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<document id="9CD29172A6C084590BC71F7807511988" ID-CLB-Dataset="27463" ID-DOI="10.11646/zootaxa.4559.1.9" ID-GBIF-Dataset="8e4bae52-8e8a-4ce8-acde-a1134438bcc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="2626740" ID-ZooBank="6C0BB3B3-DD29-4917-9309-2BB9543653CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554321167055" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03DF87DFFFCFA41806ADEFF1FBE5FE93" docLanguage="en" docName="zootaxa.4559.1.10.pdf" docOrigin="Zootaxa 4559 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmoraphes yunnanensis Jałoszyński 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="183" masterDocId="FFE6FFA7FFCFA41A063AE91BFFD9FFEC" masterDocTitle="The first species of Scydmoraphes (Coleoptera, Staphylinidae, Scydmaeninae) in China" masterLastPageNumber="184" masterPageNumber="182" pageNumber="182" updateTime="1698717214664" updateUser="ExternalLinkService">
<mods:mods id="6C45BB6C034E2C4E891B21E2F2C777C7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C371827EFB5A719F049A1523BA61B6D">
<mods:title id="72E3F35DF92DC64AAAF8FEF93E0F6EC8">The first species of Scydmoraphes (Coleoptera, Staphylinidae, Scydmaeninae) in China</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="2FBD07F8A974CCF09601C6C9108A736A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="895C6E7D8E0AB557076A39724F3E5328">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="45439FA88D787D213D61878E54BB5D42">text</mods:typeOfResource>
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<mods:title id="83FF6884C147C002D953CFB63AFB21F6">Zootaxa</mods:title>
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<mods:part id="1B132F216BCF387CF6D3317F706F4186">
<mods:date id="413DCDECFF3C0847454A5EDAABD859D7">2019</mods:date>
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<mods:number id="1855AA203B2111A1747542EBA71AA1A7">2019-02-19</mods:number>
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<mods:number id="C88F3FC7FE8F2E87174E47E7F47CA6BC">4559</mods:number>
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<mods:classification id="C549678FBAE33A73C0E26A89033E3627">journal article</mods:classification>
<mods:identifier id="671E586750C093A53781530107D3BC19" type="CLB-Dataset">27463</mods:identifier>
<mods:identifier id="B5681BFFC48F667AFAE5139C05C2B229" type="DOI">10.11646/zootaxa.4559.1.9</mods:identifier>
<mods:identifier id="B6859224EDE4138022B3BC2177540991" type="GBIF-Dataset">8e4bae52-8e8a-4ce8-acde-a1134438bcc0</mods:identifier>
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</mods:mods>
<treatment id="03DF87DFFFCFA41806ADEFF1FBE5FE93" ID-DOI="http://doi.org/10.5281/zenodo.5940439" ID-GBIF-Taxon="154550582" ID-Zenodo-Dep="5940439" LSID="urn:lsid:plazi:treatment:03DF87DFFFCFA41806ADEFF1FBE5FE93" httpUri="http://treatment.plazi.org/id/03DF87DFFFCFA41806ADEFF1FBE5FE93" lastPageId="2" lastPageNumber="183" pageId="0" pageNumber="182">
<subSubSection id="C36C6542FFCFA41A06ADEFF1FD6BF8E9" box="[151,690,1770,1797]" pageId="0" pageNumber="182" type="nomenclature">
<paragraph id="8BC936C9FFCFA41A06ADEFF1FD6BF8E9" blockId="0.[151,1435,1736,2036]" box="[151,690,1770,1797]" pageId="0" pageNumber="182">
<heading id="D08181A5FFCFA41A06ADEFF1FD6BF8E9" bold="true" box="[151,690,1770,1797]" fontSize="11" level="1" pageId="0" pageNumber="182" reason="1">
<emphasis id="B902EADBFFCFA41A06ADEFF1FD6BF8E9" bold="true" box="[151,690,1770,1797]" pageId="0" pageNumber="182">
