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<document id="3CCBB80260B022077E55555B56BCD423" ID-DOI="10.1080/00222930310001618895" ID-ISSN="1464-5262" ID-Zenodo-Dep="4676180" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617981093153" checkinUser="carolina" docAuthor="Marino, Pablo I. &amp; Spinelli, Gustavo R." docDate="2004" docId="03DF2D33AB49FFB5FE09265BFE90E557" docLanguage="en" docName="JNATHIST.38.2251-2262.pdf" docOrigin="Journal of Natural History 38 (17)" docSource="http://dx.doi.org/10.1080/00222930310001618895" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Forcipomyia (Trichohelea) tehuelche Marino &amp; Spinelli 2004, sp. n." docType="treatment" docVersion="3" lastPageNumber="2261" masterDocId="FFE6554BAB42FFB9FF90241EFFF9E057" masterDocTitle="Descriptions of the Patagonian species of the subgenus Trichohelea of Forcipomyia, with a key to the Neotropical species (Diptera: Ceratopogonidae)" masterLastPageNumber="2262" masterPageNumber="2251" pageNumber="2260" updateTime="1698903867452" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DE639236F9620C30753EF2B9EF930AE2">Descriptions of the Patagonian species of the subgenus Trichohelea of Forcipomyia, with a key to the Neotropical species (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="7A5816BAC04CAFBCC0EF36F8840342BF">Marino, Pablo I.</mods:namePart>
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<mods:namePart id="5F91A67C1707D560195063BDF1E5D241">Spinelli, Gustavo R.</mods:namePart>
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<mods:title id="FAE8996814D2BA01EB240A3752723D8D">Journal of Natural History</mods:title>
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<mods:part id="B6AEFE1F223ECFD153796215CB951778">
<mods:date id="545C949CD1E12110F22118EEAFF6FBB7">2004</mods:date>
<mods:detail id="85DB6FC6EDA5D557A370D12F824B1433" type="pubDate">
<mods:number id="72F4B673D155F8DC78B06840C2DE052B">2010-05-25</mods:number>
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<mods:number id="B28718462A71831522A12B348D2442A7">38</mods:number>
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<mods:number id="8ED3F39CF17E6059231EE67EA4DAB5FC">17</mods:number>
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<mods:classification id="CE69DE466A37E30EA44D8408DE89CACE">journal article</mods:classification>
<mods:identifier id="3C055C3736207CE2E5635C2B030EE128" type="DOI">10.1080/00222930310001618895</mods:identifier>
<mods:identifier id="14CEB084B2E816A1115B3FFD007D4B17" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D9406DE743F2A5F6A62B0CCA0B2B234C" type="Zenodo-Dep">4676180</mods:identifier>
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<treatment id="03DF2D33AB49FFB5FE09265BFE90E557" LSID="urn:lsid:plazi:treatment:03DF2D33AB49FFB5FE09265BFE90E557" httpUri="http://treatment.plazi.org/id/03DF2D33AB49FFB5FE09265BFE90E557" lastPageId="12" lastPageNumber="2261" pageId="11" pageNumber="2260">
<subSubSection id="C36CCFAEAB49FFB2FE09265BFD27E22A" pageId="11" pageNumber="2260" type="nomenclature">
<paragraph id="8BC99C25AB49FFB2FE09265BFC86E20A" blockId="11.[409,895,581,605]" box="[409,895,581,605]" pageId="11" pageNumber="2260">
<heading id="D0812B49AB49FFB2FE09265BFC86E20A" box="[409,895,581,605]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="2260" reason="5">
<taxonomicName id="4C76E7A6AB49FFB2FE09265BFCCCE20A" ID-CoL="8C75Q" authority="Marino &amp; Spinelli, 2004" authorityName="Marino &amp; Spinelli" authorityYear="2004" box="[409,821,581,605]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="2260" phylum="Arthropoda" rank="species" species="tehuelche" status="sp. nov." subGenus="Trichohelea">
<emphasis id="B9024037AB49FFB2FE09265BFCCCE20A" bold="true" box="[409,821,581,605]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FE09265BFDDCE20A" bold="true" box="[409,549,581,605]" italics="true" pageId="11" pageNumber="2260">Forcipomyia</emphasis>
(
<emphasis id="B9024037AB49FFB2FDA8265BFD43E20A" bold="true" box="[568,698,581,605]" italics="true" pageId="11" pageNumber="2260">Trichohelea</emphasis>
