516 lines
90 KiB
XML
516 lines
90 KiB
XML
<document id="3CDF633C5EAE97160E3D0BF07D1443DD" ID-DOI="10.6620/ZS.2018.57-57" ID-ISSN="1810-522X" ID-PMC="PMC6409488" ID-PubMed="31966297" ID-Zenodo-Dep="13316528" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721917816406" checkinUser="felipe" docAuthor="Chen, Ssu-Li & Shih, Hsi-Te" docDate="2018" docId="03D587E31804AB3AF5F7FC0AFDDDFE30" docLanguage="en" docName="ZoolStud.57.57.1-16.pdf" docOrigin="Zoological Studies 57 (57)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Metadentobunus brevispinus Chen & Shih, 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="FFECFF9B180CAB36F51BFFB7FF9DFFC6" masterDocTitle="A Review of the Harvestman Genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with A Description of A New Species From Taiwan" masterLastPageNumber="16" masterPageNumber="1" pageNumber="9" updateTime="1723600132761" updateUser="ExternalLinkService">
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<mods:mods id="C895AAE295F31FF59344BCC051D825CD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A77F178302D2BDCBB38BF452630467CC">
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<mods:title id="5F7C89D78D4317A9EC7F1E5D60CD39DA">A Review of the Harvestman Genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with A Description of A New Species From Taiwan</mods:title>
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</mods:titleInfo>
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<mods:name id="0A4B7D0107C85550382E23FFB8960E24" type="personal">
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<mods:role id="F98E7BCFA6DBEBAD46275BB90A269A9F">
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<mods:roleTerm id="7564012EF01C7AE150251F497EBC8F51">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6550843F4B6B6DB979BEFF1664E28D60">Chen, Ssu-Li</mods:namePart>
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<mods:affiliation id="4C3530EEBB4E64B097960F1F42FCA906">Department of Life Science, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan</mods:affiliation>
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</mods:name>
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<mods:name id="818B5626C561D8EA8FCB959909E80711" type="personal">
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<mods:role id="FB20E2DB3EA2CDBB429377BA3028F496">
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<mods:roleTerm id="4C06639A82D6D9AC0F4E1D5645782DE3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B682D25999684A0C977C2849FDC4E7F7">Shih, Hsi-Te</mods:namePart>
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<mods:affiliation id="501732A2ED6DA68A565348DBC9D295F3">Department of Life Science, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan & Research Center for Global Change Biology, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="400B646F89B9645A00F6AC9AD32E525D">text</mods:typeOfResource>
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<mods:relatedItem id="11FED0463D2B2AE36884014498124A1D" type="host">
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<mods:titleInfo id="EF668F5FA6A3A60AC46FC9D5736C8588">
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<mods:title id="541C7AF4EB3E847CE0E5CFCD1438F1F7">Zoological Studies</mods:title>
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</mods:titleInfo>
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<mods:part id="7B3A1E04C4D0EA9FAA3ACF3146BB9CA7">
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<mods:date id="52FFDD1CA9BC5502F39217F17974A5EB">2018</mods:date>
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<mods:detail id="3F7CC7F90A9B9F802442C3AD16C491EB" type="pubDate">
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<mods:number id="553871160554A861ACA14B0123954278">2018-12-12</mods:number>
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</mods:detail>
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<mods:detail id="FE15D779789BD12EC04AB9D2929213B6" type="volume">
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<mods:number id="2C14E74915F2710ED9057E4B62B44B96">57</mods:number>
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</mods:detail>
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<mods:detail id="56DE560EEBA426D6F2C8D278EF324A72" type="issue">
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<mods:number id="8219A79BE35ADB2A469C0CBDCE17B13D">57</mods:number>
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</mods:detail>
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<mods:extent id="73F0F901DD3A2D4E874FB78B3A11000D" unit="page">
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<mods:start id="1465C71C62B6F791483A2D104B3D0611">1</mods:start>
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<mods:end id="A74446866777926D438D97D63200DA14">16</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="33D70B0132137FC3AADAA5743E2AF935">journal article</mods:classification>
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<mods:identifier id="F6ABACB9516831CE89E3ECD5BBFE9776" type="DOI">10.6620/ZS.2018.57-57</mods:identifier>
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<mods:identifier id="2B10D1D976DCB3BC88D905C097DE243B" type="ISSN">1810-522X</mods:identifier>
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<mods:identifier id="D319835728DB13F9088B938848A482EB" type="PMC">PMC6409488</mods:identifier>
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<mods:identifier id="664CFC07844092AAA847112EB8027E3F" type="PubMed">31966297</mods:identifier>
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<mods:identifier id="7845313AAEA1C0E4C254014ABF538AFE" type="Zenodo-Dep">13316528</mods:identifier>
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</mods:mods>
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<treatment id="03D587E31804AB3AF5F7FC0AFDDDFE30" ID-DOI="http://doi.org/10.5281/zenodo.12866464" ID-Zenodo-Dep="12866464" LSID="urn:lsid:plazi:treatment:03D587E31804AB3AF5F7FC0AFDDDFE30" httpUri="http://treatment.plazi.org/id/03D587E31804AB3AF5F7FC0AFDDDFE30" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
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<subSubSection id="C366657E1804AB3EF5F7FC0AFD55FC10" box="[236,712,957,982]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BC336F51804AB3EF5F7FC0AFD55FC10" blockId="8.[180,768,957,1067]" box="[236,712,957,982]" pageId="8" pageNumber="9">
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<heading id="D08B81991804AB3EF5F7FC0AFD55FC10" bold="true" box="[236,712,957,982]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
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<emphasis id="B908EAE71804AB3EF5F7FC0AFD55FC10" bold="true" box="[236,712,957,982]" pageId="8" pageNumber="9">
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<taxonomicName id="4C7C4D761804AB3EF5F7FC0AFDC2FC10" ID-CoL="7X938" authorityName="Chen & Shih" authorityYear="2018" box="[236,607,957,982]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brevispinus" status="sp. nov.">
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<emphasis id="B908EAE71804AB3EF5F7FC0AFDC2FC10" bold="true" box="[236,607,957,982]" italics="true" pageId="8" pageNumber="9">Metadentobunus brevispinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B579C1804AB3EF77DFC0AFD55FC10" box="[614,712,957,982]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C366657E1804AB3AF485FC6AFDDDFE30" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BC336F51804AB3EF485FC6AFD8AFC30" blockId="8.[180,768,957,1067]" box="[414,535,989,1014]" pageId="8" pageNumber="9">
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(
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<figureCitation id="13472A701804AB3EF4BEFC6AFD8DFC30" box="[421,528,989,1014]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[136,170,1925,1945]" captionStartId-1="10.[174,208,1812,1832]" captionTargetBox-0="[265,1357,257,1899]" captionTargetBox-1="[262,1362,245,1787]" captionTargetId-0="figure-6@9.[105,1445,196,1982]" captionTargetId-1="figure-6@10.[176,1450,165,1863]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 4. Metadentobunus brevispinus sp. nov. A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm." captionText-1="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13316538" figureDoi-1="http://doi.org/10.5281/zenodo.13316542" httpUri-0="https://zenodo.org/record/13316538/files/figure.png" httpUri-1="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Figs. 4-5</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC336F51804AB3EF5AFFC4FFDACFBED" blockId="8.[180,768,957,1067]" pageId="8" pageNumber="9">
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<uri id="FFED3AF71804AB3EF5AFFC4FFDACFBED" pageId="8" pageNumber="9">
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urn:lsid:zoobank.org:act:
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<uuid id="FFDA0C201804AB3EF4A9FC4FFDACFBED" pageId="8" pageNumber="9">0CBDAE16-E48C-4753-A40A- 0E09999E4DEF</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BC336F51804AB3EF5F1FBE7FC9AF9CF" blockId="8.[174,776,1104,1962]" pageId="8" pageNumber="9">
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<emphasis id="B908EAE71804AB3EF5F1FBE7FE0EFBAF" box="[234,403,1104,1129]" italics="true" pageId="8" pageNumber="9">Type material</emphasis>
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:
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<typeStatus id="54C788571804AB3EF4BDFBE7FD85FBAF" box="[422,536,1104,1129]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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: 1 S (NCHUZOOL 14599), Sitou, Lugu,
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<collectingRegion id="49B8F8171804AB3EF712FBC7FDE8FB4F" box="[521,629,1136,1161]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<collectingCountry id="F36B76651804AB3EF789FBC7FD66FB4F" box="[658,763,1136,1161]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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,
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<geoCoordinate id="EE4850321804AB3EF5B5FB27FEEEFB6F" box="[174,371,1168,1193]" degrees="23" direction="north" minutes="40" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="27.92" value="23.674421">23°40'27.92"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF49FFB27FDCAFB6F" box="[388,599,1168,1193]" degrees="120" direction="east" minutes="47" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="45.19" value="120.79588">120°47'45.19"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF5B5FB07FE96FB0F" box="[174,267,1200,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.107" pageId="8" pageNumber="9" unit="m" value="1107.0">1107 m</quantity>
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, coll. S.-L. Chen,
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<date id="FFC210351804AB3EF4F8FB07FDE6FB0F" box="[483,635,1200,1225]" pageId="8" pageNumber="9" value="2013-12-01">1 Dec. 2013</date>
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.
