treatments-xml/data/03/DD/87/03DD8798FFE06A64FF42DEA5A532FDC8.xml

<document id="303CBE1FBD9782ADF0EE3761539A707E" ID-DOI="10.11646/phytotaxa.340.3.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="13719193" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725656262042" checkinUser="felipe" docAuthor="Rana, Hum Kala, Sun, Hang, Paudel, Asha &amp; Ghimire, Suresh Kumar" docDate="2018" docId="03DD8798FFE06A64FF42DEA5A532FDC8" docLanguage="en" docName="phytotaxa.340.3.7.pdf" docOrigin="Phytotaxa 340 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.340.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Saussurea ramchaudharyi S. K. Ghimire &amp; H. K. Rana 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="274" masterDocId="FFE4FFE0FFE16A67FFCADE53A059FF45" masterDocTitle="Saussurea ramchaudharyi (Asteraceae), a new species from Nepal" masterLastPageNumber="276" masterPageNumber="271" pageNumber="272" updateTime="1725667032461" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B4EC0E34BC5F88BEDED80E15D80DA040" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="753AD15052DE881F2435FD51F24F2101">
<mods:title id="6AF7EDEC99C6B1F6BCE507DC762382FB">Saussurea ramchaudharyi (Asteraceae), a new species from Nepal</mods:title>
</mods:titleInfo>
<mods:name id="2E3919040FE29FBDB09C9B3A2F8936C3" type="personal">
<mods:role id="1C8D8E870024F60B1A3CF6886BCEB81E">
<mods:roleTerm id="2657ED968F25A6DB47262485561AFAC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA9788ACE71FB20278F3808ABD97F86E">Rana, Hum Kala</mods:namePart>
<mods:affiliation id="CF96A30309C9ADC0FE2E3B63AFBD1E88">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China &amp; Central Department of Botany, Tribhuvan University, Kirtipur, Kathmandu, Nepal</mods:affiliation>
</mods:name>
<mods:name id="D54725CBA792F48E88F43A456D52E19B" type="personal">
<mods:role id="F4525124FC15109BE49BF760A537EB2C">
<mods:roleTerm id="3B0BCDD417D9E0A16DC2C0C28C172E95">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFE84E332FB5FCF2FA6DCF865C73B03F">Sun, Hang</mods:namePart>
<mods:affiliation id="AF1C9DCB825416F63212368406C8F218">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name id="EF7798A224CB687CAFB095115DA9C746" type="personal">
<mods:role id="B8B3042C8F34CB83AA5CCF64D617C869">
<mods:roleTerm id="3BC6AD5212BABE774ADCCBC1087FC0EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BE780247A8F6BED474058897B0A3548">Paudel, Asha</mods:namePart>
<mods:affiliation id="4D56870BF9BAC871D78BECB5E23D9102">Central Department of Botany, Tribhuvan University, Kirtipur, Kathmandu, Nepal</mods:affiliation>
</mods:name>
<mods:name id="8D27E09D2FB1FBE598D9E973F10FA319" type="personal">
<mods:role id="4B8BC45E7E2E97817E9F60E105F5CDFF">
<mods:roleTerm id="B060A02122B78C8083EC2B197E89B838">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="580630C786BF72F768C340143FBE5974">Ghimire, Suresh Kumar</mods:namePart>
<mods:affiliation id="24BF3A8C0169737C6A01E2160CE7F791">Central Department of Botany, Tribhuvan University, Kirtipur, Kathmandu, Nepal</mods:affiliation>
</mods:name>
<mods:typeOfResource id="87182E30D63B117E8C5921989BE40600">text</mods:typeOfResource>
<mods:relatedItem id="7E1E9997945E2785C560A8F7F140E158" type="host">
<mods:titleInfo id="2662BBC31A34A1B807C08B5B7328101A">
<mods:title id="CD10F534695CE65EF9ED078C4E6D0907">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="22941712276D6740E90DCCB93D007E76">
<mods:date id="5697C4BE20078A0D5BEC545F79AAD585">2018</mods:date>
<mods:detail id="9D5034905CFFF136841B51D3B0D8BF30" type="pubDate">
<mods:number id="DC739DFF71C531EEADA90D2C0AC2E085">2018-02-27</mods:number>
</mods:detail>
<mods:detail id="04338C6067492F27B806BC76FF5B16A1" type="volume">
<mods:number id="E7251F084DF8A4C79503937E80B12A17">340</mods:number>
</mods:detail>
<mods:detail id="3A00F879DBA5E97AC5064FA02CBEB4EF" type="issue">
<mods:number id="C2833FE0A36A90C4C9D97A58A3D49571">3</mods:number>
</mods:detail>
<mods:extent id="A2F2A11C65DB624A93B27A92D511874B" unit="page">
<mods:start id="1E04A901B00D0E6EFE5FD1EE0E9D63E5">271</mods:start>
<mods:end id="252F349E805D23B68EA69077E39119E2">276</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="81A0EBB0EFF00B97C2551E7E85C7A316">
<mods:url id="6E5EE78AF04840A36F89F9A105FF1F8D">http://dx.doi.org/10.11646/phytotaxa.340.3.7</mods:url>
</mods:location>
<mods:classification id="085AB92304878332E222E72E090DC354">journal article</mods:classification>
