214 lines
37 KiB
XML
214 lines
37 KiB
XML
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<mods:title id="D5BF1873FE5A4CAA9BBD64CC573ADCB1">Rhomeocalpsua torosa gen. et sp. nov., a unique lineage of Endomychidae from mid-Cretaceous Burmese amber (Coleoptera: Coccinelloidea)</mods:title>
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<mods:namePart id="3A60F07BC338840EB465DD64A3E38963">Li, Yan-da</mods:namePart>
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<mods:namePart id="F3549EE1F1518E3BA78DC631806B19BF">Tomaszewska, K. Wioletta</mods:namePart>
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<mods:namePart id="8E95592922721F71D27B801BB0E67E5F">Huang, Di-ying</mods:namePart>
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<mods:namePart id="C55188C72F971A30E284666CDCA27D6F">Cai, Chen-yang</mods:namePart>
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<mods:title id="37AD08B9C781D7644AEC0C03BBCF1372">Palaeoentomology</mods:title>
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<mods:date id="3B5790335E7A0CC1C8E01E7C4717A87D">2022</mods:date>
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<mods:number id="31863823B642CB8C53CA9D9006380110">2022-04-29</mods:number>
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<treatment id="03C887E8C45D7C3BDAB0FC53BA2AF88D" ID-DOI="http://doi.org/10.5281/zenodo.6533049" ID-GBIF-Taxon="195277873" ID-Zenodo-Dep="6533049" LSID="urn:lsid:plazi:treatment:03C887E8C45D7C3BDAB0FC53BA2AF88D" httpUri="http://treatment.plazi.org/id/03C887E8C45D7C3BDAB0FC53BA2AF88D" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148">
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<subSubSection id="C37B6575C45D7C3ADAB0FC53BBA4FBE8" pageId="2" pageNumber="148" type="nomenclature">
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<paragraph id="8BDE36FEC45D7C3ADAB0FC53BBA4FBE8" blockId="2.[136,777,971,1072]" pageId="2" pageNumber="148">
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<heading id="D0968192C45D7C3ADAB0FC53BBA4FBE8" bold="true" centered="true" fontSize="10" level="2" pageId="2" pageNumber="148" reason="3">
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<emphasis id="B915EAECC45D7C3ADAB0FC53BBA4FBE8" bold="true" pageId="2" pageNumber="148">
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<taxonomicName id="4C614D7DC45D7C3ADAB0FC53B9C8FC06" authority="Li, Tomaszewska & Cai" authorityName="Li, Tomaszewska & Cai" authorityYear="2022" box="[136,725,971,998]" class="Insecta" family="Endomychidae" genus="Rhomeocalpsua" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="148" phylum="Arthropoda" rank="species" species="torosa" status="sp. nov.">
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<emphasis id="B915EAECC45D7C3ADAB0FC53BA8AFC05" bold="true" box="[136,407,971,998]" italics="true" pageId="2" pageNumber="148">Rhomeocalpsua torosa</emphasis>
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Li, Tomaszewska & Cai
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2265797C45D7C3AD8D0FC54BBA4FBE8" pageId="2" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C37B6575C45D7C3ADAB0FB8EBBE1FBD3" box="[136,252,1045,1072]" pageId="2" pageNumber="148" type="description">
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<paragraph id="8BDE36FEC45D7C3ADAB0FB8EBBE1FBD3" blockId="2.[136,777,971,1072]" box="[136,252,1045,1072]" pageId="2" pageNumber="148">
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(
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<figureCitation id="135A2A7BC45D7C3ADAA8FB8EBBEFFBD3" box="[144,242,1045,1072]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[136,235,2015,2039]" captionStartId-1="2.[136,235,2009,2033]" captionStartId-2="3.[136,235,1157,1181]" captionStartId-3="4.[136,235,1533,1557]" captionStartId-4="5.[136,235,1428,1452]" captionTargetBox-0="[215,1372,1047,1992]" captionTargetBox-1="[215,1372,1328,1985]" captionTargetBox-2="[215,1372,189,1134]" captionTargetBox-3="[145,1443,198,1509]" captionTargetBox-4="[151,1436,189,1405]" captionTargetId-0="figure-334@1.[215,1372,1047,1992]" captionTargetId-1="figure-390@2.[215,1372,1328,1985]" captionTargetId-2="figure-312@3.[215,1372,189,1134]" captionTargetId-3="figure-132@4.[145,1443,198,1509]" captionTargetId-4="figure-217@5.[151,1436,189,1405]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="FIGURE 1. General habitus of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 500 μm." captionText-1="FIGURE 2. General habitus of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars = 500 μm." captionText-2="FIGURE 3. General habitus of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars = 500 μm." captionText-3="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." captionText-4="FIGURE 5. X-ray microtomographic reconstruction of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054. A, Dorsal view. B, Ventral view. C, Anterodorsal view. D, Lateral view. E, Posterior view. F, Antenna, rendered under “Sum along Ray” mode. Scale bars = 500 μm in A–E, 200 μm in F." figureDoi-0="http://doi.org/10.5281/zenodo.6530505" figureDoi-1="http://doi.org/10.5281/zenodo.6530509" figureDoi-2="http://doi.org/10.5281/zenodo.6530511" figureDoi-3="http://doi.org/10.5281/zenodo.6530513" figureDoi-4="http://doi.org/10.5281/zenodo.6530517" httpUri-0="https://zenodo.org/record/6530505/files/figure.png" httpUri-1="https://zenodo.org/record/6530509/files/figure.png" httpUri-2="https://zenodo.org/record/6530511/files/figure.png" httpUri-3="https://zenodo.org/record/6530513/files/figure.png" httpUri-4="https://zenodo.org/record/6530517/files/figure.png" pageId="2" pageNumber="148">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6575C45D7C3ADAB0FBC7B9A9FB99" box="[136,692,1119,1146]" pageId="2" pageNumber="148" type="materials_examined">