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<typeStatus id="54C788571804AB3EF79CFB07FC9EFB0F" box="[647,771,1200,1225]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
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: 1 $ (NCHUZOOL 14600), 1 S (NCHUZOOL 14586), 1 $ (NCHUZOOL 14587), 1 S (NMNS 7926-001) and 1 $ (NMNS 7926-002), same data as
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<typeStatus id="54C788571804AB3EF408FA87FEE0FA8F" box="[275,381,1328,1353]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
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; 1 S (NCHUZOOL 14585), Sun Moon Lake (= Rihyuetan), Yuchih,
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<collectingRegion id="49B8F8171804AB3EF753FAE7FD3FFAAF" box="[584,674,1360,1385]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<collectingCountry id="F36B76651804AB3EF7B7FAE7FC9EFAAF" box="[684,771,1360,1385]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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,
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<geoCoordinate id="EE4850321804AB3EF5B5FAC7FEAAFA4F" box="[174,311,1392,1417]" degrees="23" direction="north" minutes="51" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="26" value="23.857224">23°51'26"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF45FFAC7FE46FA4F" box="[324,475,1392,1417]" degrees="120" direction="east" minutes="56" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="42" value="120.945">120°56'42"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF760FAC7FD57FA4F" box="[635,714,1392,1417]" metricMagnitude="2" metricUnit="m" metricValue="7.64" pageId="8" pageNumber="9" unit="m" value="764.0">764 m</quantity>
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, coll. H.-Y. Chang,
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<date id="FFC210351804AB3EF444FA27FD8DFA6F" box="[351,528,1424,1449]" pageId="8" pageNumber="9" value="2014-01-19">19 Jan. 2014</date>
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; 1 S (NCHUZOOL 14595) and 1 S (NCHUZOOL 14596), Tongfu, Sinyi,
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<collectingRegion id="49B8F8171804AB3EF5E7FA67FEC4FA2F" box="[252,345,1488,1513]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<collectingCountry id="F36B76651804AB3EF47DFA67FE5DFA2F" box="[358,448,1488,1513]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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,
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<geoCoordinate id="EE4850321804AB3EF4D6FA67FDC7FA2F" box="[461,602,1488,1513]" degrees="23" direction="north" minutes="31" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="23" value="23.523056">23°31'23"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF773FA67FC9EFA2F" box="[616,771,1488,1513]" degrees="120" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="60" value="120.9">120°53'60"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF426FA47FE04F9CF" box="[317,409,1520,1545]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="8" pageNumber="9" unit="m" value="2250.0">2250 m</quantity>
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, coll. S.-L. Chen,
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<date id="FFC210351804AB3EF773FA47FC9EF9CF" box="[616,771,1520,1545]" pageId="8" pageNumber="9" value="2015-07-15">15 July 2015</date>
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.
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</paragraph>
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<paragraph id="8BC336F51804AB3EF5F1F9A7FB5CFD30" blockId="8.[174,776,1104,1962]" lastBlockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
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<emphasis id="B908EAE71804AB3EF5F1F9A7FD60F9EF" box="[234,765,1552,1577]" italics="true" pageId="8" pageNumber="9">
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Other material examined from
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<collectingCountry id="F36B76651804AB3EF785F9A7FD60F9EF" box="[670,765,1552,1577]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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</emphasis>
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: 1 $ (NCHUZOOL 14503), Sitou, Lugu,
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<collectingRegion id="49B8F8171804AB3EF7BDF987FC9EF98F" box="[678,771,1584,1609]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<geoCoordinate id="EE4850321804AB3EF5B5F9E7FEEEF9AF" box="[174,371,1616,1641]" degrees="23" direction="north" minutes="40" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="29.15" value="23.674763">23°40'29.15"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF49FF9E7FDC5F9AF" box="[388,600,1616,1641]" degrees="120" direction="east" minutes="47" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="48.55" value="120.79682">120°47'48.55"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF5B5F9C7FE91F94F" box="[174,268,1648,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.107" pageId="8" pageNumber="9" unit="m" value="1107.0">1107 m</quantity>
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, coll. S.-L. Chen,
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<date id="FFC210351804AB3EF4F7F9C7FD1AF94F" box="[492,647,1648,1673]" pageId="8" pageNumber="9" value="2013-12-01">1 Dec. 2013</date>
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; 4 S 3 $ (NCHUZOOL 14504), Tongfu, Sinyi,
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<collectingRegion id="49B8F8171804AB3EF784F927FC9FF96F" box="[671,770,1680,1705]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<geoCoordinate id="EE4850321804AB3EF5B5F907FEDDF90F" box="[174,320,1712,1737]" degrees="23" direction="north" minutes="31" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="23" value="23.523056">23°31'23"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF454F907FE6DF90F" box="[335,496,1712,1737]" degrees="120" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="60" value="120.9">120°53'60"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF784F907FC9FF90F" box="[671,770,1712,1737]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="8" pageNumber="9" unit="m" value="2250.0">2250 m</quantity>
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, coll. S.-L. Chen,
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<date id="FFC210351804AB3EF467F967FD83F92F" box="[380,542,1744,1769]" pageId="8" pageNumber="9" value="2015-07-15">15 July 2015</date>
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; 1 $ (NCHUZOOL 14588), Huisun Forest Area, Ren-ai,
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<collectingRegion id="49B8F8171804AB3EF7BAF947FC9FF8CF" box="[673,770,1776,1801]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
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,
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<geoCoordinate id="EE4850321804AB3EF5B5F8A7FEC7F8EF" box="[174,346,1808,1833]" degrees="24" direction="north" minutes="05" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="26.55" value="24.09071">24°05'26.55"N</geoCoordinate>
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,
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<geoCoordinate id="EE4850321804AB3EF47EF8A7FD83F8EF" box="[357,542,1808,1833]" degrees="121" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="58.48" value="121.03291">121°01'58.48"E</geoCoordinate>
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, elevation of
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<quantity id="4C849B101804AB3EF7ADF8A7FC9EF8EF" box="[694,771,1808,1833]" metricMagnitude="2" metricUnit="m" metricValue="6.88" pageId="8" pageNumber="9" unit="m" value="688.0">688 m</quantity>
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, coll. S.-L. Chen,
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<date id="FFC210351804AB3EF461F887FD82F88F" box="[378,543,1840,1865]" pageId="8" pageNumber="9" value="2014-11-14">14 Nov. 2014</date>
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; 1 $ (NCHUZOOL 14589), Furong Waterfall, Guosing,
