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<document id="5D1263448AB4E88EE77652CE1419E0F7" ID-DOI="10.11646/palaeoentomology.4.5.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="5530149" ID-ZooBank="50F1EB3E-450C-4B2C-A4F8-11E113C0FD02" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632739847826" checkinUser="plazi" docAuthor="BASHKUEV, ALEXEY S. &amp; SUKATSHEVA, IRINA D." docDate="2021" docId="03D8B42BF231FF9AFCC7FB73C9F6F979" docLanguage="en" docName="palaeoentomology.4.5.11.pdf" docOrigin="Palaeoentomology 4 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.5.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Kamopanorpa sivchikovi BASHKUEV &amp; SUKATSHEVA 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="457" masterDocId="FFE1CC53F232FF9EFFEDFFF8C913FFD7" masterDocTitle="New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)" masterLastPageNumber="461" masterPageNumber="453" pageNumber="456" updateTime="1699415050900" updateUser="plazi">
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<mods:title id="E41A82FC81C2F0E56AF06B434DAE5E08">New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)</mods:title>
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<mods:name id="4D84F2095C852BC8A5A7D34520B5630F" type="personal">
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<mods:namePart id="D606B818CF262770370C9A52A69B6C0A">BASHKUEV, ALEXEY S.</mods:namePart>
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<mods:namePart id="C709F2BF93B5F63E8433EE0C163C2BE0">SUKATSHEVA, IRINA D.</mods:namePart>
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<mods:title id="AF5E15B176FB75DFC28200746FE9FC81">Palaeoentomology</mods:title>
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<mods:part id="37A622B4BD0B700060B32BFFDA040DE9">
<mods:date id="C51F4D246449D784BDF0A6089652121E">2021</mods:date>
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<mods:number id="A5A4E30EFE649E12A851585D43E4E266">2021-09-24</mods:number>
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<mods:identifier id="E4072E2C4CBD59DA967206185022B653" type="ISSN">2624-2834</mods:identifier>
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<treatment id="03D8B42BF231FF9AFCC7FB73C9F6F979" ID-DOI="http://doi.org/10.5281/zenodo.5530173" ID-Zenodo-Dep="5530173" LSID="urn:lsid:plazi:treatment:03D8B42BF231FF9AFCC7FB73C9F6F979" httpUri="http://treatment.plazi.org/id/03D8B42BF231FF9AFCC7FB73C9F6F979" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456">
<subSubSection id="C36B56B6F231FF9DFCC7FB73CDBAFB72" box="[810,1193,1163,1190]" pageId="3" pageNumber="456" type="nomenclature">
<paragraph id="8BCE053DF231FF9DFCC7FB73CDBAFB72" blockId="3.[810,1193,1163,1225]" box="[810,1193,1163,1190]" pageId="3" pageNumber="456">
<heading id="D086B251F231FF9DFCC7FB73CDBAFB72" bold="true" box="[810,1193,1163,1190]" fontSize="10" level="2" pageId="3" pageNumber="456" reason="3">
<emphasis id="B905D92FF231FF9DFCC7FB73CDBAFB72" bold="true" box="[810,1193,1163,1190]" pageId="3" pageNumber="456">
<taxonomicName id="4C717EBEF231FF9DFCC7FB73CD5AFB71" authority="BASHKUEV &amp; SUKATSHEVA, 2021" authorityName="BASHKUEV &amp; SUKATSHEVA" authorityYear="2021" box="[810,1097,1163,1190]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="sivchikovi" status="sp. nov.">
<emphasis id="B905D92FF231FF9DFCC7FB73CD5AFB71" bold="true" box="[810,1097,1163,1190]" italics="true" pageId="3" pageNumber="456">Kamopanorpa sivchikovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2366454F231FF9DFBBDFB73CDBAFB72" box="[1104,1193,1163,1189]" pageId="3" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B56B6F231FF9DFCC7FB57CAD7FB1E" box="[810,964,1199,1225]" pageId="3" pageNumber="456" type="description">
<paragraph id="8BCE053DF231FF9DFCC7FB57CAD7FB1E" blockId="3.[810,1193,1163,1225]" box="[810,964,1199,1225]" pageId="3" pageNumber="456">
(
<figureCitation id="134A19B8F231FF9DFCDFFB57CA9DFB1E" box="[818,910,1199,1225]" captionStart="FIGURE 1" captionStartId="3.[136,229,1027,1049]" captionTargetBox="[222,1353,246,987]" captionTargetId="figure-428@3.[215,1371,223,1003]" captionTargetPageId="3" captionText="FIGURE 1. A, B, Kamopanorpa rasnitsyni sp. nov. A, Holotype PIN 5386/9. B, Paratype PIN 5386/3. C, K. rotunda sp. nov., holotype PIN 5386/5. D, K. sivchikovi sp. nov., holotype PIN 5386/1. E, F, Kamopanorpa spp. E, 5386/8. F, 5386/11.All to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5530151" httpUri="https://zenodo.org/record/5530151/files/figure.png" pageId="3" pageNumber="456">Figs 1D</figureCitation>
,
<figureCitation id="134A19B8F231FF9DFC77FB57CAAFFB1E" box="[922,956,1199,1225]" captionStart="FIGURE 2" captionStartId="4.[136,229,1083,1105]" captionTargetBox="[137,776,190,1059]" captionTargetId="figure-479@4.[137,776,190,1059]" captionTargetPageId="4" captionText="FIGURE 2. A, Kamopanorpa rasnitsyni sp. nov. Holotype PIN 5386/9. B, K. sivchikovi sp. nov., holotype PIN 5386/1. C, K. rotunda sp. nov., holotype PIN 5386/5. SEM images." figureDoi="http://doi.org/10.5281/zenodo.5530153" httpUri="https://zenodo.org/record/5530153/files/figure.png" pageId="3" pageNumber="456">2B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36B56B6F231FF9DFCC7FB0FCC49FAC6" box="[810,1370,1271,1298]" pageId="3" pageNumber="456" type="materials_examined">
<paragraph id="8BCE053DF231FF9DFCC7FB0FCC49FAC6" blockId="3.[810,1452,1271,2060]" box="[810,1370,1271,1298]" pageId="3" pageNumber="456">
<emphasis id="B905D92FF231FF9DFCC7FB0FCACBFAC6" bold="true" box="[810,984,1271,1297]" pageId="3" pageNumber="456">Type material.</emphasis>
<materialsCitation id="3B190F60F231FF9DFC33FB0FCC49FAC6" box="[990,1370,1271,1298]" collectionCode="PIN" pageId="3" pageNumber="456" specimenCode="PIN 5386" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CABB9FF231FF9DFC33FB0FCD55FAC6" box="[990,1094,1271,1297]" pageId="3" pageNumber="456" type="holotype">Holotype</typeStatus>
<specimenCode id="DBD7AD46F231FF9DFBA3FB00CDACFAC5" box="[1102,1215,1271,1298]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="3" pageNumber="456">PIN 5386</specimenCode>
/1±, forewing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B56B6F231FF9DFCB2FAE3CDFEFA76" pageId="3" pageNumber="456" type="etymology">
<paragraph id="8BCE053DF231FF9DFCB2FAE3CDFEFA76" blockId="3.[810,1452,1271,2060]" pageId="3" pageNumber="456">
<emphasis id="B905D92FF231FF9DFCB2FAE3CAF5FAE2" bold="true" box="[863,998,1307,1333]" pageId="3" pageNumber="456">Etymology.</emphasis>
Named in honour of late geologist and palaeobotanist Victor E. Sivchikov, who made a great contribution to the upper Paleozoic stratigraphy of Siberia and particularly of the Minusinsk Basin.
</paragraph>
</subSubSection>
<subSubSection id="C36B56B6F231FF9DFCB2FA53CCB7F9AE" pageId="3" pageNumber="456" type="diagnosis">
<paragraph id="8BCE053DF231FF9DFCB2FA53CC4DF9DA" blockId="3.[810,1452,1271,2060]" pageId="3" pageNumber="456">
<emphasis id="B905D92FF231FF9DFCB2FA53CACAFA12" bold="true" box="[863,985,1451,1477]" pageId="3" pageNumber="456">Diagnosis.</emphasis>
SC without apical fork, ending at C at level of RS
<subScript id="17F50778F231FF9DFC36FA1ACAF0FA27" attach="left" box="[987,995,1506,1520]" fontSize="6" pageId="3" pageNumber="456">1</subScript>
fork, with three subcostal crossveins; subcostal space narrow with no visible crossveins.
</paragraph>
<paragraph id="8BCE053DF231FF9DFCB2F9EFCCB7F9AE" blockId="3.[810,1452,1271,2060]" pageId="3" pageNumber="456">
<emphasis id="B905D92FF231FF9DFCB2F9EFCD7FF9E6" bold="true" box="[863,1132,1559,1585]" pageId="3" pageNumber="456">Locality and horizon.</emphasis>
Izykh Coal Mines, quarry no.1, south of Belyi Yar,
<collectingRegion id="49B5CBDFF231FF9DFB85F9C3CDCDF982" box="[1128,1246,1595,1621]" country="Russia" name="Khakasiya" pageId="3" pageNumber="456">Khakassia</collectingRegion>
, Minusinsk Coal Basin, South Siberia; Izykh Formation, Middle Permian.
</paragraph>
</subSubSection>
<subSubSection id="C36B56B6F231FF9AFCB2F97BC9F6F979" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456" type="description">
<paragraph id="8BCE053DF231FF9AFCB2F97BC85CF9B3" blockId="3.[810,1452,1271,2060]" lastBlockId="4.[136,778,1239,1710]" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456">
<emphasis id="B905D92FF231FF9DFCB2F97BCAE2F94A" bold="true" box="[863,1009,1667,1693]" pageId="3" pageNumber="456">Description.</emphasis>
Wing ovate (L:W ratio 1:2.5), slightly broadened at 2/3 of wing length, maximal width at level of M
<subScript id="17F50778F231FF9DFC4EF926CAB8F93B" attach="left" box="[931,939,1758,1772]" fontSize="6" pageId="3" pageNumber="456">4</subScript>
apex. Anterior margin slightly convex, apex at RS
<subScript id="17F50778F231FF9DFC83F8FACA65F8C7" attach="right" box="[878,886,1794,1808]" fontSize="6" pageId="3" pageNumber="456">3</subScript>
RS
<subScript id="17F50778F231FF9DFC4BF8FACABDF8C7" attach="left" box="[934,942,1794,1808]" fontSize="6" pageId="3" pageNumber="456">4</subScript>
. SC reaching deep into pterostigmal area, meeting wing margin without apical fork; with three short subcostal crossveins. Subcostal space half as wide as costal space, with no visible crossveins except for pterostigmal one. Pterostigma not preserved. R nearly straight, only slightly curved apically, without pterostigmal branches. RS forking well before middle of wing, with six branches reaching wing margin; fork on RS
<subScript id="17F50778F231FF9DFB56F806CDD0F7DB" attach="left" box="[1211,1219,2046,2060]" fontSize="6" pageId="3" pageNumber="456">2</subScript>
very short. Cell DC closed by crossvein rs
<subScript id="17F50778F236FF9AFE6FFB12C899FB2F" attach="left" box="[386,394,1258,1272]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
rs
<subScript id="17F50778F236FF9AFE40FB12C8A6FB2F" attach="left" box="[429,437,1258,1272]" fontSize="6" pageId="4" pageNumber="457">3</subScript>
located nearly at level of RS
<subScript id="17F50778F236FF9AFCEFFB12CA19FB2F" attach="right" box="[770,778,1258,1272]" fontSize="6" pageId="4" pageNumber="457">1</subScript>
fork. Cell MC closed. M five-branched. Base of CuA long, strongly oblique, M
<subScript id="17F50778F236FF9AFE2DFACBC8DBFA96" attach="left" box="[448,456,1331,1345]" fontSize="6" pageId="4" pageNumber="457">5</subScript>
much shorter. Anal veins forming double-Y loop typical for the genus, free distal portion of A
<subScript id="17F50778F236FF9AFEFDFA86C80BFA5B" attach="none" box="[272,280,1406,1420]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
distinct. Crossveins in cubito-anal area quite distinct, but in distal wing half mostly faint and can be traced only by subtle breaks at longitudinal veins. There are rrs
<subScript id="17F50778F236FF9AFF31FA15C9F7FA2C" attach="both" box="[220,228,1517,1531]" fontSize="6" pageId="4" pageNumber="457">1</subScript>
a, rs
<subScript id="17F50778F236FF9AFEFEFA15C808FA2C" attach="left" box="[275,283,1517,1531]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
rs
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; two crossveins in RSM space; m
<subScript id="17F50778F236FF9AFD23FA15CBC5FA2C" attach="left" box="[718,726,1517,1531]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
m
<subScript id="17F50778F236FF9AFD17FA15CA11FA2C" attach="left" box="[762,770,1517,1531]" fontSize="6" pageId="4" pageNumber="457">3</subScript>
; long m
<subScript id="17F50778F236FF9AFF3AF9EAC9E3F9F7" attach="left" box="[215,240,1554,1568]" fontSize="6" pageId="4" pageNumber="457">3+4</subScript>
cua and possibly additional crossvein m
<subScript id="17F50778F236FF9AFD50F9EACBD6F9F7" attach="left" box="[701,709,1554,1568]" fontSize="6" pageId="4" pageNumber="457">4</subScript>
cua
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; strong transverse cuacup located distal to apex of A
<subScript id="17F50778F236FF9AFD17F9CFCA11F992" attach="left" box="[762,770,1591,1605]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
; cupa
<subScript id="17F50778F236FF9AFF26F9A4C9C0F9BD" attach="left" box="[203,211,1628,1642]" fontSize="6" pageId="4" pageNumber="457">1</subScript>
, and a
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a
<subScript id="17F50778F236FF9AFEADF9A4C85BF9BD" attach="left" box="[320,328,1628,1642]" fontSize="6" pageId="4" pageNumber="457">2</subScript>
.
</paragraph>
<caption id="DF0E55B5F236FF9AFF65FBC3CB99FB4D" ID-DOI="http://doi.org/10.5281/zenodo.5530153" ID-Zenodo-Dep="5530153" httpUri="https://zenodo.org/record/5530153/files/figure.png" pageId="4" pageNumber="457" startId="4.[136,229,1083,1105]" targetBox="[137,776,190,1059]" targetPageId="4">
<paragraph id="8BCE053DF236FF9AFF65FBC3CB99FB4D" blockId="4.[136,778,1083,1178]" pageId="4" pageNumber="457">
<emphasis id="B905D92FF236FF9AFF65FBC3C808FB87" bold="true" box="[136,283,1083,1105]" pageId="4" pageNumber="457">FIGURE 2. A</emphasis>
,
<taxonomicName id="4C717EBEF236FF9AFEC6FBC4CB0DFB86" authority="BASHKUEV &amp; SUKATSHEVA, 2021" authorityName="BASHKUEV &amp; SUKATSHEVA" authorityYear="2021" box="[299,542,1084,1105]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
<emphasis id="B905D92FF236FF9AFEC6FBC4CB0DFB86" box="[299,542,1084,1105]" italics="true" pageId="4" pageNumber="457">Kamopanorpa rasnitsyni</emphasis>
</taxonomicName>
<emphasis id="B905D92FF236FF9AFDCAFBC4CB64FB86" bold="true" box="[551,631,1084,1105]" pageId="4" pageNumber="457">
<taxonomicNameLabel id="A2366454F236FF9AFDCAFBC4CB64FB86" box="[551,631,1084,1105]" pageId="4" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype PIN 5386/9.
<emphasis id="B905D92FF236FF9AFF31FB98C9FFFBA2" bold="true" box="[220,236,1120,1141]" pageId="4" pageNumber="457">B</emphasis>
,
<taxonomicName id="4C717EBEF236FF9AFF10FB98C86DFBA2" authority="BASHKUEV &amp; SUKATSHEVA, 2021" authorityName="BASHKUEV &amp; SUKATSHEVA" authorityYear="2021" box="[253,382,1120,1141]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="sivchikovi" status="sp. nov.">
<emphasis id="B905D92FF236FF9AFF10FB98C81EFBA2" box="[253,269,1120,1141]" italics="true" pageId="4" pageNumber="457">K</emphasis>
.
<emphasis id="B905D92FF236FF9AFEF3FB98C86DFBA2" box="[286,382,1120,1141]" italics="true" pageId="4" pageNumber="457">sivchikovi</emphasis>
</taxonomicName>
<emphasis id="B905D92FF236FF9AFE64FB98C8C8FBA2" bold="true" box="[393,475,1120,1141]" pageId="4" pageNumber="457">
<taxonomicNameLabel id="A2366454F236FF9AFE64FB98C8C8FBA2" box="[393,475,1120,1141]" pageId="4" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype PIN 5386/1.
<emphasis id="B905D92FF236FF9AFD3CFBA7CBF1FBA2" bold="true" box="[721,738,1119,1141]" pageId="4" pageNumber="457">C</emphasis>
,
<taxonomicName id="4C717EBEF236FF9AFD19FB98C9C0FB4E" authority="BASHKUEV &amp; SUKATSHEVA, 2021" authorityName="BASHKUEV &amp; SUKATSHEVA" authorityYear="2021" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="rotunda" status="sp. nov.">
<emphasis id="B905D92FF236FF9AFD19FB98CA17FBA2" box="[756,772,1120,1141]" italics="true" pageId="4" pageNumber="457">K</emphasis>
.
<emphasis id="B905D92FF236FF9AFF65FB7CC9C0FB4E" box="[136,211,1156,1177]" italics="true" pageId="4" pageNumber="457">rotunda</emphasis>
</taxonomicName>
<emphasis id="B905D92FF236FF9AFF34FB7CC835FB4E" bold="true" box="[217,294,1156,1177]" pageId="4" pageNumber="457">
<taxonomicNameLabel id="A2366454F236FF9AFF34FB7CC835FB4E" box="[217,294,1156,1177]" pageId="4" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype PIN 5386/5. SEM images.
</paragraph>
</caption>
<paragraph id="8BCE053DF236FF9AFF50F997C9F6F979" blockId="4.[136,778,1239,1710]" pageId="4" pageNumber="457">
<emphasis id="B905D92FF236FF9AFF50F997C867F95E" bold="true" box="[189,372,1647,1673]" pageId="4" pageNumber="457">Measurements.</emphasis>
Forewing length,
<quantity id="4C89A8D8F236FF9AFDA6F996CBA7F95E" box="[587,692,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" pageId="4" pageNumber="457" unit="mm" value="4.35">4.35 mm</quantity>
, width,
<quantity id="4C89A8D8F236FF9AFF65F96CC9F2F979" box="[136,225,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="457" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>