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<paragraph id="8BDE36FEC45D7C3ADAB0FBC7B9A9FB99" blockId="2.[136,778,1119,1294]" box="[136,692,1119,1146]" pageId="2" pageNumber="148">
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<emphasis id="B915EAECC45D7C3ADAB0FBC7BBEAFB9A" bold="true" box="[136,247,1119,1145]" pageId="2" pageNumber="148">Material.</emphasis>
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<materialsCitation id="3B093CA3C45D7C3ADAC6FBF8B9A9FB99" ID-GBIF-Occurrence="3763883301" box="[254,692,1119,1146]" collectionCode="NIGP" pageId="2" pageNumber="148" specimenCode="NIGP180054" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54DA885CC45D7C3ADAC6FBF8BA74FB99" box="[254,361,1120,1146]" pageId="2" pageNumber="148" type="holotype">Holotype</typeStatus>
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, sex unknown,
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<specimenCode id="DBC79E85C45D7C3AD82FFBC7B9ADFB99" box="[535,688,1119,1146]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="2" pageNumber="148">NIGP180054</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6575C45D7C3ADA85FB1DBE48FF3E" pageId="2" pageNumber="148" type="etymology">
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<paragraph id="8BDE36FEC45D7C3ADA85FB1DBA90FB27" blockId="2.[136,778,1119,1294]" pageId="2" pageNumber="148">
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<emphasis id="B915EAECC45D7C3ADA85FB1DBA59FB7C" bold="true" box="[189,324,1157,1183]" pageId="2" pageNumber="148">Etymology.</emphasis>
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The specific name refers to its distinctly enlarged antennal club.
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</paragraph>
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<paragraph id="8BDE36FEC45D7C3ADA85FB56BE48FF3E" blockId="2.[136,778,1119,1294]" lastBlockId="2.[810,1452,158,1294]" pageId="2" pageNumber="148">
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<emphasis id="B915EAECC45D7C3ADA85FB56BAD1FB0B" bold="true" box="[189,460,1230,1257]" pageId="2" pageNumber="148">Locality and horizon.</emphasis>
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Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
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<collectingRegion id="49A5F81CC45D7C3AD912FF06B8CDFF5B" box="[810,976,158,184]" country="Myanmar" name="Kachin" pageId="2" pageNumber="148">Kachin State</collectingRegion>
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,
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<collectingCountry id="F376766EC45D7C3AD9D7FF07BF7DFF5A" box="[1007,1120,159,185]" name="Myanmar" pageId="2" pageNumber="148">Myanmar</collectingCountry>
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; unnamed horizon, mid- Cretaceous, Upper Albian to Lower Cenomanian.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6575C45D7C3AD967FF70BF83FEE0" box="[863,1182,232,259]" pageId="2" pageNumber="148" type="diagnosis">
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<paragraph id="8BDE36FEC45D7C3AD967FF70BF83FEE0" blockId="2.[810,1452,158,1294]" box="[863,1182,232,259]" pageId="2" pageNumber="148">
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<emphasis id="B915EAECC45D7C3AD967FF70B8C4FEE1" bold="true" box="[863,985,232,258]" pageId="2" pageNumber="148">Diagnosis.</emphasis>
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As for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6575C45D7C3BD967FE95BA2AF88D" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148" type="description">
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<paragraph id="8BDE36FEC45D7C3AD967FE95B8CFFE91" blockId="2.[810,1452,158,1294]" pageId="2" pageNumber="148">
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<emphasis id="B915EAECC45D7C3AD967FE95B8ECFEC4" bold="true" box="[863,1009,269,295]" pageId="2" pageNumber="148">Description.</emphasis>
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Body relatively elongate, rather flattened, about
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<quantity id="4C999B1BC45D7C3AD9DBFEAABF26FEAE" box="[995,1083,306,333]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="148" unit="mm" value="1.3">1.3 mm</quantity>
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long,
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<quantity id="4C999B1BC45D7C3ADEBEFEAABFC3FEAE" box="[1158,1246,306,333]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="148" unit="mm" value="0.5">0.5 mm</quantity>
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wide, moderately densely setose.
