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<document id="8AAD7564351D571291EF8A35BBFD85E5" ID-ISSN="1536-9307" ID-ZooBank="07C4C2D6-E904-41DF-9B26-B7C80C689CDC" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638264206251" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2019" docId="03DD87EBFFC7FFF74680FF3930D4FA0A" docLanguage="en" docName="Euscorpius.294.1-16.pdf" docOrigin="Euscorpius 294" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Neobuthus haeckeli Kovařík 2019, sp. n." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="FFE4FF93FFC3FFFF455CFFD33374FFEA" masterDocTitle="Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXII. Two new species of Neobuthus from Somaliland (Buthidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="3" updateTime="1699436607717" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="80C67859A36997A9C780BA836C87E854" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E18B890734FFA2A28D58B8272DC06B87">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXII. Two new species of Neobuthus from Somaliland (Buthidae)</mods:title>
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<mods:name id="2B63D29341E57BD33114C6F08D21389F" type="personal">
<mods:role id="67ACCAF18C6C3E8AB6408FB67CF95E8B">
<mods:roleTerm id="15EACA19663F0EC60B229E3E6F665280">Author</mods:roleTerm>
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<mods:namePart id="115DFB3316F5496AE29030D9FF03B984">Kovařík, František</mods:namePart>
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<mods:typeOfResource id="F203B0DDD183EAFC7A74F82F9AFC7246">text</mods:typeOfResource>
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<mods:title id="4080DB3A32D62A50079ABF7D4733695B">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="E3FC928D7C0B09126A4AF052D39DADB0">
<mods:date id="9DDC25E579846A316CCAB71E382E3541">2019</mods:date>
<mods:detail id="724B2B456520E60A1426E0B9BDC5237C" type="volume">
<mods:number id="7A413807572C4322F856ABA37506B0D3">294</mods:number>
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<mods:start id="545FD36C717FCE11167BAEB55FF591F0">1</mods:start>
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<mods:classification id="BD820BD1A57F9C4A3BCE16F12BAD73B3">journal article</mods:classification>
<mods:identifier id="64C696760355FA7728D1204DD1EEBE5E" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="55AC94F41F8B647D1664AC3961600C84" type="ZooBank">07C4C2D6-E904-41DF-9B26-B7C80C689CDC</mods:identifier>
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<treatment id="03DD87EBFFC7FFF74680FF3930D4FA0A" LSID="urn:lsid:plazi:treatment:03DD87EBFFC7FFF74680FF3930D4FA0A" httpUri="http://treatment.plazi.org/id/03DD87EBFFC7FFF74680FF3930D4FA0A" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3">
<subSubSection id="C36E6576FFC7FFFB4680FF393662FEEF" box="[988,1302,234,261]" pageId="4" pageNumber="3" type="nomenclature">
<paragraph id="8BCB36FDFFC7FFFB4680FF393662FEEF" blockId="4.[820,1471,234,356]" box="[988,1302,234,261]" pageId="4" pageNumber="3">
<heading id="D0838191FFC7FFFB4680FF393662FEEF" box="[988,1302,234,261]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="3" reason="2">
<taxonomicName id="4C744D7EFFC7FFFB4680FF3937B9FEEF" ID-CoL="46C25" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[988,1229,234,261]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">Neobuthus haeckeli</taxonomicName>
<taxonomicNameLabel id="A2335794FFC7FFFB4188FF383662FEEF" box="[1236,1302,235,261]" pageId="4" pageNumber="3" rank="species">
<emphasis id="B900EAEFFFC7FFFB4188FF383784FEEF" bold="true" box="[1236,1264,235,261]" pageId="4" pageNumber="3">sp</emphasis>
.
<emphasis id="B900EAEFFFC7FFFB41A2FF38367AFEEF" bold="true" box="[1278,1294,235,261]" pageId="4" pageNumber="3">n</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFC7FFFB46B4FEDF3634FE8E" pageId="4" pageNumber="3" type="description">
<paragraph id="8BCB36FDFFC7FFFB46B4FEDF36CBFEAE" blockId="4.[820,1471,234,356]" pageId="4" pageNumber="3">
(
<figureCitation id="134F2A78FFC7FFFB46ACFEDF370BFECE" box="[1008,1151,268,292]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–4" captionStart-2="Figures 5–18" captionStart-3="Figures 19–26" captionStartId-0="3.[114,191,2015,2037]" captionStartId-1="5.[113,190,1516,1538]" captionStartId-2="6.[115,192,1928,1950]" captionStartId-3="7.[114,191,1981,2003]" captionTargetBox-0="[114,1475,234,2007]" captionTargetBox-1="[119,1452,260,1502]" captionTargetBox-2="[121,1470,243,1893]" captionTargetBox-3="[113,1474,240,1941]" captionTargetId-0="figure-28@3.[114,1475,234,2009]" captionTargetId-1="figure-9@5.[113,1474,234,1508]" captionTargetId-2="figure-103@6.[113,1474,238,1901]" captionTargetId-3="figure-41@7.[113,1474,240,1956]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Figures 1–2. Neobuthus haeckeli sp. n., paratype male in vivo habitus (1) and type locality (2)." captionText-1="Figures 3–4. Neobuthus haeckeli sp. n., holotype male, dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-2="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." captionText-3="Figures 19–26: Neobuthus haeckeli sp. n., holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26)." pageId="4" pageNumber="3">Figures 1–26</figureCitation>
,
<figureCitation id="134F2A78FFC7FFFB41D7FEDE37DCFECF" box="[1163,1192,269,293]" captionStart="Figure 70" captionStartId="15.[112,180,1521,1543]" captionTargetBox="[117,1473,239,1511]" captionTargetId="figure-276@15.[113,1476,234,1515]" captionTargetPageId="15" captionText="Figure 70: Map showing distribution of Neobuthus spp." pageId="4" pageNumber="3">70</figureCitation>
,
<tableCitation id="C6F60346FFC7FFFB41EEFEDF3674FECF" box="[1202,1280,268,293]" captionStart="Table 1" captionStartId="8.[113,169,953,974]" captionText="Table 1. Comparative measurements of adults of Neobuthus haeckeli sp. n. and N. solegladi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="4" pageNumber="3">Table 1</tableCitation>
)
<emphasis id="B900EAEFFFC7FFFB4668FEFF36CBFEAE" bold="true" box="[820,1471,300,324]" pageId="4" pageNumber="3">http://zoobank.org/urn:lsid:zoobank.org:act:CC1873AB-</emphasis>
</paragraph>
<paragraph id="8BCB36FDFFC7FFFB46EFFE9F3634FE8E" blockId="4.[820,1471,234,356]" box="[947,1344,332,356]" pageId="4" pageNumber="3">
<emphasis id="B900EAEFFFC7FFFB46EFFE9F3634FE8E" bold="true" box="[947,1344,332,356]" pageId="4" pageNumber="3">069D-49BD-8E2C-98CE9BA9672F</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFC7FFFB466CFE5E36FCFD8F" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="8BCB36FDFFC7FFFB466CFE5E3624FE2F" blockId="4.[816,1474,396,453]" pageId="4" pageNumber="3">
<materialsCitation id="3B1C3CA0FFC7FFFB466CFE5E3777FE4F" box="[816,1027,397,421]" collectionCode="LOCALITY, AND" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF885FFFC7FFFB466CFE5E3012FE4E" box="[816,870,397,420]" pageId="4" pageNumber="3">TYPE</typeStatus>
<collectionCode id="ED65AE38FFC7FFFB4637FE5E3777FE4F" box="[875,1027,397,421]" pageId="4" pageNumber="3">LOCALITY AND</collectionCode>
</materialsCitation>
<materialsCitation id="3B1C3CA0FFC7FFFB4154FE5E3788FE2E" collectionCode="DEPOSITORY" elevation="1056" latitude="10.047389" longLatPrecision="1" longitude="43.78575" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CF885FFFC7FFFB4154FE5E374EFE4E" box="[1032,1082,397,420]" pageId="4" pageNumber="3">TYPE</typeStatus>
<collectionCode id="ED65AE38FFC7FFFB4163FE5E37C8FE4F" box="[1087,1212,397,421]" pageId="4" pageNumber="3">DEPOSITORY</collectionCode>
. Somaliland, Cali Haidh,
<geoCoordinate id="EE40503AFFC7FFFB466CFE7E30B6FE2E" box="[816,962,428,453]" degrees="10" direction="north" minutes="02" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="50.6" value="10.047389">10°02'50.6&quot;N</geoCoordinate>
<geoCoordinate id="EE40503AFFC7FFFB4695FE7E372EFE2F" box="[969,1114,429,453]" degrees="43" direction="east" minutes="47" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="08.7" value="43.78575">43°47'08.7&quot;E</geoCoordinate>
,
<elevation id="0059D1CEFFC7FFFB4139FE7F3788FE2E" box="[1125,1276,428,452]" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" pageId="4" pageNumber="3" unit="m" value="1056.0">
<quantity id="4C8C9B18FFC7FFFB4139FE7F37C2FE2E" box="[1125,1206,428,452]" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" pageId="4" pageNumber="3" unit="m" value="1056.0">1056 m</quantity>
a. s. l.