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<collectingRegion id="49B8F8171804AB3EF784F8E7FC9FF8AF" box="[671,770,1872,1897]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF5B5F8C7FEC7F84F" box="[174,346,1904,1929]" degrees="24" direction="north" minutes="05" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="31.14" value="24.091984">24°05'31.14"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF47EF8C7FD83F84F" box="[357,542,1904,1929]" degrees="120" direction="east" minutes="56" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="13.95" value="120.93721">120°56'13.95"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF7ADF8C7FC9EF84F" box="[694,771,1904,1929]" metricMagnitude="2" metricUnit="m" metricValue="6.36" pageId="8" pageNumber="9" unit="m" value="636.0">636 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF462F827FDBDF86F" box="[377,544,1936,1962]" pageId="8" pageNumber="9" value="2014-09-14">14 Sep. 2014</date>
|
|
; 1 S (NCHUZOOL 14590), Aowanda National Forest Recreation Area, Ren-ai,
|
|
<collectingRegion id="49B8F8171804AB3EF6ADFEAAFB8EFEF0" box="[950,1043,285,310]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF139FEAAFB4BFEF0" box="[1058,1238,285,310]" degrees="23" direction="north" minutes="56" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="43.20" value="23.945332">23°56'43.20"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF1FEFEAAFA3AFEF0" box="[1253,1447,285,310]" degrees="121" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="58.01" value="121.18278">121°10'58.01"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF6FEFE8AFBDEFE90" box="[997,1091,317,342]" metricMagnitude="3" metricUnit="m" metricValue="1.2429999999999999" pageId="8" pageNumber="9" unit="m" value="1243.0">1243 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF003FE8AFA35FE90" box="[1304,1448,317,343]" pageId="8" pageNumber="9" value="2013-10-01">1 Oct. 2013</date>
|
|
; 1 S (NCHUZOOL 14505), Dasyueshan Forest Recreation Area, Heping,
|
|
<collectingRegion id="49B8F8171804AB3EF19AFECAFB6DFE50" box="[1153,1264,381,406]" country="Taiwan" name="Taichung" pageId="8" pageNumber="9">Taichung</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF1E1FECAFA35FE50" box="[1274,1448,381,406]" degrees="24" direction="north" minutes="15" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="22.32" value="24.2562">24°15'22.32"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF649FE2AFB8AFE70" box="[850,1047,413,438]" degrees="121" direction="east" minutes="00" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="24.82" value="121.0069">121°00'24.82"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF1D9FE2AFAB9FE70" box="[1218,1316,413,438]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="8" pageNumber="9" unit="m" value="2240.0">2240 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF6BCFE0AFBCEFE10" box="[935,1107,445,470]" pageId="8" pageNumber="9" value="2013-11-26">26 Nov. 2013</date>
|
|
; 1 $ (NCHUZOOL 14591), Dasyueshan Forest Recreation Area, Heping,
|
|
<collectingRegion id="49B8F8171804AB3EF649FE4AFC52FDD0" box="[850,975,509,534]" country="Taiwan" name="Taichung" pageId="8" pageNumber="9">Taichung</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF6FCFE4AFB2DFDD0" box="[999,1200,509,534]" degrees="24" direction="north" minutes="15" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="22. 32" value="24.2562">24°15'22.32"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF1D2FE4AFA3FFDD0" box="[1225,1442,509,534]" degrees="121" direction="east" minutes="00" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="24. 82" value="121.0069">121°00'24.82"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF6EEFDAAFBC7FDF0" box="[1013,1114,541,566]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="8" pageNumber="9" unit="m" value="2240.0">2240 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF053FDAAFC0CFD90" pageId="8" pageNumber="9" value="2014-08-23">23 Aug. 2014</date>
|
|
; 1 S (NCHUZOOL 14592), Cingjing, Ren-ai,
|
|
<collectingRegion id="49B8F8171804AB3EF649FDEAFC30FDB0" box="[850,941,605,630]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF6A1FDEAFBF7FDB0" box="[954,1130,605,630]" degrees="24" direction="north" minutes="03" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="26.19" value="24.057274">24°03'26.19"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF16DFDEAFAAEFDB0" box="[1142,1331,605,630]" degrees="121" direction="east" minutes="09" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="47.48" value="121.16319">121°09'47.48"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF66EFDCAFC47FD50" box="[885,986,637,662]" metricMagnitude="3" metricUnit="m" metricValue="1.891" pageId="8" pageNumber="9" unit="m" value="1891.0">1891 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF1DDFDCAFAFAFD50" box="[1222,1383,637,662]" pageId="8" pageNumber="9" value="2014-08-03">3 Aug. 2014</date>
|
|
; 1 $ (NCHUZOOL 14594), Tunyuan, Ren-ai,
|
|
<collectingRegion id="49B8F8171804AB3EF051FD2AFA3AFD70" box="[1354,1447,669,694]" country="Taiwan" name="Nantou" pageId="8" pageNumber="9">Nantou</collectingRegion>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF649FD0AFC6DFD10" box="[850,1008,701,726]" degrees="24" direction="north" minutes="03" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="1.16" value="24.050322">24°03'1.16"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EE4850321804AB3EF6E0FD0AFB29FD10" box="[1019,1204,701,726]" degrees="121" direction="east" minutes="12" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="54.48" value="121.21513">121°12'54.48"E</geoCoordinate>
|
|
, elevation of
|
|
<quantity id="4C849B101804AB3EF057FD0AFA35FD10" box="[1356,1448,701,726]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="8" pageNumber="9" unit="m" value="1590.0">1590 m</quantity>
|
|
, coll. S.-L. Chen,
|
|
<date id="FFC210351804AB3EF103FD6AFB23FD30" box="[1048,1214,733,758]" pageId="8" pageNumber="9" value="2015-06-16">16 June 2015</date>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BC336F51804AB3EF694FD4AFA1FFC50" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
|
|
<emphasis id="B908EAE71804AB3EF694FD4AFB90FCD0" box="[911,1037,765,790]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
|
|
: The name
|
|
<taxonomicName id="4C7C4D761804AB3EF1B8FD4AFAB6FCD0" authorityName="Chen & Shih" authorityYear="2018" box="[1187,1323,765,790]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brevispinus">
|
|
<emphasis id="B908EAE71804AB3EF1B8FD4AFAB6FCD0" box="[1187,1323,765,790]" italics="true" pageId="8" pageNumber="9">brevispinus</emphasis>
|
|
</taxonomicName>
|
|
is derived from the Latin
|
|
<emphasis id="B908EAE71804AB3EF6E5FCAAFBA5FCF0" box="[1022,1080,797,822]" italics="true" pageId="8" pageNumber="9">brevi</emphasis>
|
|
and
|
|
<emphasis id="B908EAE71804AB3EF16FFCAAFB5CFCF0" box="[1140,1217,797,822]" italics="true" pageId="8" pageNumber="9">spinus</emphasis>
|
|
, which refers to the one pair frontal spines on ocularium of this species are shorter (
|
|
<figureCitation id="13472A701804AB3EF6F0FCEAFBA9FCB0" box="[1003,1076,861,886]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
|
D-E) than the other species of the genus
|
|
<taxonomicName id="4C7C4D761804AB3EF6D4FCCAFB0BFC50" authorityName="Roewer" authorityYear="1915" box="[975,1174,893,918]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B908EAE71804AB3EF6D4FCCAFB0BFC50" box="[975,1174,893,918]" italics="true" pageId="8" pageNumber="9">Metadentobunus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="13472A701804AB3EF1BFFCCAFB72FC51" box="[1188,1263,893,919]" captionStart="Fig" captionStartId="4.[174,208,1896,1916]" captionTargetBox="[198,1427,203,1863]" captionTargetId="figure-6@4.[46,1511,156,2228]" captionTargetPageId="4" captionText="Fig. 1. Metadentobunus formosae Roewer, 1915 (holotype, SMF 9801162-RI/1162-66). A, B: dorsal and lateral views of female (body length: 6 mm); C, D: dorsal and lateral views of ocularium; E, F: outer and inner side of distal segment of left chelicera; G: dorsal view of supra-chelicera lamellae; H: ventral view of female genital operculum." figureDoi="http://doi.org/10.5281/zenodo.13316530" httpUri="https://zenodo.org/record/13316530/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
|
|
C-D; 3D-E).
|
|
</paragraph>
|
|
<paragraph id="8BC336F51804AB3EF694FC2AFAAAFBB0" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
|
|
<emphasis id="B908EAE71804AB3EF694FC2AFB9AFC70" box="[911,1031,925,950]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
|
|
: Male: ocularium with one pair of short frontal spines (
|
|
<figureCitation id="13472A701804AB3EF15FFC0AFB17FC10" box="[1092,1162,957,982]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
|
D-E). Patella of pedipalp with inner apophysis at tip (
|
|
<figureCitation id="13472A701804AB3EF180FC6AFB7CFC31" box="[1179,1249,989,1015]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
|
N-O). Shaft long, constricted at base (
|
|
<figureCitation id="13472A701804AB3EF146FC4AFB22FBD0" box="[1117,1215,1021,1046]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5F</figureCitation>
|
|
). Female: seminal receptacle at 5th-6th segment of ovipositor, with two ampullas terminally (
|
|
<figureCitation id="13472A701804AB3EF1C0FB8AFAA9FB90" box="[1243,1332,1085,1110]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5I</figureCitation>
|
|
). Genital operculum flat at frontal margin (
|
|
<figureCitation id="13472A701804AB3EF1CAFBEAFAB6FBB0" box="[1233,1323,1117,1142]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5P</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BC336F51804AB3EF694FBCAFC5AF9D0" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
|
|
<emphasis id="B908EAE71804AB3EF694FBCAFB81FB50" box="[911,1052,1149,1174]" italics="true" pageId="8" pageNumber="9">Description</emphasis>
|
|
: Male
|
|
<typeStatus id="54C788571804AB3EF16DFBCAFB78FB50" box="[1142,1253,1149,1174]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
|
. Dorsal surface of body granulated. Supra-cheliceral lamellae tipped with 5 small tubercles (
|
|
<figureCitation id="13472A701804AB3EF1DDFB0AFABFFB10" box="[1222,1314,1213,1238]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5L</figureCitation>
|
|
). Abdomen with 1 short median spine on the second tergite. Ocularium rectangle (
|
|
<figureCitation id="13472A701804AB3EF16AFB4AFB47FAD0" box="[1137,1242,1277,1302]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5D</figureCitation>
|
|
), length slightly larger than width, constricted basally, with short spine at frontal margin (
|
|
<figureCitation id="13472A701804AB3EF19AFA8AFB7FFA90" box="[1153,1250,1341,1366]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5E</figureCitation>
|
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). Frontal spines of ocularium smooth except sparse hairs. All free sternites and genital operculum smooth. Surface of all coxae with scattered coarse granules. Coxa I and IV with a row of humps anteriorly and posteriorly; Coxa II and III with a row of humps anteriorly.
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</paragraph>
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<paragraph id="8BC336F51804AB3EF694F9AAFB71F950" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
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Chelicera (
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|
<figureCitation id="13472A701804AB3EF10FF9AAFBC0F9F0" box="[1044,1117,1565,1590]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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J-K). Basal segment with 7 tubercles and setae above, with a slender spur ventrally. Distal segment with setae above, outer side smooth, inner side with setae.
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</paragraph>
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<paragraph id="8BC336F51804AB3DF694F92AFF60FED0" blockId="8.[850,1452,253,1975]" lastBlockId="11.[136,738,253,1558]" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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Pedipalp (
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<figureCitation id="13472A701804AB3EF117F92AFBC9F970" box="[1036,1108,1693,1718]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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A-B). All segments with lots of setae, excepting trochanter. Trochanter with 1 conspicuous tubercle. Femur with conspicuous tubercles ventrally, with 11 tubercles on outer side (
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<figureCitation id="13472A701804AB3EF641F8AAFC21F8F0" box="[858,956,1821,1846]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5B</figureCitation>
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) and a row of tubercles on inner side (
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<figureCitation id="13472A701804AB3EF641F88AFC24F890" box="[858,953,1853,1878]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5A</figureCitation>
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). Patella thickened distally, with an inner apophysis at tip. Tibia covered with many simple setae especially in inner tip. Tarsus elongated, with a row of tubercles at inner side (
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<figureCitation id="13472A701804AB3EF1EEF82AFAC5F870" box="[1269,1368,1949,1974]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="8" pageNumber="9">Fig. 5A</figureCitation>
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). Claw pectinate.
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</paragraph>
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<footnote id="E8672AFB1804AB3EF187F86AFA36F829" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">
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<paragraph id="8BC336F51804AB3EF187F86AFA36F829" blockId="8.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">© 2018 Academia Sinica, Taiwan</paragraph>
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</footnote>
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<caption id="DF03667D1805AB3FF593F832FBE6F873" ID-DOI="http://doi.org/10.5281/zenodo.13316538" ID-Zenodo-Dep="13316538" httpUri="https://zenodo.org/record/13316538/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,170,1925,1945]" targetBox="[265,1357,257,1899]" targetPageId="9" targetType="figure">
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<paragraph id="8BC336F51805AB3FF593F832FBE6F873" blockId="9.[136,1414,1925,1973]" pageId="9" pageNumber="10">
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<emphasis id="B908EAE71805AB3FF593F832FF5AF85F" bold="true" box="[136,199,1925,1945]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
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<taxonomicName id="4C7C4D761805AB3FF5CDF832FE74F85F" authorityName="Chen & Shih" authorityYear="2018" box="[214,489,1925,1945]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevispinus" status="sp. nov.">
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<emphasis id="B908EAE71805AB3FF5CDF832FE74F85F" box="[214,489,1925,1945]" italics="true" pageId="9" pageNumber="10">Metadentobunus brevispinus</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="A23B579C1805AB3FF4EBF832FDA1F85F" box="[496,572,1925,1945]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<footnote id="E8672AFB1805AB3FF593F86AFE05F829" box="[136,408,2013,2031]" pageId="9" pageNumber="10">
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<paragraph id="8BC336F51805AB3FF593F86AFE05F829" blockId="9.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="9" pageNumber="10">© 2018 Academia Sinica, Taiwan</paragraph>
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</footnote>
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<caption id="DF03667D1806AB3CF5B5F8A3FC71F872" ID-DOI="http://doi.org/10.5281/zenodo.13316542" ID-Zenodo-Dep="13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,208,1812,1832]" targetBox="[262,1362,245,1787]" targetPageId="10" targetType="figure">
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<paragraph id="8BC336F51806AB3CF5B5F8A3FC71F872" blockId="10.[174,1452,1812,1972]" pageId="10" pageNumber="11">
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|
<emphasis id="B908EAE71806AB3CF5B5F8A3FF73F8EE" bold="true" box="[174,238,1812,1832]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
|
|
<taxonomicName id="4C7C4D761806AB3CF5E5F8A3FD8CF8EE" authorityName="Chen & Shih" authorityYear="2018" box="[254,529,1812,1832]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevispinus" status="sp. nov.">
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<emphasis id="B908EAE71806AB3CF5E5F8A3FD8CF8EE" box="[254,529,1812,1832]" italics="true" pageId="10" pageNumber="11">Metadentobunus brevispinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B579C1806AB3CF701F8A3FDFAF8EE" box="[538,615,1812,1832]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm.
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</paragraph>
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</caption>
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<footnote id="E8672AFB1806AB3CF187F86AFA36F829" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">
|
|
<paragraph id="8BC336F51806AB3CF187F86AFA36F829" blockId="10.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">© 2018 Academia Sinica, Taiwan</paragraph>
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</footnote>
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<paragraph id="8BC336F51807AB3DF5DEFEAAFDA6FEB0" blockId="11.[136,738,253,1558]" pageId="11" pageNumber="12">
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|
Legs. All legs slim and elongate. Trochanters mound-shaped (
|
|
<figureCitation id="13472A701807AB3DF452FE8AFE10FE90" box="[329,397,317,342]" captionStart="Fig" captionStartId="9.[136,170,1925,1945]" captionTargetBox="[265,1357,257,1899]" captionTargetId="figure-6@9.[105,1445,196,1982]" captionTargetPageId="9" captionText="Fig. 4. Metadentobunus brevispinus sp. nov. A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316538" httpUri="https://zenodo.org/record/13316538/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
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A-B), femora II with two false articulations, tarsus longer than tibia.
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</paragraph>
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<paragraph id="8BC336F51807AB3DF5DEFECAFEA6FDD0" blockId="11.[136,738,253,1558]" pageId="11" pageNumber="12">
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Penis (
|
|
<figureCitation id="13472A701807AB3DF403FECAFEC2FE51" box="[280,351,381,407]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
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F-G). Shaft 2.28 long, 0.21 wide at base, 0.29 wide at middle; glans 0.2 long, 0.1 wide at base. Shaft long, constricted at base; alate part with sacs (
|
|
<figureCitation id="13472A701807AB3DF454FE6AFE2CFE30" box="[335,433,477,502]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5F</figureCitation>
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). Glans widest at base. Stylus acicular.
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</paragraph>
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<paragraph id="8BC336F51807AB3DF5DEFDAAFEFEFBD0" blockId="11.[136,738,253,1558]" pageId="11" pageNumber="12">
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|
<emphasis id="B908EAE71807AB3DF5DEFDAAFE21FDF0" box="[197,444,541,566]" italics="true" pageId="11" pageNumber="12">
|
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Female
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<typeStatus id="54C788571807AB3DF42DFDAAFE21FDF0" box="[310,444,541,566]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
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</emphasis>
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(NCHUZOOL 14600, NMNS 7926-002): Dorsal surface of body similar to male, excepting abdomen of female longer than male (
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<figureCitation id="13472A701807AB3DF414FDCAFEC8FD51" box="[271,341,637,663]" captionStart="Fig" captionStartId="9.[136,170,1925,1945]" captionTargetBox="[265,1357,257,1899]" captionTargetId="figure-6@9.[105,1445,196,1982]" captionTargetPageId="9" captionText="Fig. 4. Metadentobunus brevispinus sp. nov. A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316538" httpUri="https://zenodo.org/record/13316538/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
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C-D). Frontal spines of ocularium smooth except sparse hairs. Supra-cheliceral lamellae tipped with 2-4 small tubercles (
|
|
<figureCitation id="13472A701807AB3DF7B5FD0AFF2FFD30" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5M</figureCitation>
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). Trochanter of pedipalps with 1 conspicuous tubercle ventrally. Pedipalpal femur with a row of black-tipped tubercles at inner side, tibia covered with simple setae entirely (
|
|
<figureCitation id="13472A701807AB3DF4EAFC8AFDC6FC91" box="[497,603,829,855]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5C</figureCitation>
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). Inner-tip apophysis of pedipalpal patella longer than male. Pedipalpal tibia of female with small apophysis at inner tip (
|
|
<figureCitation id="13472A701807AB3DF5E3FC2AFECAFC71" box="[248,343,925,951]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5O</figureCitation>
|
|
). Genital operculum flat at frontal margin (
|
|
<figureCitation id="13472A701807AB3DF5EFFC0AFEC7FC10" box="[244,346,957,982]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5P</figureCitation>
|
|
). Seminal receptacle (
|
|
<figureCitation id="13472A701807AB3DF79AFC0AFD4AFC10" box="[641,727,957,982]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5I</figureCitation>
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) located in 5th-6th segment of ovipositor, terminal with two ampullas.
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF5DEFBAAFEC5FB30" blockId="11.[136,738,253,1558]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF5DEFBAAFE13FBF0" box="[197,398,1053,1078]" italics="true" pageId="11" pageNumber="12">Measurements</emphasis>
|
|
:
|
|
<typeStatus id="54C788571807AB3DF4BEFBAAFD83FBF0" box="[421,542,1053,1078]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
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|
male (female
|
|
<typeStatus id="54C788571807AB3DF593FB8AFF6EFB90" box="[136,243,1085,1110]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
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|
, NCHUZOOL 14600): Cephalothorax 1.15 (1.68) long, 2.62 (3.0) wide; abdomen 2.16 (3.78) long, 2.41 (3.39) wide. Total length of body 3.31 (5.46). Measurements of left pedipalp and legs are shown in table 3. Female: similar with male, but darker than male.
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF5DEFB4AFAF1FE91" blockId="11.[136,738,253,1558]" lastBlockId="11.[813,1414,253,1110]" pageId="11" pageNumber="12">
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|
<emphasis id="B908EAE71807AB3DF5DEFB4AFEA2FAD0" box="[197,319,1277,1302]" italics="true" pageId="11" pageNumber="12">Coloration</emphasis>
|
|
: Male. Dorsum golden yellow with granules, spine on second tergite black (
|
|
<figureCitation id="13472A701807AB3DF765FAAAFD58FAF0" box="[638,709,1309,1334]" captionStart="Fig" captionStartId="9.[136,170,1925,1945]" captionTargetBox="[265,1357,257,1899]" captionTargetId="figure-6@9.[105,1445,196,1982]" captionTargetPageId="9" captionText="Fig. 4. Metadentobunus brevispinus sp. nov. A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316538" httpUri="https://zenodo.org/record/13316538/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
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|
A- B). Carapace golden yellow with small light brown flecks along frontal and lateral edge. Ocularium golden yellow, with white eye rings. Coxa of all legs golden yellow. Genital operculum golden yellow, light yellow base, distal edge light yellow. Sternites pale yellow, whiten posterior. Chelicerae pale yellow, basal segment with brown spots. Pedipalps pale yellow, with femora and patellae brown. All legs brown with golden yellow rings. Female similar with male, but darker (
|
|
<figureCitation id="13472A701807AB3DF1F3FE8AFAB0FE91" box="[1256,1325,317,343]" captionStart="Fig" captionStartId="9.[136,170,1925,1945]" captionTargetBox="[265,1357,257,1899]" captionTargetId="figure-6@9.[105,1445,196,1982]" captionTargetPageId="9" captionText="Fig. 4. Metadentobunus brevispinus sp. nov. A, B: dorsal and lateral views of male (holotype, NCHUZOOL 14599); C, D: dorsal and lateral views of female (paratype, NCHUZOOL 14600); E: color in live, male and female. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316538" httpUri="https://zenodo.org/record/13316538/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
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C-D).
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</paragraph>
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<paragraph id="8BC336F51807AB3DF672FEEAFC01FDF0" blockId="11.[813,1414,253,1110]" pageId="11" pageNumber="12">
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|
<emphasis id="B908EAE71807AB3DF672FEEAFC4CFEB0" box="[873,977,349,374]" italics="true" pageId="11" pageNumber="12">Variation</emphasis>
|
|
:
|
|
<specimenCount id="9D7AFD7C1807AB3DF6C4FEEAFBF4FEB0" box="[991,1129,349,375]" count="1" pageId="11" pageNumber="12" type="female">One female</specimenCount>
|
|
specimen (NCHUZOOL 14588) has frontal spines with finely granulated tip on ocularium. In addition, the female specimen (NCHUZOOL 14591) from the Dasyueshan Forest Recreation Area has slender and long penile shaft and the different morphology of penile alate part (
|
|
<figureCitation id="13472A701807AB3DF62FFDAAFC0DFDF0" box="[820,912,541,566]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5H</figureCitation>
|
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).
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|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF672FD8AFA8DFD90" blockId="11.[813,1414,253,1110]" box="[873,1296,573,599]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF672FD8AFC73FD90" box="[873,1006,573,598]" italics="true" pageId="11" pageNumber="12">Distribution</emphasis>
|
|
: Central
|
|
<collectingCountry id="F36B76651807AB3DF141FD8AFB30FD90" box="[1114,1197,573,598]" name="Taiwan" pageId="11" pageNumber="12">Taiwan</collectingCountry>
|
|
(
|
|
<figureCitation id="13472A701807AB3DF1A7FD8AFA99FD90" box="[1212,1284,573,598]" captionStart="Fig" captionStartId="13.[136,170,1897,1917]" captionTargetBox="[137,1411,281,1859]" captionTargetId="figure-6@13.[55,1478,55,2047]" captionTargetPageId="13" captionText="Fig. 7. The collection sites for the species of Metadentobunus in Taiwan. Numbers beside the symbols correspond to the collection sites in table 1 and figure 6. In addition, the numbers 12 and 13 mean the sites for Cingjing, Nantou and Lugang (= Rokko), Changhua, respectively." figureDoi="http://doi.org/10.5281/zenodo.13316546" httpUri="https://zenodo.org/record/13316546/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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|
).
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF672FDEAFC27FD70" blockId="11.[813,1414,253,1110]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF672FDEAFC5FFDB0" box="[873,962,605,630]" italics="true" pageId="11" pageNumber="12">Habitat</emphasis>
|
|
: This species can be found on the tree trunks, sometimes on the artificial wooden guideboard.
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF672FD0AFC25FB90" blockId="11.[813,1414,253,1110]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF672FD0AFC49FD10" box="[873,980,701,726]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
|
|
: This new species can be separated from other congeneric species by having one pair of shorter (than other congeners) spines at frontal margin of the ocularium (
|
|
<figureCitation id="13472A701807AB3DF17AFCAAFB34FCF0" box="[1121,1193,797,822]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
|
D-E, Q). It can be also distinguished from
|
|
<taxonomicName id="4C7C4D761807AB3DF147FC8AFAB1FC90" authorityName="Suzuki" authorityYear="1944" box="[1116,1324,829,854]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="garampiensis">
|
|
<emphasis id="B908EAE71807AB3DF147FC8AFAB1FC90" box="[1116,1324,829,854]" italics="true" pageId="11" pageNumber="12">M. garampiensis</emphasis>
|
|
</taxonomicName>
|
|
by the number of tubercles on the dorsal basal segment of the chelicera (7 vs. 14-15) (
|
|
<figureCitation id="13472A701807AB3DF194FCCAFB4CFC50" box="[1167,1233,893,918]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
|
J-K vs.
|
|
<figureCitation id="13472A701807AB3DF037FCCAFAF2FC50" box="[1324,1391,893,918]" captionStart="Fig" captionStartId="7.[136,170,1840,1860]" captionTargetBox="[220,1322,246,1812]" captionTargetId="figure-6@7.[149,1401,197,1867]" captionTargetPageId="7" captionText="Fig. 3. Metadentobunus garampiensis Suzuki, 1944. Male (neotype, NCHUZOOL 14584). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; G, H: ventral and lateral views of penis; I: different morphology of penis from Tengjhih, ventral view (NCHUZOOL 14598); J: inner side of left chelicera; K: outer side of left chelicera. Female (NCHUZOOL 14583). C: inner side of female left pedipalp; F: lateral view of ocularium; L: female seminal receptacle (NCHUZOOL 14501); M: ventral view of female genital operculum (NCHUZOOL 14583). s: stylus; g: glans; ap: alate part. Scale bars: A-C, G-K, M = 1 mm; D, E, F = 0.5 mm; L = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316536" httpUri="https://zenodo.org/record/13316536/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
|
J- K), the basal shape of the penile shaft (constricted vs. parallel) (
|
|
<figureCitation id="13472A701807AB3DF6D5FC0AFBBAFC10" box="[974,1063,957,982]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5F</figureCitation>
|
|
vs.
|
|
<figureCitation id="13472A701807AB3DF144FC0AFB5DFC11" box="[1119,1216,957,983]" captionStart="Fig" captionStartId="7.[136,170,1840,1860]" captionTargetBox="[220,1322,246,1812]" captionTargetId="figure-6@7.[149,1401,197,1867]" captionTargetPageId="7" captionText="Fig. 3. Metadentobunus garampiensis Suzuki, 1944. Male (neotype, NCHUZOOL 14584). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; G, H: ventral and lateral views of penis; I: different morphology of penis from Tengjhih, ventral view (NCHUZOOL 14598); J: inner side of left chelicera; K: outer side of left chelicera. Female (NCHUZOOL 14583). C: inner side of female left pedipalp; F: lateral view of ocularium; L: female seminal receptacle (NCHUZOOL 14501); M: ventral view of female genital operculum (NCHUZOOL 14583). s: stylus; g: glans; ap: alate part. Scale bars: A-C, G-K, M = 1 mm; D, E, F = 0.5 mm; L = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316536" httpUri="https://zenodo.org/record/13316536/files/figure.png" pageId="11" pageNumber="12">Fig. 3G</figureCitation>
|
|
), location of the seminal receptacle in ovipositor (5th-6th segment vs. 6th-7th segment) and number of ampullas in the terminal of seminal receptacle (2 vs. 3) (
|
|
<figureCitation id="13472A701807AB3DF022FBAAFA1BFBF0" box="[1337,1414,1053,1078]" captionStart="Fig" captionStartId="10.[174,208,1812,1832]" captionTargetBox="[262,1362,245,1787]" captionTargetId="figure-6@10.[176,1450,165,1863]" captionTargetPageId="10" captionText="Fig. 5. Metadentobunus brevispinus sp. nov. Male (holotype, NCHUZOOL 14599). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; F, G: ventral and lateral views of penis; J: inner side of left chelicera; K: outer side of left chelicera; L: supra-chelicera lamellae of male; H: different morphology of penis from Dasyueshan Forest Recreation Area, ventral view (NCHUZOOL 14505). Female (paratype, NCHUZOOL 14600). C: inner side of female left pedipalp; I: female seminal receptacle; M: supra-chelicera lamellae of female; N: dorsal view of left pedipalp; O: dorsal view of left pedipalp; P: ventral view of female genital operculum. s: stylus; g: glans; ap: alate part. Scale bars: A-C, J-K, F-H, L-P = 1 mm; D, E = 0.5 mm; I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316542" httpUri="https://zenodo.org/record/13316542/files/figure.png" pageId="11" pageNumber="12">Fig. 5I</figureCitation>
|
|
vs.
|
|
<figureCitation id="13472A701807AB3DF64DFB8AFC31FB90" box="[854,940,1085,1110]" captionStart="Fig" captionStartId="7.[136,170,1840,1860]" captionTargetBox="[220,1322,246,1812]" captionTargetId="figure-6@7.[149,1401,197,1867]" captionTargetPageId="7" captionText="Fig. 3. Metadentobunus garampiensis Suzuki, 1944. Male (neotype, NCHUZOOL 14584). A, B: inner and outer side of left pedipalp; D, E: dorsal and lateral views of ocularium; G, H: ventral and lateral views of penis; I: different morphology of penis from Tengjhih, ventral view (NCHUZOOL 14598); J: inner side of left chelicera; K: outer side of left chelicera. Female (NCHUZOOL 14583). C: inner side of female left pedipalp; F: lateral view of ocularium; L: female seminal receptacle (NCHUZOOL 14501); M: ventral view of female genital operculum (NCHUZOOL 14583). s: stylus; g: glans; ap: alate part. Scale bars: A-C, G-K, M = 1 mm; D, E, F = 0.5 mm; L = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13316536" httpUri="https://zenodo.org/record/13316536/files/figure.png" pageId="11" pageNumber="12">Fig. 3L</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3DF636FBCAFC43FB50" blockId="11.[813,990,1149,1174]" box="[813,990,1149,1174]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF636FBCAFC43FB50" bold="true" box="[813,990,1149,1174]" pageId="11" pageNumber="12">DNA analyses</emphasis>
|
|
</paragraph>
|
|
<paragraph id="8BC336F51807AB3AF672FB0AFDA7FEB0" blockId="11.[813,1414,1212,1558]" lastBlockId="12.[174,775,253,502]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
|
A 451 base-pair (bp) segment of 16S and 624 bp segment of COI from
|
|
<specimenCount id="9D7AFD7C1807AB3DF1ABFB6AFAFFFB30" box="[1200,1378,1245,1270]" count="19" pageId="11" pageNumber="12" type="generic">19 specimens</specimenCount>
|
|
of the genus
|
|
<taxonomicName id="4C7C4D761807AB3DF6A8FB4AFB1CFAD0" authorityName="Roewer" authorityYear="1915" box="[947,1153,1277,1302]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B908EAE71807AB3DF6A8FB4AFB1CFAD0" box="[947,1153,1277,1302]" italics="true" pageId="11" pageNumber="12">Metadentobunus</emphasis>
|
|
</taxonomicName>
|
|
and outgroups were amplified and aligned (
|
|
<tableCitation id="C6FE034E1807AB3DF142FAAAFB26FAF0" box="[1113,1211,1309,1334]" captionStart="Table 1" captionStartId="2.[174,244,991,1016]" captionText="Table 1. Haplotypes of 16S rRNA and COI genes of Metadentobunus garampiensis Suzuki, 1944, M. brevispinus sp. nov. and outgroups from Taiwan. The numbers behind localities are the collecting localities in figure 7" pageId="11" pageNumber="12">Table 1</tableCitation>
|
|
). No specimen of
|
|
<taxonomicName id="4C7C4D761807AB3DF654FA8AFC76FA90" authorityName="Roewer" authorityYear="1915" box="[847,1003,1341,1366]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="formosae">
|
|
<emphasis id="B908EAE71807AB3DF654FA8AFC76FA90" box="[847,1003,1341,1366]" italics="true" pageId="11" pageNumber="12">M. formosae</emphasis>
|
|
</taxonomicName>
|
|
was collected and therefore the species was not be included in the molecular analysis. The phylogenetic tree of the combined genes was constructed by BI (
|
|
<figureCitation id="13472A701807AB3DF1A1FA2AFA96FA70" box="[1210,1291,1437,1462]" captionStart="Fig" captionStartId="12.[174,208,1897,1917]" captionTargetBox="[173,1453,1012,1871]" captionTargetId="figure-226@12.[85,1540,957,1985]" captionTargetPageId="12" captionText="Fig. 6. A Bayesian inference (BI) tree for the species of Metadentobunus from Taiwan and outgroups, based on the combined dataset of 16S and COI genes. Probability values at the nodes represent the support values for BI and maximum likelihood (ML). Numbers behind the collection sites correspond to the sites in figure 7." figureDoi="http://doi.org/10.5281/zenodo.13316544" httpUri="https://zenodo.org/record/13316544/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
|
), and the bootstrap support values from ML analysis was also shown on the tree. It is clear that the genus
|
|
<taxonomicName id="4C7C4D761807AB3DF636FA4AFC69F9D0" authorityName="Roewer" authorityYear="1915" box="[813,1012,1533,1558]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B908EAE71807AB3DF636FA4AFC69F9D0" box="[813,1012,1533,1558]" italics="true" pageId="11" pageNumber="12">Metadentobunus</emphasis>
|
|
</taxonomicName>
|
|
is monophyletic with high support by both BI and ML methods. There are two clades within
|
|
<taxonomicName id="4C7C4D761800AB3AF5E5FEAAFE55FEF0" authorityName="Roewer" authorityYear="1915" box="[254,456,285,310]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B908EAE71800AB3AF5E5FEAAFE55FEF0" box="[254,456,285,310]" italics="true" pageId="12" pageNumber="13">Metadentobunus</emphasis>
|
|
</taxonomicName>
|
|
with high BI support, and they are referred to as
|
|
<taxonomicName id="4C7C4D761800AB3AF4CDFE8AFD38FE90" authorityName="Suzuki" authorityYear="1944" box="[470,677,317,342]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="garampiensis">
|
|
<emphasis id="B908EAE71800AB3AF4CDFE8AFD38FE90" box="[470,677,317,342]" italics="true" pageId="12" pageNumber="13">M. garampiensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C7C4D761800AB3AF7F2FE8AFEA8FEB0" authorityName="Chen & Shih" authorityYear="2018" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brevispinus" status="sp. nov.">
|
|
<emphasis id="B908EAE71800AB3AF7F2FE8AFEA8FEB0" italics="true" pageId="12" pageNumber="13">M. brevispinus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A23B579C1800AB3AF427FEEAFE01FEB0" box="[316,412,349,374]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, respectively.
|
|
</paragraph>
|
|
<caption id="DF03667D1807AB3DF593F9EAFE86F97A" pageId="11" pageNumber="12" startId="11.[136,204,1629,1654]" targetType="table">
|
|
<paragraph id="8BC336F51807AB3DF593F9EAFE86F97A" blockId="11.[136,1413,1629,1724]" pageId="11" pageNumber="12">
|
|
<emphasis id="B908EAE71807AB3DF593F9EAFF71F9B0" bold="true" box="[136,236,1629,1654]" pageId="11" pageNumber="12">Table 3.</emphasis>
|
|
Pedipalp (left) and legs (left) measurements (in mm) of the holotype male (NCHUZOOL 14599) of
|
|
<taxonomicName id="4C7C4D761807AB3DF5B1F937FD89F95F" authorityName="Chen & Shih" authorityYear="2018" box="[170,532,1664,1689]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brevispinus" status="sp. nov.">
|
|
<emphasis id="B908EAE71807AB3DF5B1F937FD89F95F" box="[170,532,1664,1689]" italics="true" pageId="11" pageNumber="12">Metadentobunus brevispinus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A23B579C1807AB3DF704F937FD1FF95F" box="[543,642,1664,1689]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
|
Measurements of female (NCHUZOOL 14600) are shown in parentheses
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BC336F51807AB3DF458F952FAFAF862" pageId="11" pageNumber="12">
|
|
<table id="F97CC455180754C9F597F952FAFAF862" box="[140,1383,1765,1956]" gridcols="7" gridrows="6" pageId="11" pageNumber="12">
|
|
<tr id="354C34B7180754C9F597F952FAFAF93F" box="[140,1383,1765,1785]" gridrow="0" pageId="11" pageNumber="12" rowspan-0="1">
|
|
<th id="769D5DCB180754C9F458F952FE34F93F" box="[323,425,1765,1785]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">Trochanter</th>
|
|
<th id="769D5DCB180754C9F4EFF952FDEFF93F" box="[500,626,1765,1785]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">Femur</th>
|
|
<th id="769D5DCB180754C9F7A7F952FCB9F93F" box="[700,804,1765,1785]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">Patella</th>
|
|
<th id="769D5DCB180754C9F675F952FC71F93F" box="[878,1004,1765,1785]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">Tibia</th>
|
|
<th id="769D5DCB180754C9F130F952FB34F93F" box="[1067,1193,1765,1785]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">Tarsus</th>
|
|
<th id="769D5DCB180754C9F1F3F952FAFAF93F" box="[1256,1383,1765,1785]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">Total</th>
|
|
</tr>
|
|
<tr id="354C34B7180754C9F597F8AAFAFAF8F7" box="[140,1383,1821,1841]" gridrow="1" pageId="11" pageNumber="12">
|
|
<th id="769D5DCB180754C9F597F8AAFF42F8F7" box="[140,223,1821,1841]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Pedipalp</th>
|
|
<td id="769D5DCB180754C9F458F8AAFE34F8F7" box="[323,425,1821,1841]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">0.32 (0.37)</td>
|
|
<td id="769D5DCB180754C9F4EFF8AAFDEFF8F7" box="[500,626,1821,1841]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">1.08 (1.18)</td>
|
|
<td id="769D5DCB180754C9F7A7F8AAFCB9F8F7" box="[700,804,1821,1841]" gridcol="3" gridrow="1" pageId="11" pageNumber="12">0.53 (0.56)</td>
|
|
<td id="769D5DCB180754C9F675F8AAFC71F8F7" box="[878,1004,1821,1841]" gridcol="4" gridrow="1" pageId="11" pageNumber="12">0.85 (0.8)</td>
|
|
<td id="769D5DCB180754C9F130F8AAFB34F8F7" box="[1067,1193,1821,1841]" gridcol="5" gridrow="1" pageId="11" pageNumber="12">1.17 (1.41)</td>
|
|
<td id="769D5DCB180754C9F1F3F8AAFAFAF8F7" box="[1256,1383,1821,1841]" gridcol="6" gridrow="1" pageId="11" pageNumber="12">3.95 (4.32)</td>
|
|
</tr>
|
|
<tr id="354C34B7180754C9F597F88EFAFAF88B" box="[140,1383,1849,1869]" gridrow="2" pageId="11" pageNumber="12">
|
|
<th id="769D5DCB180754C9F597F88EFF42F88B" box="[140,223,1849,1869]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Leg I</th>
|
|
<td id="769D5DCB180754C9F458F88EFE34F88B" box="[323,425,1849,1869]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">0.4 (0.44)</td>
|
|
<td id="769D5DCB180754C9F4EFF88EFDEFF88B" box="[500,626,1849,1869]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">7.7 (6.13)</td>
|
|
<td id="769D5DCB180754C9F7A7F88EFCB9F88B" box="[700,804,1849,1869]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">0.93 (1.02)</td>
|
|
<td id="769D5DCB180754C9F675F88EFC71F88B" box="[878,1004,1849,1869]" gridcol="4" gridrow="2" pageId="11" pageNumber="12">6.34 (5.31)</td>
|
|
<td id="769D5DCB180754C9F130F88EFB34F88B" box="[1067,1193,1849,1869]" gridcol="5" gridrow="2" pageId="11" pageNumber="12">17.52 (15.75)</td>
|
|
<td id="769D5DCB180754C9F1F3F88EFAFAF88B" box="[1256,1383,1849,1869]" gridcol="6" gridrow="2" pageId="11" pageNumber="12">32.89 (28.65)</td>
|
|
</tr>
|
|
<tr id="354C34B7180754C9F597F8E1FAFAF8AC" box="[140,1383,1878,1898]" gridrow="3" pageId="11" pageNumber="12">
|
|
<th id="769D5DCB180754C9F597F8E1FF42F8AC" box="[140,223,1878,1898]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Leg II</th>
|
|
<td id="769D5DCB180754C9F458F8E1FE34F8AC" box="[323,425,1878,1898]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">0.42 (0.44)</td>
|
|
<td id="769D5DCB180754C9F4EFF8E1FDEFF8AC" box="[500,626,1878,1898]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">14.22 (13.09)</td>
|
|
<td id="769D5DCB180754C9F7A7F8E1FCB9F8AC" box="[700,804,1878,1898]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">1.01 (1.05)</td>
|
|
<td id="769D5DCB180754C9F675F8E1FC71F8AC" box="[878,1004,1878,1898]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">13.85 (13.26)</td>
|
|
<td id="769D5DCB180754C9F130F8E1FB34F8AC" box="[1067,1193,1878,1898]" gridcol="5" gridrow="3" pageId="11" pageNumber="12">42.42 (39.74)</td>
|
|
<td id="769D5DCB180754C9F1F3F8E1FAFAF8AC" box="[1256,1383,1878,1898]" gridcol="6" gridrow="3" pageId="11" pageNumber="12">71.92 (67.58)</td>
|
|
</tr>
|
|
<tr id="354C34B7180754C9F597F8C4FAFAF841" box="[140,1383,1907,1927]" gridrow="4" pageId="11" pageNumber="12">
|
|
<th id="769D5DCB180754C9F597F8C4FF42F841" box="[140,223,1907,1927]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Leg III</th>
|
|
<td id="769D5DCB180754C9F458F8C4FE34F841" box="[323,425,1907,1927]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">0.41 (0.46)</td>
|
|
<td id="769D5DCB180754C9F4EFF8C4FDEFF841" box="[500,626,1907,1927]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">5.81 (5.62)</td>
|
|
<td id="769D5DCB180754C9F7A7F8C4FCB9F841" box="[700,804,1907,1927]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">0.92 (1.03)</td>
|
|
<td id="769D5DCB180754C9F675F8C4FC71F841" box="[878,1004,1907,1927]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">5.31 (4.46)</td>
|
|
<td id="769D5DCB180754C9F130F8C4FB34F841" box="[1067,1193,1907,1927]" gridcol="5" gridrow="4" pageId="11" pageNumber="12">17.29 (15.14)</td>
|
|
<td id="769D5DCB180754C9F1F3F8C4FAFAF841" box="[1256,1383,1907,1927]" gridcol="6" gridrow="4" pageId="11" pageNumber="12">29.74 (26.71)</td>
|
|
</tr>
|
|
<tr id="354C34B7180754C9F597F827FAFAF862" box="[140,1383,1936,1956]" gridrow="5" pageId="11" pageNumber="12">
|
|
<th id="769D5DCB180754C9F597F827FF42F862" box="[140,223,1936,1956]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Leg IV</th>
|
|
<td id="769D5DCB180754C9F458F827FE34F862" box="[323,425,1936,1956]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">0.42 (0.5)</td>
|
|
<td id="769D5DCB180754C9F4EFF827FDEFF862" box="[500,626,1936,1956]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">9.5 (9.11)</td>
|
|
<td id="769D5DCB180754C9F7A7F827FCB9F862" box="[700,804,1936,1956]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">1.28 (1.08)</td>
|
|
<td id="769D5DCB180754C9F675F827FC71F862" box="[878,1004,1936,1956]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">8.28 (6.87)</td>
|
|
<td id="769D5DCB180754C9F130F827FB34F862" box="[1067,1193,1936,1956]" gridcol="5" gridrow="5" pageId="11" pageNumber="12">27.45 (25.35)</td>
|
|
<td id="769D5DCB180754C9F1F3F827FAFAF862" box="[1256,1383,1936,1956]" gridcol="6" gridrow="5" pageId="11" pageNumber="12">46.93 (42.91)</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<footnote id="E8672AFB1807AB3DF593F86AFE05F829" box="[136,408,2013,2031]" pageId="11" pageNumber="12">
|
|
<paragraph id="8BC336F51807AB3DF593F86AFE05F829" blockId="11.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="11" pageNumber="12">© 2018 Academia Sinica, Taiwan</paragraph>
|
|
</footnote>
|
|
<paragraph id="8BC336F51800AB3AF5F1FECAFDDDFE30" blockId="12.[174,775,253,502]" pageId="12" pageNumber="13">
|
|
With regard to genetic divergence, the maximum intraspecific and the minimum interspecific COI distances of
|
|
<taxonomicName id="4C7C4D761800AB3AF464FE0AFDDBFE10" authorityName="Roewer" authorityYear="1915" box="[383,582,445,470]" class="Arachnida" family="Sclerosomatidae" genus="Metadentobunus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B908EAE71800AB3AF464FE0AFDDBFE10" box="[383,582,445,470]" italics="true" pageId="12" pageNumber="13">Metadentobunus</emphasis>
|
|
</taxonomicName>
|
|
are 2.63-5.03% and 6.79%, respectively (
|
|
<tableCitation id="C6FE034E1800AB3AF4CDFE6AFDAEFE30" box="[470,563,477,502]" captionStart="Table 4" captionStartId="12.[174,242,623,648]" captionText="Table 4. Matrix of percentage pairwise nucleotide divergences with K2P distance based on 16S rRNA and cytochrome c oxidase I (COI) genes of M. garampiensis Suzuki, 1944 and M. brevispinus sp. nov. In the right half, the lower left values are 16S and the upper right ones are COI. Values of range are shown in parentheses" pageId="12" pageNumber="13">Table 4</tableCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |