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<document id="7F287D5C702DFBA4D95B29AF37D009E1" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627005272297" checkinUser="felipe" docAuthor="Published, First" docDate="2008" docId="03DF87D2FF57ABC99EC8134244903A39" docLanguage="en" docName="zt01804p456part1.pdf" docOrigin="Zootaxa 1804" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Corethrella (Corethrella) atricornis Borkent 2008, new species" docType="treatment" docVersion="3" lastPageNumber="189" masterDocId="FFE6FFAAFFECAB759E5F1025465D3823" masterDocTitle="The Frog-Biting Midges of the World (Corethrellidae: Diptera)" masterLastPageNumber="456" masterPageNumber="1" pageNumber="188" updateTime="1699014769975" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B7A2C01E854B21C7DEB6A73EFE96017D">The Frog-Biting Midges of the World (Corethrellidae: Diptera)</mods:title>
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<mods:date id="AEEA565E5E1C5707E52159A194677142">2008</mods:date>
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<treatment id="03DF87D2FF57ABC99EC8134244903A39" ID-DOI="http://doi.org/10.5281/zenodo.5126415" ID-Zenodo-Dep="5126415" LSID="urn:lsid:plazi:treatment:03DF87D2FF57ABC99EC8134244903A39" httpUri="http://treatment.plazi.org/id/03DF87D2FF57ABC99EC8134244903A39" lastPageId="188" lastPageNumber="189" pageId="187" pageNumber="188">
<subSubSection id="C36C654FFF57ABCE9EC8134245123BA2" box="[151,847,871,897]" pageId="187" pageNumber="188" type="nomenclature">
<paragraph id="8BC936C4FF57ABCE9EC8134245123BA2" blockId="187.[151,847,871,897]" box="[151,847,871,897]" pageId="187" pageNumber="188">
<heading id="D08181A8FF57ABCE9EC8134245123BA2" bold="true" box="[151,847,871,897]" fontSize="11" level="1" pageId="187" pageNumber="188" reason="1">
<emphasis id="B902EAD6FF57ABCE9EC8134245123BA2" bold="true" box="[151,847,871,897]" pageId="187" pageNumber="188">
<taxonomicName id="4C764D47FF57ABCE9EC8134244EF3BA2" ID-CoL="8BL4Q" authority="Borkent" authorityName="Borkent" authorityYear="2008" box="[151,690,871,897]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="187" pageNumber="188" phylum="Arthropoda" rank="species" species="atricornis" status="sp. nov." subGenus="Corethrella">
<emphasis id="B902EAD6FF57ABCE9EC81342477C3BA2" bold="true" box="[151,289,871,897]" italics="true" pageId="187" pageNumber="188">Corethrella</emphasis>
(
<emphasis id="B902EAD6FF57ABCE9F6D134247E13BA2" bold="true" box="[306,444,871,897]" italics="true" pageId="187" pageNumber="188">Corethrella</emphasis>
)
<emphasis id="B902EAD6FF57ABCE9F911342441E3BA2" bold="true" box="[462,579,871,897]" italics="true" pageId="187" pageNumber="188">atricornis</emphasis>
Borkent
</taxonomicName>
,
<taxonomicNameLabel id="A23157ADFF57ABCE9CE0134245123BA2" box="[703,847,871,897]" pageId="187" pageNumber="188" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF57ABCE9EC8139742AE3C9F" pageId="187" pageNumber="188" type="diagnosis">
<paragraph id="8BC936C4FF57ABCE9EC8139742AE3C9F" blockId="187.[151,1437,946,2015]" pageId="187" pageNumber="188">
DIAGNOSIS:
<emphasis id="B902EAD6FF57ABCE9F0E1397478E3BE8" box="[337,467,946,971]" italics="true" pageId="187" pageNumber="188">Male adult</emphasis>
: unknown.
<emphasis id="B902EAD6FF57ABCE9C301397454A3BE8" box="[623,791,946,971]" italics="true" pageId="187" pageNumber="188">Female adult:</emphasis>
only extant species of
<taxonomicName id="4C764D47FF57ABCE9A6D139742E43BE8" box="[1074,1209,946,971]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="187" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF57ABCE9A6D139742E43BE8" box="[1074,1209,946,971]" italics="true" pageId="187" pageNumber="188">Corethrella</emphasis>
</taxonomicName>
in the New World with the clypeus wider than long (
<figureCitation id="134D2A41FF57ABCE9C6913FF44D13BD7" box="[566,652,986,1012]" captionStart="FIGURE 18" captionStartId="324.[151,255,1947,1971]" captionTargetBox="[177,1410,299,1907]" captionTargetId="figure-33@324.[177,1411,197,1921]" captionTargetPageId="324" captionText="FIGURE 18. Clypei of female Corethrella. Lateral setae, even if present, are not shown." pageId="187" pageNumber="188">Fig. 18</figureCitation>
AM), palpal segment 3 elongate and slightly swollen subapically (
<figureCitation id="134D2A41FF57ABCE9EC01427475A3C3F" box="[159,263,1026,1052]" captionStart="FIGURE 35" captionStartId="341.[151,255,1947,1971]" captionTargetBox="[177,1375,197,1921]" captionTargetId="figure-21@341.[177,1411,197,1921]" captionTargetPageId="341" captionText="FIGURE 35. Right palpi of female Corethrella." pageId="187" pageNumber="188">Fig. 35S</figureCitation>
), flagellomeres 13 each elongate (
<figureCitation id="134D2A41FF57ABCE9CF3142745473C3F" box="[684,794,1026,1052]" captionStart="FIGURE 32" captionStartId="338.[151,255,1947,1971]" captionTargetBox="[190,1408,238,1917]" captionTargetId="figure-25@338.[177,1411,197,1921]" captionTargetPageId="338" captionText="FIGURE 32. Antennae of female Corethrella." pageId="187" pageNumber="188">Fig. 32D</figureCitation>
), flagellomere 3, 912 each with one sensillum coeloconicum, a distinct midlength wing band, with broad subbasal band extending posterior to CuA (Fig. 72D), thorax dark brown (
<figureCitation id="134D2A41FF57ABCE9F20147747B43C4F" box="[383,489,1106,1132]" captionStart="FIGURE 58" captionStartId="364.[151,255,2012,2036]" captionTargetBox="[176,1410,197,1980]" captionTargetId="figure-29@364.[176,1411,197,1985]" captionTargetPageId="364" captionText="FIGURE 58. Thoraces and legs of female Corethrella, lateral view." pageId="187" pageNumber="188">Fig. 58B</figureCitation>
), halter light brown, much paler than scutellum, midfemur dark brown and equal to that of base of hind femur, base of hind tibia darkly pigmented (contrasting with pale apex of hind femur), hind tibia with basal and apical pigmentation discrete (
<figureCitation id="134D2A41FF57ABCE9D47148745DE3C9F" box="[792,899,1186,1212]" captionStart="FIGURE 58" captionStartId="364.[151,255,2012,2036]" captionTargetBox="[176,1410,197,1980]" captionTargetId="figure-29@364.[176,1411,197,1985]" captionTargetPageId="364" captionText="FIGURE 58. Thoraces and legs of female Corethrella, lateral view." pageId="187" pageNumber="188">Fig. 58B</figureCitation>
), and midfemur without scales.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF57ABC99E9914EF478E3891" lastPageId="188" lastPageNumber="189" pageId="187" pageNumber="188" type="description">
<paragraph id="8BC936C4FF57ABCE9E9914EF47783FFC" blockId="187.[151,1437,946,2015]" pageId="187" pageNumber="188">
DESCRIPTION:
<emphasis id="B902EAD6FF57ABCE9FC214EE44423CC7" box="[413,543,1227,1252]" italics="true" pageId="187" pageNumber="188">Male adult</emphasis>
. Unknown.
<emphasis id="B902EAD6FF57ABCE9CE314EE45073CC7" box="[700,858,1227,1252]" italics="true" pageId="187" pageNumber="188">Female adult</emphasis>
. Descriptive statistics: see
<tableCitation id="C6F4037FFF57ABCE9AF614EF43663CC7" box="[1193,1339,1226,1252]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStart-2="TABLE 8" captionStart-3="TABLE 9" captionStart-4="TABLE 10" captionStart-5="TABLE 11" captionStartId-0="277.[151,241,153,177]" captionStartId-1="280.[151,241,153,177]" captionStartId-2="283.[151,241,153,177]" captionStartId-3="286.[151,241,151,175]" captionStartId-4="289.[151,239,153,177]" captionStartId-5="292.[151,241,153,177]" captionText-0="TABLE 6. Length of wing of females of species of Corethrella. Values are in mm." captionText-1="TABLE 7. Ratio of length of R1/wing length of females of species of Corethrella." captionText-2="TABLE 8. Ratio of head capsule width/height of females of species of Corethrella." captionText-3="TABLE 9. Number of setae on the clypeus of females of species of Corethrella. The number does not include those" captionText-4="TABLE 10. Number of posterior anepisternal setae of females of species of Corethrella." captionText-5="TABLE 11. Number of anepimeral setae of females of species of Corethrella." pageId="187" pageNumber="188">Tables 611</tableCitation>
.
<emphasis id="B902EAD6FF57ABCE9B0E14EF43C93CC7" bold="true" box="[1361,1428,1226,1252]" pageId="187" pageNumber="188">Head</emphasis>
: Outline in anterior view laterally elongate (
<figureCitation id="134D2A41FF57ABCE9CD214D744AA3D2F" box="[653,759,1266,1292]" captionStart="FIGURE 14" captionStartId="320.[151,255,1947,1971]" captionTargetBox="[177,1410,241,1904]" captionTargetId="figure-28@320.[177,1411,197,1921]" captionTargetPageId="320" captionText="FIGURE 14. Head capsules of female Corethrella, anterior view." pageId="187" pageNumber="188">Fig. 14E</figureCitation>
). Coronal suture short, extending ventrally about midway along area between ommatidia (as in
<figureCitation id="134D2A41FF57ABCE9C38153F448E3D17" box="[615,723,1306,1332]" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="187" pageNumber="188">Fig. 16F</figureCitation>
). Two large setae on frons between ventromedial area of ommatida (as in
<figureCitation id="134D2A41FF57ABCE9F3E156747973D7F" box="[353,458,1346,1372]" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="187" pageNumber="188">Fig. 16F</figureCitation>
). Antennal flagellomeres 111 dark brown (1213 missing); pedicel with at least one distinctive, more elongate, stout, dorsal or dorsolateral seta; flagellomeres as in
<figureCitation id="134D2A41FF57ABCE9A25154F42BB3DA7" box="[1146,1254,1386,1412]" captionStart="FIGURE 32" captionStartId="338.[151,255,1947,1971]" captionTargetBox="[190,1408,238,1917]" captionTargetId="figure-25@338.[177,1411,197,1921]" captionTargetPageId="338" captionText="FIGURE 32. Antennae of female Corethrella." pageId="187" pageNumber="188">Fig. 32D</figureCitation>
, sensilla coeloconica distributed as in
<tableCitation id="C6F4037FFF57ABCE9FE715B7444E3D8F" box="[440,531,1426,1452]" captionStart="TABLE 1" captionStartId="265.[151,241,151,175]" captionText="TABLE 1. Pattern of sensilla coeloconica (= s.c.) on antennal flagellomeres of species of Corethrella. Numbers refer to the flagellomere with sensilla coeloconica, with those in brackets referring to flagellomeres with or without sensilla coeloconica. Mis. (missing) refers to damaged specimens, unknown refers to missing stages, uncertain refers to specimens in which visibility was too poor to be certain of whether sensilla coeloconica were present or not. Those species names with a “†” are those of fossils. Corethrella librata, known only as immature stages, is not included." pageId="187" pageNumber="188">Table 1</tableCitation>
. Clypeus (
<figureCitation id="134D2A41FF57ABCE9CC015B744A93D8F" box="[671,756,1426,1452]" captionStart="FIGURE 18" captionStartId="324.[151,255,1947,1971]" captionTargetBox="[177,1410,299,1907]" captionTargetId="figure-33@324.[177,1411,197,1921]" captionTargetPageId="324" captionText="FIGURE 18. Clypei of female Corethrella. Lateral setae, even if present, are not shown." pageId="187" pageNumber="188">Fig. 18</figureCitation>
AM) wide. Mandible with small, pointed teeth. Palpus (
<figureCitation id="134D2A41FF57ABCE9EC0159F47593DF7" box="[159,260,1466,1492]" captionStart="FIGURE 35" captionStartId="341.[151,255,1947,1971]" captionTargetBox="[177,1375,197,1921]" captionTargetId="figure-21@341.[177,1411,197,1921]" captionTargetPageId="341" captionText="FIGURE 35. Right palpi of female Corethrella." pageId="187" pageNumber="188">Fig. 35S</figureCitation>
) pale, with segment 5 light brown; segment 3 elongate, swollen apically.
<emphasis id="B902EAD6FF57ABCE9A2C159F428D3DF7" bold="true" box="[1139,1232,1466,1492]" pageId="187" pageNumber="188">Thorax</emphasis>
(
<figureCitation id="134D2A41FF57ABCE9ABF159F43103DF7" box="[1248,1357,1466,1492]" captionStart="FIGURE 58" captionStartId="364.[151,255,2012,2036]" captionTargetBox="[176,1410,197,1980]" captionTargetId="figure-29@364.[176,1411,197,1985]" captionTargetPageId="364" captionText="FIGURE 58. Thoraces and legs of female Corethrella, lateral view." pageId="187" pageNumber="188">Fig. 58B</figureCitation>
): Dark brown, pale sclerites around base of wing. Posterior portion of dorsocentral row with group of about 5 elongate setae. Prescutal suture short, not extending more than half way to dorsocentral row of setae. Anterior anepisternum divided diagonally by sinuous suture, dorsal portion about equal to ventral portion. Ventral portion of posterior anepisternum triangular, uniformly brown, with anterodorsal margin thick.
<emphasis id="B902EAD6FF57ABCE9A91167F434F3E57" bold="true" box="[1230,1298,1626,1652]" pageId="187" pageNumber="188">Wing</emphasis>
(Fig. 72D): Apex of R
<subScript id="17F23481FF57ABCE9F4816B747423E83" attach="left" box="[279,287,1682,1696]" fontSize="6" pageId="187" pageNumber="188">2</subScript>
basal to apex of M
<subScript id="17F23481FF57ABCE9C4E16B744443E83" attach="left" box="[529,537,1682,1696]" fontSize="6" pageId="187" pageNumber="188">1</subScript>
. Anterior margin with differently, discretely pigmented scales (indicating anterior margin of midlength band), broad subbasal band extending posteriorly to CuA, with midlength band; veins (other than costa and wing margin) with well-developed scales. Halter light brown, lighter than scutellum.
<emphasis id="B902EAD6FF57ABCE9E8116D847443F34" bold="true" box="[222,281,1789,1815]" pageId="187" pageNumber="188">Legs</emphasis>
(
<figureCitation id="134D2A41FF57ABCE9F7516D847C53F34" box="[298,408,1789,1815]" captionStart="FIGURE 58" captionStartId="364.[151,255,2012,2036]" captionTargetBox="[176,1410,197,1980]" captionTargetId="figure-29@364.[176,1411,197,1985]" captionTargetPageId="364" captionText="FIGURE 58. Thoraces and legs of female Corethrella, lateral view." pageId="187" pageNumber="188">Fig. 58B</figureCitation>
): Dark brown with knees of fore-, midlegs pale, hind femur with apical 0.4 pale, hind tibia pale with basal and apical discrete dark brown pigmentation. With only slender setae, lacking scales (except for some in patch of whip-like setae on posterior portion of hind tibia). Midleg with thick, subapical setae on each of at least tarsomeres 13. Claws of each leg equal to those of others; equal on each leg, simple (without inner teeth). Empodia slender.
<emphasis id="B902EAD6FF57ABCE9CDF17B844A73F94" bold="true" box="[640,762,1949,1975]" pageId="187" pageNumber="188">Abdomen</emphasis>
: Medium brown with segments 8-9 dark brown. Cercus dark brown.
</paragraph>
<paragraph id="8BC936C4FF50ABC99E9910BD478E3891" blockId="188.[151,1436,152,538]" box="[198,467,152,178]" pageId="188" pageNumber="189">
<emphasis id="B902EAD6FF50ABC99E9910BD471F3892" box="[198,322,152,177]" italics="true" pageId="188" pageNumber="189">Immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF50ABC99E9910E5476A3971" pageId="188" pageNumber="189" type="distribution">
<paragraph id="8BC936C4FF50ABC99E9910E5420E3921" blockId="188.[151,1436,152,538]" pageId="188" pageNumber="189">
<materialsCitation id="3B1E3C99FF50ABC99E9910E542123921" country="Brazil" elevation="603" location="Brazil" pageId="188" pageNumber="189" specimenCount="603" typeStatus="holotype">
DISTRIBUTION AND BIONOMICS:
<taxonomicName id="4C764D47FF50ABC99CF910E545F738FA" authorityName="Borkent" authorityYear="2008" box="[678,938,192,217]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="B902EAD6FF50ABC99CF910E545F738FA" box="[678,938,192,217]" italics="true" pageId="188" pageNumber="189">Corethrella atricornis</emphasis>
</taxonomicName>
is known only the type locality in
<collectingCountry id="F3617654FF50ABC99B0B10E543C138F9" box="[1364,1436,192,218]" name="Brazil" pageId="188" pageNumber="189">Brazil</collectingCountry>
(Fig. 137) at an altitude of
<specimenCount id="9D70FD4DFF50ABC99F8D10CD447E3921" box="[466,547,232,258]" pageId="188" pageNumber="189" type="generic">
<quantity id="4C8E9B21FF50ABC99F8D10CD44423921" box="[466,543,232,258]" metricMagnitude="2" metricUnit="m" metricValue="6.029999999999999" pageId="188" pageNumber="189" unit="m" value="603.0">
<elevation id="005BD1F7FF50ABC99F8D10CD44423921" box="[466,543,232,258]" metricMagnitude="2" metricUnit="m" metricValue="6.029999999999999" pageId="188" pageNumber="189" unit="m" value="603.0">603 m</elevation>
</quantity>
.
</specimenCount>
It is uncertain how the
<typeStatus id="54CD8866FF50ABC99D6010CD45FB3921" box="[831,934,232,258]" pageId="188" pageNumber="189" type="holotype">holotype</typeStatus>
was collected
</materialsCitation>
.
</paragraph>
<paragraph id="8BC936C4FF50ABC99E991135476A3971" blockId="188.[151,1436,152,538]" pageId="188" pageNumber="189">
TAXONOMIC DISCUSSION: The
<typeStatus id="54CD8866FF50ABC99C31113544893909" box="[622,724,272,298]" pageId="188" pageNumber="189" type="holotype">holotype</typeStatus>
was originally on a pin but was placed on a microscope slide for this study.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF50ABC99E99114544903A39" pageId="188" pageNumber="189" type="materials_examined">
<paragraph id="8BC936C4FF50ABC99E991145475A39E9" blockId="188.[151,1436,152,538]" pageId="188" pageNumber="189">
<materialsCitation id="3B1E3C99FF50ABC99E991145475A39E9" collectionCode="USNM" collectorName="S. Paulo" pageId="188" pageNumber="189" specimenCount="2" typeStatus="holotype">
TYPE:
<typeStatus id="54CD8866FF50ABC99F68114547FA3959" box="[311,423,352,378]" pageId="188" pageNumber="189" type="holotype">Holotype</typeStatus>
, female adult on microscope slide, labeled &quot;
<typeStatus id="54CD8866FF50ABC99DBB114542DE3959" box="[996,1155,352,378]" pageId="188" pageNumber="189" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C764D47FF50ABC99ACD114546A63981" authority="Borkent" authorityName="Borkent" authorityYear="2008" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="B902EAD6FF50ABC99ACD114543C1395A" box="[1170,1436,352,377]" italics="true" pageId="188" pageNumber="189">Corethrella atricornis</emphasis>
Borkent
</taxonomicName>
”, “Paratipo”, “
<taxonomicName id="4C764D47FF50ABC99FF211AD44B73981" authority="Lane" authorityName="Lane" authorityYear="1942" box="[429,746,392,418]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="cardosoi">
<emphasis id="B902EAD6FF50ABC99FF211AD44FE3982" box="[429,675,392,417]" italics="true" pageId="188" pageNumber="189">Corethrella cardosoi</emphasis>
Lane
</taxonomicName>
, 41”, “
<collectorName id="26835312FF50ABC99D1E11AD45F53981" box="[833,936,392,418]" pageId="188" pageNumber="189">S. Paulo</collectorName>
, Casa Grande, Bar. &amp; Galv. Col. XII-40” (
<collectionCode id="ED67AE01FF50ABC99EC0119546A739E9" box="[159,250,432,458]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="188" pageNumber="189" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC936C4FF50ABC99E9911FD44903A39" blockId="188.[151,1436,152,538]" pageId="188" pageNumber="189">
DERIVATION OF SPECIFIC EPITHET: The name
<taxonomicName id="4C764D47FF50ABC99D6911FD45F739D2" authorityName="Borkent" authorityYear="2008" box="[822,938,472,497]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="B902EAD6FF50ABC99D6911FD45F739D2" box="[822,938,472,497]" italics="true" pageId="188" pageNumber="189">atricornis</emphasis>
</taxonomicName>
(black, horn) refers to the dark antenna of this species (also present in some other species).
</paragraph>
</subSubSection>
</treatment>
</document>