</elevation>
</materialsCitation>
;
<collectionCode id="ED65AE38FFC7FFFB4055FE7E3638FE2F" box="[1289,1356,429,453]" pageId="4" pageNumber="3">FKCP</collectionCode>
.
</paragraph>
<paragraph id="8BCB36FDFFC7FFFB466CFE3E36FCFD8F" blockId="4.[816,1475,492,613]" pageId="4" pageNumber="3">
<materialsCitation id="3B1C3CA0FFC7FFFB466CFE3E37D4FD8E" collectingDate="2019-07-08" collectionCode="TYPE, MATERIAL, EXAMINED" collectorName="Locality No. &amp; F. Kovarik" elevation="1056" latitude="10.047389" longLatPrecision="1" longitude="43.78575" pageId="4" pageNumber="3" specimenCount="7" specimenCount-male="7" typeStatus="holotype">
<collectionCode id="ED65AE38FFC7FFFB466CFE3E3705FDEF" box="[816,1137,493,517]" pageId="4" pageNumber="3">TYPE MATERIAL EXAMINED</collectionCode>
.
<emphasis id="B900EAEFFFC7FFFB41D3FE3F3664FDEE" bold="true" box="[1167,1296,492,516]" pageId="4" pageNumber="3">Somaliland</emphasis>
, Cali Haidh,
<geoCoordinate id="EE40503AFFC7FFFB466CFDDE30B6FDCE" box="[816,962,524,549]" degrees="10" direction="north" minutes="02" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="50.6" value="10.047389">10°02'50.6&quot;N</geoCoordinate>
<geoCoordinate id="EE40503AFFC7FFFB4693FDDE3714FDCF" box="[975,1120,525,549]" degrees="43" direction="east" minutes="47" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="08.7" value="43.78575">43°47'08.7&quot;E</geoCoordinate>
,
<elevation id="0059D1CEFFC7FFFB412DFDDF366BFDCE" box="[1137,1311,524,548]" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" pageId="4" pageNumber="3" unit="m" value="1056.0">
<quantity id="4C8C9B18FFC7FFFB412DFDDF37BCFDCE" box="[1137,1224,524,548]" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" pageId="4" pageNumber="3" unit="m" value="1056.0">1056 m</quantity>
a. s. l.
</elevation>
(
<collectorName id="2681532BFFC7FFFB406FFDDF36B7FDCE" box="[1331,1475,524,548]" pageId="4" pageNumber="3">Locality No.</collectorName>
<emphasis id="B900EAEFFFC7FFFB466CFDFF3019FDAE" bold="true" box="[816,877,556,580]" pageId="4" pageNumber="3">19SN</emphasis>
),
<date id="FFCA103DFFC7FFFB46D8FDFE3088FDAF" box="[900,1020,557,581]" pageId="4" pageNumber="3" value="2019-07-08">
<collectingDate id="EF8EE9D5FFC7FFFB46D8FDFE3088FDAF" box="[900,1020,557,581]" pageId="4" pageNumber="3" value="2019-07-08">8.VII.2019</collectingDate>
</date>
,
<specimenCount id="9D72FD74FFC7FFFB4154FDFE375EFDAE" box="[1032,1066,557,580]" pageId="4" pageNumber="3" type="male">7♂</specimenCount>
(
<typeStatus id="54CF885FFFC7FFFB4164FDFF37E3FDAE" box="[1080,1175,556,580]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CF885FFFC7FFFB4192FDFE3643FDAF" box="[1230,1335,557,581]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
, 1697, 1712, 1713, 1715, 1716), leg.
<collectorName id="2681532BFFC7FFFB4173FD9E37D4FD8E" box="[1071,1184,588,612]" pageId="4" pageNumber="3">F. Kovařík</collectorName>
</materialsCitation>
&amp;
<materialsCitation id="3B1C3CA0FFC7FFFB419EFD9E3643FD8E" box="[1218,1335,588,612]" collectionCode="T" collectorName="Mazuch" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED65AE38FFC7FFFB419EFD9E37A0FD8E" box="[1218,1236,589,612]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="3">T</collectionCode>
.
<collectorName id="2681532BFFC7FFFB4182FD9F3643FD8E" box="[1246,1335,588,612]" pageId="4" pageNumber="3">Mazuch</collectorName>
</materialsCitation>
,
<collectionCode id="ED65AE38FFC7FFFB401DFD9E36F0FD8F" box="[1345,1412,589,613]" pageId="4" pageNumber="3">FKCP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFC7FFFB466CFD5E37E4FD0E" pageId="4" pageNumber="3" type="etymology">
<paragraph id="8BCB36FDFFC7FFFB466CFD5E37E4FD0E" blockId="4.[816,1474,652,741]" pageId="4" pageNumber="3">ETYMOLOGY. The name honors Czech entomologist, my friend and physician Martin Häckel who visited Somaliland with me and knows my body from inside.</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFC7FFFB466CFCDE30E5FA6E" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="8BCB36FDFFC7FFFB466CFCDE30E5FA6E" blockId="4.[816,1475,780,1412]" pageId="4" pageNumber="3">
DIAGNOSIS. Total length of adult males
<quantity id="4C8C9B18FFC7FFFB41B3FCDE361EFCCE" box="[1263,1386,781,805]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="4" pageNumber="3" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">17–20 mm</quantity>
; female unknown; carapace with area between anterior median carinae pale, densely granular; tergites with 3 dark stripes; pedipalp relatively slender, males with femur L/
<geoCoordinate id="EE40503AFFC7FFFB4181FCBE3644FC6E" box="[1245,1328,876,901]" degrees="2.22" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="555" value="-2.22">W 2.22</geoCoordinate>
–2.36, patella L/
<geoCoordinate id="EE40503AFFC7FFFB4614FC5E30EDFC4E" box="[840,921,908,933]" degrees="2.28" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="555" value="-2.28">W 2.28</geoCoordinate>
–2.36, chela L/
<geoCoordinate id="EE40503AFFC7FFFB4163FC5E37E5FC4E" box="[1087,1169,908,933]" degrees="4.44" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="555" value="-4.44">W 4.44</geoCoordinate>
–4.58; chela movable finger with 5 subrows of primary denticles, 4 external accessory denticles flanking proximal end of each subrow; trichobothria
<emphasis id="B900EAEFFFC7FFFB466CFC3F3049FBEE" box="[816,829,1004,1028]" italics="true" pageId="4" pageNumber="3">d</emphasis>
2 present on femur and patella; dorsoexternal and ventroexternal carinae on pedipalp patella present, smooth; posterior margins of tergites usually lacking macrosetae; pedipalps, legs, metasoma and telson with moderately short, stout macrosetae in males; males with coxae sparsely granulated, sternites III–VI shagreened to smooth medially and granulate laterally, sternite VII densely granulated with 4 well-defined, granulated carinae; metasoma I–II with median lateral carinae present in males; lateral surface of metasoma V in males densely granulated, with granules separated; soles of telotarsi with relatively sparse setation, leg III of adults with 10–12 ventral macrosetae on telotarsus; pectine teeth
<quantity id="4C8C9B18FFC7FFFB4023FA9F3031FA6E" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="4" pageNumber="3" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">16–19 in</quantity>
males.
