448 lines
60 KiB
XML
448 lines
60 KiB
XML
<document id="73611CC1CDF68853C01D6480CA02063D" ID-CLB-Dataset="22393" ID-DOI="10.5852/ejt.2018.438" ID-GBIF-Dataset="34ad2255-e191-4870-83e8-74819ca0fe06" ID-Zenodo-Dep="3785478" ID-ZooBank="00B57E96-225C-4646-9CB8-B3514C4EC7E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622038275" checkinUser="valdenar" docAuthor="Jia, Fenglong & Tang, Yu-dan" docDate="2018" docId="03D587848177FFB1FDEBFAD3FDB2FA00" docLanguage="en" docName="ejt-438_jia_tang.pdf.imf" docOrigin="European Journal of Taxonomy 438" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helochares atropiceus Regimbart 1903" docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="FFECFFFC817FFFBAFFC7FF81FFF9FFFF" masterDocTitle="A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="9" updateTime="1698838436565" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="9165BA92A87E0FFBA2713BE074938736">
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<mods:title id="D0BF91E565478B5348B57D201546E103">A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)</mods:title>
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<mods:name id="FFE902BB70C98B68E556EE29793A96A4" type="personal">
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<mods:namePart id="209645B0460ACE0B09AFAC818272151C">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="45A79AE59A8EF1DD6C2B0DC3F66405CB">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<mods:nameIdentifier id="1C7AE29B6760455DDDFFEB35637D2E38" type="email">lssjfl@mail.sysu.edu.cn;fenglongjia@aliyun.com</mods:nameIdentifier>
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<mods:namePart id="1997B19EA97CDED1A349FE18F5F9F6EB">Tang, Yu-dan</mods:namePart>
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<mods:affiliation id="55B0594D89F0801CD16A592547883E89">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<mods:nameIdentifier id="2155FD84F7A28DC6D51945B7F20FDA61" type="email">804586571@qq.com</mods:nameIdentifier>
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<mods:title id="6BDA48E4E999A7BD3DA9C48E6F58D3F1">European Journal of Taxonomy</mods:title>
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<mods:part id="2C5F01C31080A0727137183EAC997681">
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<mods:date id="16D57835292DF63F50D0F74D36E5E3CE">2018</mods:date>
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<mods:number id="229BEC4ED0455D9401B088346451166A">2018-05-24</mods:number>
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<mods:number id="59C89DD64EB8E45C7954107603701803">438</mods:number>
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<treatment id="03D587848177FFB1FDEBFAD3FDB2FA00" ID-DOI="http://doi.org/10.5281/zenodo.3794611" ID-GBIF-Taxon="163943954" ID-Zenodo-Dep="3794611" LSID="urn:lsid:plazi:treatment:03D587848177FFB1FDEBFAD3FDB2FA00" httpUri="http://treatment.plazi.org/id/03D587848177FFB1FDEBFAD3FDB2FA00" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<subSubSection id="C36665198177FFB2FDEBFAD3FBF1FA92" box="[556,1032,1362,1389]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BC336928177FFB2FDEBFAD3FBF1FA92" blockId="8.[556,1032,1362,1425]" box="[556,1032,1362,1389]" pageId="8" pageNumber="9">
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<heading id="D08B81FE8177FFB2FDEBFAD3FBF1FA92" box="[556,1032,1362,1389]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="4C7C4D118177FFB2FDEBFAD3FBF1FA92" ID-CoL="9P2B7" authority="Regimbart, 1903" authorityName="Regimbart" authorityYear="1903" box="[556,1032,1362,1389]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="atropiceus">
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<emphasis id="B908EA808177FFB2FDEBFAD3FCC2FA92" bold="true" box="[556,827,1362,1389]" italics="true" pageId="8" pageNumber="9">Helochares atropiceus</emphasis>
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<bibRefCitation id="EFED4B638177FFB2FC85FAD2FBF1FA92" author="Regimbart M." box="[834,1032,1363,1389]" pageId="8" pageNumber="9" pagination="52 - 64" refId="ref16408" refString="Regimbart M. 1903. Contribution a la faune Indo-Chinoise. 19 e memoire. Hydrophilidae. Annales de la Societe entomologique de France 72: 52 - 64. Available from https: // biodiversitylibrary. org / page / 8247940 [accessed 6 Apr. 2018]." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36665198177FFB2FD6BFAF7FC7EFA6E" box="[684,903,1398,1425]" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BC336928177FFB2FD6BFAF7FC7EFA6E" blockId="8.[556,1032,1362,1425]" box="[684,903,1398,1425]" pageId="8" pageNumber="9">
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<figureCitation id="13472A178177FFB2FD6BFAF7FCCBFA6E" box="[684,818,1398,1425]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Figs 18–25</figureCitation>
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,
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<figureCitation id="13472A178177FFB2FCF9FAF7FC7EFA6E" box="[830,903,1398,1425]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="8" pageNumber="9">50–51</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36665198177FFB2FF7AFA36FB43F9E7" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8BC336928177FFB2FF7AFA36FBEEFA0A" blockId="8.[189,1398,1463,1560]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB6C11BC8177FFB2FF7AFA36FBEEFA0A" pageId="8" pageNumber="9">
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<treatmentCitation id="0ADD10838177FFB2FF7AFA36FCE3FA2D" author="Regimbart M." box="[189,794,1463,1490]" page="53" pageId="8" pageNumber="9" year="1903">
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<taxonomicName id="4C7C4D118177FFB2FF7AFA36FCE3FA2D" ID-CoL="9P2B7" authority="Regimbart, 1903: 53" authorityName="Regimbart" authorityPageNumber="53" authorityYear="1903" box="[189,794,1463,1490]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="atropiceus">
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<emphasis id="B908EA808177FFB2FF7AFA36FEBAFA2E" box="[189,323,1463,1489]" italics="true" pageId="8" pageNumber="9">Helochares</emphasis>
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(s. str.)
