="Asterales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="B902D301E023FFCAFE44D000FDDCFBB9" box="[439,609,1050,1077]" italics="true" pageId="63" pageNumber="64">Gazania rigida</emphasis>
</taxonomicName>
(Burm. f.) Roessler;
<taxonomicName id="4C767490E023FFCAFCA2D000FC4AFBB9" box="[849,1015,1051,1077]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="erosus">
<emphasis id="B902D301E023FFCAFCA2D000FC4AFBB9" box="[849,1015,1051,1077]" italics="true" pageId="63" pageNumber="64">Senecio erosus</emphasis>
</taxonomicName>
L. f.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E023FFCAFF64D079FEB1FB2C" pageId="63" pageNumber="64" type="etymology">
<paragraph id="8BC90F13E023FFCAFF64D079FEB1FB2C" blockId="63.[151,1436,1122,1185]" pageId="63" pageNumber="64">
<emphasis id="B902D301E023FFCAFF64D079FEA3FBF1" bold="true" box="[151,286,1122,1148]" pageId="63" pageNumber="64">Etymology.</emphasis>
The name derives from the black setae, especially on the pronotum [Latin: nigr- = black; Latin: set = a bristles].
</paragraph>
</subSubSection>
</treatment>
</document>