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFC7FFF7466CFA7E3630FAEA" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3" type="description">
<paragraph id="8BCB36FDFFC7FFFB466CFA7E37ECF90E" blockId="4.[816,1475,1452,2020]" pageId="4" pageNumber="3">
DESCRIPTION. Total length of adult males
<quantity id="4C8C9B18FFC7FFFB41AAFA7E3619FA2E" box="[1270,1389,1453,1477]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="4" pageNumber="3" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">17–20 mm</quantity>
, female unknown; measurements of carapace, telson, segments of metasoma and pedipalps given in
<tableCitation id="C6F60346FFC7FFFB419CFA3F366CF9EF" box="[1216,1304,1516,1541]" captionStart="Table 1" captionStartId="8.[113,169,953,974]" captionText="Table 1. Comparative measurements of adults of Neobuthus haeckeli sp. n. and N. solegladi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="4" pageNumber="3">Table 1</tableCitation>
; positions and distribution of trichobothria on pedipalps shown in
<figureCitation id="134F2A78FFC7FFFB4004F9DF36C9F9CE" box="[1368,1469,1548,1572]" captionStart="Figures 5–18" captionStartId="6.[115,192,1928,1950]" captionTargetBox="[121,1470,243,1893]" captionTargetId="figure-103@6.[113,1474,238,1901]" captionTargetPageId="6" captionText="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." pageId="4" pageNumber="3">Figs. 6–9</figureCitation>
, and 12–13; trichobothrium
<emphasis id="B900EAEFFFC7FFFB4106F9FF3713F9AE" box="[1114,1127,1580,1604]" italics="true" pageId="4" pageNumber="3">d</emphasis>
<subScript id="17F034B8FFC7FFFB413BF9EE371BF9A1" attach="left" box="[1127,1135,1597,1611]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
present on pedipalp femur and patella; base color pale yellow to light orange with variable fuscous pigmentation (
<figureCitation id="134F2A78FFC7FFFB4172F9BF3708F96F" box="[1070,1148,1644,1669]" captionStart="Figures 1–2" captionStartId="3.[114,191,2015,2037]" captionTargetBox="[114,1475,234,2007]" captionTargetId="figure-28@3.[114,1475,234,2009]" captionTargetPageId="3" captionText="Figures 1–2. Neobuthus haeckeli sp. n., paratype male in vivo habitus (1) and type locality (2)." pageId="4" pageNumber="3">Figs. 1</figureCitation>
,
<figureCitation id="134F2A78FFC7FFFB41D3F9BE37CCF96F" box="[1167,1208,1645,1669]" captionStart="Figures 3–4" captionStartId="5.[113,190,1516,1538]" captionTargetBox="[119,1452,260,1502]" captionTargetId="figure-9@5.[113,1474,234,1508]" captionTargetPageId="5" captionText="Figures 3–4. Neobuthus haeckeli sp. n., holotype male, dorsal (3) and ventral (4) views. Scale bar: 10 mm." pageId="4" pageNumber="3">3–4</figureCitation>
) and extensive patterns of dark maculation on pedipalps, mesosoma, metasoma and partially on legs; chelicerae yellow with dark reticulation on anterior manus, dentition reddish.
</paragraph>
<paragraph id="8BCB36FDFFC7FFFB466CF93F360CF80E" blockId="4.[816,1475,1452,2020]" pageId="4" pageNumber="3">
<emphasis id="B900EAEFFFC7FFFB466CF93F30E1F8EE" bold="true" box="[816,917,1772,1796]" pageId="4" pageNumber="3">Pedipalp</emphasis>
(
<figureCitation id="134F2A78FFC7FFFB46F1F93F375CF8EE" box="[941,1064,1772,1796]" captionStart="Figures 5–18" captionStartId="6.[115,192,1928,1950]" captionTargetBox="[121,1470,243,1893]" captionTargetId="figure-103@6.[113,1474,238,1901]" captionTargetPageId="6" captionText="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." pageId="4" pageNumber="3">Figs. 5–15</figureCitation>
). Pedipalp mostly sparsely hirsute; finely granulated in males; femur with five conspicuously granulose carinae, more strongly developed in males; patella with seven granulose carinae in males, and five smooth carinae in females; chela with dorsal and external carinae present. The chela movable and fixed fingers with 5 subrows of primary denticles, 4 external accessory denticles on movable and 4–5 on fixed fingers flanking proximal end of each subrow.
</paragraph>
<caption id="DF0B6675FFC6FFFA452DFA3F3703F9EB" box="[113,1143,1516,1538]" pageId="5" pageNumber="4" startId="5.[113,190,1516,1538]" targetBox="[119,1452,260,1502]" targetPageId="5">
<paragraph id="8BCB36FDFFC6FFFA452DFA3F3703F9EB" blockId="5.[113,1143,1516,1538]" box="[113,1143,1516,1538]" pageId="5" pageNumber="4">
<emphasis id="B900EAEFFFC6FFFA452DFA3F339CF9EB" bold="true" box="[113,232,1516,1538]" pageId="5" pageNumber="4">Figures 3–4</emphasis>
.
<taxonomicName id="4C744D7EFFC6FFFA45A8FA3F32C5F9EB" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[244,433,1516,1537]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFC6FFFA45A8FA3F32C5F9EB" box="[244,433,1516,1537]" italics="true" pageId="5" pageNumber="4">Neobuthus haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFC6FFFA44EBFA3F3298F9EB" box="[439,492,1516,1537]" pageId="5" pageNumber="4" rank="species">
<emphasis id="B900EAEFFFC6FFFA44EBFA3F32BAF9EB" bold="true" box="[439,462,1516,1537]" pageId="5" pageNumber="4">sp</emphasis>
.
<emphasis id="B900EAEFFFC6FFFA4486FA3F3293F9EB" bold="true" box="[474,487,1516,1537]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
, holotype male, dorsal (3) and ventral (4) views. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="8BCB36FDFFC6FFFA452DF9CA36CCF9BB" blockId="5.[112,1474,1561,2033]" pageId="5" pageNumber="4">
<emphasis id="B900EAEFFFC6FFFA452DF9CA33ABF9DB" bold="true" box="[113,223,1561,1585]" pageId="5" pageNumber="4">Carapace</emphasis>
(
<figureCitation id="134F2A78FFC6FFFA445EF9CA3212F9DB" box="[258,358,1561,1585]" captionStart="Figures 19–26" captionStartId="7.[114,191,1981,2003]" captionTargetBox="[113,1474,240,1941]" captionTargetId="figure-41@7.[113,1474,240,1956]" captionTargetPageId="7" captionText="Figures 19–26: Neobuthus haeckeli sp. n., holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26)." pageId="5" pageNumber="4">Fig. 19</figureCitation>
). Strongly trapezoidal (narrower distal denticle, medium-sized subdistal and medial, and 2 small, anteriorly), approximately wide as long (L/
<geoCoordinate id="EE40503AFFC6FFFA473EF9EA31CCF9BB" box="[610,696,1593,1617]" degrees="0.96" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="555" value="-0.96">W 0.96</geoCoordinate>
–0.99); partially fused basal denticles; ventral margin with 2 denticles.
