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<document id="9CB31B51BBCFB29BB27955F451AF7C7C" ID-DOI="10.11646/zootaxa.3539.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10096759" ID-ZooBank="0564EF02-FCF9-49A8-BDAC-F432876C16B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629821597637" checkinUser="felipe" docAuthor="Ramos, Magno S. &amp; Azevedo, Celso O." docDate="2012" docId="03D887D08A1CFFBD1399FAEEFB43FAB8" docLanguage="en" docName="zt03539p080.pdf" docOrigin="Zootaxa 3539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3539.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eupsenella karanja Ramos &amp; Azevedo, sp. nov." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="FFE1FFA88A33FF8D130EFFB1FFFCFFEC" masterDocTitle="Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="48" updateTime="1699581041301" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="43DD6DF065E9B84B6F6F1B40E70DBBC9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="30FFB88625F05AF8B3DA7531A11535F6">Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)</mods:title>
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<mods:name id="48A5413A0F822ACA57A1E69E56E0DE95" type="personal">
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<mods:namePart id="8DE965E5B04128A91056D13A11FB4487">Ramos, Magno S.</mods:namePart>
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<mods:namePart id="7DA6F15F813881615CE6284BDC5E72A2">Azevedo, Celso O.</mods:namePart>
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<mods:title id="BC8C9FFD822CCCA097C303EEFE5BBAF4">Zootaxa</mods:title>
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<mods:date id="4C08C66BF5671F08C2F4536304DA5084">2012</mods:date>
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<mods:number id="92285784366E1DCE3FB0E7ECE549B038">2012-11-05</mods:number>
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<mods:number id="8F5E550E0E7ADED04FC38EB29D97A543">3539</mods:number>
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<mods:classification id="898144650DF42BFF49E00FFDCE30FEB1">journal article</mods:classification>
<mods:identifier id="00608BBD7A26EA41E65647B4EF37B2E6" type="DOI">10.11646/zootaxa.3539.1.1</mods:identifier>
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<treatment id="03D887D08A1CFFBD1399FAEEFB43FAB8" ID-DOI="http://doi.org/10.5281/zenodo.5461877" ID-Zenodo-Dep="5461877" LSID="urn:lsid:plazi:treatment:03D887D08A1CFFBD1399FAEEFB43FAB8" httpUri="http://treatment.plazi.org/id/03D887D08A1CFFBD1399FAEEFB43FAB8" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<subSubSection id="C36B654D8A1CFFA21399FAEEFD22FA95" box="[151,734,1375,1401]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BCE36C68A1CFFA21399FAEEFD22FA95" blockId="47.[151,734,1375,1434]" box="[151,734,1375,1401]" pageId="47" pageNumber="48">
<heading id="D08681AA8A1CFFA21399FAEEFD22FA95" bold="true" box="[151,734,1375,1401]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="B905EAD48A1CFFA21399FAEEFD22FA95" bold="true" box="[151,734,1375,1401]" pageId="47" pageNumber="48">
<taxonomicName id="4C714D458A1CFFA21399FAEEFD8FFA95" authority="Ramos &amp; Azevedo" authorityName="Ramos &amp; Azevedo" box="[151,627,1375,1401]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="karanja" status="sp. nov.">
<emphasis id="B905EAD48A1CFFA21399FAEEFE7FFA95" bold="true" box="[151,387,1375,1401]" italics="true" pageId="47" pageNumber="48">Eupsenella karanja</emphasis>
Ramos &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A23657AF8A1CFFA2118FFAEEFD22FA95" box="[641,734,1375,1401]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A1CFFBD1399FA33FE66FBE3" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="8BCE36C68A1CFFA21399FA33FEA2FA76" blockId="47.[151,734,1375,1434]" box="[151,350,1410,1434]" pageId="47" pageNumber="48">
(
<figureCitation id="134A2A438A1CFFA21391FA33FF09FA76" box="[159,245,1410,1434]" captionStart="FIGURES 27–38" captionStartId="64.[151,264,1847,1870]" captionTargetBox="[151,1436,193,1826]" captionTargetId="figure-88@64.[151,1436,193,1826]" captionTargetPageId="64" captionText="FIGURES 27–38. Head in dorsal view. 27. Eupsenella ingura sp. nov. 28. E. iwaidja sp. nov. 29. E. jaadwa sp. nov. 30. E. jaara sp. nov. 31. E. jaburara sp. nov. 32. E. jadira sp. nov. 33. E. jagara sp. nov. 34. E. janda sp. nov. 35. E. kabalbara sp. nov. 36. E. kaibara sp. nov. 37. E. karanja sp. nov. 38. E. karawa sp. nov. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096767" httpUri="https://zenodo.org/record/10096767/files/figure.png" pageId="47" pageNumber="48">Figs 37</figureCitation>
, 82, 127)
</paragraph>
<paragraph id="8BCE36C68A1CFFA21399FA7BFC24FA0E" blockId="47.[151,1437,1482,1939]" box="[151,984,1482,1507]" pageId="47" pageNumber="48">
<emphasis id="B905EAD48A1CFFA21399FA7BFED4FA0F" bold="true" box="[151,296,1482,1507]" inLineHeading="true" pageId="47" pageNumber="48" reason="1">Description.</emphasis>
FEMALE (
<typeStatus id="54CA88648A1CFFA212A0FA7BFDEBFA0E" box="[430,535,1482,1506]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
). Body length
<quantity id="4C899B238A1CFFA211B7FA7BFCDCFA0E" box="[697,800,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="7.07" pageId="47" pageNumber="48" unit="mm" value="7.07">7.07 mm</quantity>
. LFW
<quantity id="4C899B238A1CFFA21063FA7BFC28FA0E" box="[877,980,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.58" pageId="47" pageNumber="48" unit="mm" value="3.58">3.58 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36C68A1CFFA213C9FA5FFB65F99E" blockId="47.[151,1437,1482,1939]" pageId="47" pageNumber="48">
<emphasis id="B905EAD48A1CFFA213C9FA5FFEB5F9EA" box="[199,329,1518,1542]" italics="true" pageId="47" pageNumber="48">Coloration.</emphasis>
Head dark castaneous almost black; scape, pedicel, flagellum, mandible and palpi light castaneous; mesosoma dark castaneous; tegula light castaneous; wing venation dark castaneous; procoxa dark castaneous almost black; mesocoxa, metacoxa, trochanters castaneous; profemur and mesofemur castaneous; metafemur dark castaneous; tarsi light castaneous; metasoma dark castaneous almost black.
</paragraph>
<paragraph id="8BCE36C68A1CFFBD13C9F9CFFDB2FF1B" blockId="47.[151,1437,1482,1939]" lastBlockId="48.[151,1436,151,1364]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B905EAD48A1CFFA213C9F9CFFEF8F97A" box="[199,260,1662,1686]" italics="true" pageId="47" pageNumber="48">Head</emphasis>
(
<figureCitation id="134A2A438A1CFFA2121BF9CFFE96F97A" box="[277,362,1662,1686]" captionStart="FIGURES 27–38" captionStartId="64.[151,264,1847,1870]" captionTargetBox="[151,1436,193,1826]" captionTargetId="figure-88@64.[151,1436,193,1826]" captionTargetPageId="64" captionText="FIGURES 27–38. Head in dorsal view. 27. Eupsenella ingura sp. nov. 28. E. iwaidja sp. nov. 29. E. jaadwa sp. nov. 30. E. jaara sp. nov. 31. E. jaburara sp. nov. 32. E. jadira sp. nov. 33. E. jagara sp. nov. 34. E. janda sp. nov. 35. E. kabalbara sp. nov. 36. E. kaibara sp. nov. 37. E. karanja sp. nov. 38. E. karawa sp. nov. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096767" httpUri="https://zenodo.org/record/10096767/files/figure.png" pageId="47" pageNumber="48">Fig. 37</figureCitation>
). Head wider than long, subrectangular in profile; gena weakly concave in profile. Mandible evenly wide, two lowermost teeth larger than upper ones, lower tooth curved inward, lower margin strongly setose. Clypeus coriaceous, median clypeal lobe subangulate, median concavity of median clypeal lobe absent, 0.02 LH, well marked off from frons, median carina present, 0.1 LH. Malar space conspicuous, 0.33 mandibular base. Antenna
<quantity id="4C899B238A1CFFA2120CF8BFFE92F8CA" box="[258,366,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="47" pageNumber="48" unit="mm" value="1.58">1.58 mm</quantity>
; first five antennomeres in ratio of about 10:9:9:9:8; flagellomere 1 1.0 longer than wide, flagellar pubescence dense, with many outstanding erect setae. Region between torulus and eye concave. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.14 LH. Eye protuberant, 0.57 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons coriaceous-punctate. WH DAO, anterior ocellus not surpassing imaginary top eye line, posterior ocellus reaching vertex. Vertex concave, long setae, 0.22 LH, temple profile slightly convergent in dorsal view, with corner convex, not projected posterolaterad. Hipostomal carina arched, low.
</paragraph>
<paragraph id="8BCE36C68A03FFBD13C9FEB4FE9BFD20" blockId="48.[151,1436,151,1364]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FEB4FEC3FEF0" box="[199,319,261,284]" italics="true" pageId="48" pageNumber="49">Mesosoma</emphasis>
(Fig. 82). Pronotal disc coriaceous-punctate, 0.42 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow inconspicuous. Notauli convergent posterad. Mesoscutum 1.4 length pronotal disc. Scutellum coriaceous-punctate, 1.1 length pronotal disc, almost flat in profile, posterior margin convex medially, scutellar pit elliptical, narrowed, distant each other 11.0 its own diameter. Dorsal axillar surface with fovea oval. Metanotum with median region high, coriaceous-puncticulate, as high as scutellum in profile, with fovea oval laterally. Propodeal disc wider than long, anterior margin of propodeal disc with a transverse furrow, dilated ends; median carina 0.49 length propodeal disc, incomplete posterad, located within a median furrow; discal carina 0.15 length propodeal disc, converging posterad, slightly arched; space between median and discal carina confused-rugulose. Coriaceous. Lateral of propodeum predominantly coriaceous. Mesopleuron with subtegular furrow elliptical; mesepimeral suture almost inconspicuous posterad; mesopleural epicoxal lobe inconspicuous. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression composed by two large foveae almost fused; median suture not evident.
</paragraph>
<paragraph id="8BCE36C68A03FFBD13C9FD69FAD2FC93" blockId="48.[151,1436,151,1364]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FD69FEEFFD03" box="[199,275,728,751]" italics="true" pageId="48" pageNumber="49">Wings.</emphasis>
Macropterous. Forewing (
<figureCitation id="134A2A438A03FFBD1133FD66FD61FD03" box="[573,669,727,751]" captionStart="FIGURES 123–132" captionStartId="74.[151,263,1911,1934]" captionTargetBox="[151,1436,193,1890]" captionTargetId="figure-77@74.[151,1436,193,1890]" captionTargetPageId="74" captionText="FIGURES 123–132. Right forewing. 123. Eupsenella jagara sp. nov. 124. E. janda sp. nov. 125. E. kabalbara sp. nov. 126. E." figureDoi="http://doi.org/10.5281/zenodo.10096781" httpUri="https://zenodo.org/record/10096781/files/figure.png" pageId="48" pageNumber="49">Fig. 127</figureCitation>
) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell widening distally;1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs
<emphasis id="B905EAD48A03FFBD1780FD4DFB60FCF8" box="[1166,1180,764,788]" italics="true" pageId="48" pageNumber="49">b</emphasis>
convex; 2R1 cell 0.22 LFW; distal margin of Rs
<emphasis id="B905EAD48A03FFBD12B0FC90FE36FCD4" box="[446,458,801,824]" italics="true" pageId="48" pageNumber="49">c</emphasis>
convex; r-rs vein inclined; R1
<emphasis id="B905EAD48A03FFBD1024FC91FCC4FCD4" box="[810,824,800,824]" italics="true" pageId="48" pageNumber="49">b</emphasis>
entirely on anterior margin of forewing; Cu
<emphasis id="B905EAD48A03FFBD163BFC91FABFFCD4" box="[1333,1347,800,824]" italics="true" pageId="48" pageNumber="49">b</emphasis>
absent; Rs+M vein 0.91 Rs
<emphasis id="B905EAD48A03FFBD1278FCF4FE78FCB0" box="[374,388,837,860]" italics="true" pageId="48" pageNumber="49">a</emphasis>
; M vein 1.5 Rs
<emphasis id="B905EAD48A03FFBD1137FCF4FDBBFCB0" box="[569,583,837,860]" italics="true" pageId="48" pageNumber="49">a</emphasis>
; Rs
<emphasis id="B905EAD48A03FFBD1178FCF4FD78FCB0" box="[630,644,837,860]" italics="true" pageId="48" pageNumber="49">a</emphasis>
subperpendicular to anterior margin of forewing; stigma rectangular; distal stigmal margin truncate. Hind wing with four distal hamuli, last hamulus discontinuous each other.
</paragraph>
<paragraph id="8BCE36C68A03FFBD13C9FC3CFC87FC2B" blockId="48.[151,1436,151,1364]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FC3CFEFFFC48" box="[199,259,909,932]" italics="true" pageId="48" pageNumber="49">Legs.</emphasis>
Profemur very swollen, 0.63 as wide as long. Metacoxa with median region 0.59 as wide as long. Metacoxal projection in dorsal region conspicuous, apex rounded.
</paragraph>
<paragraph id="8BCE36C68A03FFBD13C9FC64FC6BFC07" blockId="48.[151,1436,151,1364]" box="[199,919,980,1004]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FC64FEBFFC00" box="[199,323,981,1004]" italics="true" pageId="48" pageNumber="49">Metasoma.</emphasis>
0.2 as wide as long, ventral carina of petiole present.
</paragraph>
<paragraph id="8BCE36C68A03FFBD13C9FC46FE66FBE3" blockId="48.[151,1436,151,1364]" box="[199,410,1015,1039]" pageId="48" pageNumber="49">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A03FFBD13C9FBAAFBC8FBBB" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8BCE36C68A03FFBD13C9FBAAFBC8FBBB" blockId="48.[151,1436,151,1364]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FBAAFE4DFBD8" bold="true" box="[199,433,1051,1076]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">Material examined.</emphasis>
<materialsCitation id="3B193C9B8A03FFBD12B7FBADFBC8FBBB" collectionCode="ANIC" collectorName="Km N Landsdowne &amp; G. Williams" country="Australia" location="AUSTRALIA" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B905EAD48A03FFBD12B7FBADFDDCFBD8" box="[441,544,1052,1076]" italics="true" pageId="48" pageNumber="49">
<typeStatus id="54CA88648A03FFBD12B7FBADFDDCFBD8" box="[441,544,1052,1076]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D77FD4F8A03FFBD1129FBADFDC0FBDF" box="[551,572,1052,1075]" pageId="48" pageNumber="49" type="female">♀</specimenCount>
.
<collectingCountry id="F36676568A03FFBD1145FBADFD11FBD8" box="[587,749,1052,1076]" name="Australia" pageId="48" pageNumber="49">AUSTRALIA</collectingCountry>
, N[ew] S[outh] W[ales], 3
<collectorName id="268453108A03FFBD172FFBADFAFDFBD8" box="[1057,1281,1052,1076]" pageId="48" pageNumber="49">Km N Landsdowne</collectorName>
, n[ea]r Taree, 20–27 Dec[ember] 1990,
<collectorName id="268453108A03FFBD12B8FB8EFDCBFBBB" box="[438,567,1087,1111]" pageId="48" pageNumber="49">G.Williams</collectorName>
, rainf[orest]/wet sclerophyll, Malaise,
<collectionCode id="ED60AE038A03FFBD10E6FB8EFBCCFBBB" box="[1000,1072,1087,1111]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="48" pageNumber="49">ANIC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A03FFBD13C9FBD2FD4BFB90" box="[199,695,1123,1148]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BCE36C68A03FFBD13C9FBD2FD4BFB90" blockId="48.[151,1436,151,1364]" box="[199,695,1123,1148]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FBD2FE9CFB90" bold="true" box="[199,352,1123,1148]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">Distribution.</emphasis>
<collectingCountry id="F36676568A03FFBD1269FBD5FE32FB90" box="[359,462,1124,1148]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
(
<collectingRegion id="49B5F8248A03FFBD12D3FBD5FD57FB90" box="[477,683,1124,1148]" country="Australia" name="New South Wales" pageId="48" pageNumber="49">New South Wales</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A03FFBD13C9FB39FB43FAB8" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8BCE36C68A03FFBD13C9FB39FB43FAB8" blockId="48.[151,1436,151,1364]" pageId="48" pageNumber="49">
<emphasis id="B905EAD48A03FFBD13C9FB39FEC7FB4C" bold="true" box="[199,315,1160,1184]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">Remarks.</emphasis>
This specie species is similar to
<taxonomicName id="4C714D458A03FFBD11C3FB38FC86FB4C" authorityName="Dodd" authorityYear="1916" box="[717,890,1160,1184]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diemenensis">
<emphasis id="B905EAD48A03FFBD11C3FB38FC86FB4C" box="[717,890,1160,1184]" italics="true" pageId="48" pageNumber="49">E. diemenensis</emphasis>
</taxonomicName>
by having eye large, the median clypeal lobe angulate, the posterior ocelli near vertex crest, the 2R1 cell of forewing slightly elongated. However this species has the median clypeal lobe short, median clypeal lobe with median carina absent, the propodeal disc with median carina short, the anterior margin of propodeal disc with transverse furrow, dilated ends, whereas
<taxonomicName id="4C714D458A03FFBD17C7FB44FA8CFAE0" authorityName="Dodd" authorityYear="1916" box="[1225,1392,1268,1292]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diemenensis">
<emphasis id="B905EAD48A03FFBD17C7FB44FA8CFAE0" box="[1225,1392,1268,1292]" italics="true" pageId="48" pageNumber="49">E. diemenensis</emphasis>
</taxonomicName>
has the median clypeal lobe clearly higher, median clypeal lobe with median carina present, the median carina of propodeal disc long, the anterior margin of propodeal disc with transverse furrow, dilated ends.
</paragraph>
</subSubSection>
</treatment>
</document>
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