)
<emphasis id="B9024037AB49FFB2FD5E265BFCCCE20A" bold="true" box="[718,821,581,605]" italics="true" pageId="11" pageNumber="2260">tehuelche</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231FD4CAB49FFB2FCAF265BFC86E20A" box="[831,895,581,605]" pageId="11" pageNumber="2260" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BC99C25AB49FFB2FDAA267BFD27E22A" blockId="11.[570,734,613,637]" box="[570,734,613,637]" pageId="11" pageNumber="2260">(figures 1823)</paragraph>
</subSubSection>
<subSubSection id="C36CCFAEAB49FFB5FF0A2688FE90E557" lastPageId="12" lastPageNumber="2261" pageId="11" pageNumber="2260" type="diagnosis">
<paragraph id="8BC99C25AB49FFB2FF0A2688FEFEE2F9" blockId="11.[154,1110,662,888]" box="[154,263,662,686]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF0A2688FEFEE2F9" box="[154,263,662,686]" italics="true" pageId="11" pageNumber="2260">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC99C25AB49FFB2FF5226A8FCE3E32F" blockId="11.[154,1110,662,888]" pageId="11" pageNumber="2260">A medium-sized, brown species; flagellomeres 18 distinctly broader than long; third segment of maxillary palpi broadly swollen, sensory pit deep opening by a small pore; CR 0.55; tarsal claws equal, small, bifid at extreme apex; female genital sclerotization open, subquadrangular; two pyriform spermathecae; aedeagus stout with distal medial process, basal arch extending to one-tenth of total length; parameres inverted U-shaped, lateral arms subparallel.</paragraph>
<paragraph id="8BC99C25AB49FFB2FF0A27B2FF13E393" blockId="11.[154,1111,940,1733]" box="[154,234,940,964]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF0A27B2FF13E393" box="[154,234,940,964]" italics="true" pageId="11" pageNumber="2260">Female</emphasis>
</paragraph>
<paragraph id="8BC99C25AB49FFB2FF5227D2FBABE3B3" blockId="11.[154,1111,940,1733]" box="[194,1106,972,996]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF5227D2FEFFE3B3" box="[194,262,972,996]" italics="true" pageId="11" pageNumber="2260">Wing.</emphasis>
Length
<quantity id="4C8E31C0AB49FFB2FEFB27D2FE32E3B3" box="[363,459,972,996]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="11" pageNumber="2260" unit="mm" value="1.26">1.26 mm</quantity>
(1.221.30,
<emphasis id="B9024037AB49FFB2FDCC27D2FD89E3B3" box="[604,624,972,996]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2); breadth
<quantity id="4C8E31C0AB49FFB2FC8627D2FC8CE3B3" box="[790,885,972,996]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="2260" unit="mm" value="0.56">0.56 mm</quantity>
(0.540.58,
<emphasis id="B9024037AB49FFB2FB9627D2FBE3E3B3" box="[1030,1050,972,996]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2).
</paragraph>
<paragraph id="8BC99C25AB49FFB2FF5227F2FE07E563" blockId="11.[154,1111,940,1733]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF5227F2FEFFE453" box="[194,262,1004,1028]" italics="true" pageId="11" pageNumber="2260">Head.</emphasis>
Dark brown. Vertex pilose. Eyes bare, contiguous by a distance equal to diameter of three ommatidia. Flagellum of antenna with flagellomeres 18 distinctly broader than long; 912 elongated, cylindrical, approximately 2.5 times longer than greatest breadth; distal flagellomere slightly longer than 912; AR 1.68 (1.621.74,
<emphasis id="B9024037AB49FFB2FEF7206CFE82E4DD" box="[359,379,1138,1162]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2). Palpus short; third segment broadly swollen at midportion, nearly as long as the combined length of fourth and fifth segments, with deep sensory pit opening by a small rounded pore; fifth segment slightly narrower than 4;
<collectingRegion id="49B252C7AB49FFB2FEEE20C6FE5AE4A7" box="[382,419,1240,1264]" country="Brazil" name="Para" pageId="11" pageNumber="2260">PR</collectingRegion>
1.70 (1.591.81,
<emphasis id="B9024037AB49FFB2FD1120C6FD6CE4A7" box="[641,661,1240,1264]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2). Mandible abruptly tapering to pointed apex, armed with 10 very small teeth. Maxilla slender, nearly straight, with six pairs of minute teeth.