</paragraph>
<paragraph id="8BCB36FDFFC6FFF7452DF98A31C7FBAA" blockId="5.[112,1474,1561,2033]" lastBlockId="8.[113,772,1032,1824]" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4">
posterior median postocular area flat, anterior median
<emphasis id="B900EAEFFFC6FFFA466CF98A30D2F99B" bold="true" box="[816,934,1625,1649]" pageId="5" pageNumber="4">Mesosoma</emphasis>
(
<figureCitation id="134F2A78FFC6FFFA46E7F98A3737F99B" box="[955,1091,1625,1649]" captionStart="Figures 19–26" captionStartId="7.[114,191,1981,2003]" captionTargetBox="[113,1474,240,1941]" captionTargetId="figure-41@7.[113,1474,240,1956]" captionTargetPageId="7" captionText="Figures 19–26: Neobuthus haeckeli sp. n., holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26)." pageId="5" pageNumber="4">Figs. 19–20</figureCitation>
). Tergites I–VI bear three carinae preocular area gently sloped downwards towards anterior of which the lateral pair may be less conspicuous mainly margin; lateral flanks steeply sloped; ocular tubercle broad, on tergites I–IV; tergite VII bears five well-defined carinae prominent, located slightly anterior to middle of carapace; (median, submedians and laterals); tergites I–VI densely anterior margin straight, finely microdenticulate, with coarser granular, with coarser granules on posterior lateral areas; tergite granules overlapping edge, bearing 6–8 macrosetae; anterior VII densely granular; sternites III–VI smooth medially and median carinae weak, coarsely granular, other carinae granulate laterally in males; sternite VII with four well-defined indistinct; dense granulation covering most of carapace. carinae, densely, finely granulated in both sexes; sternum
<emphasis id="B900EAEFFFC6FFFA452DF88A33A8F89B" bold="true" box="[113,220,1881,1905]" pageId="5" pageNumber="4">Chelicera</emphasis>
. Fingers with typical buthid dentition (
<bibRefCitation id="EFE54B0CFFC6FFFA4737F88A318AF89B" author="VACHON" box="[619,766,1881,1905]" pageId="5" pageNumber="4" pagination="161 - 166" refId="ref6172" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
;
<typeStatus id="54CF885FFFC6FFFA4673F88A3028F89B" box="[815,860,1881,1905]" pageId="5" pageNumber="4">type</typeStatus>
1, triangular in shape, smooth, with deep posteromedian Lowe &amp; Kovařík, 2016;
<bibRefCitation id="EFE54B0CFFC6FFFA442DF8AA3137F87B" author="KOVARIK" box="[369,579,1913,1937]" pageId="5" pageNumber="4" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
); fixed finger with invagination; genital opercula smooth; genital papillae present; large distal denticle, 1 subdistal denticle and 2 basal denticles pectines extending to around a quarter of sternite V in male; fused into bicusp, single denticle on ventral surface at level of pectine teeth 16–19 (2 x 16, 7 x 17, 3 x 18, 2 x 19) in males; bicusp; dorsal margin of movable finger with 5 denticles: 1 large combs with 3 marginal lamellae and 6–8 middle lamellae; marginal lamellae, middle lamellae and fulcra with dense cover of short dark reddish macrosetae; fulcra with 2–4 setae.
</paragraph>
<caption id="DF0B6675FFC5FFF9452FF85B3728F81E" pageId="6" pageNumber="5" startId="6.[115,192,1928,1950]" targetBox="[121,1470,243,1893]" targetPageId="6">
<paragraph id="8BCB36FDFFC5FFF9452FF85B3728F81E" blockId="6.[115,1473,1928,2036]" pageId="6" pageNumber="5">
<emphasis id="B900EAEFFFC5FFF9452FF85B3381F877" bold="true" box="[115,245,1928,1950]" pageId="6" pageNumber="5">Figures 5–18</emphasis>
:
<taxonomicName id="4C744D7EFFC5FFF9445DF85B32C8F877" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[257,444,1928,1949]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFC5FFF9445DF85B32C8F877" box="[257,444,1928,1949]" italics="true" pageId="6" pageNumber="5">Neobuthus haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFC5FFF9449EF85B3282F877" box="[450,502,1928,1949]" pageId="6" pageNumber="5" rank="species">
<emphasis id="B900EAEFFFC5FFF9449EF85B32ADF877" bold="true" box="[450,473,1928,1949]" pageId="6" pageNumber="5">sp</emphasis>
.
<emphasis id="B900EAEFFFC5FFF944BFF85B3284F877" bold="true" box="[483,496,1928,1949]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
, holotype male.
<emphasis id="B900EAEFFFC5FFF947C9F85B3063F877" bold="true" box="[661,791,1928,1950]" pageId="6" pageNumber="5">Figures 5–15</emphasis>
. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles).
<emphasis id="B900EAEFFFC5FFF9402FF81233DBF819" bold="true" pageId="6" pageNumber="5">Figures 16–18</emphasis>
. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18).
</paragraph>
</caption>
<caption id="DF0B6675FFC4FFF8452EF86E311DF81A" pageId="7" pageNumber="6" startId="7.[114,191,1981,2003]" targetBox="[113,1474,240,1941]" targetPageId="7">
<paragraph id="8BCB36FDFFC4FFF8452EF86E311DF81A" blockId="7.[114,1472,1981,2033]" pageId="7" pageNumber="6">
<emphasis id="B900EAEFFFC4FFF8452EF86E3276F839" bold="true" box="[114,258,1981,2003]" pageId="7" pageNumber="6">Figures 19–26</emphasis>
:
<taxonomicName id="4C744D7EFFC4FFF8444CF86D32BAF839" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[272,462,1982,2003]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFC4FFF8444CF86D32BAF839" box="[272,462,1982,2003]" italics="true" pageId="7" pageNumber="6">Neobuthus haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFC4FFF8448AF86D3178F839" box="[470,524,1982,2003]" pageId="7" pageNumber="6" rank="species">
<emphasis id="B900EAEFFFC4FFF8448AF86D3299F839" bold="true" box="[470,493,1982,2003]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="B900EAEFFFC4FFF844A6F86D3173F839" bold="true" box="[506,519,1982,2003]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
, holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26).
</paragraph>
</caption>
<paragraph id="8BCB36FDFFCBFFF74743FF273643FC4C" pageId="8" pageNumber="7">
<table id="F974C45DFFCB00004526FF2036F3FC43" box="[122,1415,243,937]" gridcols="5" gridrows="19" pageId="8" pageNumber="7">
<tr id="354434BFFFCB00004526FF2036F3FEE6" box="[122,1415,243,268]" gridrow="0" pageId="8" pageNumber="7" rowspan-0="1" rowspan-1="1">
<th id="76955DC3FFCB0000474CFF20319EFEE6" box="[528,746,243,268]" gridcol="2" gridrow="0" pageId="8" pageNumber="7">
<taxonomicName id="4C744D7EFFCBFFF74743FF2731EDFEE1" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[543,665,243,267]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF74743FF2731EDFEE1" bold="true" box="[543,665,243,267]" italics="true" pageId="8" pageNumber="7">N. haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF747F8FF2731AAFEE1" box="[676,734,244,268]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF747F8FF2731C8FEE6" bold="true" box="[676,700,244,268]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF74795FF2731A3FEE6" bold="true" box="[713,727,244,268]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="76955DC3FFCB00004603FF20374DFEE6" box="[863,1081,243,268]" gridcol="3" gridrow="0" pageId="8" pageNumber="7">