<mods:identifier id="0F68FA0E7E238CF5A07C6B59E4FF7426" type="DOI">10.11646/phytotaxa.340.3.7</mods:identifier>
<mods:identifier id="117275822D2AA72802D20A0DC56DCB34" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="3F53BFCAA465923912ECF3416A72AE9D" type="Zenodo-Dep">13719193</mods:identifier>
</mods:mods>
<treatment id="03DD8798FFE06A64FF42DEA5A532FDC8" LSID="urn:lsid:plazi:treatment:03DD8798FFE06A64FF42DEA5A532FDC8" httpUri="http://treatment.plazi.org/id/03DD8798FFE06A64FF42DEA5A532FDC8" lastPageId="3" lastPageNumber="274" pageId="1" pageNumber="272">
<subSubSection id="C36E6505FFE06A66FF42DEA5A385FE4A" box="[136,988,246,271]" pageId="1" pageNumber="272" type="nomenclature">
<paragraph id="8BCB368EFFE06A66FF42DEA5A385FE4A" blockId="1.[136,988,246,271]" box="[136,988,246,271]" pageId="1" pageNumber="272">
<heading id="D08381E2FFE06A66FF42DEA5A385FE4A" box="[136,988,246,271]" fontSize="11" level="2" pageId="1" pageNumber="272" reason="3">
<taxonomicName id="4C744D0DFFE06A66FF42DEA5A2ADFE4A" ID-CoL="6Y3DJ" authority="S.K. Ghimire &amp; H.K. Rana" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[136,756,246,271]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="272" phylum="Tracheophyta" rank="species" species="ramchaudharyi" status="sp. nov.">
<emphasis id="B900EA9CFFE06A66FF42DEA5A1E3FE4A" bold="true" box="[136,442,246,271]" italics="true" pageId="1" pageNumber="272">Saussurea ramchaudharyi</emphasis>
S.K. Ghimire &amp; H.K. Rana
</taxonomicName>
, 
<emphasis id="B900EA9CFFE06A66FCCADEA5A385FE4A" bold="true" box="[768,988,246,271]" pageId="1" pageNumber="272">
<taxonomicNameLabel id="A23357E7FFE06A66FCCADEA5A300FE4A" box="[768,857,246,271]" pageId="1" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="134F2A0BFFE06A66FCA2DEA5A38CFE4A" box="[872,981,246,271]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[136,229,1311,1332]" captionStartId-1="2.[136,229,1112,1133]" captionStartId-2="2.[136,229,2011,2032]" captionTargetBox-0="[174,1433,377,1270]" captionTargetBox-1="[453,1133,196,1080]" captionTargetBox-2="[477,1108,1227,1963]" captionTargetId-0="figure-65@1.[151,1437,374,1283]" captionTargetId-1="figure-13@2.[448,1139,190,1086]" captionTargetId-2="figure-39@2.[448,1139,1220,1973]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Distribution of Saussurea ramchaudharyi. (A, B) Distribution map showing type locality in Nepal (A) and Humla district (B). (C, D) Type locality habitat at Upper Chungsa valley, Humla district (C–D by S.K. Ghimire)." captionText-1="FIGURE 2. Saussurea ramchaudharyi. (A, B) Reproductive plant. (C) Vegetative plant (A–C by S.K. Ghimire). (D) Synflorescence exposed at anthesis (by H.K. Rana)." captionText-2="FIGURE 3. Saussurea ramchaudharyi (based on the holotype). (A) Habit. (B) Leaf (B1: upper leaf, and B2: lower leaf). (C) Floret with pappus. (D) Phyllaries (outer to inner from right→left). (E) Stamens. (F) Achene with pappus (drawn by H.K. Rana)." figureDoi-0="http://doi.org/10.5281/zenodo.13719195" figureDoi-1="http://doi.org/10.5281/zenodo.13719197" figureDoi-2="http://doi.org/10.5281/zenodo.13719199" httpUri-0="https://zenodo.org/record/13719195/files/figure.png" httpUri-1="https://zenodo.org/record/13719197/files/figure.png" httpUri-2="https://zenodo.org/record/13719199/files/figure.png" pageId="1" pageNumber="272">Figs. 1–3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E6505FFE06A64FF42DF4FA532FDC8" lastPageId="3" lastPageNumber="274" pageId="1" pageNumber="272" type="materials_examined">
<paragraph id="8BCB368EFFE06A66FF42DF4FA5F1FE10" blockId="1.[136,1451,284,341]" pageId="1" pageNumber="272">
<emphasis id="B900EA9CFFE06A66FF42DF4FA0ECFE74" box="[136,181,284,305]" italics="true" pageId="1" pageNumber="272">Type</emphasis>
:— 
<materialsCitation id="3B1C3CD3FFE06A66FF1BDF4FA5FDFE10" collectingDate="2012-09-10" collectionCode="KATH" collectorName="S. K. Ghimire &amp; A. Poudel &amp; C. Thapa &amp; L. R. Joshi &amp; S. Lo &amp; P. Subedi" country="Nepal" county="Seliman Lake" elevation="4650" latitude="30.143717" location="Humla" longLatPrecision="8" longitude="81.71567" municipality="Chungsa valley" pageId="1" pageNumber="272" specimenCount="2" typeStatus="holotype">
<collectingCountry id="F363761EFFE06A66FF1BDF4FA17BFE74" box="[209,290,284,305]" name="Nepal" pageId="1" pageNumber="272">NEPAL</collectingCountry>
. 