<taxonomicName id="4C764D4AFFCFA41A06ADEFF1FE3EF8E8" ID-CoL="8Y9K2" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,487,1770,1796]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="182" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B902EADBFFCFA41A06ADEFF1FE3EF8E8" bold="true" box="[151,487,1770,1796]" italics="true" pageId="0" pageNumber="182">Scydmoraphes yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A0FFCFA41A07D4EFF0FDE9F8E9" box="[494,560,1771,1797]" pageId="0" pageNumber="182" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="134D2A4CFFCFA41A0405EFF0FD70F8E9" box="[575,681,1771,1797]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1787,1809]" captionTargetBox="[173,1421,1110,1744]" captionTargetId="figure@1.[151,1436,1095,1763]" captionTargetPageId="1" captionText="FIGURES 1–5. Scydmoraphes yunnanensis sp. n., habitus of holotype male (1); aedeagus in ventral (2, 4) and lateral (3, 5) views." httpUri="https://zenodo.org/record/2626742/files/figure.png" pageId="0" pageNumber="182">Figs 1–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C6542FFCFA41A06ADEE16FB24F885" pageId="0" pageNumber="182" type="materials_examined">
<paragraph id="8BC936C9FFCFA41A06ADEE16FB24F885" blockId="0.[151,1435,1736,2036]" pageId="0" pageNumber="182">
<emphasis id="B902EADBFFCFA41A06ADEE16FD42F8C8" bold="true" box="[151,667,1805,1828]" pageId="0" pageNumber="182">
Type material.
<materialsCitation id="3B1E3C94FFCFA41A0773EE16FD42F8C8" ID-GBIF-Occurrence="2234682623" box="[329,667,1805,1828]" collectingDate="1996-10-14" collectionCode="NSMT" collectorName="S. Nomura" country="China" determinerName="P. Jaloszynski" elevation="1950" location="Huanxipo" pageId="0" pageNumber="182" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54CD886BFFCFA41A0773EE16FE6CF8C8" box="[329,437,1805,1828]" pageId="0" pageNumber="182" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3617659FFCFA41A07FEEE16FDC4F8C8" box="[452,541,1805,1828]" name="China" pageId="0" pageNumber="182">CHINA</collectingCountry>
(
<collectingRegion id="49B2F82BFFCFA41A0414EE16FD56F8C8" box="[558,655,1805,1828]" country="China" name="Yunnan" pageId="0" pageNumber="182">Yunnan</collectingRegion>
):
</materialsCitation>
</emphasis>
<specimenCount id="9D70FD40FFCFA41A049EEE16FD63F8C8" box="[676,698,1805,1828]" pageId="0" pageNumber="182" type="male">♂</specimenCount>
, three labels: &quot;
<location id="8EA96012FFCFA41A055AEE16FC15F8C8" LSID="urn:lsid:plazi:treatment:03DF87DFFFCFA41806ADEFF1FBE5FE93:8EA96012FFCFA41A055AEE16FC15F8C8" box="[864,972,1805,1828]" country="China" name="Huanxipo" pageId="0" pageNumber="182" stateProvince="Yunnan">Huanxipo</location>
(
<quantity id="4C8E9B2CFFCFA41A05E7EE16FBE8F8C8" box="[989,1073,1805,1828]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="0" pageNumber="182" unit="m" value="1950.0">
<elevation id="005BD1FAFFCFA41A05E7EE16FBE8F8C8" box="[989,1073,1805,1828]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="0" pageNumber="182" unit="m" value="1950.0">1,950m</elevation>
</quantity>
) / Tengchong X.,
<collectingRegion id="49B2F82BFFCFA41A02C4EE16FA96F8C8" box="[1278,1359,1805,1828]" country="China" name="Yunnan" pageId="0" pageNumber="182">Yunnan</collectingRegion>
/ [SE-
<collectingCountry id="F3617659FFCFA41A06ADEE2BFF28F8AB" box="[151,241,1840,1863]" name="China" pageId="0" pageNumber="182">CHINA</collectingCountry>
]&quot; [white, printed]; &quot;
<date id="FFC81009FFCFA41A07F3EE2BFDEEF8AB" box="[457,567,1840,1863]" pageId="0" pageNumber="182" value="1996-10-14">
<collectingDate id="EF8CE9E1FFCFA41A07F3EE2BFDEEF8AB" box="[457,567,1840,1863]" pageId="0" pageNumber="182" value="1996-10-14">14.x.1996</collectingDate>
</date>
,
<collectorName id="2683531FFFCFA41A047FEE2BFD64F8AB" box="[581,701,1840,1863]" pageId="0" pageNumber="182">S. Nomura</collectorName>
&quot; and locality name repeated in Chinese characters [white, printed], &quot;
<taxonomicName id="4C764D4AFFCFA41A0698EE49FEB7F885" authorityName="Reitter" authorityYear="1891" box="[162,366,1874,1897]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADBFFCFA41A0698EE49FEB7F885" box="[162,366,1874,1897]" italics="true" pageId="0" pageNumber="182">SCYDMORAPHES</emphasis>
</taxonomicName>
\
<taxonomicName id="4C764D4AFFCFA41A07B9EE49FDDFF885" authorityName="Jałoszyński" authorityYear="2019" box="[387,518,1874,1897]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="182" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B902EADBFFCFA41A07B9EE49FDDFF885" box="[387,518,1874,1897]" italics="true" pageId="0" pageNumber="182">yunnanensis</emphasis>
</taxonomicName>
m. \ det.