<taxonomicName id="4C744D7EFFCBFFF74636FF27309EFEE1" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[874,1002,243,267]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF74636FF27309EFEE1" bold="true" box="[874,1002,243,267]" italics="true" pageId="8" pageNumber="7">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF746A9FF27375BFEE1" box="[1013,1071,244,268]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF746A9FF273779FEE6" bold="true" box="[1013,1037,244,268]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF74146FF27375CFEE6" bold="true" box="[1050,1064,244,268]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="76955DC3FFCB000041F1FF2036F3FEE6" box="[1197,1415,243,268]" gridcol="4" gridrow="0" pageId="8" pageNumber="7">
<taxonomicName id="4C744D7EFFCBFFF741E4FF27364CFEE1" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[1208,1336,243,267]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF741E4FF27364CFEE1" bold="true" box="[1208,1336,243,267]" italics="true" pageId="8" pageNumber="7">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF7401FFF273609FEE1" box="[1347,1405,244,268]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF7401FFF27362FFEE6" bold="true" box="[1347,1371,244,268]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF74035FF273603FEE6" bold="true" box="[1385,1399,244,268]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
</th>
</tr>
<tr id="354434BFFFCB00004526FEC936F3FEDF" box="[122,1415,282,309]" gridrow="1" pageId="8" pageNumber="7" rowspan-1="1">
<th id="76955DC3FFCB00004526FEC93237FEDF" box="[122,323,282,309]" gridcol="0" gridrow="1" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF74526FEC93237FED8" bold="true" box="[122,323,282,306]" pageId="8" pageNumber="7">Dimensions (MM)</emphasis>
</th>
<td id="76955DC3FFCB0000474CFEC9319EFEDF" box="[528,746,282,309]" gridcol="2" gridrow="1" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF7471EFECF31C8FED9" bold="true" box="[578,700,283,309]" pageId="8" pageNumber="7">♂ holotype</emphasis>
</td>
<td id="76955DC3FFCB00004603FEC9374DFEDF" box="[863,1081,282,309]" gridcol="3" gridrow="1" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF746CCFECF377EFED9" bold="true" box="[912,1034,283,309]" pageId="8" pageNumber="7">♂ holotype</emphasis>
</td>
<td id="76955DC3FFCB000041F1FEC936F3FEDF" box="[1197,1415,282,309]" gridcol="4" gridrow="1" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF74180FECF362EFED9" bold="true" box="[1244,1370,283,309]" pageId="8" pageNumber="7">♀ paratype</emphasis>
</td>
</tr>
<tr id="354434BFFFCB00004526FEEE36F3FEB2" box="[122,1415,317,344]" gridrow="2" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FEEE3237FEB2" box="[122,323,317,344]" gridcol="0" gridrow="2" pageId="8" pageNumber="7">Carapace</th>
<td id="76955DC3FFCB00004438FEEE32BCFEB2" box="[356,456,317,344]" gridcol="1" gridrow="2" pageId="8" pageNumber="7">L / W</td>
<td id="76955DC3FFCB0000474CFEEE319EFEB2" box="[528,746,317,344]" gridcol="2" gridrow="2" pageId="8" pageNumber="7">2.569 / 2.570</td>
<td id="76955DC3FFCB00004603FEEE374DFEB2" box="[863,1081,317,344]" gridcol="3" gridrow="2" pageId="8" pageNumber="7">2.404 / 2.699</td>
<td id="76955DC3FFCB000041F1FEEE36F3FEB2" box="[1197,1415,317,344]" gridcol="4" gridrow="2" pageId="8" pageNumber="7">3.348 / 3.914</td>
</tr>
<tr id="354434BFFFCB00004526FEB036F3FE96" box="[122,1415,355,380]" gridrow="3" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FEB03237FE96" box="[122,323,355,380]" gridcol="0" gridrow="3" pageId="8" pageNumber="7">Mesosoma</th>
<td id="76955DC3FFCB00004438FEB032BCFE96" box="[356,456,355,380]" gridcol="1" gridrow="3" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFEB0319EFE96" box="[528,746,355,380]" gridcol="2" gridrow="3" pageId="8" pageNumber="7">4.308</td>
<td id="76955DC3FFCB00004603FEB0374DFE96" box="[863,1081,355,380]" gridcol="3" gridrow="3" pageId="8" pageNumber="7">3.598</td>
<td id="76955DC3FFCB000041F1FEB036F3FE96" box="[1197,1415,355,380]" gridcol="4" gridrow="3" pageId="8" pageNumber="7">5.953</td>
</tr>
<tr id="354434BFFFCB00004526FE5B36F3FE48" box="[122,1415,392,418]" gridrow="4" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FE5B3237FE48" box="[122,323,392,418]" gridcol="0" gridrow="4" pageId="8" pageNumber="7">Tergite VII</th>
<td id="76955DC3FFCB00004438FE5B32BCFE48" box="[356,456,392,418]" gridcol="1" gridrow="4" pageId="8" pageNumber="7">L / W</td>
<td id="76955DC3FFCB0000474CFE5B319EFE48" box="[528,746,392,418]" gridcol="2" gridrow="4" pageId="8" pageNumber="7">1.211 / 2.524</td>
<td id="76955DC3FFCB00004603FE5B374DFE48" box="[863,1081,392,418]" gridcol="3" gridrow="4" pageId="8" pageNumber="7">1.077 / 2.524</td>
<td id="76955DC3FFCB000041F1FE5B36F3FE48" box="[1197,1415,392,418]" gridcol="4" gridrow="4" pageId="8" pageNumber="7">1.504 / 3.811</td>
</tr>
<tr id="354434BFFFCB00004526FE7E36F3FE2D" box="[122,1415,429,455]" gridrow="5" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FE7E3237FE2D" box="[122,323,429,455]" gridcol="0" gridrow="5" pageId="8" pageNumber="7">Metasoma + telson</th>
<td id="76955DC3FFCB00004438FE7E32BCFE2D" box="[356,456,429,455]" gridcol="1" gridrow="5" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFE7E319EFE2D" box="[528,746,429,455]" gridcol="2" gridrow="5" pageId="8" pageNumber="7">12.731</td>
<td id="76955DC3FFCB00004603FE7E374DFE2D" box="[863,1081,429,455]" gridcol="3" gridrow="5" pageId="8" pageNumber="7">11.724</td>
<td id="76955DC3FFCB000041F1FE7E36F3FE2D" box="[1197,1415,429,455]" gridcol="4" gridrow="5" pageId="8" pageNumber="7">16.390</td>
</tr>
<tr id="354434BFFFCB00004526FE0136F3FE06" box="[122,1415,466,492]" gridrow="6" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FE013237FE06" box="[122,323,466,492]" gridcol="0" gridrow="6" pageId="8" pageNumber="7">Segment I</th>
<td id="76955DC3FFCB00004438FE0132BCFE06" box="[356,456,466,492]" gridcol="1" gridrow="6" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFE01319EFE06" box="[528,746,466,492]" gridcol="2" gridrow="6" pageId="8" pageNumber="7">1.612 / 1.681 / 1.428</td>
<td id="76955DC3FFCB00004603FE01374DFE06" box="[863,1081,466,492]" gridcol="3" gridrow="6" pageId="8" pageNumber="7">1.478 / 1.627 / 1.334</td>
<td id="76955DC3FFCB000041F1FE0136F3FE06" box="[1197,1415,466,492]" gridcol="4" gridrow="6" pageId="8" pageNumber="7">2.119 / 2.220 / 1.982</td>
</tr>
<tr id="354434BFFFCB00004526FE2436F3FDFB" box="[122,1415,503,529]" gridrow="7" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FE243237FDFB" box="[122,323,503,529]" gridcol="0" gridrow="7" pageId="8" pageNumber="7">Segment II</th>
<td id="76955DC3FFCB00004438FE2432BCFDFB" box="[356,456,503,529]" gridcol="1" gridrow="7" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFE24319EFDFB" box="[528,746,503,529]" gridcol="2" gridrow="7" pageId="8" pageNumber="7">1.875 / 1.520 / 1.482</td>
<td id="76955DC3FFCB00004603FE24374DFDFB" box="[863,1081,503,529]" gridcol="3" gridrow="7" pageId="8" pageNumber="7">1.574 / 1.435 / 1.386</td>
<td id="76955DC3FFCB000041F1FE2436F3FDFB" box="[1197,1415,503,529]" gridcol="4" gridrow="7" pageId="8" pageNumber="7">2.342 / 1.978 / 1.981</td>
</tr>
<tr id="354434BFFFCB00004526FDCF36F3FDDC" box="[122,1415,540,566]" gridrow="8" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FDCF3237FDDC" box="[122,323,540,566]" gridcol="0" gridrow="8" pageId="8" pageNumber="7">Segment III</th>
<td id="76955DC3FFCB00004438FDCF32BCFDDC" box="[356,456,540,566]" gridcol="1" gridrow="8" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFDCF319EFDDC" box="[528,746,540,566]" gridcol="2" gridrow="8" pageId="8" pageNumber="7">1.888 / 1.484 / 1.435</td>
<td id="76955DC3FFCB00004603FDCF374DFDDC" box="[863,1081,540,566]" gridcol="3" gridrow="8" pageId="8" pageNumber="7">1.713 / 1.335 / 1.372</td>
<td id="76955DC3FFCB000041F1FDCF36F3FDDC" box="[1197,1415,540,566]" gridcol="4" gridrow="8" pageId="8" pageNumber="7">2.580 / 1.960 / 1.900</td>
</tr>
<tr id="354434BFFFCB00004526FD9236F3FDB1" box="[122,1415,577,603]" gridrow="9" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FD923237FDB1" box="[122,323,577,603]" gridcol="0" gridrow="9" pageId="8" pageNumber="7">Segment IV</th>
<td id="76955DC3FFCB00004438FD9232BCFDB1" box="[356,456,577,603]" gridcol="1" gridrow="9" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFD92319EFDB1" box="[528,746,577,603]" gridcol="2" gridrow="9" pageId="8" pageNumber="7">2.222 / 1.457 / 1.295</td>
<td id="76955DC3FFCB00004603FD92374DFDB1" box="[863,1081,577,603]" gridcol="3" gridrow="9" pageId="8" pageNumber="7">2.081 / 1.319 / 1.258</td>
<td id="76955DC3FFCB000041F1FD9236F3FDB1" box="[1197,1415,577,603]" gridcol="4" gridrow="9" pageId="8" pageNumber="7">2.887 / 1.938 / 1.703</td>
</tr>
<tr id="354434BFFFCB00004526FDB536F3FD6A" box="[122,1415,614,640]" gridrow="10" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FDB53237FD6A" box="[122,323,614,640]" gridcol="0" gridrow="10" pageId="8" pageNumber="7">Segment V</th>
<td id="76955DC3FFCB00004438FDB532BCFD6A" box="[356,456,614,640]" gridcol="1" gridrow="10" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFDB5319EFD6A" box="[528,746,614,640]" gridcol="2" gridrow="10" pageId="8" pageNumber="7">2.737 / 1.330 / 1.145</td>
<td id="76955DC3FFCB00004603FDB5374DFD6A" box="[863,1081,614,640]" gridcol="3" gridrow="10" pageId="8" pageNumber="7">2.539 / 1.255 / 1.090</td>
<td id="76955DC3FFCB000041F1FDB536F3FD6A" box="[1197,1415,614,640]" gridcol="4" gridrow="10" pageId="8" pageNumber="7">3.437 / 1.849 / 1.478</td>
</tr>
<tr id="354434BFFFCB00004526FD5836F3FD4C" box="[122,1415,651,678]" gridrow="11" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FD583237FD4C" box="[122,323,651,678]" gridcol="0" gridrow="11" pageId="8" pageNumber="7">Telson</th>
<td id="76955DC3FFCB00004438FD5832BCFD4C" box="[356,456,651,678]" gridcol="1" gridrow="11" pageId="8" pageNumber="7">L / W / D</td>
<td id="76955DC3FFCB0000474CFD58319EFD4C" box="[528,746,651,678]" gridcol="2" gridrow="11" pageId="8" pageNumber="7">2.397 / 0.937 / 0.814</td>
<td id="76955DC3FFCB00004603FD58374DFD4C" box="[863,1081,651,678]" gridcol="3" gridrow="11" pageId="8" pageNumber="7">2.339 / 0.877 / 0.751</td>
<td id="76955DC3FFCB000041F1FD5836F3FD4C" box="[1197,1415,651,678]" gridcol="4" gridrow="11" pageId="8" pageNumber="7">3.025 / 1.349 / 1.151</td>
</tr>
<tr id="354434BFFFCB00004526FD6336F3FD20" box="[122,1415,688,714]" gridrow="12" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FD633237FD20" box="[122,323,688,714]" gridcol="0" gridrow="12" pageId="8" pageNumber="7">Pedipalp</th>
<td id="76955DC3FFCB00004438FD6332BCFD20" box="[356,456,688,714]" gridcol="1" gridrow="12" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFD63319EFD20" box="[528,746,688,714]" gridcol="2" gridrow="12" pageId="8" pageNumber="7">6.410</td>
<td id="76955DC3FFCB00004603FD63374DFD20" box="[863,1081,688,714]" gridcol="3" gridrow="12" pageId="8" pageNumber="7">5.755</td>
<td id="76955DC3FFCB000041F1FD6336F3FD20" box="[1197,1415,688,714]" gridcol="4" gridrow="12" pageId="8" pageNumber="7">8.443</td>
</tr>
<tr id="354434BFFFCB00004526FD0536F3FD1A" box="[122,1415,726,752]" gridrow="13" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FD053237FD1A" box="[122,323,726,752]" gridcol="0" gridrow="13" pageId="8" pageNumber="7">Femur</th>
<td id="76955DC3FFCB00004438FD0532BCFD1A" box="[356,456,726,752]" gridcol="1" gridrow="13" pageId="8" pageNumber="7">L / W</td>
<td id="76955DC3FFCB0000474CFD05319EFD1A" box="[528,746,726,752]" gridcol="2" gridrow="13" pageId="8" pageNumber="7">1.578 / 0.694</td>
<td id="76955DC3FFCB00004603FD05374DFD1A" box="[863,1081,726,752]" gridcol="3" gridrow="13" pageId="8" pageNumber="7">1.428 / 0.605</td>
<td id="76955DC3FFCB000041F1FD0536F3FD1A" box="[1197,1415,726,752]" gridcol="4" gridrow="13" pageId="8" pageNumber="7">2.051 / 0.896</td>
</tr>
<tr id="354434BFFFCB00004526FD2836F3FCFF" box="[122,1415,763,789]" gridrow="14" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FD283237FCFF" box="[122,323,763,789]" gridcol="0" gridrow="14" pageId="8" pageNumber="7">Patella</th>
<td id="76955DC3FFCB00004438FD2832BCFCFF" box="[356,456,763,789]" gridcol="1" gridrow="14" pageId="8" pageNumber="7">L / W</td>
<td id="76955DC3FFCB0000474CFD28319EFCFF" box="[528,746,763,789]" gridcol="2" gridrow="14" pageId="8" pageNumber="7">2.159 / 0.949</td>
<td id="76955DC3FFCB00004603FD28374DFCFF" box="[863,1081,763,789]" gridcol="3" gridrow="14" pageId="8" pageNumber="7">1.960 / 0.858</td>
<td id="76955DC3FFCB000041F1FD2836F3FCFF" box="[1197,1415,763,789]" gridcol="4" gridrow="14" pageId="8" pageNumber="7">2.821 / 1.269</td>
</tr>
<tr id="354434BFFFCB00004526FCF336F3FCD0" box="[122,1415,800,826]" gridrow="15" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FCF33237FCD0" box="[122,323,800,826]" gridcol="0" gridrow="15" pageId="8" pageNumber="7">Chela</th>
<td id="76955DC3FFCB00004438FCF332BCFCD0" box="[356,456,800,826]" gridcol="1" gridrow="15" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFCF3319EFCD0" box="[528,746,800,826]" gridcol="2" gridrow="15" pageId="8" pageNumber="7">2.673</td>
<td id="76955DC3FFCB00004603FCF3374DFCD0" box="[863,1081,800,826]" gridcol="3" gridrow="15" pageId="8" pageNumber="7">2.367</td>
<td id="76955DC3FFCB000041F1FCF336F3FCD0" box="[1197,1415,800,826]" gridcol="4" gridrow="15" pageId="8" pageNumber="7">3.571</td>
</tr>
<tr id="354434BFFFCB00004526FC9636F3FCB5" box="[122,1415,837,863]" gridrow="16" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FC963237FCB5" box="[122,323,837,863]" gridcol="0" gridrow="16" pageId="8" pageNumber="7">Manus</th>
<td id="76955DC3FFCB00004438FC9632BCFCB5" box="[356,456,837,863]" gridcol="1" gridrow="16" pageId="8" pageNumber="7">W / D</td>
<td id="76955DC3FFCB0000474CFC96319EFCB5" box="[528,746,837,863]" gridcol="2" gridrow="16" pageId="8" pageNumber="7">0.585 / 0.592</td>
<td id="76955DC3FFCB00004603FC96374DFCB5" box="[863,1081,837,863]" gridcol="3" gridrow="16" pageId="8" pageNumber="7">0.554 / 0.534</td>
<td id="76955DC3FFCB000041F1FC9636F3FCB5" box="[1197,1415,837,863]" gridcol="4" gridrow="16" pageId="8" pageNumber="7">0.888 / 0.791</td>
</tr>
<tr id="354434BFFFCB00004526FCB936F3FC6E" box="[122,1415,874,900]" gridrow="17" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FCB93237FC6E" box="[122,323,874,900]" gridcol="0" gridrow="17" pageId="8" pageNumber="7">Movable finger</th>
<td id="76955DC3FFCB00004438FCB932BCFC6E" box="[356,456,874,900]" gridcol="1" gridrow="17" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFCB9319EFC6E" box="[528,746,874,900]" gridcol="2" gridrow="17" pageId="8" pageNumber="7">1.751</td>
<td id="76955DC3FFCB00004603FCB9374DFC6E" box="[863,1081,874,900]" gridcol="3" gridrow="17" pageId="8" pageNumber="7">1.581</td>
<td id="76955DC3FFCB000041F1FCB936F3FC6E" box="[1197,1415,874,900]" gridcol="4" gridrow="17" pageId="8" pageNumber="7">2.349</td>
</tr>
<tr id="354434BFFFCB00004526FC5D36F3FC43" box="[122,1415,910,937]" gridrow="18" pageId="8" pageNumber="7">
<th id="76955DC3FFCB00004526FC5D3237FC43" box="[122,323,910,937]" gridcol="0" gridrow="18" pageId="8" pageNumber="7">Total</th>
<td id="76955DC3FFCB00004438FC5D32BCFC43" box="[356,456,910,937]" gridcol="1" gridrow="18" pageId="8" pageNumber="7">L</td>
<td id="76955DC3FFCB0000474CFC5D319EFC43" box="[528,746,910,937]" gridcol="2" gridrow="18" pageId="8" pageNumber="7">19.61</td>
<td id="76955DC3FFCB00004603FC5D374DFC43" box="[863,1081,910,937]" gridcol="3" gridrow="18" pageId="8" pageNumber="7">17.73</td>
<td id="76955DC3FFCB000041F1FC5D36F3FC43" box="[1197,1415,910,937]" gridcol="4" gridrow="18" pageId="8" pageNumber="7">25.69</td>
</tr>
</table>
</paragraph>
<caption id="DF0B6675FFCBFFF7452DFC6A310FFC06" pageId="8" pageNumber="7" startId="8.[113,169,953,974]">
<paragraph id="8BCB36FDFFCBFFF7452DFC6A310FFC06" blockId="8.[113,1471,953,1004]" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF7452DFC6A33C9FC24" bold="true" box="[113,189,953,974]" pageId="8" pageNumber="7">Table 1</emphasis>
. Comparative measurements of adults of
<taxonomicName id="4C744D7EFFCBFFF7470AFC6A3060FC24" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[598,788,953,974]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF7470AFC6A3060FC24" box="[598,788,953,974]" italics="true" pageId="8" pageNumber="7">Neobuthus haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF74640FC6A3027FC24" box="[796,851,953,974]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF74640FC6A3047FC24" bold="true" box="[796,819,953,974]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF7461CFC6A3039FC24" bold="true" box="[832,845,953,974]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C744D7EFFCBFFF746D9FC6A308EFC24" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[901,1018,953,974]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="solegladi" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF746D9FC6A308EFC24" box="[901,1018,953,974]" italics="true" pageId="8" pageNumber="7">N. solegladi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF7415DFC6A374DFC24" box="[1025,1081,953,974]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF7415DFC6A376CFC24" bold="true" box="[1025,1048,953,974]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF74179FC6A3746FC24" bold="true" box="[1061,1074,953,974]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<paragraph id="8BCB36FDFFCBFFF7452DFB9B31FCFA8A" blockId="8.[113,772,1032,1824]" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF7452DFB9B33D2FB8A" bold="true" box="[113,166,1096,1120]" pageId="8" pageNumber="7">Legs</emphasis>
(
<figureCitation id="134F2A78FFCBFFF745E7FB9B3230FB8A" box="[187,324,1096,1120]" captionStart="Figures 19–26" captionStartId="7.[114,191,1981,2003]" captionTargetBox="[113,1474,240,1941]" captionTargetId="figure-41@7.[113,1474,240,1956]" captionTargetPageId="7" captionText="Figures 19–26: Neobuthus haeckeli sp. n., holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26)." pageId="8" pageNumber="7">Figs. 22–26</figureCitation>
). Coxa, femora, patella and tibia of all legs bearing variable numbers of medium length, straight, dark-reddish macrosetae; tarsi with mix of short and longer, dark-reddish macrosetae; basitarsi I–III slightly compressed with flat retrolateral surfaces, with bristle combs consisting of retrosuperior series of longer macrosetae, plus retroinferior and proinferior series of shorter macrosetae; telotarsi with two rows of short setae on ventral aspect, 10–12 macrosetae on telotarsus III; tibial spurs moderate on legs III–IV.
</paragraph>
<paragraph id="8BCB36FDFFCBFFF7452DFABB318FF8CA" blockId="8.[113,772,1032,1824]" pageId="8" pageNumber="7">
<emphasis id="B900EAEFFFCBFFF7452DFABB3207FA6A" bold="true" box="[113,371,1384,1408]" pageId="8" pageNumber="7">Metasoma and telson</emphasis>
(
<figureCitation id="134F2A78FFCBFFF744D6FABB3167FA6A" box="[394,531,1384,1408]" captionStart="Figures 5–18" captionStartId="6.[115,192,1928,1950]" captionTargetBox="[121,1470,243,1893]" captionTargetId="figure-103@6.[113,1474,238,1901]" captionTargetPageId="6" captionText="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." pageId="8" pageNumber="7">Figs. 16–18</figureCitation>
,
<figureCitation id="134F2A78FFCBFFF7477AFABB3130FA6A" box="[550,580,1384,1408]" captionStart="Figures 19–26" captionStartId="7.[114,191,1981,2003]" captionTargetBox="[113,1474,240,1941]" captionTargetId="figure-41@7.[113,1474,240,1956]" captionTargetPageId="7" captionText="Figures 19–26: Neobuthus haeckeli sp. n., holotype male, carapace and tergites I–V (19), coxosternal area and sternites (20), telson lateral (21), and right legs I–IV, retrolateral aspect (22–26)." pageId="8" pageNumber="7">21</figureCitation>
). Metasoma and telson sparsely hirsute, macrosetae moderately short in male, straight and reddish to black; metasomal segments I–II with 10 carinae, III with 8 carinae, IV with 6–8 carinae, V with 2 carinae; segments I–III with well-defined, granulate carinae; segment IV with weakly indicated dorsolateral carinae; segment V with strong, granulate to dentate-lobate ventrolateral carinae; segments I–IV with dense granulation on all intercarinal surfaces including dorsal surfaces; segment V densely granular on lateral and ventral surfaces, more coarsely so on ventral surface, granules not arranged along any traces of carinae; telson smooth, ventral surface sparsely, weakly granular; vesicle slightly elongated; aculeus stout, shorter than vesicle, tip of aculeus almost vertically directed.
</paragraph>
<paragraph id="8BCB36FDFFCBFFF7452DF89B3630FAEA" blockId="8.[113,772,1864,2016]" lastBlockId="8.[816,1475,1032,1280]" pageId="8" pageNumber="7">
AFFINITIES. The described features distinguish
<taxonomicName id="4C744D7EFFCBFFF7470CF89A31B0F88A" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[592,708,1864,1888]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF7470CF89A31B0F88A" box="[592,708,1864,1888]" italics="true" pageId="8" pageNumber="7">N. haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF74794F89B3077F88A" box="[712,771,1864,1888]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF74794F89B3195F88A" bold="true" box="[712,737,1864,1888]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF747B2F89B3189F88A" bold="true" box="[750,765,1864,1888]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. According to the characters used in the key published in
<bibRefCitation id="EFE54B0CFFCBFFF744E8F85B31EAF84A" author="KOVARIK" box="[436,670,1928,1952]" pageId="8" pageNumber="7" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al. (2018)</bibRefCitation>
, the new species is most similar to
<taxonomicName id="4C744D7EFFCBFFF744D4F87A3070F82A" authority="Kovarik et al., 2018" authorityName="Kovarik" authorityYear="2018" box="[392,772,1960,1984]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="amoudensis">
<emphasis id="B900EAEFFFCBFFF744D4F87A3151F82A" box="[392,549,1960,1984]" italics="true" pageId="8" pageNumber="7">N. amoudensis</emphasis>
<bibRefCitation id="EFE54B0CFFCBFFF74771F87B3070F82A" author="KOVARIK" box="[557,772,1960,1984]" pageId="8" pageNumber="7" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C744D7EFFCBFFF745C0F81A3177F80A" authority="Kovarik et al., 2018" authorityName="Kovarik" authorityYear="2018" box="[156,515,1992,2016]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="B900EAEFFFCBFFF745C0F81A3244F80A" box="[156,304,1992,2016]" italics="true" pageId="8" pageNumber="7">N. gubanensis</emphasis>
<bibRefCitation id="EFE54B0CFFCBFFF74468F81B3177F80A" author="KOVARIK" box="[308,515,1992,2016]" pageId="8" pageNumber="7" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
</taxonomicName>
, which is also confirmed by DNA phylogeny (paper in preparation). All examined specimens of
<taxonomicName id="4C744D7EFFCBFFF746E1FBFA3723FBAA" box="[957,1111,1064,1088]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="amoudensis">
<emphasis id="B900EAEFFFCBFFF746E1FBFA3723FBAA" box="[957,1111,1064,1088]" italics="true" pageId="8" pageNumber="7">N. amoudensis</emphasis>
</taxonomicName>
have pedipalp movable finger with 6 subrows of denticles (figs. 42 and
<quantity id="4C8C9B18FFCBFFF741F1FB9B3796FB8A" box="[1197,1250,1096,1120]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" pageId="8" pageNumber="7" unit="in" value="52.0">52 in</quantity>
<bibRefCitation id="EFE54B0CFFCBFFF741B5FB9B303EFB6A" author="KOVARIK" pageId="8" pageNumber="7" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018: 11</bibRefCitation>
) while all
<typeStatus id="54CF885FFFCBFFF746EBFBBA3090FB6B" box="[951,996,1129,1153]" pageId="8" pageNumber="7">type</typeStatus>
specimens of
<taxonomicName id="4C744D7EFFCBFFF7412BFBBA3798FB6A" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[1143,1260,1128,1152]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF7412BFBBA3798FB6A" box="[1143,1260,1128,1152]" italics="true" pageId="8" pageNumber="7">N. haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF741ADFBBB3658FB6A" box="[1265,1324,1128,1152]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF741ADFBBB367EFB6A" bold="true" box="[1265,1290,1128,1152]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF7404AFBBB3651FB6A" bold="true" box="[1302,1317,1128,1152]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
have pedipalp movable finger with 5 of these subrows (
<figureCitation id="134F2A78FFCBFFF7418DFB5B366AFB4A" box="[1233,1310,1160,1184]" captionStart="Figures 5–18" captionStartId="6.[115,192,1928,1950]" captionTargetBox="[121,1470,243,1893]" captionTargetId="figure-103@6.[113,1474,238,1901]" captionTargetPageId="6" captionText="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." pageId="8" pageNumber="7">Fig. 14</figureCitation>
).
<taxonomicName id="4C744D7EFFCBFFF74073FB5A36B6FB4A" box="[1327,1474,1160,1184]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="B900EAEFFFCBFFF74073FB5A36B6FB4A" box="[1327,1474,1160,1184]" italics="true" pageId="8" pageNumber="7">N. gubanensis</emphasis>
</taxonomicName>
could be characterized by the presence of a median lateral carina on metasoma III (figs 230 and
<quantity id="4C8C9B18FFCBFFF7412DFB1B37C7FB0A" box="[1137,1203,1224,1248]" metricMagnitude="0" metricUnit="m" metricValue="5.9182" pageId="8" pageNumber="7" unit="in" value="233.0">233 in</quantity>
<bibRefCitation id="EFE54B0CFFCBFFF741E6FB1B36C2FB0A" author="KOVARIK" box="[1210,1462,1224,1248]" pageId="8" pageNumber="7" pagination="1 - 82" refId="ref5785" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI &amp; A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">Kovařík et al., 2018: 49</bibRefCitation>
). This carina is absent in
<taxonomicName id="4C744D7EFFCBFFF7417AFB3A37EFFAEA" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[1062,1179,1256,1280]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF7417AFB3A37EFFAEA" box="[1062,1179,1256,1280]" italics="true" pageId="8" pageNumber="7">N. haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF741FDFB3B37A9FAEA" box="[1185,1245,1256,1280]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF741FDFB3B37CEFAEA" bold="true" box="[1185,1210,1256,1280]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF7419BFB3B37A2FAEA" bold="true" box="[1223,1238,1256,1280]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="134F2A78FFCBFFF741B6FB3B364CFAEA" box="[1258,1336,1256,1280]" captionStart="Figures 5–18" captionStartId="6.[115,192,1928,1950]" captionTargetBox="[121,1470,243,1893]" captionTargetId="figure-103@6.[113,1474,238,1901]" captionTargetPageId="6" captionText="Figures 5–18: Neobuthus haeckeli sp. n., holotype male. Figures 5–15. Pedipalp chela, dorsal (5), external (6), and ventral (7) views, pedipalp patella, dorsal (8), external (9) and ventral (10) views, pedipalp femur and trochanter ventral (11), internal (12), and dorsal (13) views, pedipalp movable (14) and fixed (15) fingers dentate margins. The trichobothrial pattern is indicated in Figures 6–10, 12–13 (white circles). Figures 16–18. Metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bar: 10 mm (16–18)." pageId="8" pageNumber="7">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E6576FFCBFFF7466CFAFB30D4FA0A" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8BCB36FDFFCBFFF7466CFAFB30D4FA0A" blockId="8.[816,1475,1320,1504]" pageId="8" pageNumber="7">
COMMENTS ON LOCALITIES AND LIFE STRATEGY.
<taxonomicName id="4C744D7EFFCBFFF7407FFAFA36EFFAAA" authority="Kovařík, 2019" authorityName="Kovařík" authorityYear="2019" box="[1315,1435,1320,1344]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="B900EAEFFFCBFFF7407FFAFA36EFFAAA" box="[1315,1435,1320,1344]" italics="true" pageId="8" pageNumber="7">N. haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335794FFCBFFF740FFFAFB3032FA8A" pageId="8" pageNumber="7" rank="species">
<emphasis id="B900EAEFFFCBFFF740FFFAFB36C8FAAA" bold="true" box="[1443,1468,1320,1344]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B900EAEFFFCBFFF7466CFA9B304BFA8A" bold="true" box="[816,831,1352,1376]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
inhabits rocky mountain area in central Somaliland (
<figureCitation id="134F2A78FFCBFFF740CBFA9B3034FA6A" captionStart="Figures 1–2" captionStartId="3.[114,191,2015,2037]" captionTargetBox="[114,1475,234,2007]" captionTargetId="figure-28@3.[114,1475,234,2009]" captionTargetPageId="3" captionText="Figures 1–2. Neobuthus haeckeli sp. n., paratype male in vivo habitus (1) and type locality (2)." pageId="8" pageNumber="7">Fig. 2</figureCitation>
). The
<typeStatus id="54CF885FFFCBFFF746D5FABA30B5FA6B" box="[905,961,1385,1409]" pageId="8" pageNumber="7">types</typeStatus>
were collected at night in open terrain by UV detection together with
<taxonomicName id="4C744D7EFFCBFFF74175FA5B36CAFA4A" authority="Pocock, 1901" authorityName="Pocock" authorityYear="1901" box="[1065,1470,1416,1440]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="B900EAEFFFCBFFF74175FA5B3652FA4A" box="[1065,1318,1416,1440]" italics="true" pageId="8" pageNumber="7">Parabuthus abyssinicus</emphasis>
Pocock, 1901
</taxonomicName>
,
<taxonomicName id="4C744D7EFFCBFFF7466CFA7A37ACFA2A" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[816,1240,1448,1472]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="polystictus">
<emphasis id="B900EAEFFFCBFFF7466CFA7A3752FA2A" box="[816,1062,1448,1472]" italics="true" pageId="8" pageNumber="7">Hottentotta polystictus</emphasis>
(Pocock, 1896)
</taxonomicName>
,
<taxonomicName id="4C744D7EFFCBFFF741BAFA7B36F9FA2A" box="[1254,1421,1448,1472]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAEFFFCBFFF741BAFA7B3616FA2A" box="[1254,1378,1448,1472]" italics="true" pageId="8" pageNumber="7">Pandinurus</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="4C744D7EFFCBFFF7466CFA1B30D4FA0A" box="[816,928,1480,1504]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B900EAEFFFCBFFF7466CFA1B300EFA0A" box="[816,890,1480,1504]" italics="true" pageId="8" pageNumber="7">Buthus</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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