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</paragraph>
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<paragraph id="8BDE36FEC45D7C3AD967FEE4B88CFC94" blockId="2.[810,1452,158,1294]" pageId="2" pageNumber="148">
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Head prognathous, not retracted into prothorax. Eyes well developed, moderately protuberant, coarsely facetted, without interfacetal setae (
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<figureCitation id="135A2A7BC45D7C3ADE6CFE5EBFCFFE02" box="[1108,1234,454,481]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Fig. 4A, D</figureCitation>
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). Antennal grooves absent (
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<figureCitation id="135A2A7BC45D7C3AD9B9FE73B8E0FDE5" box="[897,1021,491,518]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Fig. 4B, D</figureCitation>
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). Antennal sockets visible from above (
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<figureCitation id="135A2A7BC45D7C3AD90AFD88B890FDC8" box="[818,909,528,555]" captionStart="FIGURE 5" captionStartId="5.[136,235,1428,1452]" captionTargetBox="[151,1436,189,1405]" captionTargetId="figure-217@5.[151,1436,189,1405]" captionTargetPageId="5" captionText="FIGURE 5. X-ray microtomographic reconstruction of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054. A, Dorsal view. B, Ventral view. C, Anterodorsal view. D, Lateral view. E, Posterior view. F, Antenna, rendered under “Sum along Ray” mode. Scale bars = 500 μm in A–E, 200 μm in F." figureDoi="http://doi.org/10.5281/zenodo.6530517" httpUri="https://zenodo.org/record/6530517/files/figure.png" pageId="2" pageNumber="148">Fig. 5C</figureCitation>
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). Antennae (
|
||
<figureCitation id="135A2A7BC45D7C3ADE26FD88BFBCFDC8" box="[1054,1185,528,555]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Figs 4C, D</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A7BC45D7C3ADE97FD88BFD0FDC8" box="[1199,1229,528,555]" captionStart="FIGURE 5" captionStartId="5.[136,235,1428,1452]" captionTargetBox="[151,1436,189,1405]" captionTargetId="figure-217@5.[151,1436,189,1405]" captionTargetPageId="5" captionText="FIGURE 5. X-ray microtomographic reconstruction of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054. A, Dorsal view. B, Ventral view. C, Anterodorsal view. D, Lateral view. E, Posterior view. F, Antenna, rendered under “Sum along Ray” mode. Scale bars = 500 μm in A–E, 200 μm in F." figureDoi="http://doi.org/10.5281/zenodo.6530517" httpUri="https://zenodo.org/record/6530517/files/figure.png" pageId="2" pageNumber="148">5F</figureCitation>
|
||
) composed of nine antennomeres; antennomere 2 oblong; antennomeres 3–6 moniliform; antennomeres 7–9 forming a distinct club; antennomere 7 strongly asymmetrical, short and strongly transverse; antennomere 8 strongly enlarged. Maxillary palps (
|
||
<figureCitation id="135A2A7BC45D7C3AD947FD51B8C2FD00" box="[895,991,713,739]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Fig. 4B</figureCitation>
|
||
) 4-segmented, palpomere 1 smallest; palpomeres 2–4 narrowing progressively; palpomere 4 about as long as 2 and 3 combined, conical. Labial palps (
|
||
<figureCitation id="135A2A7BC45D7C3AD90AFCA0B894FCB1" box="[818,905,824,850]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Fig. 4B</figureCitation>
|
||
) 3-segmented, with apical palpomere apparently enlarged.