<determinerName id="7E15DA0DFFCFA41A0451EE49FCDBF885" box="[619,770,1874,1897]" pageId="0" pageNumber="182">P. Jałoszyński</determinerName>
, 2018 \
<typeStatus id="54CD886BFFCFA41A056DEE49FC21F885" box="[855,1016,1874,1897]" pageId="0" pageNumber="182" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (NSMT).
</paragraph>
</subSubSection>
<subSubSection id="C36C6542FFCFA41A06FDEE6EFED2F818" pageId="0" pageNumber="182" type="diagnosis">
<paragraph id="8BC936C9FFCFA41A06FDEE6EFED2F818" blockId="0.[151,1435,1736,2036]" pageId="0" pageNumber="182">
<emphasis id="B902EADBFFCFA41A06FDEE6EFEE5F860" bold="true" box="[199,316,1909,1932]" pageId="0" pageNumber="182">Diagnosis.</emphasis>
Frons conspicuously densely but finely punctate; aedeagus with parameres strongly curved ventrad, so that in ventral view their distal 3/4 are in front of median lobe; median lobe with a distinct, broad submedian constriction; ventral apical plate bifurcate; endophallus with four lightly sclerotized elongate bunches of sclerites, two long and two short ones.
</paragraph>
</subSubSection>
<subSubSection id="C36C6542FFCEA41B06FDE983FE4AFC20" pageId="1" pageNumber="183" type="description">
<paragraph id="8BC936C9FFCEA41B06FDE983FEA6FF3D" blockId="1.[151,1436,152,1041]" pageId="1" pageNumber="183">
<emphasis id="B902EADBFFCEA41B06FDE983FE8BFF43" bold="true" box="[199,338,152,175]" pageId="1" pageNumber="183">Description.</emphasis>
BL
<quantity id="4C8E9B2CFFCEA41B07BAE983FE3BFF43" box="[384,482,152,175]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="183" unit="mm" value="0.88">0.88 mm</quantity>
. Body of male (
<figureCitation id="134D2A4CFFCEA41B04B1E983FD10FF43" box="[651,713,152,175]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1787,1809]" captionTargetBox="[173,1421,1110,1744]" captionTargetId="figure@1.[151,1436,1095,1763]" captionTargetPageId="1" captionText="FIGURES 1–5. Scydmoraphes yunnanensis sp. n., habitus of holotype male (1); aedeagus in ventral (2, 4) and lateral (3, 5) views." httpUri="https://zenodo.org/record/2626742/files/figure.png" pageId="1" pageNumber="183">Fig. 1</figureCitation>
) strongly elongate and slightly flattened, yellowish-brown, covered with yellow vestiture.
