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<document id="6D950D24AC61C9A9C2EE1A76B4B6D273" ID-CLB-Dataset="54709" ID-DOI="10.5852/ejt.2023.875.2147" ID-GBIF-Dataset="3351d966-cbeb-42a8-ac49-627b5fc261c7" ID-ISSN="2118-9773" ID-Zenodo-Dep="8083773" ID-ZooBank="32FA008C-B35D-483C-9DBE-1DCCD0868FAC" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687788869828" checkinUser="felipe" docAuthor="Fasbender, Andrew" docDate="2023" docId="03D887BE816E36362A07FDC043460E84" docLanguage="en" docName="ejt-875_159-202.pdf" docOrigin="European Journal of Taxonomy 875" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratoculicoides pacificus Fasbender 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="187" masterDocId="FFE1FFC68175362A284DFF9741030970" masterDocTitle="Revision of the New World Ceratoculicoides Wirth &amp; Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)" masterLastPageNumber="202" masterPageNumber="159" pageNumber="186" updateTime="1687806523842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BF52B60593F51A0D7983AA6E648921DD">Revision of the New World Ceratoculicoides Wirth &amp; Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)</mods:title>
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<mods:namePart id="9624363C12D9DE2D69317E1BDEF179CA">Fasbender, Andrew</mods:namePart>
<mods:affiliation id="953186B49B42B75493730C5D13B10CB5">Rhithron Associates Inc., 33 Fort Missoula Road, Missoula, MT 59847, USA.</mods:affiliation>
<mods:nameIdentifier id="60FD116A7ACD3C014DB41E49DDCE097A" type="email">afasbender@rhithron.com</mods:nameIdentifier>
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<mods:title id="396800993364D4C94CD833D4D2487096">European Journal of Taxonomy</mods:title>
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<mods:date id="27DF56E7F7C7A5BE8B08120027A6343D">2023</mods:date>
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<subSubSection id="C36B6523816E36312A07FDC042EA0B02" box="[586,1001,599,626]" pageId="27" pageNumber="186" type="nomenclature">
<paragraph id="8BCE36A8816E36312A07FDC042EA0B02" blockId="27.[377,1212,599,700]" box="[586,1001,599,626]" pageId="27" pageNumber="186">
<heading id="D08681C4816E36312A07FDC042EA0B02" box="[586,1001,599,626]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="186" reason="2">
<taxonomicName id="4C714D2B816E36312A07FDC0428B0B01" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[586,904,599,626]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="186" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
<emphasis id="B905EABA816E36312A07FDC0428B0B01" bold="true" box="[586,904,599,626]" italics="true" pageId="27" pageNumber="186">Ceratoculicoides pacificus</emphasis>
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<taxonomicNameLabel id="A23657C1816E36312BC2FDCF42EA0B02" box="[911,1001,600,626]" pageId="27" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCE36A8816E36312A22FD3642C70BCB" blockId="27.[377,1212,599,700]" box="[623,964,673,700]" pageId="27" pageNumber="186">
<figureCitation id="134A2A2D816E36312A22FD3643D40BCC" box="[623,727,673,700]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="4.[189,232,1998,2024]" captionStartId-1="5.[189,232,1998,2024]" captionStartId-2="6.[189,232,1998,2024]" captionTargetBox-0="[225,1363,1044,1945]" captionTargetBox-1="[225,1362,976,1941]" captionTargetBox-2="[262,1322,620,1924]" captionTargetId-0="figure-234@4.[225,1363,1044,1945]" captionTargetId-1="figure-163@5.[225,1362,976,1941]" captionTargetId-2="figure-47@6.[262,1322,620,1924]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 1. Ceratoculicoides pacificus sp. nov., male habitus (CNCI). Scale bar = 100 μm." captionText-1="Fig. 2. Ceratoculicoides pacificus sp. nov. (CNCI), female habitus. Scale bar = 100 μm." captionText-2="Fig. 3. Ceratoculicoides pacificus sp. nov. (CNCI), male thorax. Scale bar = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.8083775" figureDoi-1="http://doi.org/10.5281/zenodo.8083779" figureDoi-2="http://doi.org/10.5281/zenodo.8083783" httpUri-0="https://zenodo.org/record/8083775/files/figure.png" httpUri-1="https://zenodo.org/record/8083779/files/figure.png" httpUri-2="https://zenodo.org/record/8083783/files/figure.png" pageId="27" pageNumber="186">Figs 13</figureCitation>
,
<figureCitation id="134A2A2D816E36312AA9FD3642000BCB" box="[740,771,673,699]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[195,1398,668,1908]" captionTargetId="figure-118@9.[195,1398,668,1908]" captionTargetPageId="9" captionText="Fig. 4. Male heads I. a. Ceratoculicoides confusus sp. nov. b. C. pacificus sp. nov. c. C. confusus (Colombia). d. C.virginianus (Wirth, 1951). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083785" httpUri="https://zenodo.org/record/8083785/files/figure.png" pageId="27" pageNumber="186">4b</figureCitation>
,
<figureCitation id="134A2A2D816E36312B5DFD36422E0BCB" box="[784,813,673,699]" captionStart="Fig" captionStartId="11.[189,232,1963,1989]" captionTargetBox="[222,1365,830,1930]" captionTargetId="figure-172@11.[222,1365,830,1930]" captionTargetPageId="11" captionText="Fig. 6. Female heads I. a. Ceratoculicoidesconfusus sp. nov. b. Ceratoculicoides sp. F2 (Colombia). c. C. pacificus sp. nov. d. C.virginianus (Wirth, 1951). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083791" httpUri="https://zenodo.org/record/8083791/files/figure.png" pageId="27" pageNumber="186">6c</figureCitation>
,
<figureCitation id="134A2A2D816E36312B77FD3642730BCB" box="[826,880,673,699]" captionStart="Fig" captionStartId="14.[189,232,1787,1813]" captionTargetBox="[259,1269,265,1731]" captionTargetId="figure-119@14.[259,1269,265,1731]" captionTargetPageId="14" captionText="Fig. 8. Male genitalia I. Ceratoculicoides confusus sp. nov.: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.confusus (Colombia): d. Genitalia, aedeagus and parameres removed. e. Parameres. f. Aedeagus.C. pacificus sp. nov: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083798" httpUri="https://zenodo.org/record/8083798/files/figure.png" pageId="27" pageNumber="186">
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i
</figureCitation>
,
<figureCitation id="134A2A2D816E36312B30FD3642C70BCB" box="[893,964,673,699]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="27" pageNumber="186">12ab</figureCitation>
</paragraph>
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<subSubSection id="C36B6523816E363128F0FD7340DB0D06" pageId="27" pageNumber="186" type="diagnosis">
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<emphasis id="B905EABA816E363128F0FD7340350B8E" bold="true" box="[189,310,740,766]" pageId="27" pageNumber="186">Diagnosis</emphasis>
</heading>
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<emphasis id="B905EABA816E363128A7FC8240290A5F" bold="true" box="[234,298,789,815]" pageId="27" pageNumber="186">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A8816E363128F0FCAD435C0AB3" blockId="27.[189,1399,789,963]" pageId="27" pageNumber="186">
<taxonomicName id="4C714D2B816E363128F0FCAD40FA0A24" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[189,505,826,852]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="186" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
<emphasis id="B905EABA816E363128F0FCAD40FA0A24" box="[189,505,826,852]" italics="true" pageId="27" pageNumber="186">Ceratoculicoides pacificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C1816E36312A48FCAD43670A24" box="[517,612,826,852]" pageId="27" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from congeners by the following combination of characters: femora and tibiae brown; apices of parameres acute, tapering distally; aedeagus lateral margins convex, medial apex of aedeagus with hyaline incision, posterolateral spines elongate, 0.5 width of apex of aedeagus (
<figureCitation id="134A2A2D816E36312A4CFC3E43520AB3" box="[513,593,937,963]" captionStart="Fig" captionStartId="14.[189,232,1787,1813]" captionTargetBox="[259,1269,265,1731]" captionTargetId="figure-119@14.[259,1269,265,1731]" captionTargetPageId="14" captionText="Fig. 8. Male genitalia I. Ceratoculicoides confusus sp. nov.: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.confusus (Colombia): d. Genitalia, aedeagus and parameres removed. e. Parameres. f. Aedeagus.C. pacificus sp. nov: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083798" httpUri="https://zenodo.org/record/8083798/files/figure.png" pageId="27" pageNumber="186">Fig. 8i</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A8816E363128A7FC7B40460D76" blockId="27.[189,1398,1004,1142]" box="[234,325,1004,1030]" pageId="27" pageNumber="186">
<emphasis id="B905EABA816E363128A7FC7B40460D76" bold="true" box="[234,325,1004,1030]" pageId="27" pageNumber="186">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A8816E363128F0FB8640DB0D06" blockId="27.[189,1398,1004,1142]" pageId="27" pageNumber="186">
Only species of
<taxonomicName id="4C714D2B816E36312933FB8643450D5B" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[382,582,1041,1067]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816E36312933FB8643450D5B" box="[382,582,1041,1067]" italics="true" pageId="27" pageNumber="186">Ceratoculicoides</emphasis>
</taxonomicName>
with the following combination of characters: FR 1.621.72; femora and tibiae brown, wing length
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; 2 major spermathecae, largest 8896; medial margin of 9
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sternite deeply concave.
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816E363128F0FB0945980C7D" pageId="27" pageNumber="186" type="etymology">
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<emphasis id="B905EABA816E363128F0FB0940470DC8" bold="true" box="[189,324,1182,1208]" pageId="27" pageNumber="186">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8816E363128F0FB5945980C7D" blockId="27.[189,1398,1230,1294]" pageId="27" pageNumber="186">
The specific epithet
<taxonomicName id="4C714D2B816E363129F7FB5843210D99" authorityName="Fasbender" authorityYear="2023" box="[442,546,1231,1257]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="186" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B905EABA816E363129F7FB5843210D99" box="[442,546,1231,1257]" italics="true" pageId="27" pageNumber="186">pacificus</emphasis>
</taxonomicName>
is a Latin adjective referring to peacemaking or peacefulness. This epithet is in reference to this species inhabiting the coastal Pacific Northwest region.
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816E363628F0FAA145D108E1" lastPageId="28" lastPageNumber="187" pageId="27" pageNumber="186" type="materials_examined">
<paragraph id="8BCE36A8816E363128F0FAA140A90C20" blockId="27.[189,426,1334,1360]" box="[189,426,1334,1360]" pageId="27" pageNumber="186">
<heading id="D08681C4816E363128F0FAA140A90C20" bold="true" box="[189,426,1334,1360]" fontSize="11" level="3" pageId="27" pageNumber="186" reason="3">
<emphasis id="B905EABA816E363128F0FAA140A90C20" bold="true" box="[189,426,1334,1360]" pageId="27" pageNumber="186">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B193CF5816E363128A7FAF042400CBB" collectingDate="1991-06-21" collectingDateMax="1991-07-03" collectingDateMin="1991-06-21" collectionCode="CNCI" collectorName="N. Winchester" country="Canada" location="Upper Carmanah Valley" municipality="Cowichan District" pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="British Columbia" typeStatus="holotype">
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<emphasis id="B905EABA816E363128A7FAF0405F0CF1" bold="true" box="[234,348,1383,1409]" pageId="27" pageNumber="186">
<typeStatus id="54CA880A816E363128A7FAF0405F0CF1" box="[234,348,1383,1409]" pageId="27" pageNumber="186">Holotype</typeStatus>
</emphasis>
</paragraph>
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<collectingCountry id="F3667638816E363128F0FA1B403A0CD6" box="[189,313,1420,1446]" name="Canada" pageId="27" pageNumber="186">CANADA</collectingCountry>
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;
<collectingRegion id="49B5F84A816E36312932FA1B43530CD6" box="[383,592,1420,1446]" country="Canada" name="British Columbia" pageId="27" pageNumber="186">British Columbia</collectingRegion>
,
<collectingMunicipality id="6BAAACD2816E36312A13FA1B42340CD6" box="[606,823,1420,1446]" pageId="27" pageNumber="186">Cowichan District</collectingMunicipality>
,
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NW of Port Renfrew
</locationDeviation>
,
<location id="8EAE6073816E36312CE6FA1A400F0CBB" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312CE6FA1A400F0CBB" country="Canada" municipality="Cowichan District" name="Upper Carmanah Valley" pageId="27" pageNumber="186" stateProvince="British Columbia">Upper Carmanah Valley</location>
;
<date id="FFCF1068816E3631295AFA2643030CBB" box="[279,512,1457,1483]" pageId="27" pageNumber="186" value="1991-06-21" valueMax="1991-07-03" valueMin="1991-06-21">
<collectingDate id="EF8BE980816E3631295AFA2643030CBB" box="[279,512,1457,1483]" pageId="27" pageNumber="186" value="1991-06-21" valueMax="1991-07-03" valueMin="1991-06-21">21 Jun.3 Jul. 1991</collectingDate>
</date>
;
<collectorName id="2684537E816E36312A41FA2543B00CBB" box="[524,691,1457,1484]" pageId="27" pageNumber="186">N. Winchester</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312ABBFA2642430CBB" box="[758,832,1457,1483]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="3B193CF5816E363128A7FA6345130F43" collectingDate="1991-06-21" collectingDateMax="1991-07-03" collectingDateMin="1991-06-21" collectionCode="CNCI" collectorName="N. Winchester" country="Canada" location="CANADA" municipality="Cowichan District" pageId="27" pageNumber="186" specimenCount="12" specimenCount-female="2" specimenCount-male="10" stateProvince="British Columbia" typeStatus="paratype">
<paragraph id="8BCE36A8816E363128A7FA6340640F7E" blockId="27.[189,1399,1524,1661]" box="[234,359,1524,1550]" pageId="27" pageNumber="186">
<emphasis id="B905EABA816E363128A7FA6340640F7E" bold="true" box="[234,359,1524,1550]" pageId="27" pageNumber="186">
<typeStatus id="54CA880A816E363128A7FA6340640F7E" box="[234,359,1524,1550]" pageId="27" pageNumber="186">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCE36A8816E363128F0F98E40950F0D" blockId="27.[189,1399,1524,1661]" pageId="27" pageNumber="186">
<collectingCountry id="F3667638816E363128F0F98E403A0F43" box="[189,313,1561,1587]" name="Canada" pageId="27" pageNumber="186">CANADA</collectingCountry>
<specimenCount id="9D77FD21816E36312918F98E40AA0F44" box="[341,425,1561,1588]" pageId="27" pageNumber="186" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D77FD21816E363129FAF98E40F80F44" box="[439,507,1561,1588]" pageId="27" pageNumber="186" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED60AE6D816E36312B87F98E45130F43" box="[970,1040,1561,1587]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
<materialsCitation id="3B193CF5816E36312C61F98E43C50F28" collectingDate="1991-07-04" collectingDateMax="1991-07-15" collectingDateMin="1991-07-04" collectionCode="CNCI" collectorName="N. Winchester" country="Canada" location="Upper Carmanah Valley" municipality="Cowichan District" pageId="27" pageNumber="186" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="British Columbia" typeStatus="paratype">
<specimenCount id="9D77FD21816E36312C61F98E45720F44" box="[1068,1137,1561,1588]" pageId="27" pageNumber="186" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D77FD21816E36312C32F98E45AD0F44" box="[1151,1198,1561,1588]" pageId="27" pageNumber="186" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<date id="FFCF1068816E363129F0F9A943710F28" box="[445,626,1598,1625]" pageId="27" pageNumber="186" value="1991-07-04" valueMax="1991-07-15" valueMin="1991-07-04">
<collectingDate id="EF8BE980816E363129F0F9A943710F28" box="[445,626,1598,1625]" pageId="27" pageNumber="186" value="1991-07-04" valueMax="1991-07-15" valueMin="1991-07-04">415 Jul. 1991</collectingDate>
</date>
;
<collectionCode id="ED60AE6D816E36312ACDF9A943C50F28" box="[640,710,1598,1624]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312AAEF9A940910F0D" collectingDate="1991-07-31" collectingDateMax="1991-08-11" collectingDateMin="1991-07-31" collectionCode="CNCI" collectorName="N. Winchester" country="Canada" location="Upper Carmanah Valley" municipality="Cowichan District" pageId="27" pageNumber="186" specimenCount="3" specimenCount-female="3" stateProvince="British Columbia" typeStatus="paratype">
<specimenCount id="9D77FD21816E36312AAEF9A9422B0F29" box="[739,808,1598,1625]" pageId="27" pageNumber="186" type="female">3 ♀♀</specimenCount>
; same collection data as for holotype;
<date id="FFCF1068816E36312CB6F9A9403F0F0D" pageId="27" pageNumber="186" value="1991-07-31" valueMax="1991-08-11" valueMin="1991-07-31">
<collectingDate id="EF8BE980816E36312CB6F9A9403F0F0D" pageId="27" pageNumber="186" value="1991-07-31" valueMax="1991-08-11" valueMin="1991-07-31">31 Jul.11 Aug. 1991</collectingDate>
</date>
;
<collectionCode id="ED60AE6D816E36312905F9F440910F0D" box="[328,402,1635,1661]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
.
</paragraph>
</materialsCitation>
<paragraph id="8BCE36A8816E363128A7F93140A50FB0" blockId="27.[189,1399,1702,2025]" box="[234,422,1702,1728]" pageId="27" pageNumber="186">
<emphasis id="B905EABA816E363128A7F93140A50FB0" bold="true" box="[234,422,1702,1728]" pageId="27" pageNumber="186">Other material</emphasis>
</paragraph>
<paragraph id="8BCE36A8816E363628F0F95B45D108E1" blockId="27.[189,1399,1702,2025]" lastBlockId="28.[189,1399,265,402]" lastPageId="28" lastPageNumber="187" pageId="27" pageNumber="186">
<materialsCitation id="3B193CF5816E363128F0F95B40DB0E7B" collectingDate="1984-07-31" collectingDateMax="1984-08-14" collectingDateMin="1984-07-31" collectionCode="CNCI" collectorName="R. A. Cannings" country="Canada" location="East Sooke Pk." pageId="27" pageNumber="186" specimenCount="1" specimenCount-female="1" stateProvince="British Columbia">
<collectingCountry id="F3667638816E363128F0F95B403A0F96" box="[189,313,1740,1766]" name="Canada" pageId="27" pageNumber="186">CANADA</collectingCountry>
<emphasis id="B905EABA816E3631291AF95C43330F95" bold="true" box="[343,560,1739,1765]" pageId="27" pageNumber="186">
<collectingRegion id="49B5F84A816E3631291AF95C43330F95" box="[343,560,1739,1765]" country="Canada" name="British Columbia" pageId="27" pageNumber="186">British Columbia</collectingRegion>
</emphasis>
<specimenCount id="9D77FD21816E36312A04F95B43740F96" box="[585,631,1740,1766]" pageId="27" pageNumber="186" type="female">1 ♀</specimenCount>
;
<location id="8EAE6073816E36312AC8F95B42340F95" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312AC8F95B42340F95" box="[645,823,1739,1766]" country="Canada" name="East Sooke Pk." pageId="27" pageNumber="186" stateProvince="British Columbia">East Sooke Pk.</location>
<locationDeviation id="9A7C2F8A816E36312B73F95B45470F95" box="[830,1092,1739,1766]" location="Victoria" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E36312B73F95B42840F95" box="[830,903,1739,1766]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="186" unit="km" value="20.0">20 km</quantity>
SW of Victoria
</locationDeviation>
;
<date id="FFCF1068816E36312C1DF95B444C0F96" box="[1104,1359,1739,1766]" pageId="27" pageNumber="186" value="1984-07-31" valueMax="1984-08-14" valueMin="1984-07-31">
<collectingDate id="EF8BE980816E36312C1DF95B444C0F96" box="[1104,1359,1739,1766]" pageId="27" pageNumber="186" value="1984-07-31" valueMax="1984-08-14" valueMin="1984-07-31">31 Jul.14 Aug. 1984</collectingDate>
</date>
;
<collectorName id="2684537E816E36312D11F95B40530E7A" pageId="27" pageNumber="186">R. A. Cannings</collectorName>
leg.;
<collectionCode id="ED60AE6D816E363129DFF96640DB0E7B" box="[402,472,1777,1803]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E363129BFF966403D0E40" collectingDate="1989-06-29" collectingDateMax="1989-07-18" collectingDateMin="1989-06-29" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="East Sooke Pk." pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="British Columbia">
<specimenCount id="9D77FD21816E363129BFF96643220E7B" box="[498,545,1777,1803]" pageId="27" pageNumber="186" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<date id="FFCF1068816E36312BBFF96645EA0E7B" box="[1010,1257,1776,1803]" pageId="27" pageNumber="186" value="1989-06-29" valueMax="1989-07-18" valueMin="1989-06-29">
<collectingDate id="EF8BE980816E36312BBFF96645EA0E7B" box="[1010,1257,1776,1803]" pageId="27" pageNumber="186" value="1989-06-29" valueMax="1989-07-18" valueMin="1989-06-29">29 Jun.18 Jul. 1989</collectingDate>
</date>
;
<collectorName id="2684537E816E36312CB9F96644740E7A" box="[1268,1399,1776,1803]" pageId="27" pageNumber="186">A. Borkent</collectorName>
leg.;
<collectionCode id="ED60AE6D816E363128B5F881403D0E40" box="[248,318,1814,1840]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E3631291AF88145690E40" box="[343,1130,1813,1840]" collectingDate="1989-07-18" collectingDateMax="1989-07-26" collectingDateMin="1989-07-18" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="East Sooke Pk." pageId="27" pageNumber="186" specimenCount="1" specimenCount-female="1" stateProvince="British Columbia">
<specimenCount id="9D77FD21816E3631291AF88140870E40" box="[343,388,1814,1840]" pageId="27" pageNumber="186" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<date id="FFCF1068816E36312B1AF88245140E40" box="[855,1047,1813,1840]" pageId="27" pageNumber="186" value="1989-07-18" valueMax="1989-07-26" valueMin="1989-07-18">
<collectingDate id="EF8BE980816E36312B1AF88245140E40" box="[855,1047,1813,1840]" pageId="27" pageNumber="186" value="1989-07-18" valueMax="1989-07-26" valueMin="1989-07-18">1826 Jul. 1989</collectingDate>
</date>
;
<collectionCode id="ED60AE6D816E36312C69F88145690E40" box="[1060,1130,1814,1840]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312CCEF88145CD0E25" collectingDate="1991-07-18" collectingDateMax="1989-07-26" collectingDateMin="1989-07-18" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="Gold River" pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="British Columbia">
<specimenCount id="9D77FD21816E36312CCEF88145B10E40" box="[1155,1202,1814,1840]" pageId="27" pageNumber="186" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<locationDeviation id="9A7C2F8A816E36312988F8AD435D0E25" box="[453,606,1850,1877]" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E36312988F8AD43260E24" box="[453,549,1850,1877]" metricMagnitude="4" metricUnit="m" metricValue="1.25" pageId="27" pageNumber="186" unit="km" value="12.5">12.5 km</quantity>
NW
</locationDeviation>
of
<location id="8EAE6073816E36312ACAF8AD420E0E24" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312ACAF8AD420E0E24" box="[647,781,1850,1876]" country="Canada" name="Gold River" pageId="27" pageNumber="186" stateProvince="British Columbia">Gold River</location>
, coll.
<collectorName id="2684537E816E36312B1EF8AC42DA0E24" box="[851,985,1850,1877]" pageId="27" pageNumber="186">A. Borkent</collectorName>
;
<date id="FFCF1068816E36312BABF8AC45780E25" box="[998,1147,1850,1877]" pageId="27" pageNumber="186" value="1991-07-18">
<collectingDate id="EF8BE980816E36312BABF8AC45780E25" box="[998,1147,1850,1877]" pageId="27" pageNumber="186" value="1991-07-18">18 Jul. 1991</collectingDate>
</date>
;
<collectionCode id="ED60AE6D816E36312CC5F8AC45CD0E25" box="[1160,1230,1851,1877]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312CA4F8AC43C70E09" collectingDate="1979-07-26" collectionCode="CNCI" collectorName="J. Smith" country="Canada" location="Lake Cowichan" pageId="27" pageNumber="186" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D77FD21816E36312CA4F8AC442E0E25" box="[1257,1325,1851,1877]" pageId="27" pageNumber="186" type="female">2 ♀♀</specimenCount>
;
<location id="8EAE6073816E36312D71F8AD403B0E09" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312D71F8AD403B0E09" country="Canada" name="Lake Cowichan" pageId="27" pageNumber="186">Lake Cowichan</location>
;
<date id="FFCF1068816E3631290FF8C840D70E09" box="[322,468,1887,1913]" pageId="27" pageNumber="186" value="1979-07-26">
<collectingDate id="EF8BE980816E3631290FF8C840D70E09" box="[322,468,1887,1913]" pageId="27" pageNumber="186" value="1979-07-26">26 Jul. 1979</collectingDate>
</date>
;
<collectorName id="2684537E816E36312992F8F7433D0E09" box="[479,574,1887,1914]" pageId="27" pageNumber="186">J. Smith</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312A33F8C843C70E09" box="[638,708,1887,1913]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312A96F8C8403F0EEE" collectingDate="1991-07-08" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="Lake Cowichan" pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816E36312A96F8C8420B0E0A" box="[731,776,1887,1914]" pageId="27" pageNumber="186" type="male">1 ♂</specimenCount>
;
<locationDeviation id="9A7C2F8A816E36312B59F8C8427D0E0A" box="[788,894,1887,1914]" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E36312B59F8C8425F0E09" box="[788,860,1887,1914]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="186" unit="km" value="25.0">25 km</quantity>
W
</locationDeviation>
of
<location id="8EAE6073816E36312BEEF8C8455C0E09" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312BEEF8C8455C0E09" box="[931,1119,1887,1913]" country="Canada" name="Lake Cowichan" pageId="27" pageNumber="186">Lake Cowichan</location>
;
<date id="FFCF1068816E36312C24F8C845EF0E09" box="[1129,1260,1887,1913]" pageId="27" pageNumber="186" value="1991-07-08">
<collectingDate id="EF8BE980816E36312C24F8C845EF0E09" box="[1129,1260,1887,1913]" pageId="27" pageNumber="186" value="1991-07-08">8 Jul. 1991</collectingDate>
</date>
;
<collectorName id="2684537E816E36312CB8F8F744740E09" box="[1269,1399,1887,1914]" pageId="27" pageNumber="186">A. Borkent</collectorName>
leg.;
<collectionCode id="ED60AE6D816E363128BBF813403F0EEE" box="[246,316,1924,1950]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E3631291EF81345890EEE" box="[339,1162,1924,1951]" collectingDate="1991-07-09" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="Lake Cowichan" pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816E3631291EF81340820EEF" box="[339,385,1924,1951]" pageId="27" pageNumber="186" type="male">1 ♂</specimenCount>
;
<locationDeviation id="9A7C2F8A816E363129C1F813430F0EEF" box="[396,524,1924,1951]" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E363129C1F81340D60EEE" box="[396,469,1924,1950]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="186" unit="km" value="33.0">33 km</quantity>
NW
</locationDeviation>
of
<location id="8EAE6073816E36312A7CF81343EE0EEE" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312A7CF81343EE0EEE" box="[561,749,1924,1950]" country="Canada" name="Lake Cowichan" pageId="27" pageNumber="186">Lake Cowichan</location>
;
<date id="FFCF1068816E36312ABAF813427A0EEE" box="[759,889,1924,1950]" pageId="27" pageNumber="186" value="1991-07-09">
<collectingDate id="EF8BE980816E36312ABAF813427A0EEE" box="[759,889,1924,1950]" pageId="27" pageNumber="186" value="1991-07-09">9 Jul. 1991</collectingDate>
</date>
;
<collectorName id="2684537E816E36312BCEF81245070EEE" box="[899,1028,1924,1951]" pageId="27" pageNumber="186">A. Borkent</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312C09F81345890EEE" box="[1092,1162,1924,1950]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312CEDF81345590EB3" collectingDate="1985-07-19" collectingDateMax="1985-07-28" collectingDateMin="1985-07-19" collectionCode="CNCI" collectorName="I. M. Smith" country="Canada" location="Lake Cowichan" pageId="27" pageNumber="186" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D77FD21816E36312CEDF81345E10EEF" box="[1184,1250,1924,1951]" pageId="27" pageNumber="186" type="female">2 ♀♀</specimenCount>
;
<locationDeviation id="9A7C2F8A816E36312CBDF81344750EEF" box="[1264,1398,1924,1951]" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E36312CBDF813443C0EEE" box="[1264,1343,1924,1950]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="186" unit="km" value="2.3">2.3 km</quantity>
NW
</locationDeviation>
of
<location id="8EAE6073816E36312893F83E40990EB3" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312893F83E40990EB3" box="[222,410,1961,1987]" country="Canada" name="Lake Cowichan" pageId="27" pageNumber="186">Lake Cowichan</location>
,
<location id="8EAE6073816E363129EAF83E436F0EB3" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E363129EAF83E436F0EB3" box="[423,620,1961,1987]" country="Canada" name="South Shore Rd." pageId="27" pageNumber="186">South Shore Rd.</location>
;
<date id="FFCF1068816E36312A37F83E42380EB4" box="[634,827,1961,1988]" pageId="27" pageNumber="186" value="1985-07-19" valueMax="1985-07-28" valueMin="1985-07-19">
<collectingDate id="EF8BE980816E36312A37F83E42380EB4" box="[634,827,1961,1988]" pageId="27" pageNumber="186" value="1985-07-19" valueMax="1985-07-28" valueMin="1985-07-19">1928 Jul. 1985</collectingDate>
</date>
;
<collectorName id="2684537E816E36312B05F83D42D30EB3" box="[840,976,1961,1988]" pageId="27" pageNumber="186">I. M. Smith</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312C59F83E45590EB3" box="[1044,1114,1961,1987]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312C39F83E43C80E98" collectingDate="1991-07-08" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" location="Port Renfrew" pageId="27" pageNumber="186" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D77FD21816E36312C39F83E45A10EB4" box="[1140,1186,1961,1988]" pageId="27" pageNumber="186" type="female">1 ♀</specimenCount>
;
<locationDeviation id="9A7C2F8A816E36312CFCF83E441E0EB4" box="[1201,1309,1961,1988]" pageId="27" pageNumber="186">
<quantity id="4C899B4D816E36312CFCF83E45EE0EB3" box="[1201,1261,1961,1988]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="186" unit="km" value="5.0">5 km</quantity>
NE
</locationDeviation>
of
<location id="8EAE6073816E36312D0BF83D40250E98" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312D0BF83D40250E98" country="Canada" name="Port Renfrew" pageId="27" pageNumber="186">Port Renfrew</location>
;
<date id="FFCF1068816E3631297CF85940B50E98" box="[305,438,1998,2024]" pageId="27" pageNumber="186" value="1991-07-08">
<collectingDate id="EF8BE980816E3631297CF85940B50E98" box="[305,438,1998,2024]" pageId="27" pageNumber="186" value="1991-07-08">8 Jul. 1991</collectingDate>
</date>
;
<collectorName id="2684537E816E3631298CF85843470E98" box="[449,580,1998,2025]" pageId="27" pageNumber="186">A. Borkent</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312AC8F85943C80E98" box="[645,715,1998,2024]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816E36312AA9F85944710E98" box="[740,1394,1998,2025]" collectingDate="1979-07-15" collectingDateMax="1979-07-20" collectingDateMin="1979-07-15" collectionCode="CNCI" collectorName="I. M. Smith" country="Canada" location="Ucluelet" pageId="27" pageNumber="186" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816E36312AA9F85942100E99" box="[740,787,1998,2025]" pageId="27" pageNumber="186" type="male">1 ♂</specimenCount>
;
<location id="8EAE6073816E36312B6DF85942840E98" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816E36312B6DF85942840E98" box="[800,903,1998,2024]" country="Canada" name="Ucluelet" pageId="27" pageNumber="186">Ucluelet</location>
;
<date id="FFCF1068816E36312BDEF85945510E98" box="[915,1106,1998,2025]" pageId="27" pageNumber="186" value="1979-07-15" valueMax="1979-07-20" valueMin="1979-07-15">
<collectingDate id="EF8BE980816E36312BDEF85945510E98" box="[915,1106,1998,2025]" pageId="27" pageNumber="186" value="1979-07-15" valueMax="1979-07-20" valueMin="1979-07-15">1520 Jul. 1979</collectingDate>
</date>
;
<collectorName id="2684537E816E36312C13F85845E50E98" box="[1118,1254,1998,2025]" pageId="27" pageNumber="186">I. M. Smith</collectorName>
leg.;
<collectionCode id="ED60AE6D816E36312D65F85944710E98" box="[1320,1394,1998,2024]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="186">CNCI</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B193CF58169363628F0FE9E40410838" collectingDate="1985-07-19" collectionCode="CNCI" collectorName="A. Borkent" country="United States of America" county="Benton Co." location="Mary's Peak" pageId="28" pageNumber="187" specimenCount="1" specimenCount-male="1" stateProvince="Oregon">
<collectingCountry id="F36676388169363628F0FE9E41F50853" box="[189,246,265,291]" name="United States of America" pageId="28" pageNumber="187">USA</collectingCountry>
<specimenCount id="9D77FD2181693636295AFE9E40490853" box="[279,330,265,291]" pageId="28" pageNumber="187" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49B5F84A816936362916FE9E40B50853" box="[347,438,265,291]" country="United States of America" name="Oregon" pageId="28" pageNumber="187">Oregon</collectingRegion>
,
<collectingCounty id="62AF4E2481693636298BFE9E43510853" box="[454,594,265,291]" pageId="28" pageNumber="187">Benton Co.</collectingCounty>
,
<location id="8EAE6073816936362A2FFE9E43FE0853" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816936362A2FFE9E43FE0853" box="[610,765,265,291]" country="United States of America" county="Benton Co." name="Mary's Peak" pageId="28" pageNumber="187" stateProvince="Oregon">Marys Peak</location>
,
<locationDeviation id="9A7C2F8A816936362B40FE9E42940853" box="[781,919,265,291]" pageId="28" pageNumber="187">
<quantity id="4C899B4D816936362B40FE9E42580853" box="[781,859,265,291]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="28" pageNumber="187" unit="km" value="21.0">21 km</quantity>
NW
</locationDeviation>
of
<location id="8EAE6073816936362B8BFE9E45360853" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816936362B8BFE9E45360853" box="[966,1077,265,291]" country="United States of America" county="Benton Co." name="Corvallis" pageId="28" pageNumber="187" stateProvince="Oregon">Corvallis</location>
;
<date id="FFCF1068816936362C08FE9E45E20854" box="[1093,1249,265,292]" pageId="28" pageNumber="187" value="1985-07-19">
<collectingDate id="EF8BE980816936362C08FE9E45E20854" box="[1093,1249,265,292]" pageId="28" pageNumber="187" value="1985-07-19">19 Jul. 1985</collectingDate>
</date>
;
<collectorName id="2684537E816936362CBDFE9E44740853" box="[1264,1399,265,291]" pageId="28" pageNumber="187">A. Borkent</collectorName>
leg.;
<collectionCode id="ED60AE6D8169363628B1FEB940410838" box="[252,322,302,328]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="28" pageNumber="187">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF581693636292EFEB9401108E1" collectingDate="1971-08-11" collectingMethod="light trap" collectionCode="USNM" collectorName="Goeden &amp; Gurney" country="United States of America" county="Kittatas Co." location="DeRoux Forest Campground" municipality="Okanagan-Wenatchee National Forest" pageId="28" pageNumber="187" specimenCount="1" specimenCount-male="1" stateProvince="Washington">
<specimenCount id="9D77FD2181693636292EFEB940950838" box="[355,406,302,328]" pageId="28" pageNumber="187" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49B5F84A8169363629EBFEB943360838" box="[422,565,302,328]" country="United States of America" name="Washington" pageId="28" pageNumber="187">Washington</collectingRegion>
,
<collectingCounty id="62AF4E24816936362A09FEB943D40838" box="[580,727,302,328]" pageId="28" pageNumber="187">Kittatas Co.</collectingCounty>
,
<collectingMunicipality id="6BAAACD2816936362AAAFEB945B30838" box="[743,1200,302,328]" pageId="28" pageNumber="187">Okanagan-Wenatchee National Forest</collectingMunicipality>
,
<location id="8EAE6073816936362C8DFEB94059081C" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816936362C8DFEB94059081C" country="United States of America" county="Kittatas Co." municipality="Okanagan-Wenatchee National Forest" name="DeRoux Forest Campground" pageId="28" pageNumber="187" stateProvince="Washington">DeRoux Forest Campground</location>
;
<date id="FFCF1068816936362928FEC44306081D" box="[357,517,339,365]" pageId="28" pageNumber="187" value="1971-08-11">
<collectingDate id="EF8BE980816936362928FEC44306081D" box="[357,517,339,365]" pageId="28" pageNumber="187" value="1971-08-11">11 Aug. 1971</collectingDate>
</date>
;
<collectorName id="2684537E816936362A5CFEC5436F081C" box="[529,620,338,364]" pageId="28" pageNumber="187">Goeden</collectorName>
and
<collectorName id="2684537E816936362AE9FEC443FF081D" box="[676,764,339,365]" pageId="28" pageNumber="187">Gurney</collectorName>
leg.;
<collectingMethod id="52304EBF816936362B70FEC542A9081D" box="[829,938,338,365]" pageId="28" pageNumber="187">light trap</collectingMethod>
; originally identified as
<taxonomicName id="4C714D2B816936362C89FEC4446D081C" baseAuthorityName="Wirth" baseAuthorityYear="1952" box="[1220,1390,338,365]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B905EABA816936362C89FEC4446D081C" box="[1220,1390,338,365]" italics="true" pageId="28" pageNumber="187">C. longipennis</emphasis>
</taxonomicName>
;
<collectionCode id="ED60AE6D8169363628F0FEE0401108E1" box="[189,274,375,401]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="187" type="Museum">USNM</collectionCode>
</materialsCitation>
<materialsCitation id="3B193CF5816936362966FEE045CD08E1" box="[299,1230,375,402]" collectingDate="1971-08-11" collectionCode="CNCI" collectorName="Goeden &amp; Gurney" country="United States of America" county="Kittatas Co." location="Olympic National Park" municipality="Okanagan-Wenatchee National Forest" pageId="28" pageNumber="187" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Washington">
<specimenCount id="9D77FD21816936362966FEE0405A08E2" box="[299,345,375,402]" pageId="28" pageNumber="187" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D77FD21816936362928FEE0409108E2" box="[357,402,375,402]" pageId="28" pageNumber="187" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<location id="8EAE6073816936362B29FEE0457B08E1" LSID="urn:lsid:plazi:treatment:03D887BE816E36362A07FDC043460E84:8EAE6073816936362B29FEE0457B08E1" box="[868,1144,375,401]" country="United States of America" county="Kittatas Co." municipality="Okanagan-Wenatchee National Forest" name="Olympic National Park" pageId="28" pageNumber="187" stateProvince="Washington">Olympic National Park</location>
;
<collectionCode id="ED60AE6D816936362CC9FEE045CD08E1" box="[1156,1230,375,401]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="28" pageNumber="187">CNCI</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B65238169363628F0FE2D436F0CEE" pageId="28" pageNumber="187" type="description">
<paragraph id="8BCE36A88169363628F0FE2D404D08A4" blockId="28.[189,334,442,468]" box="[189,334,442,468]" pageId="28" pageNumber="187">
<heading id="D08681C48169363628F0FE2D404D08A4" bold="true" box="[189,334,442,468]" fontSize="11" level="3" pageId="28" pageNumber="187" reason="3">
<emphasis id="B905EABA8169363628F0FE2D404D08A4" bold="true" box="[189,334,442,468]" pageId="28" pageNumber="187">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88169363628A7FE7D40290B74" blockId="28.[189,1399,490,700]" box="[234,298,490,516]" pageId="28" pageNumber="187">
<emphasis id="B905EABA8169363628A7FE7D40290B74" bold="true" box="[234,298,490,516]" pageId="28" pageNumber="187">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A88169363628F0FD9840710BCC" blockId="28.[189,1399,490,700]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FD98406D0B57" baselines="546,546" box="[189,366,527,553]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="28" pageNumber="187">MEASUREMENTS</smallCapsWord>
(n = 5). Head width 327349(339); flagellomeres 4953(51), 2936(32), 2630(27), 2133(29), 2529(28), 2730(30), 2531(29), 2730(30), 3035(32), 2937(33), 84102(96), 66 75(71), 7988(81); AR 0.891.02(1.00); FR (n = 1) 1.67; wing length 1.141.34(1.22) mm; wing width 0.370.45(0.42) mm; costal ratio (n = 1) 0.50; GCR 2.12.2(2.1); GSR 1.01.1(1.1); aedeagus ratio 0.831.0(0.91).
</paragraph>
<paragraph id="8BCE36A88169363628F0FD7243630A53" blockId="28.[189,1399,741,804]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FD72401A0B8D" baselines="760,760" box="[189,281,741,767]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="28" pageNumber="187">THORAX</smallCapsWord>
. Dorsocentral punctations prominent and interspersed in posterior half of dorsocentral setae. Legs with femora and tibiae brown.
</paragraph>
<paragraph id="8BCE36A88169363628F0FCDB44090AA4" blockId="28.[189,1399,844,981]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FCDB40320A15" baselines="863,864" box="[189,305,844,870]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="28" pageNumber="187">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D816936362909FCDB40B60A16" box="[324,437,844,870]" captionStart="Fig" captionStartId="14.[189,232,1787,1813]" captionTargetBox="[259,1269,265,1731]" captionTargetId="figure-119@14.[259,1269,265,1731]" captionTargetPageId="14" captionText="Fig. 8. Male genitalia I. Ceratoculicoides confusus sp. nov.: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.confusus (Colombia): d. Genitalia, aedeagus and parameres removed. e. Parameres. f. Aedeagus.C. pacificus sp. nov: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083798" httpUri="https://zenodo.org/record/8083798/files/figure.png" pageId="28" pageNumber="187">
Fig.
<quantity id="4C899B4D816936362930FCDB40980A16" box="[381,411,844,870]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="28" pageNumber="187" unit="g" value="8.0">8g</quantity>
i
</figureCitation>
). Distal portion of parameres tapering gradually to acute apex. Aedeagus lateral margins convex, rounded, posterior margin convex, tapering posteriorly to medial notch and acute hyaline incision; elongate acute posterolateral point at transition of posterior and lateral margin, spine length 0.5 times width of aedeagus; posteriorly directed apical point adjacent to medial notch.
</paragraph>
<paragraph id="8BCE36A88169363628A7FC6940460D68" blockId="28.[189,1399,1022,1231]" box="[234,325,1022,1048]" pageId="28" pageNumber="187">
<emphasis id="B905EABA8169363628A7FC6940460D68" bold="true" box="[234,325,1022,1048]" pageId="28" pageNumber="187">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A88169363628F0FBB4454E0DBF" blockId="28.[189,1399,1022,1231]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FBB4406D0D4B" baselines="1078,1078" box="[189,366,1059,1085]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="28" pageNumber="187">MEASUREMENTS</smallCapsWord>
(n = 5). Head width (n = 3) 310360(343); flagellomeres (n = 3) 3743(40), 2327(23), 2729(29), 2932(30), 3236(36), 3236(33), 3438(37), 3638(38), 5157(56), 5053(53), 5863(60), 5562(56), 6976(72); AR 0.730.82(0.78); FR 1.621.72(1.65); wing length 1.571.84(1.63) mm; wing width 0.60.64(0.62) mm; costal ratio (n = 1) 0.53; spermathecal length 8896(93); spermathecal width 8091(81); spermathecal neck 1626(23); spermatheca/neck ratio 0.24.
</paragraph>
<paragraph id="8BCE36A88169363628F0FB6E43C80C62" blockId="28.[189,715,1272,1299]" box="[189,715,1272,1299]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FB6E401A0C61" baselines="1292,1292" box="[189,281,1273,1299]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="28" pageNumber="187">THORAX</smallCapsWord>
. Legs with femora and tibiae brown.
</paragraph>
<paragraph id="8BCE36A88169363628F0FAAC436F0CEE" blockId="28.[189,1398,1338,1439]" pageId="28" pageNumber="187">
<smallCapsWord id="8D28A0748169363628F0FAAC40320C24" baselines="1358,1359" box="[189,305,1339,1365]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="28" pageNumber="187">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D81693636290EFAAC40C40C25" box="[323,455,1339,1365]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="28" pageNumber="187">Fig. 12ab</figureCitation>
). 9
<superScript id="7C049BE08169363629A7FAAD40F40C38" attach="left" box="[490,503,1338,1352]" fontSize="6" pageId="28" pageNumber="187">th</superScript>
sternite anterior branch truncate, apices widely separated; base of posterior branch directed posteriorly, distal half curving medially, tip usually rounded, sometimes with a minute hook at apex. 2 major spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C36B65238169363628F0FA5042DD0F62" box="[189,990,1479,1554]" pageId="28" pageNumber="187" type="distribution">
<paragraph id="8BCE36A88169363628F0FA5040550C91" blockId="28.[189,342,1479,1505]" box="[189,342,1479,1505]" pageId="28" pageNumber="187">
<heading id="D08681C48169363628F0FA5040550C91" bold="true" box="[189,342,1479,1505]" fontSize="11" level="3" pageId="28" pageNumber="187" reason="3">
<emphasis id="B905EABA8169363628F0FA5040550C91" bold="true" box="[189,342,1479,1505]" pageId="28" pageNumber="187">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88169363628F0FA6042DD0F62" blockId="28.[189,990,1527,1554]" box="[189,990,1527,1554]" pageId="28" pageNumber="187">
<collectingRegion id="49B5F84A8169363628F0FA60408B0F61" box="[189,392,1527,1553]" country="Canada" name="British Columbia" pageId="28" pageNumber="187">British Columbia</collectingRegion>
(
<collectingCountry id="F36676388169363629DAFA6040F60F61" box="[407,501,1527,1553]" name="Canada" pageId="28" pageNumber="187">Canada</collectingCountry>
),
<collectingRegion id="49B5F84A816936362A44FA6043800F61" box="[521,643,1527,1553]" country="United States of America" name="California" pageId="28" pageNumber="187">California</collectingRegion>
,
<collectingRegion id="49B5F84A816936362AC2FA6F43E90F62" box="[655,746,1528,1554]" country="United States of America" name="Oregon" pageId="28" pageNumber="187">Oregon</collectingRegion>
,
<collectingRegion id="49B5F84A816936362AB8FA6042830F61" box="[757,896,1527,1553]" country="United States of America" name="Washington" pageId="28" pageNumber="187">Washington</collectingRegion>
(
<collectingCountry id="F3667638816936362BDDFA6F42D20F62" box="[912,977,1528,1554]" name="United States of America" pageId="28" pageNumber="187">USA</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B65238169363628F0F9AC43460E84" pageId="28" pageNumber="187" type="discussion">
<paragraph id="8BCE36A88169363628F0F9AC402C0F25" blockId="28.[189,303,1595,1621]" box="[189,303,1595,1621]" pageId="28" pageNumber="187">
<heading id="D08681C48169363628F0F9AC402C0F25" bold="true" box="[189,303,1595,1621]" fontSize="11" level="3" pageId="28" pageNumber="187" reason="3">
<emphasis id="B905EABA8169363628F0F9AC402C0F25" bold="true" box="[189,303,1595,1621]" pageId="28" pageNumber="187">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88169363628F0F9FD43460E84" blockId="28.[189,1401,1642,2036]" pageId="28" pageNumber="187">
Females of this species are similar in size to the female
<typeStatus id="54CA880A816936362B1CF9FD42BB0FF4" box="[849,952,1642,1668]" pageId="28" pageNumber="187">holotype</typeStatus>
of
<taxonomicName id="4C714D2B816936362BACF9FC458F0FF4" baseAuthorityName="Wirth" baseAuthorityYear="1952" box="[993,1164,1642,1669]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B905EABA816936362BACF9FC458F0FF4" box="[993,1164,1642,1669]" italics="true" pageId="28" pageNumber="187">C.longipennis</emphasis>
</taxonomicName>
, but the geographic disjunction between the type locality and the range of
<taxonomicName id="4C714D2B816936362B07F90742D60FD9" authorityName="Fasbender" authorityYear="2023" box="[842,981,1679,1706]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B905EABA816936362B07F90742D60FD9" box="[842,981,1679,1706]" italics="true" pageId="28" pageNumber="187">C. pacificus</emphasis>
</taxonomicName>
argues against their conspecificity. I believe that the stability of nomenclature is better served by describing this taxon under a new name rather than assigning this morphospecies to
<taxonomicName id="4C714D2B816936362A92F94E428C0F83" baseAuthorityName="Wirth" baseAuthorityYear="1952" box="[735,911,1753,1779]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B905EABA816936362A92F94E428C0F83" box="[735,911,1753,1779]" italics="true" pageId="28" pageNumber="187">C. longipennis</emphasis>
</taxonomicName>
based on spurious evidence, especially as another male morphospecies, sp. M1, has as much or more evidence suggesting it is the male of
<taxonomicName id="4C714D2B8169363628F0F8B540690E4C" baseAuthorityName="Wirth" baseAuthorityYear="1952" box="[189,362,1826,1852]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B905EABA8169363628F0F8B540690E4C" box="[189,362,1826,1852]" italics="true" pageId="28" pageNumber="187">C. longipennis</emphasis>
</taxonomicName>
. Females of this species are the only
<taxonomicName id="4C714D2B816936362B7EF8B542F80E4C" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[819,1019,1826,1852]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816936362B7EF8B542F80E4C" box="[819,1019,1826,1852]" italics="true" pageId="28" pageNumber="187">Ceratoculicoides</emphasis>
</taxonomicName>
in western North America with spermathecae&gt;80 μm long, but the other regional females are poorly known and this character may not be diagnostic. Males can be distinguished from all other
<taxonomicName id="4C714D2B816936362BCFF8FC45490EF5" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[898,1098,1899,1925]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816936362BCFF8FC45490EF5" box="[898,1098,1899,1925]" italics="true" pageId="28" pageNumber="187">Ceratoculicoides</emphasis>
</taxonomicName>
by their aedeagus having convex lateral and posterior margins, long (&gt;0.5× width of main body of the aedeagus) posterolateral points, and acute hyaline medial incision flanked by short triangular apical points well separated from the posterolateral points (
<figureCitation id="134A2A2D8169363629AAF84D43340E84" box="[487,567,2010,2036]" captionStart="Fig" captionStartId="14.[189,232,1787,1813]" captionTargetBox="[259,1269,265,1731]" captionTargetId="figure-119@14.[259,1269,265,1731]" captionTargetPageId="14" captionText="Fig. 8. Male genitalia I. Ceratoculicoides confusus sp. nov.: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.confusus (Colombia): d. Genitalia, aedeagus and parameres removed. e. Parameres. f. Aedeagus.C. pacificus sp. nov: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083798" httpUri="https://zenodo.org/record/8083798/files/figure.png" pageId="28" pageNumber="187">Fig. 8i</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>