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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FFDC73501894FF7CFEEC9272" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Rattus ringens subsp. dobodurae Troughton 1946" docType="treatment" docVersion="6" lastPageNumber="378" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="378" updateTime="1674500149280" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Divljan, Anja</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7555710" ID-Zenodo-Dep="7555710" LSID="urn:lsid:plazi:treatment:03DD87C8FFDC73501894FF7CFEEC9272" httpUri="http://treatment.plazi.org/id/03DD87C8FFDC73501894FF7CFEEC9272" lastPageNumber="378" pageId="101" pageNumber="378">
<subSubSection box="[166,704,151,178]" pageId="101" pageNumber="378" type="nomenclature">
<paragraph blockId="101.[166,704,151,178]" box="[166,704,151,178]" pageId="101" pageNumber="378">
<heading box="[166,704,151,178]" centered="true" fontSize="11" level="2" pageId="101" pageNumber="378" reason="2">
<taxonomicName authority="Troughton, 1946" authorityName="Troughton" authorityYear="1946" box="[166,704,151,178]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="101" pageNumber="378" phylum="Chordata" rank="subSpecies" species="ringens" subSpecies="dobodurae">
<emphasis bold="true" box="[166,704,151,178]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[166,486,151,178]" italics="true" pageId="101" pageNumber="378">Rattus ringens dobodurae</emphasis>
<bibRefCitation author="Troughton, E" box="[493,704,151,177]" pageId="101" pageNumber="378" pagination="406 - 410" refId="ref147877" refString="Troughton, E. L. G. 1946. Diagnoses of new rats from the New Guinea area. Records of the Australian Museum 21 (7): 406 - 410. https: // doi. org / 10.3853 / j. 0067 - 1975.21.1946.558" type="journal article" year="1946">Troughton, 1946</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,570,197,220]" pageId="101" pageNumber="378" type="reference_group">
<paragraph blockId="101.[151,570,197,220]" box="[151,570,197,220]" pageId="101" pageNumber="378">
<emphasis box="[151,302,198,219]" italics="true" pageId="101" pageNumber="378">Rec. Aust. Mus.</emphasis>
21(7): 407. (
<date box="[426,559,197,219]" pageId="101" pageNumber="378" value="1946-06-24">24 June 1946</date>
).
</paragraph>
</subSubSection>
<subSubSection box="[121,472,241,265]" pageId="101" pageNumber="378" type="vernacular_names">
<paragraph blockId="101.[121,472,241,265]" box="[121,472,241,265]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[121,295,241,265]" pageId="101" pageNumber="378">Common name</emphasis>
. Cape
<collectingRegion box="[369,421,241,265]" country="United Kingdom" name="York" pageId="101" pageNumber="378">York</collectingRegion>
Rat.
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="description">
<paragraph blockId="101.[119,749,286,369]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[121,290,286,310]" pageId="101" pageNumber="378">Current name</emphasis>
.
<taxonomicName authority="Troughton, 1946" authorityName="Troughton" authorityYear="1946" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="101" pageNumber="378" phylum="Chordata" rank="subSpecies" species="leucopus" subSpecies="dobodurae">
<emphasis box="[308,616,286,310]" italics="true" pageId="101" pageNumber="378">Rattus leucopus dobodurae</emphasis>
<bibRefCitation author="Troughton, E" pageId="101" pageNumber="378" pagination="406 - 410" refId="ref147877" refString="Troughton, E. L. G. 1946. Diagnoses of new rats from the New Guinea area. Records of the Australian Museum 21 (7): 406 - 410. https: // doi. org / 10.3853 / j. 0067 - 1975.21.1946.558" type="journal article" year="1946">Troughton, 1946</bibRefCitation>
</taxonomicName>
; following
<bibRefCitation author="Taylor, J" box="[295,498,316,340]" pageId="101" pageNumber="378" pagination="177 - 336" refId="ref144998" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History 173 (3): 177 - 336." type="journal article" year="1982">
Taylor
<emphasis box="[368,420,316,340]" italics="true" pageId="101" pageNumber="378">et al.</emphasis>
(1982)
</bibRefCitation>
and Musser &amp; Carleton (2005).