<location id="8EAB6055FFE06A66FEE1DF4FA136FE74" LSID="urn:lsid:plazi:treatment:03DD8798FFE06A64FF42DEA5A532FDC8:8EAB6055FFE06A66FEE1DF4FA136FE74" box="[299,367,284,305]" country="Nepal" county="Seliman Lake" latitude="30.143717" longLatPrecision="8" longitude="81.71567" municipality="Chungsa valley" name="Humla" pageId="1" pageNumber="272">Humla</location>
, 
<location id="8EAB6055FFE06A66FEB3DF4FA253FE74" LSID="urn:lsid:plazi:treatment:03DD8798FFE06A64FF42DEA5A532FDC8:8EAB6055FFE06A66FEB3DF4FA253FE74" box="[377,522,284,305]" country="Nepal" county="Seliman Lake" latitude="30.143717" longLatPrecision="8" longitude="81.71567" municipality="Chungsa valley" name="upper Chungsa" pageId="1" pageNumber="272">
upper 
<collectingMunicipality id="6BAFACF4FFE06A66FE7CDF4FA253FE74" box="[438,522,284,305]" pageId="1" pageNumber="272">Chungsa</collectingMunicipality>
</location>
valley, above 
<collectingCounty id="62AA4E02FFE06A66FD5DDF4FA347FE74" box="[663,798,284,305]" pageId="1" pageNumber="272">Seliman Lake</collectingCounty>
, rocky and scree slope, 
<geoCoordinate id="EE405049FFE06A66FBCCDF4FA4DCFE74" box="[1030,1157,284,305]" degrees="30" direction="north" minutes="08.623" orientation="latitude" pageId="1" pageNumber="272" precision="1" value="30.143717">30°08.623’N</geoCoordinate>
, 
<geoCoordinate id="EE405049FFE06A66FB45DF4FA559FE74" box="[1167,1280,284,305]" degrees="81" direction="east" minutes="42.94" orientation="longitude" pageId="1" pageNumber="272" precision="9" value="81.71567">81°42.94’E</geoCoordinate>
, 
<elevation id="0059D1BDFFE06A66FAC0DF4FA5D1FE74" box="[1290,1416,284,305]" metricMagnitude="3" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="272" unit="m" value="4650.0">
<quantity id="4C8C9B6BFFE06A66FAC0DF4FA50DFE74" box="[1290,1364,284,305]" metricMagnitude="3" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="272" unit="m" value="4650.0">4650 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFCA104EFFE06A66FA59DF4FA105FE10" pageId="1" pageNumber="272" value="2012-09-10">
<collectingDate id="EF8EE9A6FFE06A66FA59DF4FA105FE10" pageId="1" pageNumber="272" value="2012-09-10">10 September 2012</collectingDate>
</date>
, 
<emphasis id="B900EA9CFFE06A66FEACDF13A412FE10" box="[358,1099,320,341]" italics="true" pageId="1" pageNumber="272">
<collectorName id="26815358FFE06A66FEACDF13A1BCFE10" box="[358,485,320,341]" pageId="1" pageNumber="272">S.K. Ghimire</collectorName>
, 
<collectorName id="26815358FFE06A66FE25DF13A216FE10" box="[495,591,320,341]" pageId="1" pageNumber="272">A. Poudel</collectorName>
, 
<collectorName id="26815358FFE06A66FD93DF13A2ECFE10" box="[601,693,320,341]" pageId="1" pageNumber="272">C. Thapa</collectorName>
, 
<collectorName id="26815358FFE06A66FD75DF13A37BFE10" box="[703,802,320,341]" pageId="1" pageNumber="272">L.R. Joshi</collectorName>
, 
<collectorName id="26815358FFE06A66FCE1DF13A306FE10" box="[811,863,320,341]" pageId="1" pageNumber="272">S. Lo</collectorName>
, &amp; 
<collectorName id="26815358FFE06A66FC4BDF13A383FE10" box="[897,986,320,341]" pageId="1" pageNumber="272">P. Subedi</collectorName>
CHH-1247
</emphasis>
(
<typeStatus id="54CF882CFFE06A66FB92DF13A4E8FE10" box="[1112,1201,320,341]" pageId="1" pageNumber="272" type="holotype">holotype</typeStatus>
: TUCH; 