</paragraph>
<paragraph id="8BC99C25AB49FFB2FF522122FE66E635" blockId="11.[154,1111,940,1733]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF522122FEE5E503" box="[194,284,1340,1364]" italics="true" pageId="11" pageNumber="2260">Thorax.</emphasis>
Scutum brown, humeral areas slightly paler; scutellum pale brown, with approximately 20 strong setae and 15 slender ones. Legs pale brown, densely covered by hairs; prothoracic TR 1.93 (1.921.93,
<emphasis id="B9024037AB49FFB2FCA32160FCBEE5C1" box="[819,839,1406,1430]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2), mesothoracic TR 1.87 (1.861.87,
<emphasis id="B9024037AB49FFB2FEF421BEFE81E5EF" box="[356,376,1440,1464]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2), metathoracic TR 1.88 (1.871.88,
<emphasis id="B9024037AB49FFB2FCF321BEFC8EE5EF" box="[867,887,1440,1464]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2). Claws small, very curved, moderately stout, bifid at extreme apex; empodium well developed. Wing densely covered with macrotrichia, more abundant on anterior margin; a small pale area situated immediately distad to the second radial cell; first radial cell almost obliterated, second radial cell well formed; CR 0.55 (0.540.55,
<emphasis id="B9024037AB49FFB2FF0A2255FF57E634" box="[154,174,1611,1635]" italics="true" pageId="11" pageNumber="2260">N</emphasis>
~2). Halter whitish.
</paragraph>
<paragraph id="8BC99C25AB49FFB2FF522275FD05E693" blockId="11.[154,1111,940,1733]" pageId="11" pageNumber="2260">
<emphasis id="B9024037AB49FFB2FF522275FECBE6D4" box="[194,306,1643,1667]" italics="true" pageId="11" pageNumber="2260">Abdomen.</emphasis>
Pale brown, pilose. Genital sclerotization open, subquadrangular. Two pyriform, well-sclerotized spermathecae, subequal, measuring 0.062×
<quantity id="4C8E31C0AB49FFB2FF0A22B2FEFEE693" box="[154,263,1708,1732]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="2260" unit="mm" value="0.05">0.050 mm</quantity>
and 0.052×
<quantity id="4C8E31C0AB49FFB2FE0E22B2FDF7E693" box="[414,526,1708,1732]" metricMagnitude="-5" metricUnit="m" metricValue="4.3999999999999995" pageId="11" pageNumber="2260" unit="mm" value="0.044">0.044 mm</quantity>
, necks of
<quantity id="4C8E31C0AB49FFB2FD1822B2FD01E693" box="[648,760,1708,1732]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="2260" unit="mm" value="0.006">0.006 mm</quantity>
.
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F24B0FF23E091" blockId="12.[159,1116,174,364]" box="[159,218,174,198]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF0F24B0FF23E091" box="[159,218,174,198]" italics="true" pageId="12" pageNumber="2261">Male</emphasis>
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF5724D0FD55E0B1" blockId="12.[159,1116,174,364]" box="[199,684,206,230]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF5724D0FEF2E0B1" box="[199,267,206,230]" italics="true" pageId="12" pageNumber="2261">Wing.</emphasis>
Length
<quantity id="4C8E31C0AB4EFFB5FEE124D0FE2AE0B1" box="[369,467,206,230]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="12" pageNumber="2261" unit="mm" value="1.38">1.38 mm</quantity>
; breadth
<quantity id="4C8E31C0AB4EFFB5FDD524D0FD5EE0B1" box="[581,679,206,230]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="2261" unit="mm" value="0.46">0.46 mm</quantity>
.