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="materials_examined">
<paragraph blockId="101.[121,749,390,502]" pageId="101" pageNumber="378">
<materialsCitation collectingDate="1944-04-25" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectorName="Captain G. M. Kohls" country="United States of America" location="USA" pageId="101" pageNumber="378" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[121,227,390,414]" pageId="101" pageNumber="378">
<typeStatus box="[121,227,390,414]" pageId="101" pageNumber="378">Holotype</typeStatus>
</emphasis>
. M.6960 by original designation. Male adult, [Field no. 253], skull (
<figureCitation box="[362,442,419,444]" captionStart="Figure 46" captionStartId="101.[794,856,1533,1555]" captionTargetBox="[801,1410,163,1493]" captionTargetId="figure-478@101.[794,1421,151,1512]" captionTargetPageId="101" captionText="Figure 46.AM M.6960, holotype skull ofRattus ringens dobodurae Troughton, 1946. (Photography by Stuart Humphreys)." figureDoi="http://doi.org/10.5281/zenodo.5237898" httpUri="https://zenodo.org/record/5237898/files/figure.png" pageId="101" pageNumber="378">Fig. 46</figureCitation>
), study skin, collected on
<date pageId="101" pageNumber="378" value="1944-04-25">
<collectingDate pageId="101" pageNumber="378" value="1944-04-25">25 April 1944</collectingDate>
</date>
, presented by
<collectorName box="[386,606,449,473]" pageId="101" pageNumber="378">Captain G. M. Kohls</collectorName>
,
<collectingCountry box="[615,667,449,473]" name="United States of America" pageId="101" pageNumber="378">USA</collectingCountry>
Typhus Commission, and registered on
<date box="[459,605,478,502]" pageId="101" pageNumber="378" value="1944-06-23">
<collectingDate box="[459,605,478,502]" pageId="101" pageNumber="378" value="1944-06-23">23 June 1944</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="description">
<paragraph blockId="101.[121,749,523,577]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[121,235,523,547]" pageId="101" pageNumber="378">Condition</emphasis>
. Cranium missing jugal bone from both zygomatic arches; both dentaries complete. Study skin complete.
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="distribution">
<paragraph blockId="101.[120,748,598,652]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[121,265,598,622]" pageId="101" pageNumber="378">Type locality</emphasis>
. Dobodura district,
<collectingRegion box="[475,674,598,622]" country="Papua New Guinea" name="Northern" pageId="101" pageNumber="378">Northern Province</collectingRegion>
,
<collectingCountry name="Papua New Guinea" pageId="101" pageNumber="378">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="materials_examined">
<paragraph blockId="101.[119,750,673,961]" pageId="101" pageNumber="378">
<materialsCitation collectingDate="1944-09" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectionCode="M" collectorName="Captain G. M. Kohls" country="United States of America" location="Typhus Commission" municipality="Typhus Commission" pageId="101" pageNumber="378" specimenCode="M.6987" specimenCount="5" typeStatus="paratype">
<emphasis bold="true" box="[121,236,673,697]" pageId="101" pageNumber="378">
<typeStatus box="[121,236,673,697]" pageId="101" pageNumber="378">Paratypes</typeStatus>
</emphasis>
. (
<specimenCount box="[262,277,673,697]" pageId="101" pageNumber="378">5</specimenCount>
, 4 by subsequent determination). All with same data as holotype, except dates of collection:
<emphasis bold="true" box="[664,750,702,726]" pageId="101" pageNumber="378">
<specimenCode box="[664,750,702,726]" pageId="101" pageNumber="378">M.6987</specimenCode>
</emphasis>
(by original designation), female (allotype), skull, study skin, collection date not given, presented by
<collectingCountry box="[610,662,761,785]" name="United States of America" pageId="101" pageNumber="378">USA</collectingCountry>
<collectingMunicipality pageId="101" pageNumber="378">Typhus Commission</collectingMunicipality>
, registered
<date box="[392,574,790,814]" pageId="101" pageNumber="378" value="1944-09">
<collectingDate box="[392,574,790,814]" pageId="101" pageNumber="378" value="1944-09">September 1944</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation collectingDate="1944-09" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectionCode="M" collectorName="Captain G. M. Kohls" country="United States of America" location="All" municipality="Typhus Commission" pageId="101" pageNumber="378" specimenCode="M.6917" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[587,672,790,814]" pageId="101" pageNumber="378">
<specimenCode box="[587,672,790,814]" pageId="101" pageNumber="378">M.6917</specimenCode>