<typeStatus id="54CF882CFFE06A66FACDDF13A509FE10" box="[1287,1360,320,341]" pageId="1" pageNumber="272" type="isotype">isotype</typeStatus>
: 
<collectionCode id="ED65AE4BFFE06A66FA93DF13A5C7FE10" box="[1369,1438,320,341]" country="Nepal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12932" lsid="urn:lsid:biocol.org:col:12932" name="Department of Plant Resources" pageId="1" pageNumber="272" type="Herbarium">KATH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF0B6606FFE06A66FF42DB4CA450FA1D" ID-DOI="http://doi.org/10.5281/zenodo.13719195" ID-Zenodo-Dep="13719195" httpUri="https://zenodo.org/record/13719195/files/figure.png" pageId="1" pageNumber="272" startId="1.[136,229,1311,1332]" targetBox="[174,1433,377,1270]" targetPageId="1" targetType="figure">
<paragraph id="8BCB368EFFE06A66FF42DB4CA450FA1D" blockId="1.[136,1452,1311,1368]" pageId="1" pageNumber="272">
<emphasis id="B900EA9CFFE06A66FF42DB4CA0A4FA71" bold="true" box="[136,253,1311,1332]" pageId="1" pageNumber="272">FIGURE 1.</emphasis>
Distribution of 
<taxonomicName id="4C744D0DFFE06A66FE59DB4CA2D7FA71" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[403,654,1311,1332]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="272" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE06A66FE59DB4CA2D7FA71" box="[403,654,1311,1332]" italics="true" pageId="1" pageNumber="272">Saussurea ramchaudharyi</emphasis>
</taxonomicName>
. 
<emphasis id="B900EA9CFFE06A66FD53DB4CA28CFA71" bold="true" box="[665,725,1311,1332]" pageId="1" pageNumber="272">(A, B)</emphasis>
Distribution map showing type locality in Nepal (A) and Humla district (B). 
<emphasis id="B900EA9CFFE06A66FF42DB10A09EFA1D" bold="true" box="[136,199,1347,1368]" pageId="1" pageNumber="272">(C, D)</emphasis>
Type locality habitat at Upper Chungsa valley, Humla district (C–D by S.K. Ghimire).
</paragraph>
</caption>
<paragraph id="8BCB368EFFE06A66FF42DBDBA5C2F948" blockId="1.[136,1452,1416,2089]" pageId="1" pageNumber="272">
<emphasis id="B900EA9CFFE06A66FF42DBDBA0A3FAE4" box="[136,250,1416,1441]" italics="true" pageId="1" pageNumber="272">Diagnosis</emphasis>
:—Monocarpic herb, 
<quantity id="4C8C9B6BFFE06A66FE3BDBDBA21FFAE4" box="[497,582,1416,1441]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="272" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
tall; caudex unbranched; stem axis short 
<quantity id="4C8C9B6BFFE06A66FBE3DBDBA4C1FAE4" box="[1065,1176,1416,1441]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="272" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
; leaves almost prostrate on ground, upper leaves greenish purple, extending half of length of capitula; synflorescence 
<quantity id="4C8C9B6BFFE06A66FB68DBFFA546FA80" box="[1186,1311,1452,1477]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="1" pageNumber="272" unit="cm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 cm</quantity>
in diameter, exposed and surrounded by uppermost stem leaves; involucre broadly campanulate; phyllaries 4-seriate, inner phyllaries apically glabrous; achene much smaller ≤ 
<quantity id="4C8C9B6BFFE06A66FD90DBA7A2C2F948" box="[602,667,1524,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="272" unit="mm" value="2.0">2 mm</quantity>
with glabrous apex and inner bristles of pappus shorter than corolla.