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<emphasis id="B908EA808177FFB2FE63FA39FDE6FA2E" box="[420,543,1464,1489]" italics="true" pageId="8" pageNumber="9">atropiceus</emphasis>
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<bibRefCitation id="EFED4B638177FFB2FDE0FA36FCE3FA2D" author="Regimbart M." box="[551,794,1463,1490]" pageId="8" pageNumber="9" pagination="52 - 64" refId="ref16408" refString="Regimbart M. 1903. Contribution a la faune Indo-Chinoise. 19 e memoire. Hydrophilidae. Annales de la Societe entomologique de France 72: 52 - 64. Available from https: // biodiversitylibrary. org / page / 8247940 [accessed 6 Apr. 2018]." type="journal article" year="1903">Régimbart, 1903: 53</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(specific rank confirmed by
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<treatmentCitation id="0ADD10838177FFB2FBA9FA36FABDFA2D" author="Hebauer F." box="[1134,1348,1463,1490]" page="11" pageId="8" pageNumber="9" year="2001">
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<bibRefCitation id="EFED4B638177FFB2FBA9FA36FABDFA2D" author="Hebauer F." box="[1134,1348,1463,1490]" pageId="8" pageNumber="9" pagination="10 - 16" refId="ref15386" refString="Hebauer F. 2001. The real Helochares taprobanicus Sharp, 1890 and its allies (Coleoptera: Hydrophilidae). Lattisimus 14: 10 - 16." type="journal article" year="2001">Hebauer 2001: 11</bibRefCitation>
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||
</treatmentCitation>
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||
; not a synonym of
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||
<taxonomicName id="4C7C4D118177FFB2FE56FA5AFD1FFA0A" ID-CoL="3KGXB" authority="Sharp, 1890" authorityName="Sharp" authorityYear="1890" box="[401,742,1498,1525]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taprobanicus">
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||
<emphasis id="B908EA808177FFB2FE56FA5AFDB6FA0A" box="[401,591,1499,1525]" italics="true" pageId="8" pageNumber="9">H. taprobanicus</emphasis>
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Sharp, 1890
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</taxonomicName>
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, as in
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<treatmentCitation id="0ADD10838177FFB2FCF6FA5AFBF3FA0A" author="Hansen M." box="[817,1034,1498,1525]" page="163" pageId="8" pageNumber="9" year="1999">
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||
<bibRefCitation id="EFED4B638177FFB2FCF6FA5AFBF3FA0A" author="Hansen M." box="[817,1034,1498,1525]" pageId="8" pageNumber="9" refId="ref15262" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, Denmark." type="book" year="1999">Hansen 1999: 163</bibRefCitation>
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</treatmentCitation>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BC336928177FFB2FF7AFA7FFB43F9E7" blockId="8.[189,1398,1463,1560]" box="[189,1210,1534,1560]" pageId="8" pageNumber="9">
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||
<treatmentCitationGroup id="AB6C11BC8177FFB2FF7AFA7FFB43F9E7" box="[189,1210,1534,1560]" pageId="8" pageNumber="9">
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||
<treatmentCitation id="0ADD10838177FFB2FF7AFA7FFD01F9E7" author="Sato M." box="[189,760,1534,1560]" page="21" pageId="8" pageNumber="9" year="1976">
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<taxonomicName id="4C7C4D118177FFB2FF7AFA7FFD01F9E7" authority="Sato, 1976: 21" authorityName="Sato" authorityPageNumber="21" authorityYear="1976" box="[189,760,1534,1560]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ohkurai" subGenus="Helochares">
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<emphasis id="B908EA808177FFB2FF7AFA7FFDBFF9E7" box="[189,582,1534,1560]" italics="true" pageId="8" pageNumber="9">Helochares (Helochares) ohkurai</emphasis>
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<bibRefCitation id="EFED4B638177FFB2FD89FA7FFD01F9E7" author="Sato M." box="[590,760,1534,1560]" pageId="8" pageNumber="9" pagination="21 - 24" refId="ref16758" refString="Sato M. 1976. Two Helochares - species from the Ryukyus (Hydrophilidae). Entomological Review of Japan 29: 21 - 24." type="journal article" year="1976">Satô, 1976: 21</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(synonymyzed by
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<treatmentCitation id="0ADD10838177FFB2FC1FFA7FFB55F9E7" author="Hebauer F." box="[984,1196,1534,1560]" page="11" pageId="8" pageNumber="9" year="2001">
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||
<bibRefCitation id="EFED4B638177FFB2FC1FFA7FFB55F9E7" author="Hebauer F." box="[984,1196,1534,1560]" pageId="8" pageNumber="9" pagination="10 - 16" refId="ref15386" refString="Hebauer F. 2001. The real Helochares taprobanicus Sharp, 1890 and its allies (Coleoptera: Hydrophilidae). Lattisimus 14: 10 - 16." type="journal article" year="2001">Hebauer 2001: 11</bibRefCitation>
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</treatmentCitation>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36665198177FFB3FF7AF9BEFC4AFEB6" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BC336928177FFB2FF7AF9BEFECFF9A6" blockId="8.[189,310,1599,1625]" box="[189,310,1599,1625]" pageId="8" pageNumber="9">
|
||
<heading id="D08B81FE8177FFB2FF7AF9BEFECFF9A6" bold="true" box="[189,310,1599,1625]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
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||
<emphasis id="B908EA808177FFB2FF7AF9BEFECFF9A6" bold="true" box="[189,310,1599,1625]" pageId="8" pageNumber="9">Diagnosis</emphasis>
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</heading>
|
||
</paragraph>
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||