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF5724F0FBA5E13B" blockId="12.[159,1116,174,364]" pageId="12" pageNumber="2261">Similar to female with the usual sexual differences: eyes bare except inner margin with sparse pilosity; flagellomeres 19 similar in length, 1013 (figure 18) in proportion of 56-43-35-47; AR 1.22; palpus (figure 19) with third segment slightly shorter than the combined length of fourth and fifth segments, segments in</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F2569FBDFE184" blockId="12.[159,1116,375,599]" pageId="12" pageNumber="2261">
proportion of 11-19-30-18-18;
<collectingRegion id="49B252C7AB4EFFB5FE6B2569FDD9E1D8" box="[507,544,375,399]" country="Brazil" name="Para" pageId="12" pageNumber="2261">PR</collectingRegion>
2.5; tarsal claws equal, small, 0.5 times as long as tarsomere 5, bifid at extreme apex, empodium absent; prothoracic TR 1.44, mesothoracic TR 1.53, metathoracic TR 1.83; wing (figure 20) with CR 0.46.
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF5725C2FBA5E200" blockId="12.[159,1116,375,599]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF5725C2FED5E1A3" box="[199,300,476,500]" italics="true" pageId="12" pageNumber="2261">Genitalia</emphasis>
(figure 21). Sternite 9 stout, 0.66 times longer than greatest breadth, with narrow, shallow caudomedian excavation; tergite 9 tapering, with conspicuous, pilose apicolateral processes; sternite 10 spiculate. Gonocoxite 2.3 times longer than greatest (basal) breadth; gonostylus 0.85 as long as gonocoxite, slen-
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F267CFC0FE355" blockId="12.[159,1116,610,770]" pageId="12" pageNumber="2261">der, nearly straight, tip pointed. Aedeagus stout, slightly sclerotized; basal arms short, recurved; basal arch very low, extending to one-tenth of total length; lateral arms convex; distal portion very lightly sclerotized, tapering to pointed tip, with a pair of subapical, lateral pointed processes (figure 22). Parameres (figure 23) highly sclerotized, inverted U-shaped, lateral arms subparallel.</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F275BFEDDE30A" blockId="12.[159,1116,837,927]" box="[159,292,837,861]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF0F275BFEDDE30A" box="[159,292,837,861]" italics="true" pageId="12" pageNumber="2261">Distribution</emphasis>
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF57277BFEE1E3C8" blockId="12.[159,1116,837,927]" pageId="12" pageNumber="2261">
In subantarctic forest of southern Patagonia (
<collectingRegion id="49B252C7AB4EFFB5FD40277BFC42E32A" box="[720,955,869,893]" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="2261">Santa Cruz province</collectingRegion>
in
<collectingCountry id="F361DCB5AB4EFFB5FC70277BFBADE32A" box="[992,1108,869,893]" name="Argentina" pageId="12" pageNumber="2261">Argentina</collectingCountry>
) (
<figureCitation id="134D80A0AB4EFFB5FF372799FEF5E3C8" box="[167,268,903,927]" captionStart="FIG" captionStartId="10.[202,218,1498,1520]" captionTargetBox="[159,1117,170,1473]" captionTargetId="figure-18@10.[159,1117,170,1473]" captionTargetPageId="10" captionText="FIG. 24. Known localities for the Patagonian species of the subgenus Trichohelea." figureDoi="http://doi.org/10.5281/zenodo.4676187" httpUri="https://zenodo.org/record/4676187/files/figure.png" pageId="12" pageNumber="2261">figure 24</figureCitation>
).
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F27CAFBA2E45C" blockId="12.[159,1115,980,1035]" box="[159,1115,980,1035]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF0F27CAFF18E3BB" box="[159,225,980,1004]" italics="true" pageId="12" pageNumber="2261">Types</emphasis>
<materialsCitation id="3B1E9678AB4EFFB5FF5727EDFBA2E45C" box="[199,1115,1011,1035]" country="Argentina" location="Lake del Desierto" pageId="12" pageNumber="2261" specimenCount="2" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="54CD2287AB4EFFB5FF5727EDFECBE45C" box="[199,306,1011,1035]" pageId="12" pageNumber="2261" type="holotype">Holotype</typeStatus>
„;
<typeStatus id="54CD2287AB4EFFB5FEF027EDFE42E45C" box="[352,443,1011,1035]" pageId="12" pageNumber="2261" type="allotype">allotype</typeStatus>
”,
<collectingCountry id="F361DCB5AB4EFFB5FE7527EDFDA3E45C" box="[485,602,1011,1035]" name="Argentina" pageId="12" pageNumber="2261">Argentina</collectingCountry>
,
<collectingRegion id="49B252C7AB4EFFB5FDF927EDFCCEE45C" box="[617,823,1011,1035]" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="2261">Santa Cruz Prov.</collectingRegion>
,
<location id="8EA9CAFEAB4EFFB5FCD527EDFBE1E45C" LSID="urn:lsid:plazi:treatment:03DF2D33AB49FFB5FE09265BFE90E557:8EA9CAFEAB4EFFB5FCD527EDFBE1E45C" box="[837,1048,1011,1035]" country="Argentina" name="Lake del Desierto" pageId="12" pageNumber="2261" stateProvince="Santa Cruz">Lake del Desierto</location>
, 911
</materialsCitation>
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F2008FEA6E407" blockId="12.[159,1116,1046,1104]" pageId="12" pageNumber="2261">
December 1996, G. Spinelli, malaise trap;
<typeStatus id="54CD2287AB4EFFB5FD112008FD1CE479" box="[641,741,1046,1070]" pageId="12" pageNumber="2261" type="paratype">paratype</typeStatus>
”, same data except
<date id="FFC8BAE5AB4EFFB5FC442008FF2EE407" pageId="12" pageNumber="2261" value="1996-12-09">9 December 1996</date>
, sweep net.
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF0F209AFEEDE4CB" blockId="12.[159,1116,1156,1280]" box="[159,276,1156,1180]" pageId="12" pageNumber="2261">
<emphasis id="B9024037AB4EFFB5FF0F209AFEEDE4CB" box="[159,276,1156,1180]" italics="true" pageId="12" pageNumber="2261">Discussion</emphasis>
</paragraph>
<paragraph id="8BC99C25AB4EFFB5FF5720BAFE90E557" blockId="12.[159,1116,1156,1280]" pageId="12" pageNumber="2261">
Females of
<emphasis id="B9024037AB4EFFB5FEC620BAFE9FE4EB" box="[342,358,1188,1212]" italics="true" pageId="12" pageNumber="2261">F</emphasis>
.
<taxonomicName id="4C76E7A6AB4EFFB5FEE920BAFDEEE4EB" authorityName="Marino &amp; Spinelli" authorityYear="2004" box="[377,535,1188,1212]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="2261" phylum="Arthropoda" rank="species" species="tehuelche" subGenus="Trichohelea">
(
<emphasis id="B9024037AB4EFFB5FE1220BAFE6DE4EB" box="[386,404,1188,1212]" italics="true" pageId="12" pageNumber="2261">T</emphasis>
.)
<emphasis id="B9024037AB4EFFB5FE2020BAFDEEE4EB" box="[432,535,1188,1212]" italics="true" pageId="12" pageNumber="2261">tehuelche</emphasis>
</taxonomicName>
are barely distinguishable from those of
<emphasis id="B9024037AB4EFFB5FB9D20BAFBE4E4EB" box="[1037,1053,1188,1212]" italics="true" pageId="12" pageNumber="2261">F</emphasis>
.
<taxonomicName id="4C76E7A6AB4EFFB5FBA120BAFF0CE488" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="2261" phylum="Arthropoda" rank="species" species="limnetis" subGenus="Trichohelea">
(
<emphasis id="B9024037AB4EFFB5FBAA20BAFBB5E4EB" box="[1082,1100,1188,1212]" italics="true" pageId="12" pageNumber="2261">T</emphasis>
.)
<emphasis id="B9024037AB4EFFB5FF0F20D9FF0CE488" box="[159,245,1223,1247]" italics="true" pageId="12" pageNumber="2261">limnetis</emphasis>
</taxonomicName>
. Characters to distinguish their males may be found in the discussion under the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>