</paragraph>
<paragraph id="8BCB368EFFE06A66FF77D84BA44AF76C" blockId="1.[136,1452,1416,2089]" pageId="1" pageNumber="272">
<emphasis id="B900EA9CFFE06A66FF77D84BA118F974" box="[189,321,1560,1585]" italics="true" pageId="1" pageNumber="272">Description</emphasis>
:—Perennial monocarpic herb, 
<quantity id="4C8C9B6BFFE06A66FD61D84BA358F974" box="[683,769,1560,1585]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="272" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
tall. Caudex unbranched, covered with leaf remains. Stem erect, simple, club-shaped, hollow, shorter 
<quantity id="4C8C9B6BFFE06A66FDB6D86FA2B1F910" box="[636,744,1596,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="272" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
long with 
<quantity id="4C8C9B6BFFE06A66FCA6D86FA3B4F910" box="[876,1005,1596,1621]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="1" pageNumber="272" unit="cm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 cm</quantity>
broad apex. Leaves in rosette, almost prostrate on the ground, linear to narrowly oblanceolate; green or purplish green; midvein distinct, apex acute; margin sparsely denticulate to pinnatilobed in upper half; lobes 6–10 pairs, somewhat curving upward from half of length. Basal or lower leaves petiolate, narrowly oblanceolate, 8.5–10.2 × 
<quantity id="4C8C9B6BFFE06A66FCB1D8FBA3A5F984" box="[891,1020,1704,1729]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="1" pageNumber="272" unit="cm" value="0.7" valueMax="0.9" valueMin="0.5">0.5–0.9 cm</quantity>
, thinly pilose or glabrous; lobes 
<quantity id="4C8C9B6BFFE06A66FAA7D8FBA0EEF9A0" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="272" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
; petiole reddish purple. Upper leaves sessile, linear to narrowly oblanceolate, 5–9.5 × 
<quantity id="4C8C9B6BFFE06A66FB6ED89FA571F9A0" box="[1188,1320,1740,1765]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="1" pageNumber="272" unit="cm" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 cm</quantity>
; lower part pilose (covered with woolly hairs); lobes 
<quantity id="4C8C9B6BFFE06A66FD90D8A3A2E3F84C" box="[602,698,1776,1801]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="272" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
. Synflorescence 
<quantity id="4C8C9B6BFFE06A66FCBDD8A3A3ACF84C" box="[887,1013,1776,1801]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="1" pageNumber="272" unit="cm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 cm</quantity>
across, aggregated in the center of leaf rosette, terminal on stem, exposed and surrounded by densely pilose subtending uppermost stem leaves. Capitula more than 15, sessile, crowded at the apex. Involucre campanulate, 
<quantity id="4C8C9B6BFFE06A66FCF5D96BA3E5F814" box="[831,956,1848,1873]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="1" pageNumber="272" unit="cm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 cm</quantity>
in diameter. Involucral phyllaries 4–seriate, imbricate, 0.8–1.8 × 
<quantity id="4C8C9B6BFFE06A66FEBCD90FA24DF830" box="[374,532,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="1" pageNumber="272" unit="cm" value="0.2" valueMax="0.25" valueMin="0.15">0.15–0.25 cm</quantity>
, membranous; outer linear to lanceolate, densely pilose; inner linear or narrowly lanceolate, glabrous, apex acute. Receptacle convex, setaceous; bristles 
<quantity id="4C8C9B6BFFE06A66FC61D9D3A479F8DC" box="[939,1056,1920,1945]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="272" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
. Corolla fused at most of its length, glabrous, dark purple, 
<quantity id="4C8C9B6BFFE06A66FE4DD9F7A1ACF8F8" box="[391,501,1956,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="1" pageNumber="272" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
; tube 
<quantity id="4C8C9B6BFFE06A66FDF3D9F7A2C3F8F8" box="[569,666,1956,1981]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="272" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
; limb 
<quantity id="4C8C9B6BFFE06A66FD28D9F7A319F8F8" box="[738,832,1956,1981]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="272" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
including 
<emphasis id="B900EA9CFFE06A66FC72D9F7A38BF8F8" box="[952,978,1956,1981]" italics="true" pageId="1" pageNumber="272">ca</emphasis>
. 
<quantity id="4C8C9B6BFFE06A66FC28D9F7A40CF8F8" box="[994,1109,1956,1981]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="272" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long lobes. Stamen 
<quantity id="4C8C9B6BFFE06A66FAF1D9F7A5FEF8F8" box="[1339,1447,1956,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="272" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
, head ± tapered, tail glabrous. Carpel-1, style slender, 
<quantity id="4C8C9B6BFFE06A66FD12D99BA363F8A4" box="[728,826,1992,2017]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="272" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1-1.2 cm</quantity>
long, stigma bilobed. Achene ≤ 
<quantity id="4C8C9B6BFFE06A66FB51D99BA484F8A4" box="[1179,1245,1992,2017]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="272" unit="mm" value="2.0">2 mm</quantity>
, with curved base, glabrous apically. Pappus 2-seriate, whitish or straw-colored; outer series caducous, with scabrid bristles, 
<quantity id="4C8C9B6BFFE06A66FA86D9BFA5F5F740" box="[1356,1452,2028,2053]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="272" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long; inner series persistent, plumose, 
<quantity id="4C8C9B6BFFE06A66FDFCD643A2CAF76C" box="[566,659,2064,2089]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="272" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, bristles shorter than corolla.