<paragraph id="8BC336928177FFB3FF7AF9EFFC4AFEB6" blockId="8.[189,1399,1646,2024]" lastBlockId="9.[189,1398,267,329]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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||
Size
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||
<quantity id="4C849B778177FFB2FF31F9EFFE7FF977" box="[246,390,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.7" metricValueMin="5.4" pageId="8" pageNumber="9" unit="mm" value="6.05" valueMax="6.7" valueMin="5.4">5.4–6.7 mm</quantity>
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. Dorsum with distinctly sparser and finer punctures than in
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<taxonomicName id="4C7C4D118177FFB2FBF9F9EEFB5BF977" authority="Jia & Tang, 2018" authorityName="Jia & Tang" authorityYear="2018" box="[1086,1186,1647,1672]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
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<emphasis id="B908EA808177FFB2FBF9F9EEFB5BF977" box="[1086,1186,1647,1672]" italics="true" pageId="8" pageNumber="9">H. songi</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A23B57FB8177FFB2FB6FF9EFFAFBF977" box="[1192,1282,1646,1672]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13472A178177FFB2FAD5F9EFFA91F977" box="[1298,1384,1646,1672]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
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||
). Labrum and clypeus with distinct systematic punctures. Clypeus never impressed in front of eyes (
|
||
<figureCitation id="13472A178177FFB2FF02F935FEB6F931" box="[197,335,1716,1742]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Figs 20–21</figureCitation>
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||
). Eyes not emarginate anteriorly (
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<figureCitation id="13472A178177FFB2FD1CF935FC9CF931" box="[731,869,1716,1742]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Figs 20–21</figureCitation>
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). Maxillary palpi ca 1.6× as long as width of head, each palpomere uniformly yellowish brown (
|
||
<figureCitation id="13472A178177FFB2FCCAF956FC6DF90E" box="[781,916,1751,1777]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Figs 18, 21</figureCitation>
|
||
). Antennae with pedicel distinctly shorter than antennomeres 3–6 combined. Submentum without a small glabrous area medially and with even, dense, coarse and strong punctures except extreme posterior portion (
|
||
<figureCitation id="13472A178177FFB2FBCBF89FFB9EF8C7" box="[1036,1127,1822,1848]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Fig. 22</figureCitation>
|
||
). Pronotum and elytra finely punctate. Mesoventrite with a stout, backwardly pointing projection, rising to level of mesocoxae, apex of the projection with a few long setae (
|
||
<figureCitation id="13472A178177FFB2FD2EF8E5FCBDF881" box="[745,836,1892,1918]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Fig. 24</figureCitation>
|
||
). Metaventrite without glabrous area medially. Femora densely pubescent except apical eighth (
|
||
<figureCitation id="13472A178177FFB2FD38F806FC73F85D" box="[767,906,1927,1954]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="8" pageNumber="9">Figs 19, 25</figureCitation>
|
||
). Parameres somewhat wider than widest part of median lobe, each paramere with 3 branches: inner branch short and strongly curved inwards; median branch knife-formed, sharp apically, slightly longer than outer branch; outer branch narrower than other two branches, apical third curved inwards, sharp apically. Median lobe with broad basal half, gradually narrowed apically. Basal strut very short (
|
||
<figureCitation id="13472A178176FFB3FCDBFEAFFC5FFEB6" box="[796,934,302,329]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="9" pageNumber="10">Figs 50–51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198176FFB3FF7AFEF1FDF8FB86" pageId="9" pageNumber="10" type="materials_examined">
|
||
<paragraph id="8BC336928176FFB3FF7AFEF1FE53FE75" blockId="9.[189,426,368,394]" box="[189,426,368,394]" pageId="9" pageNumber="10">
|
||
<heading id="D08B81FE8176FFB3FF7AFEF1FE53FE75" bold="true" box="[189,426,368,394]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B908EA808176FFB3FF7AFEF1FE53FE75" bold="true" box="[189,426,368,394]" pageId="9" pageNumber="10">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928176FFB3FF7AFE1EFDF8FB86" blockId="9.[189,1399,413,1145]" pageId="9" pageNumber="10">
|
||
CHINA:
|
||
<emphasis id="B908EA808176FFB3FEEFFE1FFE42FE47" bold="true" box="[296,443,414,440]" pageId="9" pageNumber="10">Guangdong</emphasis>
|
||
: 1 ♂,
|
||
<specimenCount id="9D7AFD1B8176FFB3FDC5FE1EFDD4FE46" box="[514,557,415,441]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
, Danxiashan,
|
||
<date id="FFC210528176FFB3FD14FE1EFC9EFE46" box="[723,871,415,441]" pageId="9" pageNumber="10" value="2008-05-02">2 May. 2008</date>
|
||
, Fenglong Jia leg.; 2 ♂♂,
|
||
<specimenCount id="9D7AFD1B8176FFB3FB51FE1EFB2EFE46" box="[1174,1239,415,441]" pageId="9" pageNumber="10" type="female">3 ♀♀</specimenCount>
|
||
, Danxiashan,
|
||
<date id="FFC210528176FFB3FF7AFE43FEBFFE23" box="[189,326,450,476]" pageId="9" pageNumber="10" value="2009-06-08">8 Jun. 2009</date>
|
||
, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FDE1FE43FD80FE23" box="[550,633,450,476]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shaoguan, Danxiashan,
|
||
<date id="FFC210528176FFB3FC5AFE43FBC6FE23" box="[925,1087,450,476]" pageId="9" pageNumber="10" value="2010-05-27">27 May. 2010</date>
|
||
, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FAD8FE43FA8BFE23" box="[1311,1394,450,476]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Danxiashan,
|
||
<date id="FFC210528176FFB3FE9FFE64FE17FE00" box="[344,494,485,511]" pageId="9" pageNumber="10" value="2011-07-21">21 Jul. 2011</date>
|
||
, Shuang Zhao leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FD27FE64FCCFFE00" box="[736,822,485,511]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shaoguan, Danxiashan,
|
||
<date id="FFC210528176FFB3FBA3FE64FAFDFE00" box="[1124,1284,485,511]" pageId="9" pageNumber="10" value="2012-04-21">21 Apr. 2012</date>
|
||
, Haidong Chen leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FEFAFD89FE68FDDC" box="[317,401,520,547]" pageId="9" pageNumber="10" type="generic">6 spec.</specimenCount>
|
||
, Danxiashan,
|
||
<date id="FFC210528176FFB3FDF2FD89FD05FDDD" box="[565,764,520,546]" pageId="9" pageNumber="10" value="2013-04-23" valueMax="2013-04-26" valueMin="2013-04-23">23–26 Apr. 2013</date>
|
||
, Fenglong Jia, Shuang Zhao, Weicai Xie and Haidong Chen leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FEF9FDAAFE6AFDB9" box="[318,403,555,582]" pageId="9" pageNumber="10" type="generic">3 spec.</specimenCount>
|
||
, Danxiashan nursery,
|
||
<date id="FFC210528176FFB3FD50FDAAFCDAFDBA" box="[663,803,555,582]" pageId="9" pageNumber="10" value="2012-06-08">8 Jun. 2012</date>
|
||
, light trap, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FB44FDAAFB21FDB9" box="[1155,1240,555,582]" pageId="9" pageNumber="10" type="generic">2 spec.</specimenCount>
|
||
, Danxiashan, Yangyuanshan,
|
||
<date id="FFC210528176FFB3FEBDFDCEFDE3FD96" box="[378,538,591,617]" pageId="9" pageNumber="10" value="2011-04-10">10 Apr. 2011</date>
|
||
, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FCCDFDCEFC9BFD96" box="[778,866,591,617]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Zhuhai, campus of Sun Yat-sen University,
|
||
<date id="FFC210528176FFB3FF7AFDF0FE9AFD73" box="[189,355,625,652]" pageId="9" pageNumber="10" value="2011-07-05" valueMax="2011-07-08" valueMin="2011-07-05">5–8 Jul. 2011</date>
|
||
, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FD92FDF3FD55FD73" box="[597,684,626,652]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Fengkai, Heishiding,
|
||
<date id="FFC210528176FFB3FC79FDF3FBB7FD73" box="[958,1102,626,652]" pageId="9" pageNumber="10" value="2011-06-03">3 Jun. 2011</date>
|
||
, light trap, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FF31FD14FEB1FD50" box="[246,328,661,687]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Heshan,
|
||
<date id="FFC210528176FFB3FE7FFD14FD85FD50" box="[440,636,661,687]" pageId="9" pageNumber="10" value="2002-04-22" valueMax="2002-04-24" valueMin="2002-04-22">22–24 Apr. 2002</date>
|
||
, Haidong Chen leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FCAAFD14FC46FD50" box="[877,959,661,687]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Heyuan, Wanlvhu,
|
||
<date id="FFC210528176FFB3FB6FFD14FA8BFD4F" box="[1192,1394,661,688]" pageId="9" pageNumber="10" value="2005-08-15" valueMax="2005-08-17" valueMin="2005-08-15">15–17 Aug. 2005</date>
|
||
, Shuang Zhao leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FE50FD39FE13FD2C" box="[407,490,696,723]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shenzhen, Longgang,
|
||
<date id="FFC210528176FFB3FD3FFD39FC46FD2D" box="[760,959,696,723]" pageId="9" pageNumber="10" value="2011-05-23" valueMax="2011-05-25" valueMin="2011-05-23">23–25 May 2011</date>
|
||
, Fenglong Jia leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FB58FD39FB0BFD2C" box="[1183,1266,696,723]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shenzhen, Neilingding,
|
||
<date id="FFC210528176FFB3FE9BFD5AFE1EFD0A" box="[348,487,731,757]" pageId="9" pageNumber="10" value="1998-07-04">4 Jul. 1998</date>
|
||
, Haidong Chen leg. –
|
||
<emphasis id="B908EA808176FFB3FCC6FD5DFC6DFD09" bold="true" box="[769,916,732,758]" pageId="9" pageNumber="10">
|
||
<collectingCountry id="F36B76028176FFB3FCC6FD5DFC6DFD09" box="[769,916,732,758]" name="Hong Kong" pageId="9" pageNumber="10">Hong Kong</collectingCountry>
|
||
</emphasis>
|
||
: 2 ♂♂,
|
||
<specimenCount id="9D7AFD1B8176FFB3FBC6FD5AFBBFFD09" box="[1025,1094,731,758]" pageId="9" pageNumber="10" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D7AFD1B8176FFB3FB9EFD5AFB39FD09" box="[1113,1216,731,758]" pageId="9" pageNumber="10" type="generic">34 spec.</specimenCount>
|
||
, Qingkuaitang,
|
||
<date id="FFC210528176FFB3FF7AFD7EFEA6FCE6" box="[189,351,767,793]" pageId="9" pageNumber="10" value="2013-10-29">29 Oct. 2013</date>
|
||
, light trap, Y.M. Lee, Eric Chen and Alex Lee leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FC25FD7EFBC3FCE6" box="[994,1082,767,793]" pageId="9" pageNumber="10" type="generic">9 spec.</specimenCount>
|
||
, Fengkeng,
|
||
<geoCoordinate id="EE4850558176FFB3FB08FD7CFA8BFCE6" box="[1231,1394,765,793]" degrees="22" direction="north" minutes="31.695" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="22.52825">22°31.695′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4850558176FFB3FF7AFCA1FE90FCC3" box="[189,361,800,828]" degrees="114" direction="east" minutes="13.450" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="114.22417">114°13.450′ E</geoCoordinate>
|
||
,
|
||
<quantity id="4C849B778176FFB3FEBEFCA3FE41FCC3" box="[377,440,802,828]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="10" unit="m" value="13.0">13 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC210528176FFB3FDCBFCA3FD67FCC3" box="[524,670,802,828]" pageId="9" pageNumber="10" value="2014-06-09">9 Jun. 2014</date>
|
||
, Fenglong Jia, Weicai Xie and Huangjia Chen leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FADDFCA3FA8BFCC3" box="[1306,1394,802,828]" pageId="9" pageNumber="10" type="generic">7 spec.</specimenCount>
|
||
, Dabujiao (Hexi),
|
||
<geoCoordinate id="EE4850558176FFB3FE57FCC2FDCBFCA0" box="[400,562,835,863]" degrees="22" direction="north" minutes="25.697" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="22.428284">22°25.697′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4850558176FFB3FD87FCC2FD15FCA0" box="[576,748,835,863]" degrees="114" direction="east" minutes="10.834" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="114.180565">114°10.834′ E</geoCoordinate>