</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDE9C6FE38FE4D" blockId="1.[151,1436,152,1041]" pageId="1" pageNumber="183">
Head broadest at large eyes, HL
<quantity id="4C8E9B2CFFCEA41B047BE9C6FD7EFF18" box="[577,679,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="183" unit="mm" value="0.13">0.13 mm</quantity>
, HW
<quantity id="4C8E9B2CFFCEA41B04D5E9C6FC8EFF18" box="[751,855,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="183" unit="mm" value="0.2">0.20 mm</quantity>
; vertex weakly, evenly convex; frons between eyes distinctly flattened; supraantennal tubercles weakly raised. Punctures on frons small but distinct and conspicuously dense, separated by spaces narrower than diameters of punctures, those on vertex distinctly smaller and sparser, inconspicuous; setae short, sparse and suberect. Antennae slender and gradually thickened, AnL
<quantity id="4C8E9B2CFFCEA41B02A6E85EFB24FEB0" box="[1180,1277,325,348]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="183" unit="mm" value="0.38">0.38 mm</quantity>
, antennomeres I–II distinctly elongate, III–X slightly to distinctly transverse, XI much shorter than IX–X combined, only about 1.2 × as long as broad, with blunt apex.
</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDE8B6FE61FDA3" blockId="1.[151,1436,152,1041]" pageId="1" pageNumber="183">
Pronotum suboval, broadest near anterior third; PL
<quantity id="4C8E9B2CFFCEA41B04CEE8B6FC8EFE28" box="[756,855,429,452]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="183" unit="mm" value="0.25">0.25 mm</quantity>
, PW
<quantity id="4C8E9B2CFFCEA41B05A9E8B6FC2FFE28" box="[915,1014,429,452]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="183" unit="mm" value="0.23">0.23 mm</quantity>
; anterior and posterior margins weakly arcuate, sides strongly rounded in anterior third, indistinctly concave just in front of only slightly obtuse-angled posterior pronotal corners; pronotal base with a transverse groove deepened at each end. Punctures on pronotal disc very small but distinct, much sparser than those on frons; setae short and sparse, suberect, additionally sides of pronotum densely covered with thick bristles.
</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDEB41FD67FD5B" blockId="1.[151,1436,152,1041]" pageId="1" pageNumber="183">
Elytra oval, broadest slightly in front of middle; EL
<quantity id="4C8E9B2CFFCEA41B0536EB41FCA9FD9D" box="[780,880,602,625]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="183" unit="mm" value="0.5">0.50 mm</quantity>
, EW
<quantity id="4C8E9B2CFFCEA41B058BEB41FBCCFD9D" box="[945,1045,602,625]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="183" unit="mm" value="0.38">0.38 mm</quantity>
, EI 1.33. Humeral calli distinct but small, elongate; basal impression on each elytron shallow and short. Punctures fine and superficial, inconspicuous; setae short, sparse, suberect. Hind wings well-developed.
</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDEBD9FD67FD35" blockId="1.[151,1436,152,1041]" box="[199,702,706,729]" pageId="1" pageNumber="183">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDEBFEFB47FC45" blockId="1.[151,1436,152,1041]" pageId="1" pageNumber="183">
Aedeagus (
<figureCitation id="134D2A4CFFCEA41B077BEBFEFE7AFD10" box="[321,419,741,764]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1787,1809]" captionTargetBox="[173,1421,1110,1744]" captionTargetId="figure@1.[151,1436,1095,1763]" captionTargetPageId="1" captionText="FIGURES 1–5. Scydmoraphes yunnanensis sp. n., habitus of holotype male (1); aedeagus in ventral (2, 4) and lateral (3, 5) views." httpUri="https://zenodo.org/record/2626742/files/figure.png" pageId="1" pageNumber="183">Figs 2–5</figureCitation>
) moderately slender; AeL
<quantity id="4C8E9B2CFFCEA41B04F4EBFEFCEAFD10" box="[718,819,741,764]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="183" unit="mm" value="0.15">0.15 mm</quantity>
; in ventral view median lobe with rounded base; apical region broadly subtriangular and blunt, demarcated by a distinct but shallow submedian constriction; ventral diaphragm circular; ventral apical plate bifurcate, with a deep and broad distal emargination and lateral portions forming curved subtriangular apical lobes; endophallus with asymmetrical, elongate median component flanked by two groups of elongate sclerites, one very long and the other one half shorter; parameres slender, strongly curved ventrad, so that in ventral view their distal 3/4 are in front of median lobe, each paramere with one long apical seta.
</paragraph>
<paragraph id="8BC936C9FFCEA41B06FDEAAEFE4AFC20" blockId="1.[151,1436,152,1041]" box="[199,403,949,972]" pageId="1" pageNumber="183">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36C6542FFCEA41B06FDEAC3FDEFFC03" box="[199,566,984,1007]" pageId="1" pageNumber="183" type="distribution">
<paragraph id="8BC936C9FFCEA41B06FDEAC3FDEFFC03" blockId="1.[151,1436,152,1041]" box="[199,566,984,1007]" pageId="1" pageNumber="183">
<emphasis id="B902EADBFFCEA41B06FDEAC3FE80FC03" bold="true" box="[199,345,984,1007]" pageId="1" pageNumber="183">Distribution.</emphasis>
SE
<collectingCountry id="F3617659FFCEA41B07BFEAC3FE1CFC03" box="[389,453,984,1007]" name="China" pageId="1" pageNumber="183">China</collectingCountry>
(
<collectingRegion id="49B2F82BFFCEA41B07E9EAC3FDF2FC03" box="[467,555,984,1007]" country="China" name="Yunnan" pageId="1" pageNumber="183">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C6542FFCEA41B06FDEAE1FCD9FBFD" box="[199,768,1018,1041]" pageId="1" pageNumber="183" type="etymology">
<paragraph id="8BC936C9FFCEA41B06FDEAE1FCD9FBFD" blockId="1.[151,1436,152,1041]" box="[199,768,1018,1041]" pageId="1" pageNumber="183">
<emphasis id="B902EADBFFCEA41B06FDEAE1FE9EFBFD" bold="true" box="[199,327,1018,1041]" pageId="1" pageNumber="183">Etymology.</emphasis>
Locotypical, after the province of
<collectingCountry id="F3617659FFCEA41B0483EAE1FD25FBFD" box="[697,764,1018,1041]" name="China" pageId="1" pageNumber="183">China</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF096641FFCEA41B06ADEFE0FA59F8FD" box="[151,1408,1787,1809]" httpUri="https://zenodo.org/record/2626742/files/figure.png" pageId="1" pageNumber="183" startId="1.[151,258,1787,1809]" targetBox="[173,1421,1110,1744]" targetPageId="1">
<paragraph id="8BC936C9FFCEA41B06ADEFE0FA59F8FD" blockId="1.[151,1408,1787,1809]" box="[151,1408,1787,1809]" pageId="1" pageNumber="183">
<emphasis id="B902EADBFFCEA41B06ADEFE0FEEBF8FD" bold="true" box="[151,306,1787,1809]" pageId="1" pageNumber="183">FIGURES 1–5.</emphasis>
<taxonomicName id="4C764D4AFFCEA41B070DEFE7FDE6F8FD" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[311,575,1788,1809]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="183" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B902EADBFFCEA41B070DEFE7FDE6F8FD" box="[311,575,1788,1809]" italics="true" pageId="1" pageNumber="183">Scydmoraphes yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A0FFCEA41B047FEFE0FDA5F8FD" box="[581,636,1787,1809]" pageId="1" pageNumber="183" rank="species">
<emphasis id="B902EADBFFCEA41B047FEFE0FDA5F8FD" bold="true" box="[581,636,1787,1809]" pageId="1" pageNumber="183">sp. n.</emphasis>
</taxonomicNameLabel>
, habitus of holotype male (1); aedeagus in ventral (2, 4) and lateral (3, 5) views.