</emphasis>
,
<specimenCount box="[689,744,790,814]" pageId="101" pageNumber="378" type="male">male</specimenCount>
, skull, study skin, collection date not given;
</materialsCitation>
<materialsCitation collectingDate="1944-05-27" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectionCode="M" collectorName="Captain G. M. Kohls" country="United States of America" location="All" municipality="Typhus Commission" pageId="101" pageNumber="378" specimenCode="M.6973" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[591,675,819,843]" pageId="101" pageNumber="378">
<specimenCode box="[591,675,819,843]" pageId="101" pageNumber="378">M.6973</specimenCode>
</emphasis>
,
<specimenCount box="[690,744,820,844]" pageId="101" pageNumber="378" type="male">male</specimenCount>
, skull, study skin, collected on
<date box="[440,584,849,873]" pageId="101" pageNumber="378" value="1944-05-27">
<collectingDate box="[440,584,849,873]" pageId="101" pageNumber="378" value="1944-05-27">27 May 1944</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation collectingDate="1944-06-21" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectionCode="M" collectorName="Captain G. M. Kohls" country="United States of America" location="All" municipality="Typhus Commission" pageId="101" pageNumber="378" specimenCode="M.6974" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[591,677,849,873]" pageId="101" pageNumber="378">
<specimenCode box="[591,677,849,873]" pageId="101" pageNumber="378">M.6974</specimenCode>
</emphasis>
,
<specimenCount box="[690,744,849,873]" pageId="101" pageNumber="378" type="male">male</specimenCount>
, body in alc., collected on
<date box="[412,565,878,902]" pageId="101" pageNumber="378" value="1944-06-21">
<collectingDate box="[412,565,878,902]" pageId="101" pageNumber="378" value="1944-06-21">21 June 1944</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation collectingDate="1944-09" collectingDateMax="1944-06-23" collectingDateMin="1944-04-25" collectionCode="M" collectorName="Captain G. M. Kohls" country="United States of America" location="All" municipality="Typhus Commission" pageId="101" pageNumber="378" specimenCode="M.6988" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[580,665,878,902]" pageId="101" pageNumber="378">
<specimenCode box="[580,665,878,902]" pageId="101" pageNumber="378">M.6988</specimenCode>
</emphasis>
, young
<specimenCount box="[121,195,908,932]" pageId="101" pageNumber="378" type="female">female</specimenCount>
, skull, study skin, collection date not given, registered in
<date box="[146,324,937,961]" pageId="101" pageNumber="378" value="1944-09">
<collectingDate box="[146,324,937,961]" pageId="101" pageNumber="378" value="1944-09">September 1944</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="101" pageNumber="378" type="discussion">
<paragraph blockId="101.[119,751,982,1358]" pageId="101" pageNumber="378">
<emphasis bold="true" box="[121,250,982,1006]" pageId="101" pageNumber="378">Comments</emphasis>
. Troughton cites registration numbers for the
<typeStatus box="[164,258,1012,1036]" pageId="101" pageNumber="378">holotype</typeStatus>
and
<typeStatus box="[317,404,1012,1036]" pageId="101" pageNumber="378">allotype</typeStatus>
but does not indicate the total number in the type series. However, he commented that the
<typeStatus box="[165,261,1070,1094]" pageId="101" pageNumber="378">holotype</typeStatus>
and
<typeStatus box="[322,411,1070,1094]" pageId="101" pageNumber="378">allotype</typeStatus>
were submitted with a series from Dobodura presented by the US Typhus Commission. Another
<specimenCount box="[219,383,1129,1153]" pageId="101" pageNumber="378" type="generic">four specimens</specimenCount>
of that series are identified as
<taxonomicName authorityName="Troughton" authorityYear="1946" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="101" pageNumber="378" phylum="Chordata" rank="subSpecies" species="ringens" subSpecies="dobodurae">
<emphasis italics="true" pageId="101" pageNumber="378">R. ringens dobodurae</emphasis>
</taxonomicName>
in Troughtons handwriting in the register and these are the only specimens in the register with data matching that of the
<typeStatus box="[337,434,1217,1241]" pageId="101" pageNumber="378">holotype</typeStatus>
. Although none of the four are marked as
<typeStatus box="[239,341,1247,1271]" pageId="101" pageNumber="378">paratypes</typeStatus>
in the register or on specimen index cards, we treat them as such on the understanding that they would have been examined by Troughton during preparation of the manuscript.
</paragraph>
</subSubSection>
</treatment>
</document>