</paragraph>
<caption id="DF0B6606FFE36A65FF42DA0BA1BBFBD4" ID-DOI="http://doi.org/10.5281/zenodo.13719197" ID-Zenodo-Dep="13719197" httpUri="https://zenodo.org/record/13719197/files/figure.png" pageId="2" pageNumber="273" startId="2.[136,229,1112,1133]" targetBox="[453,1133,196,1080]" targetPageId="2" targetType="figure">
<paragraph id="8BCB368EFFE36A65FF42DA0BA1BBFBD4" blockId="2.[136,1452,1112,1169]" pageId="2" pageNumber="273">
<emphasis id="B900EA9CFFE36A65FF42DA0BA159FB28" bold="true" box="[136,256,1112,1133]" pageId="2" pageNumber="273">FIGURE 2.</emphasis>
<taxonomicName id="4C744D0DFFE36A65FEC3DA0BA25EFB28" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[265,519,1112,1133]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE36A65FEC3DA0BA25EFB28" box="[265,519,1112,1133]" italics="true" pageId="2" pageNumber="273">Saussurea ramchaudharyi</emphasis>
</taxonomicName>
. 
<emphasis id="B900EA9CFFE36A65FDDFDA0BA20CFB28" bold="true" box="[533,597,1112,1133]" pageId="2" pageNumber="273">(A, B)</emphasis>
Reproductive plant. 
<emphasis id="B900EA9CFFE36A65FCECDA0BA31EFB28" bold="true" box="[806,839,1112,1133]" pageId="2" pageNumber="273">(C)</emphasis>
Vegetative plant (A–C by S.K. Ghimire). (D) Synflorescence exposed at anthesis (by H.K. Rana).
</paragraph>
</caption>
<caption id="DF0B6606FFE36A65FF42D988A4A9F751" ID-DOI="http://doi.org/10.5281/zenodo.13719199" ID-Zenodo-Dep="13719199" httpUri="https://zenodo.org/record/13719199/files/figure.png" pageId="2" pageNumber="273" startId="2.[136,229,2011,2032]" targetBox="[477,1108,1227,1963]" targetPageId="2" targetType="figure">
<paragraph id="8BCB368EFFE36A65FF42D988A4A9F751" blockId="2.[136,1452,2011,2068]" pageId="2" pageNumber="273">
<emphasis id="B900EA9CFFE36A65FF42D988A0A7F8B5" bold="true" box="[136,254,2011,2032]" pageId="2" pageNumber="273">FIGURE 3.</emphasis>
<taxonomicName id="4C744D0DFFE36A65FECED988A259F8B5" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[260,512,2011,2032]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE36A65FECED988A259F8B5" box="[260,512,2011,2032]" italics="true" pageId="2" pageNumber="273">Saussurea ramchaudharyi</emphasis>
</taxonomicName>
(based on the holotype). 
<emphasis id="B900EA9CFFE36A65FD3ED988A34CF8B5" bold="true" box="[756,789,2011,2032]" pageId="2" pageNumber="273">(A)</emphasis>
Habit. 
<emphasis id="B900EA9CFFE36A65FC97D988A324F8B5" bold="true" box="[861,893,2011,2032]" pageId="2" pageNumber="273">(B)</emphasis>
Leaf (
<emphasis id="B900EA9CFFE36A65FC77D988A383F8B5" bold="true" box="[957,986,2011,2032]" pageId="2" pageNumber="273">B1</emphasis>
: upper leaf, and 
<emphasis id="B900EA9CFFE36A65FBB6D988A4C1F8B5" bold="true" box="[1148,1176,2011,2032]" pageId="2" pageNumber="273">B2</emphasis>
: lower leaf). 
<emphasis id="B900EA9CFFE36A65FAD0D988A562F8B5" bold="true" box="[1306,1339,2011,2032]" pageId="2" pageNumber="273">(C)</emphasis>
Floret with pappus. 
<emphasis id="B900EA9CFFE36A65FF12D9ACA0A0F751" bold="true" box="[216,249,2047,2068]" pageId="2" pageNumber="273">(D)</emphasis>
Phyllaries (outer to inner from right→left). 
<emphasis id="B900EA9CFFE36A65FD69D9ACA29BF751" bold="true" box="[675,706,2047,2068]" pageId="2" pageNumber="273">(E)</emphasis>
Stamens. 
<emphasis id="B900EA9CFFE36A65FCECD9ACA31DF751" bold="true" box="[806,836,2047,2068]" pageId="2" pageNumber="273">(F)</emphasis>
Achene with pappus (drawn by H.K. Rana).
</paragraph>
</caption>
<paragraph id="8BCB368EFFE26A64FF77DEF3A337FFFC" blockId="3.[136,1452,160,653]" box="[189,878,160,185]" pageId="3" pageNumber="274">
<emphasis id="B900EA9CFFE26A64FF77DEF3A16CFFFC" box="[189,309,160,185]" italics="true" pageId="3" pageNumber="274">Phenology</emphasis>
:—Flowering and fruiting from August to October.
</paragraph>
<paragraph id="8BCB368EFFE26A64FF77DE97A373FE60" blockId="3.[136,1452,160,653]" pageId="3" pageNumber="274">
<emphasis id="B900EA9CFFE26A64FF77DE97A18AFF98" box="[189,467,196,221]" italics="true" pageId="3" pageNumber="274">Distribution and habitat</emphasis>
:— 
<taxonomicName id="4C744D0DFFE26A64FE3DDE97A379FF98" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[503,800,196,221]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE26A64FE3DDE97A379FF98" box="[503,800,196,221]" italics="true" pageId="3" pageNumber="274">Saussurea ramchaudharyi</emphasis>
</taxonomicName>
is currently known only from the 
<typeStatus id="54CF882CFFE26A64FB7EDE97A4BDFF98" box="[1204,1252,196,221]" pageId="3" pageNumber="274">type</typeStatus>
locality in upper Chungsa Valley, Humla district, 
<collectingCountry id="F363761EFFE26A64FE3FDEBBA261FE44" box="[501,568,232,257]" name="Nepal" pageId="3" pageNumber="274">Nepal</collectingCountry>
at elevation 
<quantity id="4C8C9B6BFFE26A64FD03DEBBA346FE44" box="[713,799,232,257]" metricMagnitude="3" metricUnit="m" metricValue="4.64" pageId="3" pageNumber="274" unit="m" value="4640.0">4640 m</quantity>
a.s.l. (
<figureCitation id="134F2A0BFFE26A64FCA0DEBBA40AFE44" box="[874,1107,232,257]" captionStart="FIGURE 1" captionStartId="1.[136,229,1311,1332]" captionTargetBox="[174,1433,377,1270]" captionTargetId="figure-65@1.[151,1437,374,1283]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Saussurea ramchaudharyi. (A, B) Distribution map showing type locality in Nepal (A) and Humla district (B). (C, D) Type locality habitat at Upper Chungsa valley, Humla district (C–D by S.K. Ghimire)." figureDoi="http://doi.org/10.5281/zenodo.13719195" httpUri="https://zenodo.org/record/13719195/files/figure.png" pageId="3" pageNumber="274">Fig. 1A, 1B, 1C, 1D</figureCitation>
). It grows on alpine rocky and scree habitat with exposed slope (
<figureCitation id="134F2A0BFFE26A64FDC8DF5FA2D3FE60" box="[514,650,268,293]" captionStart="FIGURE 1" captionStartId="1.[136,229,1311,1332]" captionTargetBox="[174,1433,377,1270]" captionTargetId="figure-65@1.[151,1437,374,1283]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Saussurea ramchaudharyi. (A, B) Distribution map showing type locality in Nepal (A) and Humla district (B). (C, D) Type locality habitat at Upper Chungsa valley, Humla district (C–D by S.K. Ghimire)." figureDoi="http://doi.org/10.5281/zenodo.13719195" httpUri="https://zenodo.org/record/13719195/files/figure.png" pageId="3" pageNumber="274">Fig. 1C, 1D</figureCitation>
; 
<figureCitation id="134F2A0BFFE26A64FD5CDF5FA344FE60" box="[662,797,268,293]" captionStart="FIGURE 2" captionStartId="2.[136,229,1112,1133]" captionTargetBox="[453,1133,196,1080]" captionTargetId="figure-13@2.[448,1139,190,1086]" captionTargetPageId="2" captionText="FIGURE 2. Saussurea ramchaudharyi. (A, B) Reproductive plant. (C) Vegetative plant (A–C by S.K. Ghimire). (D) Synflorescence exposed at anthesis (by H.K. Rana)." figureDoi="http://doi.org/10.5281/zenodo.13719197" httpUri="https://zenodo.org/record/13719197/files/figure.png" pageId="3" pageNumber="274">Fig. 2A, 2B</figureCitation>
).
</paragraph>
<paragraph id="8BCB368EFFE26A64FF77DF63A0A4FED4" blockId="3.[136,1452,160,653]" pageId="3" pageNumber="274">
<emphasis id="B900EA9CFFE26A64FF77DF63A16CFE0C" box="[189,309,304,329]" italics="true" pageId="3" pageNumber="274">Etymology</emphasis>
:—The specific epithet is given in honor of Dr. Ram Prasad Chaudhary, a renowned Nepali plant taxonomist and professor of botany, who has made great contributions to the conservation of biodiversity in the 
<collectingCountry id="F363761EFFE26A64FAA3DF07A5F5FE28" box="[1385,1452,340,365]" name="Nepal" pageId="3" pageNumber="274">Nepal</collectingCountry>
Himalaya.
</paragraph>
<paragraph id="8BCB368EFFE26A64FF77DFCFA532FDC8" blockId="3.[136,1452,160,653]" pageId="3" pageNumber="274">
<emphasis id="B900EA9CFFE26A64FF77DFCFA1C2FEF0" box="[189,411,412,437]" italics="true" pageId="3" pageNumber="274">Conservation status</emphasis>
:— 
<taxonomicName id="4C744D0DFFE26A64FE75DFCFA2BDFEF0" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[447,740,412,437]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE26A64FE75DFCFA2BDFEF0" box="[447,740,412,437]" italics="true" pageId="3" pageNumber="274">Saussurea ramchaudharyi</emphasis>
</taxonomicName>
is a very locally endemic species, restricted to a single mountain (
<figureCitation id="134F2A0BFFE26A64FF5ADF93A140FE9C" box="[144,281,448,473]" captionStart="FIGURE 1" captionStartId="1.[136,229,1311,1332]" captionTargetBox="[174,1433,377,1270]" captionTargetId="figure-65@1.[151,1437,374,1283]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Saussurea ramchaudharyi. (A, B) Distribution map showing type locality in Nepal (A) and Humla district (B). (C, D) Type locality habitat at Upper Chungsa valley, Humla district (C–D by S.K. Ghimire)." figureDoi="http://doi.org/10.5281/zenodo.13719195" httpUri="https://zenodo.org/record/13719195/files/figure.png" pageId="3" pageNumber="274">Fig. 1B, 1D</figureCitation>
). We recorded less than 30 mature individuals of 
<taxonomicName id="4C744D0DFFE26A64FC8FDF93A457FE9C" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[837,1038,448,473]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE26A64FC8FDF93A457FE9C" box="[837,1038,448,473]" italics="true" pageId="3" pageNumber="274">S. ramchaudharyi</emphasis>
</taxonomicName>
in the 
<typeStatus id="54CF882CFFE26A64FB91DF93A4D2FE9C" box="[1115,1163,448,473]" pageId="3" pageNumber="274">type</typeStatus>
locality within an area of 
<emphasis id="B900EA9CFFE26A64FF42DFB7A0FBFEB8" box="[136,162,484,509]" italics="true" pageId="3" pageNumber="274">ca</emphasis>
. 
<quantity id="4C8C9B6BFFE26A64FF7BDFB7A0A1FEB8" box="[177,248,484,509]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="274" unit="m" value="500.0">500 m</quantity>
<superScript id="7C019BC6FFE26A64FF32DFB1A159FEB5" attach="left" box="[248,256,482,496]" fontSize="6" pageId="3" pageNumber="274">2</superScript>
. The main threat to the habitat and the species is related to high anthropogenic pressure, such as livestock grazing, and harvesting of caterpillar fungus and other medicinal and aromatic plants. Owing to its highly restricted distribution in a fragile and isolated alpine habitat, its small population size as well as the observed and projected decline in habitat quality due to anthropogenic pressure, 
<taxonomicName id="4C744D0DFFE26A64FCDDDC03A3BBFD2C" authorityName="S. K. Ghimire &amp; H. K. Rana" authorityYear="2018" box="[791,994,592,617]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="ramchaudharyi">
<emphasis id="B900EA9CFFE26A64FCDDDC03A3BBFD2C" box="[791,994,592,617]" italics="true" pageId="3" pageNumber="274">S. ramchaudharyi</emphasis>
</taxonomicName>
is categorized as Critically Endangered [CR; B1ab (iii), B2ab (iii) and D] based on the IUCN Red List categories and criteria version 3.1 (
<bibRefCitation id="EFE54B7FFFE26A64FB1FDC27A506FDC8" author="IUCN &amp; Species Survival Commission, IUCN &amp; Cambridge, UK" box="[1237,1375,628,653]" pageId="3" pageNumber="274" refId="ref3199" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. Second edition. IUCN Species Survival Commission, IUCN, Gland, Switzerland and Cambridge, UK, 38 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>