|
||
,
|
||
<quantity id="4C849B778176FFB3FD3CFCC4FCB1FCA0" box="[763,840,837,863]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="9" pageNumber="10" unit="m" value="144.0">144 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC210528176FFB3FC5EFCC4FBC3FCA0" box="[921,1082,837,863]" pageId="9" pageNumber="10" value="2014-04-10">10 Apr. 2014</date>
|
||
, Fenglong Jia, Weicai Xie and Huangjia Chen leg.;
|
||
<specimenCount id="9D7AFD1B8176FFB3FE21FCE9FDC2FC7C" box="[486,571,872,899]" pageId="9" pageNumber="10" type="generic">2 spec.</specimenCount>
|
||
, Rongshu’ao,
|
||
<geoCoordinate id="EE4850558176FFB3FD22FCE7FC7FFC7C" box="[741,902,870,899]" degrees="22" direction="north" minutes="32.646" orientation="latitude" pageId="9" pageNumber="10" precision="1" value="22.5441">22°32.646′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4850558176FFB3FC54FCE7FBC4FC7C" box="[915,1085,870,899]" degrees="114" direction="east" minutes="14.926" orientation="longitude" pageId="9" pageNumber="10" precision="1" value="114.248764">114°14.926′ E</geoCoordinate>
|
||
,
|
||
<quantity id="4C849B778176FFB3FB8DFCE9FB7EFC7C" box="[1098,1159,872,899]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="10.0">10 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC210528176FFB3FB11FCE9FA8BFC7D" box="[1238,1394,872,898]" pageId="9" pageNumber="10" value="2014-04-11">11 Apr. 2014</date>
|
||
, Fenglong Jia, Weicai Xie and Huangjia Chen leg. –
|
||
<emphasis id="B908EA808176FFB3FC8BFC0AFC41FC5A" bold="true" box="[844,952,907,933]" pageId="9" pageNumber="10">Guangxi</emphasis>
|
||
:
|
||
<specimenCount id="9D7AFD1B8176FFB3FC0BFC0AFBDCFC59" box="[972,1061,907,934]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shiwandashan forest park,
|
||
<quantity id="4C849B778176FFB3FF7AFC2FFEFEFC36" box="[189,263,942,969]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="10" unit="m" value="263.0">263 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC210528176FFB3FE96FC2EFE2AFC36" box="[337,467,942,969]" pageId="9" pageNumber="10" value="2011-07-07">7 Jul. 2011</date>
|
||
, light trap, Keqing Song leg. (English label);
|
||
<specimenCount id="9D7AFD1B8176FFB3FC24FC2EFBCFFC36" box="[995,1078,943,969]" pageId="9" pageNumber="10" type="generic">2 spec.</specimenCount>
|
||
, Shiwandashan forest park,
|
||
<quantity id="4C849B778176FFB3FF7AFC53FEFFFC13" box="[189,262,978,1004]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="10" unit="m" value="263.0">263 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC210528176FFB3FE97FC53FE29FC13" box="[336,464,978,1004]" pageId="9" pageNumber="10" value="2011-07-07">7 Jul. 2011</date>
|
||
, light trap, Keqing Song leg. (transcribed from Chinese);
|
||
<specimenCount id="9D7AFD1B8176FFB3FBAFFC53FB42FC13" box="[1128,1211,978,1004]" pageId="9" pageNumber="10" type="generic">1 spec.</specimenCount>
|
||
, Shiwandashan,
|
||
<date id="FFC210528176FFB3FF7AFC74FE81FBF0" box="[189,376,1013,1039]" pageId="9" pageNumber="10" value="2013-04-08" valueMax="2013-04-12" valueMin="2013-04-08">8–12 Apr. 2013</date>
|
||
, Haidong Chen leg. (English labels). –
|
||
<emphasis id="B908EA808176FFB3FC88FC74FC42FBF0" bold="true" box="[847,955,1013,1039]" pageId="9" pageNumber="10">Guizhou</emphasis>
|
||
: 1 ♂,
|
||
<specimenCount id="9D7AFD1B8176FFB3FBCEFC74FBCFFBF0" box="[1033,1078,1013,1039]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
, Yaoshan,
|
||
<date id="FFC210528176FFB3FB7EFC74FAA2FBF0" box="[1209,1371,1013,1040]" pageId="9" pageNumber="10" value="1938-05-25">25 May 1938</date>
|
||
. –
|
||
<emphasis id="B908EA808176FFB3FF7AFB99FEE1FBCD" bold="true" box="[189,280,1048,1074]" pageId="9" pageNumber="10">Jiangxi</emphasis>
|
||
: 1 ♂, Jiulianshan, Daqiutian,
|
||
<date id="FFC210528176FFB3FDBCFB99FCE6FBCD" box="[635,799,1048,1074]" pageId="9" pageNumber="10" value="2007-08-31">31 Aug. 2007</date>
|
||
, light trap, Hong Pang leg. –
|
||
<emphasis id="B908EA808176FFB3FBBBFB99FB28FBCD" bold="true" box="[1148,1233,1048,1074]" pageId="9" pageNumber="10">
|
||
<collectingCountry id="F36B76028176FFB3FBBBFB99FB28FBCD" box="[1148,1233,1048,1074]" name="Macao" pageId="9" pageNumber="10">Macao</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D7AFD1B8176FFB3FB26FB99FAF4FBCC" box="[1249,1293,1048,1075]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
||
, Ludang city, ecological reserve, section one,
|
||
<date id="FFC210528176FFB3FDB4FBBAFCFFFBAA" box="[627,774,1083,1110]" pageId="9" pageNumber="10" value="2013-05-09">9 May 2013</date>
|
||
, Weicai Xie leg. [All data transcribed from Chinese, unless otherwise specified.]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198176FFB1FF7AFB21FDE2FC69" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BC336928176FFB3FF7AFB21FE92FB45" blockId="9.[189,363,1184,1210]" box="[189,363,1184,1210]" pageId="9" pageNumber="10">
|
||
<heading id="D08B81FE8176FFB3FF7AFB21FE92FB45" bold="true" box="[189,363,1184,1210]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B908EA808176FFB3FF7AFB21FE92FB45" bold="true" box="[189,363,1184,1210]" pageId="9" pageNumber="10">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928176FFB3FF7AFB4EFC65FA67" blockId="9.[189,1399,1230,1433]" pageId="9" pageNumber="10">
|
||
FORM AND COLOUR. Body length
|
||
<quantity id="4C849B778176FFB3FD94FB4FFD13FB16" box="[595,746,1230,1257]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.7" metricValueMin="5.4" pageId="9" pageNumber="10" unit="mm" value="6.05" valueMax="6.7" valueMin="5.4">5.4–6.7 mm</quantity>
|
||
, width
|
||
<quantity id="4C849B778176FFB3FC8BFB4EFC1AFB16" box="[844,995,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="9" pageNumber="10" unit="mm" value="3.45" valueMax="3.7" valueMin="3.2">3.2–3.7 mm</quantity>
|
||
, oval, moderately convex. Head uniformly black (
|
||
<figureCitation id="13472A178176FFB3FE4FFB73FDF7FAF3" box="[392,526,1266,1292]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 18, 21</figureCitation>
|
||
); pronotum dark brown to black, usually narrowly reddish brown laterally, posterior margins usually yellow brown laterally (
|
||
<figureCitation id="13472A178176FFB3FCCEFA94FC68FAD0" box="[777,913,1301,1327]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 21, 23</figureCitation>
|
||
), elytra black, with reddish brown lateral margins (
|
||
<figureCitation id="13472A178176FFB3FEECFAB9FE7AFAAD" box="[299,387,1336,1362]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Fig. 20</figureCitation>
|
||
); antennae, maxillary and labial palps uniformly reddish or yellowish brown (
|
||
<figureCitation id="13472A178176FFB3FAD1FAB9FF27FA8A" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 18, 21</figureCitation>
|
||
). Ventral side dark brown to black, legs dark with reddish brown tarsomeres (
|
||
<figureCitation id="13472A178176FFB3FB98FADAFB1CFA8A" box="[1119,1253,1371,1397]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 19, 24</figureCitation>
|
||
). Immatures with reddish brown pronotum and elytra and dark brown head.
|
||
</paragraph>
|
||
<paragraph id="8BC336928176FFB3FF7AFA41FB36F87F" blockId="9.[189,1399,1472,1921]" pageId="9" pageNumber="10">
|
||
HEAD. Labrum with fine and dense punctures, smooth between punctures, with distinct systematic punctures, anterior margin broadly emarginate. Clypeus broad, with coarser and sparser punctures than those on labrum, smooth between punctures, systematic punctures distinct; anterior margin clearly emarginate, at most slightly expanded in front of eyes, never impressed in front of eyes (
|
||
<figureCitation id="13472A178176FFB3FB18F9A8FA93F9BB" box="[1247,1386,1577,1604]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 20–21</figureCitation>
|
||
). Eyes of moderate size, not emarginate anteriorly (
|
||
<figureCitation id="13472A178176FFB3FCD7F9CCFC63F998" box="[784,922,1613,1639]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Figs 20–21</figureCitation>
|
||
), separated by ca 4.0× the width of one eye. Frons with as coarse and strong punctures as clypeus, with clear systematic punctures. Maxillary palps long and slender, ca 1.6× as wide as head, second and third palpomeres curved inwards, apical palpomere ca 0.85 × as long as penultimate, asymmetrical. Antennae with 9 antennomeres, scape ca 1.5× as long as pedicel, pedicel ca ¾ × as long as antennomeres 3–6 combined, club loosely segmented with dense pubescence.
|
||
<collectingRegion id="49B8F8708176FFB3FE22F97CFDB1F8E8" box="[485,584,1789,1815]" country="Myanmar" name="Chin" pageId="9" pageNumber="10">Mentum</collectingRegion>
|
||
ca as wide as long, subquadrate, with strong v-shaped impression anteromedially, with coarse punctures, with wrinkles posteriorly (
|
||
<figureCitation id="13472A178176FFB3FC07F8A1FBEEF8C5" box="[960,1047,1824,1850]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Fig. 22</figureCitation>
|
||
). Submentum without a small glabrous area medially and with even, dense, coarse and strong punctures, smooth between punctures (
|
||
<figureCitation id="13472A178176FFB3FF01F8E7FEE7F87F" box="[198,286,1894,1920]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Fig. 22</figureCitation>
|
||
). Maxilla with a few punctures that are finer than those on submentum (
|
||
<figureCitation id="13472A178176FFB3FBADF8E7FB3BF87F" box="[1130,1218,1894,1920]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="9" pageNumber="10">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC336928176FFB1FF7AF829FD61FD7A" blockId="9.[189,1398,1960,2021]" lastBlockId="11.[189,1399,267,645]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
|
||
THORAX. Pronotum ca 2 × as wide as long, widest posteriorly, posterolateral margin broadly round, anterior and posterior margins smooth, with a very fine transverse groove, lateral margin with stronger groove; posterior margin slightly bisinuate, with median portion widely curved, punctures slightly coarser than those on head, with distinct setiferous systematic punctures (
|
||
<figureCitation id="13472A178174FFB1FBE4FEAFFB23FEB6" box="[1059,1242,302,329]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Figs 20–21, 23</figureCitation>
|
||
). Prosternum bulging in middle, not carinate, pubescent, with transverse groove, protruding anteriorly (
|
||
<figureCitation id="13472A178174FFB1FACBFED3FA90FE93" box="[1292,1385,338,364]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
|
||
). Mesoventrite with a stout, backwardly pointing projection, rising to level of mesocoxae, apex of projection with a few long setae (
|
||
<figureCitation id="13472A178174FFB1FD87FE19FD6EFE4D" box="[576,663,408,434]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
|
||
). Metaventrite without glabrous area medially; metepisterna ca 3.5× as long as wide, subparallel. Scutellum triangular, with similar punctures to those on elytra. Ground punctures on elytra similar to those on pronotum and clearly with three distinct rows of systematic punctures (
|
||
<figureCitation id="13472A178174FFB1FEF9FD83FE6CFDE3" box="[318,405,514,540]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
|
||
); epipleura very broad anteriorly, reaching posterior margin of first visible abdominal sternite, with sparse, coarse punctures. Femora densely pubescent with apical eighth glabrous (
|
||
<figureCitation id="13472A178174FFB1FAD1FDA4FF27FD9C" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Figs 19, 25</figureCitation>
|
||
). Metatarsomeres with dense golden hairs ventrally and some long swimming hairs dorsally. Claws of moderate size, rather strongly curved.
|
||
</paragraph>
|
||
<caption id="DF03661A8175FFB0FF7AF80BFE9AF814" ID-DOI="http://doi.org/10.5281/zenodo.3785486" ID-Zenodo-Dep="3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,241,1930,1956]" targetBox="[250,1325,267,1876]" targetPageId="10">
|
||
<paragraph id="8BC336928175FFB0FF7AF80BFE9AF814" blockId="10.[189,1399,1930,2027]" pageId="10" pageNumber="11">
|
||
<emphasis id="B908EA808175FFB0FF7AF80BFEBFF85B" bold="true" box="[189,326,1930,1956]" pageId="10" pageNumber="11">Figs 18–25.</emphasis>
|
||
<taxonomicName id="4C7C4D118175FFB0FE8BF80BFCE4F85B" authority="Regimbart, 1903" authorityName="Regimbart" authorityYear="1903" box="[332,797,1930,1956]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atropiceus">
|
||
<emphasis id="B908EA808175FFB0FE8BF80BFDABF85B" box="[332,594,1930,1956]" italics="true" pageId="10" pageNumber="11">Helochares atropiceus</emphasis>
|
||
<bibRefCitation id="EFED4B638175FFB0FD90F80BFCE4F85B" author="Regimbart M." box="[599,797,1930,1956]" pageId="10" pageNumber="11" pagination="52 - 64" refId="ref16408" refString="Regimbart M. 1903. Contribution a la faune Indo-Chinoise. 19 e memoire. Hydrophilidae. Annales de la Societe entomologique de France 72: 52 - 64. Available from https: // biodiversitylibrary. org / page / 8247940 [accessed 6 Apr. 2018]." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B908EA808175FFB0FCE1F80BFC89F85B" bold="true" box="[806,880,1930,1956]" pageId="10" pageNumber="11">18–20</emphasis>
|
||
. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral.
|
||
<emphasis id="B908EA808175FFB0FF7AF82CFF23F838" bold="true" box="[189,218,1965,1991]" pageId="10" pageNumber="11">21</emphasis>
|
||
. Head and pronotum.
|
||
<emphasis id="B908EA808175FFB0FE3EF82CFDEFF838" bold="true" box="[505,534,1965,1991]" pageId="10" pageNumber="11">22</emphasis>
|
||
. Head and prosternum.
|
||
<emphasis id="B908EA808175FFB0FC8FF82CFC9CF838" bold="true" box="[840,869,1965,1991]" pageId="10" pageNumber="11">23</emphasis>
|
||
. Pronotum, dorsolateral.
|
||
<emphasis id="B908EA808175FFB0FB65F82CFB46F838" bold="true" box="[1186,1215,1965,1991]" pageId="10" pageNumber="11">24</emphasis>
|
||
. Mesoventrite.
|
||
<emphasis id="B908EA808175FFB0FF7AF850FF23F814" bold="true" box="[189,218,2001,2027]" pageId="10" pageNumber="11">25</emphasis>
|
||
. Abdomen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC336928174FFB1FF7AFD2CFAA2FD37" blockId="11.[189,1371,685,712]" box="[189,1371,685,712]" pageId="11" pageNumber="12">
|
||
ABDOMEN. All visible abdominal ventrites with dense pubescence; first ventrite not carinate (
|
||
<figureCitation id="13472A178174FFB1FB31FD2CFAB7FD37" box="[1270,1358,685,712]" captionStart="Figs 18–25" captionStartId="10.[189,241,1930,1956]" captionTargetBox="[250,1325,267,1876]" captionTargetId="figure@10.[202,1384,253,1890]" captionTargetPageId="10" captionText="Figs 18–25. Helochares atropiceus Régimbart, 1903. 18–20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785486" httpUri="https://zenodo.org/record/3785486/files/figure.png" pageId="11" pageNumber="12">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AFD6FFDE2FC69" blockId="11.[189,1399,750,918]" pageId="11" pageNumber="12">
|
||
AEDEAGUS. Very broad. Parameres somewhat wider than widest part of median lobe, each paramere with 3 branches: inner branch short and strongly curved inwards; median branch knife-formed, sharp apically, almost as long as outer branch; outer branch narrower than other two branches, apical third curved inwards, sharp apically. Median lobe with broad basal half, gradually narrowed apically. Basal strut very short (
|
||
<figureCitation id="13472A178174FFB1FEB9FCFAFDF7FC69" box="[382,526,891,918]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="11" pageNumber="12">Figs 50– 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198174FFB1FF7AFC3DFE84FB6D" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BC336928174FFB1FF7AFC3DFE30FC29" blockId="11.[189,457,956,982]" box="[189,457,956,982]" pageId="11" pageNumber="12">
|
||
<heading id="D08B81FE8174FFB1FF7AFC3DFE30FC29" bold="true" box="[189,457,956,982]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B908EA808174FFB1FF7AFC3DFE30FC29" bold="true" box="[189,457,956,982]" pageId="11" pageNumber="12">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AFC6DFE84FB6D" blockId="11.[189,1399,1003,1170]" pageId="11" pageNumber="12">
|
||
It is very easy to distinguish this species from other species known from
|
||
<collectingCountry id="F36B76028174FFB1FBEFFC6AFB97FBFA" box="[1064,1134,1003,1029]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
|
||
by its large size, dark brown to black colour, pronotum and elytra finely punctate, mesoventrite with a stout, backwardly pointing projection, rising to the level of the mesocoxae, the apex of the projection with a few long setae, each paramere with 3 branches. For the differences between
|
||
<taxonomicName id="4C7C4D118174FFB1FC09FBD4FB95FB91" authorityName="Regimbart" authorityYear="1903" box="[974,1132,1109,1134]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atropiceus">
|
||
<emphasis id="B908EA808174FFB1FC09FBD4FB95FB91" box="[974,1132,1109,1134]" italics="true" pageId="11" pageNumber="12">H. atropiceus</emphasis>
|
||
</taxonomicName>
|
||
and allied species, see
|
||
<bibRefCitation id="EFED4B638174FFB1FF7AFBF9FE83FB6D" author="Hebauer F." box="[189,378,1144,1170]" pageId="11" pageNumber="12" pagination="10 - 16" refId="ref15386" refString="Hebauer F. 2001. The real Helochares taprobanicus Sharp, 1890 and its allies (Coleoptera: Hydrophilidae). Lattisimus 14: 10 - 16." type="journal article" year="2001">Hebauer (2001)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198174FFB1FF7AFB38FAADFAB7" pageId="11" pageNumber="12" type="biology_ecology">
|
||
<paragraph id="8BC336928174FFB1FF7AFB38FEE5FB2C" blockId="11.[189,284,1209,1235]" box="[189,284,1209,1235]" pageId="11" pageNumber="12">
|
||
<heading id="D08B81FE8174FFB1FF7AFB38FEE5FB2C" bold="true" box="[189,284,1209,1235]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B908EA808174FFB1FF7AFB38FEE5FB2C" bold="true" box="[189,284,1209,1235]" pageId="11" pageNumber="12">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AFB69FAADFAB7" blockId="11.[189,1398,1256,1353]" pageId="11" pageNumber="12">
|
||
Living in natural ponds with leaf litter or water grass, can sometimes be collected on wet ground with plenty of grass. It can be collected at light in May and June in South
|
||
<collectingCountry id="F36B76028174FFB1FC2FFA8AFBCBFADA" box="[1000,1074,1291,1317]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
|
||
. It has never been collected from the edges of rivers and streams. The female carries the egg case under the abdominal ventrites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198174FFB1FF7AFAEEFDB2FA00" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8BC336928174FFB1FF7AFAEEFEAFFA76" blockId="11.[189,342,1391,1417]" box="[189,342,1391,1417]" pageId="11" pageNumber="12">
|
||
<heading id="D08B81FE8174FFB1FF7AFAEEFEAFFA76" bold="true" box="[189,342,1391,1417]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B908EA808174FFB1FF7AFAEEFEAFFA76" bold="true" box="[189,342,1391,1417]" pageId="11" pageNumber="12">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AFA1FFDB2FA00" blockId="11.[189,1399,1438,1535]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="F36B76028174FFB1FF7AFA1FFEFAFA47" box="[189,259,1438,1464]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49B8F8708174FFB1FEDFFA1FFE5FFA47" box="[280,422,1438,1464]" country="China" name="Guangdong" pageId="11" pageNumber="12">Guangdong</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B8F8708174FFB1FE70FA1FFDE6FA47" box="[439,543,1438,1464]" country="China" name="Guangxi" pageId="11" pageNumber="12">Guangxi</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B8F8708174FFB1FDF7FA1FFD61FA47" box="[560,664,1438,1464]" country="China" name="Guizhou" pageId="11" pageNumber="12">Guizhou</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36B76028174FFB1FD6DFA1EFCC5FA46" box="[682,828,1439,1465]" name="Hong Kong" pageId="11" pageNumber="12">Hong Kong</collectingCountry>
|
||
, Jiangxi,
|
||
<collectingCountry id="F36B76028174FFB1FC71FA1EFBF3FA46" box="[950,1034,1439,1465]" name="Macao" pageId="11" pageNumber="12">Macau</collectingCountry>
|
||
),
|
||
<collectingCountry id="F36B76028174FFB1FBE3FA1FFB49FA47" box="[1060,1200,1438,1464]" name="Bangladesh" pageId="11" pageNumber="12">Bangladesh</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028174FFB1FB06FA1EFAFDFA46" box="[1217,1284,1439,1465]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
|
||
(Ryukyu Islands),
|
||
<collectingCountry id="F36B76028174FFB1FEEAFA40FE65FA24" box="[301,412,1473,1499]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028174FFB1FE6BFA40FE0CFA24" box="[428,501,1473,1499]" name="Nepal" pageId="11" pageNumber="12">Nepal</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028174FFB1FDC2FA40FD89FA24" box="[517,624,1473,1499]" name="Thailand" pageId="11" pageNumber="12">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028174FFB1FD46FA40FD1DFA24" box="[641,740,1473,1499]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFED4B638174FFB1FD3FFA40FC14FA23" author="Fikacek M. & Angus R. B. & Gentili E. & Jia F. & Minoshima Y. N. & Prokin A. & Przevozny M. & Ryndevich S. K." box="[760,1005,1473,1500]" pageId="11" pageNumber="12" pagination="1" refId="ref15018" refString="Fikacek M., Angus R. B., Gentili E., Jia F., Minoshima Y. N., Prokin A., Przevozny M. & Ryndevich S. K. 2015. Helophoridae, Hydrochidae, Hydrophilidae. In: Lobl I. & Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 (revised and updated edition), Hydrophiloidea - Staphylinoidea. Brill, Leiden, Boston." type="book chapter" year="2015">
|
||
Fikáček
|
||
<emphasis id="B908EA808174FFB1FCA4FA43FC5BFA24" box="[867,930,1473,1499]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B638174FFB1FC39FA43FB58FA24" author="Hansen M." box="[1022,1185,1473,1500]" pageId="11" pageNumber="12" refId="ref15262" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, Denmark." type="book" year="1999">Hansen 1999</bibRefCitation>
|
||
). New record for Guizhou,
|
||
<collectingCountry id="F36B76028174FFB1FEF6FA64FE43FA00" box="[305,442,1509,1535]" name="Hong Kong" pageId="11" pageNumber="12">Hong Kong</collectingCountry>
|
||
and
|
||
<collectingCountry id="F36B76028174FFB1FE34FA64FDBEFA00" box="[499,583,1509,1535]" name="Macao" pageId="11" pageNumber="12">Macau</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |