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<document id="442D91A0A7A64B90E7910D43DDBBF6EB" ID-CLB-Dataset="45889" ID-DOI="10.5281/zenodo.279337" ID-GBIF-Dataset="367d3d94-fa26-4f5d-ba39-1fb904401752" ID-ISSN="1175-5326" ID-Zenodo-Dep="279337" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414521621" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03DF87B01F5C8E0186C5FB232784FC75" docLanguage="en" docName="zt03113p052.pdf" docOrigin="Zootaxa 3113" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cephennula gemina Jałoszyński, 2011, sp. n." docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FFE6FFC81F5F8E048652FFD7274EFFD1" masterDocTitle="Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia" masterLastPageNumber="52" masterPageNumber="36" pageNumber="39" updateTime="1698328417371" updateUser="ExternalLinkService">
<mods:mods id="CD8CD6FAF22146B47378EAF90D64B9B2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="12B88BB0FB83491E2511A6F6635FB1E9">
<mods:title id="A966FD0F410E3D6BEA1A0D9D1DC00BB9">Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="C4E68AE72F7B8872913728E5E76A34E4" type="personal">
<mods:role id="579BCA98D9F9AB0D0785DA7B28686993">
<mods:roleTerm id="39F489C42A01FC6D2CB592B47C19B633">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64433BF485E8D58D86A7D2B0FFC26F1A">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="DA14B6C9FF4638A882C8C4643352B4F8">text</mods:typeOfResource>
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<mods:title id="312A7501E4FF81909DBC9AB0A05C3654">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8019EE1867B8AAEA1A5AE47AA12479E6">
<mods:date id="CE4084C8B6D107037E691D9440AA6726">2011</mods:date>
<mods:detail id="3D8FC95E0590AFFA1E772B567D6B3DA3" type="volume">
<mods:number id="B4C06AA9F074C8C36A83C88DF3C24C36">3113</mods:number>
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<mods:classification id="1AD2DD67BAA5391D713E76C33D30E3F6">journal article</mods:classification>
<mods:identifier id="FFBD2C64FD6742BD389367EA4E2D3F74" type="CLB-Dataset">45889</mods:identifier>
<mods:identifier id="0C8EA67CA21D14AB1848E34419E32A30" type="DOI">10.5281/zenodo.279337</mods:identifier>
<mods:identifier id="D5C9A40B997ACA81E03663E8C37DBE09" type="GBIF-Dataset">367d3d94-fa26-4f5d-ba39-1fb904401752</mods:identifier>
<mods:identifier id="4FF06E517A4D091D7879B10D8BB4172E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B037359D48557A7AE986AF2943C7EA8C" type="Zenodo-Dep">279337</mods:identifier>
</mods:mods>
<treatment id="03DF87B01F5C8E0186C5FB232784FC75" ID-DOI="http://doi.org/10.5281/zenodo.6185239" ID-GBIF-Taxon="119424585" ID-Zenodo-Dep="6185239" LSID="urn:lsid:plazi:treatment:03DF87B01F5C8E0186C5FB232784FC75" httpUri="http://treatment.plazi.org/id/03DF87B01F5C8E0186C5FB232784FC75" lastPageId="5" lastPageNumber="41" pageId="3" pageNumber="39">
<subSubSection id="C36C652D1F5C8E0786C5FB232604FAFF" pageId="3" pageNumber="39" type="nomenclature">
<paragraph id="8BC936A61F5C8E0786C5FB23269BFADC" blockId="3.[151,469,1267,1326]" box="[151,469,1267,1294]" pageId="3" pageNumber="39">
<heading id="D08181CA1F5C8E0786C5FB23269BFADC" bold="true" box="[151,469,1267,1294]" fontSize="11" level="1" pageId="3" pageNumber="39" reason="1">
<emphasis id="B902EAB41F5C8E0786C5FB23269BFADC" bold="true" box="[151,469,1267,1294]" pageId="3" pageNumber="39">
<taxonomicName id="4C764D251F5C8E0786C5FB2326C2FADF" ID-CoL="SFQJ" box="[151,396,1268,1294]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="gemina" status="sp. nov.">
<emphasis id="B902EAB41F5C8E0786C5FB2326C2FADF" bold="true" box="[151,396,1268,1294]" italics="true" pageId="3" pageNumber="39">Cephennula gemina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157CF1F5C8E0787C1FB24269BFADC" box="[403,469,1267,1293]" pageId="3" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936A61F5C8E0786C5FAC12604FAFF" blockId="3.[151,469,1267,1326]" box="[151,330,1302,1326]" pageId="3" pageNumber="39">
(
<figureCitation id="134D2A231F5C8E0786CDFAC127A0FAFF" box="[159,238,1302,1326]" captionStart="FIGURES 3 – 7" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[171,1393,210,1915]" captionTargetId="figure@4.[159,1403,193,1935]" captionTargetPageId="4" captionText="FIGURES 3 – 7. Habitus of males. 3 ― Cephennula densepunctata sp. n.; 4 ― Cephennula gemina sp. n.; 5 ― Cephennula minutissima sp. n.; 6 ― Cephennula penrissenana sp. n.; 7 ― Cephennula poringana sp. n. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/279339/files/figure.png" pageId="3" pageNumber="39">Figs. 4</figureCitation>
, 15, 16)
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5C8E0786C5FA88268FF99F" pageId="3" pageNumber="39" type="materials_examined">
<paragraph id="8BC936A61F5C8E0786C5FA88268FF99F" blockId="3.[151,1436,1374,2011]" pageId="3" pageNumber="39">
<emphasis id="B902EAB41F5C8E0786C5FA88269AFAA6" bold="true" box="[151,468,1374,1399]" pageId="3" pageNumber="39">
<typeStatus id="54CD88041F5C8E0786C5FA882646FAA6" box="[151,264,1375,1399]" pageId="3" pageNumber="39" type="holotype">Holotype</typeStatus>
: E
<collectingCountry id="F36176361F5C8E078766FA892680FAA6" box="[308,462,1374,1399]" name="Malaysia" pageId="3" pageNumber="39">MALAYSIA</collectingCountry>
:
</emphasis>
3, two labels: &quot;
<collectionCode id="ED67AE631F5C8E0784C6FA8925B8FAA7" box="[660,758,1374,1398]" pageId="3" pageNumber="39">SABAH</collectionCode>
\ Mt. Kinabalu \
<quantity id="4C8E9B431F5C8E078594FA89232EFAA7" box="[966,1120,1374,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="3" pageNumber="39" unit="m" value="1500.0" valueMax="1550.0" valueMin="1450.0">1450-1550 m</quantity>
\
<date id="FFC810661F5C8E078229FA8923A0FAA7" box="[1147,1262,1374,1398]" pageId="3" pageNumber="39" value="1987-05-23">23.V.1987</date>
\ Burckhardt - Löbl&quot; [white, printed]; &quot;
<emphasis id="B902EAB41F5C8E0787E6FA542526FA4A" box="[436,616,1411,1435]" italics="true" pageId="3" pageNumber="39">CEPHENNULA</emphasis>
\
<taxonomicName id="4C764D251F5C8E0784D4FA542469FA4A" box="[646,807,1411,1435]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="densepunctata">
<emphasis id="B902EAB41F5C8E0784D4FA542469FA4A" box="[646,807,1411,1435]" italics="true" pageId="3" pageNumber="39">densepunctata</emphasis>
</taxonomicName>
m. \ det. P. Jałoszyński, 2011 \
<typeStatus id="54CD88041F5C8E0782F9FA55221BFA4B" box="[1195,1365,1410,1434]" pageId="3" pageNumber="39" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED67AE631F5C8E078752FA712612FA6F" box="[256,348,1446,1470]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="39">MHNG</collectionCode>
).
<emphasis id="B902EAB41F5C8E078722FA7026A6FA6E" bold="true" box="[368,488,1447,1471]" pageId="3" pageNumber="39">
<typeStatus id="54CD88041F5C8E078722FA7026A6FA6E" box="[368,488,1447,1471]" pageId="3" pageNumber="39" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<date id="FFC810661F5C8E0787A8FA712529FA6F" box="[506,615,1446,1471]" pageId="3" pageNumber="39" value="1933-03-13">3 33, 13</date>
ƤƤ): 1 3, 3 ƤƤ, same data as
<typeStatus id="54CD88041F5C8E078588FA71230EFA6F" box="[986,1088,1446,1470]" pageId="3" pageNumber="39" type="holotype">holotype</typeStatus>
; 1 3, 4 ƤƤ, same data except
<quantity id="4C8E9B431F5C8E0786C5FA1D27A3FA33" box="[151,237,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="39" unit="m" value="1500.0">1500 m</quantity>
and
<date id="FFC810661F5C8E078777FA1D26D5FA33" box="[293,411,1482,1506]" pageId="3" pageNumber="39" value="1987-05-21">21.V.1987</date>
; 1 3, 1 Ƥ, same data except
<quantity id="4C8E9B431F5C8E0784B6FA1D2474FA33" box="[740,826,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="39" unit="m" value="1500.0">1500 m</quantity>
and
<date id="FFC810661F5C8E078520FA1D24BFFA33" box="[882,1009,1482,1506]" pageId="3" pageNumber="39" value="1987-04-30">30.IV.1987</date>
; 3 ƤƤ, same data except
<quantity id="4C8E9B431F5C8E078347FA1D2225FA33" box="[1301,1387,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="3" pageNumber="39" unit="m" value="1430.0">1430 m</quantity>
and
<date id="FFC810661F5C8E0786C5FA392643F9D7" box="[151,269,1518,1542]" pageId="3" pageNumber="39" value="1987-05-22">22.V.1987</date>
; 1 Ƥ, &quot;
<collectionCode id="ED67AE631F5C8E078731FA392687F9D7" box="[355,457,1518,1542]" pageId="3" pageNumber="39">SABAH</collectionCode>
: E Mt.Kinabalu \
<quantity id="4C8E9B431F5C8E0784C9FA3925A4F9D7" box="[667,746,1518,1542]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="39" unit="m" value="1150.0">1150m</quantity>
,rte Ranau-Kota \ Kinabalu,
<date id="FFC810661F5C8E078260FA3923EAF9D7" box="[1074,1188,1518,1542]" pageId="3" pageNumber="39" value="1987-05-24">24.V.1987</date>
\ Burckhardt - Löbl&quot; [white, printed]; 1 Ƥ, &quot;BORNEO
<collectionCode id="ED67AE631F5C8E07844EF9C52535F9FB" box="[540,635,1554,1578]" pageId="3" pageNumber="39">SABAH</collectionCode>
Mt. \ Kinabalu Nat.Pk. \ HQ Liwagu River \
<quantity id="4C8E9B431F5C8E0782D9F9C52397F9FB" box="[1163,1241,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="3" pageNumber="39" unit="m" value="1490.0">1490m</quantity>
<date id="FFC810661F5C8E0782B0F9C52229F9FB" box="[1250,1383,1554,1578]" pageId="3" pageNumber="39" value="1987-05-18">18. V. 1987</date>
\ A. Smetana&quot; (
<collectionCode id="ED67AE631F5C8E078743F9E12626F99F" box="[273,360,1590,1614]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="39">MHNG</collectionCode>
,
<collectionCode id="ED67AE631F5C8E078721F9E126FBF99F" box="[371,437,1590,1614]" pageId="3" pageNumber="39">PCPJ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5C8E078695F98D22C5F8F7" pageId="3" pageNumber="39" type="diagnosis">
<paragraph id="8BC936A61F5C8E078695F98D22C5F8F7" blockId="3.[151,1436,1374,2011]" pageId="3" pageNumber="39">
<emphasis id="B902EAB41F5C8E078695F98D260FF9A2" bold="true" box="[199,321,1626,1651]" pageId="3" pageNumber="39">Diagnosis.</emphasis>
Body reaching or nearly reaching
<quantity id="4C8E9B431F5C8E078480F98D245BF9A3" box="[722,789,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="39" unit="mm" value="1.0">1 mm</quantity>
in length; antennal club strongly flattened and compact; punctures on head unevenly distributed, those on sides of vertex, on frons and clypeus separated by spaces 0.5–
<date id="FFC810661F5C8E0783D3F9A922C0F947" box="[1409,1422,1662,1686]" pageId="3" pageNumber="39">2</date>
x as wide as puncture diameter, vertex with impunctate median area; median part of pronotum with dense punctures; sublateral pronotal carinae as long as pronotum; subhumeral carinae about as long as 1/
<date id="FFC810661F5C8E07823AF91123CBF90F" box="[1128,1157,1734,1758]" pageId="3" pageNumber="39">3x</date>
EL and distinctly longer than humeral carinae; aedeagus in ventral view drop-shaped, with triangular and pointed apex of median lobe; endophallus with four pairs of elongate sclerites, of which three pairs are curved laterally and one nearly straight.
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5C8E018695F8E52613FD1D" lastPageId="5" lastPageNumber="41" pageId="3" pageNumber="39" type="description">
<paragraph id="8BC936A61F5C8E078695F8E525A1F8BF" blockId="3.[151,1436,1374,2011]" pageId="3" pageNumber="39">
<emphasis id="B902EAB41F5C8E078695F8E52617F89A" bold="true" box="[199,345,1842,1867]" pageId="3" pageNumber="39">Description.</emphasis>
Body of male (
<figureCitation id="134D2A231F5C8E07844EF8E5252AF89B" box="[540,612,1842,1866]" captionStart="FIGURES 3 – 7" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[171,1393,210,1915]" captionTargetId="figure@4.[159,1403,193,1935]" captionTargetPageId="4" captionText="FIGURES 3 – 7. Habitus of males. 3 ― Cephennula densepunctata sp. n.; 4 ― Cephennula gemina sp. n.; 5 ― Cephennula minutissima sp. n.; 6 ― Cephennula penrissenana sp. n.; 7 ― Cephennula poringana sp. n. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/279339/files/figure.png" pageId="3" pageNumber="39">Fig. 4</figureCitation>
) externally nearly identical as
<taxonomicName id="4C764D251F5C8E07858AF8E423EEF89A" box="[984,1184,1843,1867]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="densepunctata">
<emphasis id="B902EAB41F5C8E07858AF8E423EEF89A" box="[984,1184,1843,1867]" italics="true" pageId="3" pageNumber="39">C. densepunctata</emphasis>
</taxonomicName>
, only differences are described below. BL 0.98–1.00 mm (mean
<quantity id="4C8E9B431F5C8E078429F88125ADF8BF" box="[635,739,1878,1902]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="39" unit="mm" value="0.98">0.98 mm</quantity>
).
</paragraph>
<paragraph id="8BC936A61F5C8E078695F8AD2393F80B" blockId="3.[151,1436,1374,2011]" pageId="3" pageNumber="39">
Head unevenly covered with moderately large and deep, sharply marked punctures, on sides of vertex, on frons and clypeus separated by spaces 0.5–
<date id="FFC810661F5C8E07846EF8492507F867" box="[572,585,1950,1974]" pageId="3" pageNumber="39">2</date>
x as wide as puncture diameter, vertex in middle impunctate; HL
<quantity id="4C8E9B431F5C8E07837FF849278DF80B" metricMagnitude="-4" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="3" pageNumber="39" unit="mm" value="0.12" valueMax="0.13" valueMin="0.11">0.11–0.13 mm</quantity>
(mean
<quantity id="4C8E9B431F5C8E078744F8152630F80B" box="[278,382,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="39" unit="mm" value="0.12">0.12 mm</quantity>
), HW
<quantity id="4C8E9B431F5C8E078795F8152524F80B" box="[455,618,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="3" pageNumber="39" unit="mm" value="0.22499999999999998" valueMax="0.24" valueMin="0.21">0.21–0.24 mm</quantity>
(mean
<quantity id="4C8E9B431F5C8E0784EFF815246BF80B" box="[701,805,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="39" unit="mm" value="0.22">0.22 mm</quantity>
). AnL
<quantity id="4C8E9B431F5C8E078521F8152358F80B" box="[883,1046,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="3" pageNumber="39" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.35–0.38 mm</quantity>
(mean
<quantity id="4C8E9B431F5C8E07823BF815239FF80B" box="[1129,1233,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="3" pageNumber="39" unit="mm" value="0.36">0.36 mm</quantity>
).
</paragraph>
<caption id="DF09662E1F5B8E0086C5F86E23F3F83E" httpUri="https://zenodo.org/record/279339/files/figure.png" pageId="4" pageNumber="40" targetBox="[171,1393,210,1915]" targetPageId="4">
<paragraph id="8BC936A61F5B8E0086C5F86E23F3F83E" blockId="4.[151,1436,1977,2031]" pageId="4" pageNumber="40">
<emphasis id="B902EAB41F5B8E0086C5F86E2677F81E" bold="true" box="[151,313,1977,2000]" pageId="4" pageNumber="40">FIGURES 3–7.</emphasis>
Habitus of males. 3―
<taxonomicName id="4C764D251F5B8E008449F86D2460F801" box="[539,814,1978,2000]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">
<emphasis id="B902EAB41F5B8E008449F86D2460F801" box="[539,814,1978,2000]" italics="true" pageId="4" pageNumber="40">Cephennula densepunctata</emphasis>
</taxonomicName>
<emphasis id="B902EAB41F5B8E008561F86D2422F81E" bold="true" box="[819,876,1978,1999]" pageId="4" pageNumber="40">
<taxonomicNameLabel id="A23157CF1F5B8E008561F86D2422F81E" box="[819,876,1978,1999]" pageId="4" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 4―
<taxonomicName id="4C764D251F5B8E0085CCF86D2326F801" box="[926,1128,1978,2000]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="gemina" status="sp. nov.">
<emphasis id="B902EAB41F5B8E0085CCF86D2326F801" box="[926,1128,1978,2000]" italics="true" pageId="4" pageNumber="40">Cephennula gemina</emphasis>
</taxonomicName>
<emphasis id="B902EAB41F5B8E00823DF86D23E9F81E" bold="true" box="[1135,1191,1978,1999]" pageId="4" pageNumber="40">
<taxonomicNameLabel id="A23157CF1F5B8E00823DF86D23E9F81E" box="[1135,1191,1978,1999]" pageId="4" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 5―
<taxonomicName id="4C764D251F5B8E008288F86D2796F83E" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov.">
<emphasis id="B902EAB41F5B8E008288F86D2796F83E" italics="true" pageId="4" pageNumber="40">Cephennula minutissima</emphasis>
</taxonomicName>
<emphasis id="B902EAB41F5B8E00868CF80E2659F83F" bold="true" box="[222,279,2009,2030]" pageId="4" pageNumber="40">
<taxonomicNameLabel id="A23157CF1F5B8E00868CF80E2659F83F" box="[222,279,2009,2030]" pageId="4" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 6―
<taxonomicName id="4C764D251F5B8E008718F80E251CF83E" box="[330,594,2009,2031]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="penrissenana" status="sp. nov.">
<emphasis id="B902EAB41F5B8E008718F80E251CF83E" box="[330,594,2009,2031]" italics="true" pageId="4" pageNumber="40">Cephennula penrissenana</emphasis>
</taxonomicName>
<emphasis id="B902EAB41F5B8E00840AF80E25DFF83F" bold="true" box="[600,657,2009,2030]" pageId="4" pageNumber="40">
<taxonomicNameLabel id="A23157CF1F5B8E00840AF80E25DFF83F" box="[600,657,2009,2030]" pageId="4" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 7―
<taxonomicName id="4C764D251F5B8E008496F80E24E0F83E" box="[708,942,2009,2031]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="poringana" status="sp. nov.">
<emphasis id="B902EAB41F5B8E008496F80E24E0F83E" box="[708,942,2009,2031]" italics="true" pageId="4" pageNumber="40">Cephennula poringana</emphasis>
</taxonomicName>
<emphasis id="B902EAB41F5B8E0085E7F80E24A0F83F" bold="true" box="[949,1006,2009,2030]" pageId="4" pageNumber="40">
<taxonomicNameLabel id="A23157CF1F5B8E0085E7F80E24A0F83F" box="[949,1006,2009,2030]" pageId="4" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC936A61F5A8E018695FF402645FEEE" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
Median part of pronotum covered with punctures similar to those on head and becoming gradually larger and denser towards sides, those on median part of disc separated by spaces approximately equal to puncture diameter or slightly wider, punctures between sublateral carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other; PL
<quantity id="4C8E9B431F5A8E018554FED324E7FECD" box="[774,937,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="5" pageNumber="41" unit="mm" value="0.305" valueMax="0.31" valueMin="0.3">0.30–0.31 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0185ACFED32329FECD" box="[1022,1127,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="41" unit="mm" value="0.3">0.30 mm</quantity>
), PW
<quantity id="4C8E9B431F5A8E0182FEFED32201FECD" box="[1196,1359,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="5" pageNumber="41" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0186C5FEF027B1FEEE" box="[151,255,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="41" unit="mm" value="0.43">0.43 mm</quantity>
).
</paragraph>
<paragraph id="8BC936A61F5A8E018695FE9B2611FE7D" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
Elytra covered with fine and sparse punctures, EL
<quantity id="4C8E9B431F5A8E018551FE9B24E9FEB5" box="[771,935,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="5" pageNumber="41" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0185AEFE9B232AFEB5" box="[1020,1124,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="41" unit="mm" value="0.56">0.56 mm</quantity>
), EW
<quantity id="4C8E9B431F5A8E0182F9FE9B2201FEB5" box="[1195,1359,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="5" pageNumber="41" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0186C5FEB8264EFE56" box="[151,256,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="41" unit="mm" value="0.49">0.49 mm</quantity>
), EI 1.13–1.18 (mean 1.15); subhumeral carinae about as long as 1/
<date id="FFC810661F5A8E018253FEB82351FE56" box="[1025,1055,367,391]" pageId="5" pageNumber="41">3x</date>
EL; humeral carinae less distinct and much shorter.
</paragraph>
<paragraph id="8BC936A61F5A8E018695FE602530FDED" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
Aedeagus (Figs. 15, 16) in ventral view drop-shaped with narrow, subtriangular and pointed apex; AeL
<quantity id="4C8E9B431F5A8E018339FE602788FE25" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="41" unit="mm" value="0.23">0.23 mm</quantity>
. Endophallus darkly sclerotized, with four pairs of elongate sclerites—three pairs strongly curved laterally and one median pair nearly straight and directed towards apex of aedeagus; parameres slender, not exceeding apex of median lobe, each bearing two apical setae.
</paragraph>
<paragraph id="8BC936A61F5A8E018695FD902613FD1D" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
Female. Externally indistinguishable from males. BL 1.00–
<quantity id="4C8E9B431F5A8E01852FFD9024AAFD8E" box="[893,996,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="5" pageNumber="41" unit="mm" value="1.01">1.01 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E018212FD9023E3FD8E" box="[1088,1197,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="5" pageNumber="41" unit="mm" value="1.01">1.01 mm</quantity>
); HL
<quantity id="4C8E9B431F5A8E0182A4FD9022D2FD8E" box="[1270,1436,583,607]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="5" pageNumber="41" unit="mm" value="0.12" valueMax="0.13" valueMin="0.11">0.11–0.13 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0186B4FDBB2601FD55" box="[230,335,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="41" unit="mm" value="0.12">0.12 mm</quantity>
), HW
<quantity id="4C8E9B431F5A8E0187CEFDBB250EFD55" box="[412,576,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="5" pageNumber="41" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.23–0.24 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0184CAFDBB244CFD55" box="[664,770,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="41" unit="mm" value="0.24">0.24 mm</quantity>
), AnL
<quantity id="4C8E9B431F5A8E018501FDBB24B9FD55" box="[851,1015,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="5" pageNumber="41" unit="mm" value="0.36" valueMax="0.38" valueMin="0.34">0.34–0.38 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E01821DFDBB23F6FD55" box="[1103,1208,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="41" unit="mm" value="0.35">0.35 mm</quantity>
); PL
<quantity id="4C8E9B431F5A8E0182A5FDBB22D2FD55" box="[1271,1436,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="5" pageNumber="41" unit="mm" value="0.32" valueMax="0.33" valueMin="0.31">0.31–0.33 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0186B6FD582605FD76" box="[228,331,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="41" unit="mm" value="0.32">0.32 mm</quantity>
), PW
<quantity id="4C8E9B431F5A8E0187DDFD58257FFD76" box="[399,561,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="5" pageNumber="41" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E0184D6FD5825A2FD76" box="[644,748,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="41" unit="mm" value="0.46">0.46 mm</quantity>
); EL
<quantity id="4C8E9B431F5A8E01857AFD582485FD76" box="[808,971,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="5.8" metricValueMin="5.6" pageId="5" pageNumber="41" unit="mm" value="0.57" valueMax="0.58" valueMin="0.56">0.56–0.58 mm</quantity>
(mean
<quantity id="4C8E9B431F5A8E01824CFD5823C8FD76" box="[1054,1158,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="5" pageNumber="41" unit="mm" value="0.57">0.57 mm</quantity>
), EW
<quantity id="4C8E9B431F5A8E018299FD58227FFD76" box="[1227,1329,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="41" unit="mm" value="0.5">0.50 mm</quantity>
, EI 1.13– 1.15 (mean 1.15).
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5A8E018695FD002582FD3E" box="[199,716,727,752]" pageId="5" pageNumber="41" type="distribution">
<paragraph id="8BC936A61F5A8E018695FD002582FD3E" blockId="5.[151,1437,151,932]" box="[199,716,727,752]" pageId="5" pageNumber="41">
<emphasis id="B902EAB41F5A8E018695FD00262EFD21" bold="true" box="[199,352,727,752]" pageId="5" pageNumber="41">Distribution.</emphasis>
East
<collectingCountry id="F36176361F5A8E0187CCFD00254BFD3E" box="[414,517,727,751]" name="Malaysia" pageId="5" pageNumber="41">Malaysia</collectingCountry>
(Borneo: Sabah).
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5A8E018695FD2B254AFCE9" pageId="5" pageNumber="41" type="etymology">
<paragraph id="8BC936A61F5A8E018695FD2B254AFCE9" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B902EAB41F5A8E018695FD2B2600FCC5" bold="true" box="[199,334,764,788]" pageId="5" pageNumber="41">Etymology.</emphasis>
The name is derived from Latin
<emphasis id="B902EAB41F5A8E01849EFD2A242CFCC5" box="[716,866,765,788]" italics="true" pageId="5" pageNumber="41">
soror
<taxonomicName id="4C764D251F5A8E018542FD2A242CFCC5" box="[784,866,765,788]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="gemina">gemina</taxonomicName>
</emphasis>
(a twin sister) and refers to the high similarity of this species to
<taxonomicName id="4C764D251F5A8E018769FCF726B0FCE9" box="[315,510,800,824]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="densepunctata">
<emphasis id="B902EAB41F5A8E018769FCF726B0FCE9" box="[315,510,800,824]" italics="true" pageId="5" pageNumber="41">C. densepunctata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F5A8E018695FC932784FC75" pageId="5" pageNumber="41" type="discussion">
<paragraph id="8BC936A61F5A8E018695FC932784FC75" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B902EAB41F5A8E018695FC932675FC8D" bold="true" box="[199,315,836,860]" pageId="5" pageNumber="41">Remarks.</emphasis>
This species is externally nearly identical to
<taxonomicName id="4C764D251F5A8E018516FC922349FC8D" box="[836,1031,836,860]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="densepunctata">
<emphasis id="B902EAB41F5A8E018516FC922349FC8D" box="[836,1031,836,860]" italics="true" pageId="5" pageNumber="41">C. densepunctata</emphasis>
</taxonomicName>
. In addition to the clearly different aedeagus
<taxonomicName id="4C764D251F5A8E018754FCBF2635FC51" box="[262,379,872,896]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="gemina">
<emphasis id="B902EAB41F5A8E018754FCBF2635FC51" box="[262,379,872,896]" italics="true" pageId="5" pageNumber="41">C. gemina</emphasis>
</taxonomicName>
can be distinguished also by distinctly smaller and sparser punctures on the head and pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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