</paragraph>
</caption>
<subSubSection id="C36C6542FFCEA41806FDEE27FBE5FE93" lastPageId="2" lastPageNumber="184" pageId="1" pageNumber="183" type="discussion">
<paragraph id="8BC936C9FFCEA41806FDEE27FBE5FE93" blockId="1.[151,1436,1851,2014]" lastBlockId="2.[151,1436,151,383]" lastPageId="2" lastPageNumber="184" pageId="1" pageNumber="183">
<emphasis id="B902EADBFFCEA41B06FDEE27FEECF8BF" bold="true" box="[199,309,1852,1875]" pageId="1" pageNumber="183">Remarks.</emphasis>
<taxonomicName id="4C764D4AFFCEA41B0706EE20FDBBF8BF" authorityName="Jałoszyński" authorityYear="2019" box="[316,610,1851,1875]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="183" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B902EADBFFCEA41B0706EE20FDBBF8BF" box="[316,610,1851,1875]" italics="true" pageId="1" pageNumber="183">Scydmoraphes yunnanensis</emphasis>
</taxonomicName>
clearly differs from all nominal species whose aedeagi have been illustrated in literature (some require revision, but those are restricted to the western part of Eurasia and are highly unlikely to be found in
<collectingCountry id="F3617659FFCEA41B06C2EE9AFEE0F874" box="[248,313,1921,1944]" name="China" pageId="1" pageNumber="183">China</collectingCountry>
) in a uniquely shaped aedeagus. The parameres curved so strongly that largely shifted to the ventral side of the median lobe are not known in any other
<taxonomicName id="4C764D4AFFCEA41B04B5EEB8FCF3F857" authorityName="Reitter" authorityYear="1891" box="[655,810,1955,1979]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADBFFCEA41B04B5EEB8FCF3F857" box="[655,810,1955,1979]" italics="true" pageId="1" pageNumber="183">Scydmoraphes</emphasis>
</taxonomicName>
, and also the bifurcate ventral apical plate and bunches of needle-like endophallic sclerites are unique for
<taxonomicName id="4C764D4AFFCEA41B04B5EEDCFCF4F832" authorityName="Jałoszyński" authorityYear="2019" box="[655,813,1991,2014]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="183" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B902EADBFFCEA41B04B5EEDCFCF4F832" box="[655,813,1991,2014]" italics="true" pageId="1" pageNumber="183">S. yunnanensis</emphasis>
</taxonomicName>
. The geographically closest species are
<taxonomicName id="4C764D4AFFCEA41802E2EEDCFE94FF43" authority="(Franz, 1979) (Pakistan)" authorityName="Franz" authorityYear="1979" baseAuthorityName="Franz" baseAuthorityYear="1979" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="2" lastPageNumber="184" order="Coleoptera" pageId="1" pageNumber="183" phylum="Arthropoda" rank="species" species="gartneri">
<emphasis id="B902EADBFFCEA41B02E2EEDCFA90F832" box="[1240,1353,1991,2014]" italics="true" pageId="1" pageNumber="183">S. gartneri</emphasis>
(
<bibRefCitation id="EFE74B38FFCEA418036DEEDCFF17FF43" author="Franz, H." lastPageId="2" lastPageNumber="184" pageId="1" pageNumber="183" pagination="235 - 274" refId="ref1378" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
) (
<collectingCountry id="F3617659FFCDA41806DCE983FE9FFF43" box="[230,326,152,175]" name="Pakistan" pageId="2" pageNumber="184">Pakistan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C764D4AFFCDA4180760E98CFD6EFF43" authority="Vit, 1999" authorityName="Vit" authorityYear="1999" box="[346,695,151,175]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="184" phylum="Arthropoda" rank="species" species="hastipes">
<emphasis id="B902EADBFFCDA4180760E98CFE17FF43" box="[346,462,151,175]" italics="true" pageId="2" pageNumber="184">S. hastipes</emphasis>
<bibRefCitation id="EFE74B38FFCDA41807E3E983FD99FF43" author="Vit, S." box="[473,576,152,175]" pageId="2" pageNumber="184" pagination="59 - 71" refId="ref1754" refString="Vit, S. (1999) Miscellaneous contributions to the knowledge of Scydmaenidae (Insecta: Coleoptera). Klapalekiana, 35, 59 - 71." type="journal article" year="1999">Vít, 1999</bibRefCitation>
(
<collectingCountry id="F3617659FFCDA418046BE983FD68FF43" box="[593,689,152,175]" name="Pakistan" pageId="2" pageNumber="184">Pakistan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C764D4AFFCDA41804FFE98CFCEEFF43" authorityName="Chaudoir" authorityYear="1845" box="[709,823,151,175]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="184" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B902EADBFFCDA41804FFE98CFCEEFF43" box="[709,823,151,175]" italics="true" pageId="2" pageNumber="184">S. minutus</emphasis>
</taxonomicName>
(Russian Far East and Europe) and
<taxonomicName id="4C764D4AFFCDA41802F0E98CFEFCFF3D" authority="Franz, 1976" authorityName="Franz" authorityYear="1976" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="184" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B902EADBFFCDA41802F0E98CFA96FF43" box="[1226,1359,151,175]" italics="true" pageId="2" pageNumber="184">S. japonicus</emphasis>
<bibRefCitation id="EFE74B38FFCDA4180363E983FF15FF3D" author="Franz, H." pageId="2" pageNumber="184" pagination="51 - 60" refId="ref1351" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Blatter, 72, 51 - 60." type="journal article" year="1976">Franz, 1976</bibRefCitation>
(
<collectingCountry id="F3617659FFCDA41806E0E9A1FEC5FF3D" box="[218,284,186,209]" name="Japan" pageId="2" pageNumber="184">Japan</collectingCountry>
)
</taxonomicName>
. The two Pakistani species have simple, sac-like aedeagi, which are suboval in ventral view, and each has a darkly sclerotized, transverse subapical endophallic structure (illustrated in
<bibRefCitation id="EFE74B38FFCDA418022CE9C6FBA6FF18" author="Vit, S." box="[1046,1151,221,244]" pageId="2" pageNumber="184" pagination="59 - 71" refId="ref1754" refString="Vit, S. (1999) Miscellaneous contributions to the knowledge of Scydmaenidae (Insecta: Coleoptera). Klapalekiana, 35, 59 - 71." type="journal article" year="1999">Vít 1999</bibRefCitation>
and
<bibRefCitation id="EFE74B38FFCDA41802F0E9C6FA48FF18" author="Jaloszynski, P." box="[1226,1425,221,244]" pageId="2" pageNumber="184" pagination="619 - 637" refId="ref1451" refString="Jaloszynski, P. (2008) Revision of Neuraphes Thomson of the Himalaya Mts. (Coleoptera: Scydmaenidae). Genus, 19 (4), 619 - 637." type="journal article" year="2008">Jałoszyński 2008</bibRefCitation>
).
<taxonomicName id="4C764D4AFFCDA41806ADE9E4FE54FEFB" authorityName="Chaudoir" authorityYear="1845" box="[151,397,255,279]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="184" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B902EADBFFCDA41806ADE9E4FE54FEFB" box="[151,397,255,279]" italics="true" pageId="2" pageNumber="184">Scydmoraphes minutus</emphasis>
</taxonomicName>
(aedeagus illustrated e.g., in
<bibRefCitation id="EFE74B38FFCDA41804FDE81BFC5AFEFB" author="Castellini, G." box="[711,899,256,279]" pageId="2" pageNumber="184" pagination="1 - 121" refId="ref1272" refString="Castellini, G. (2007) Pensieri sulla tassonomia e note su alcuni Scydmaenidae paleartici (Coleoptera). Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 98 (2006 - 2007), 1 - 121." type="journal article" year="2007">Castellini (2007))</bibRefCitation>
has a truncate apex of the median lobe and over a dozen elongate sclerites in the sub-basal region.
<taxonomicName id="4C764D4AFFCDA41804F7E83AFC07FED5" authorityName="Franz" authorityYear="1976" box="[717,990,289,313]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="184" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B902EADBFFCDA41804F7E83AFC07FED5" box="[717,990,289,313]" italics="true" pageId="2" pageNumber="184">Scydmoraphes japonicus</emphasis>
</taxonomicName>
(revised by
<bibRefCitation id="EFE74B38FFCDA4180242E839FA88FED5" author="Jaloszynski, P." box="[1144,1361,290,313]" pageId="2" pageNumber="184" pagination="137 - 145" refId="ref1411" refString="Jaloszynski, P. (2004) Redescriptions and new records of Japanese species of Neuraphes and Scydmoraphes (Coleoptera, Scydmaenidae). Bulletin of National Science Museum Tokyo, 30 (3), 137 - 145." type="journal article" year="2004">Jałoszyński (2004))</bibRefCitation>
has a subtriangular apex of median lobe with a pair of distolateral projections meeting at middle, and a pair of distinct 'palm leaf' structures in the endophallus (which can also be found in many European species).
</paragraph>
</subSubSection>
</treatment>
</document>
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