|
||
</paragraph>
|
||
<paragraph id="8BDE36FEC45D7C3AD967FC1AB8BCFBD3" blockId="2.[810,1452,158,1294]" pageId="2" pageNumber="148">
|
||
Pronotum (
|
||
<figureCitation id="135A2A7BC45D7C3AD9DFFC1ABF57FC7F" box="[999,1098,898,924]" captionStart="FIGURE 3" captionStartId="3.[136,235,1157,1181]" captionTargetBox="[215,1372,189,1134]" captionTargetId="figure-312@3.[215,1372,189,1134]" captionTargetPageId="3" captionText="FIGURE 3. General habitus of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.6530511" httpUri="https://zenodo.org/record/6530511/files/figure.png" pageId="2" pageNumber="148">Fig. 3A</figureCitation>
|
||
) transverse, widest near the middle, narrower at base than elytral bases; lateral pronotal carinae complete, without raised margin (
|
||
<figureCitation id="135A2A7BC45D7C3ADF46FC54B852FBEF" captionStart="FIGURE 5" captionStartId="5.[136,235,1428,1452]" captionTargetBox="[151,1436,189,1405]" captionTargetId="figure-217@5.[151,1436,189,1405]" captionTargetPageId="5" captionText="FIGURE 5. X-ray microtomographic reconstruction of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054. A, Dorsal view. B, Ventral view. C, Anterodorsal view. D, Lateral view. E, Posterior view. F, Antenna, rendered under “Sum along Ray” mode. Scale bars = 500 μm in A–E, 200 μm in F." figureDoi="http://doi.org/10.5281/zenodo.6530517" httpUri="https://zenodo.org/record/6530517/files/figure.png" pageId="2" pageNumber="148">Fig. 5D</figureCitation>
|
||
); anterior angles not produced; disc without pits, sulci or carinae.
|
||
</paragraph>
|
||
<paragraph id="8BDE36FEC45D7C3BD967FBA3BA1EF9FE" blockId="2.[810,1452,158,1294]" lastBlockId="3.[136,778,1277,1902]" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148">
|
||
Elytra about 1.8 times as long as width combined, completely covering abdomen; sides weakly rounded; disc irregularly punctate; epipleura comparatively narrow, incomplete towards apices. Mesocoxal cavities circular, moderately widely separated, laterally broadly closed by meeting of meso- and metaventrites (
|
||
<figureCitation id="135A2A7BC45D7C3ADF7DFB6CBE82FAED" box="[1349,1439,1268,1294]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="2" pageNumber="148">Fig. 4E</figureCitation>
|
||
); mesotrochantins concealed; mesoventral process narrower than coxal cavity, narrowing between coxae towards apex, without apparent carinae or grooves (
|
||
<figureCitation id="135A2A7BC45C7C3BD8A6FAD0B9E3FA81" box="[670,766,1352,1378]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4E</figureCitation>
|
||
). Mesometaventral junction as straight line with meso- and metaventrites contiguous (
|
||
<figureCitation id="135A2A7BC45C7C3BDB89FA0BB917FA4E" box="[433,522,1427,1453]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4E</figureCitation>
|
||
). Metaventrite without postcoxal lines; postcoxal pits not apparent. Metacoxae (
|
||
<figureCitation id="135A2A7BC45C7C3BDAA8FA46BBFAFA1B" box="[144,231,1502,1528]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4F</figureCitation>
|
||
) widely separated, transverse. Hind wings well developed.
|
||
</paragraph>
|
||
<caption id="DF1E6676C45D7C3ADAB0F841B8BDF7F5" ID-DOI="http://doi.org/10.5281/zenodo.6530509" ID-Zenodo-Dep="6530509" httpUri="https://zenodo.org/record/6530509/files/figure.png" pageId="2" pageNumber="148" startId="2.[136,235,2009,2033]" targetBox="[215,1372,1328,1985]" targetPageId="2">
|
||
<paragraph id="8BDE36FEC45D7C3ADAB0F841B8BDF7F5" blockId="2.[136,1452,2009,2070]" pageId="2" pageNumber="148">
|
||
<emphasis id="B915EAECC45D7C3ADAB0F841BA18F812" bold="true" box="[136,261,2009,2033]" pageId="2" pageNumber="148">FIGURE 2.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="4C614D7DC45D7C3ADBE9F841B8A1F812" authority="Li, Tomaszewska & Cai" authorityName="Li, Tomaszewska & Cai" authorityYear="2022" box="[465,956,2009,2034]" class="Insecta" family="Endomychidae" genus="Rhomeocalpsua" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="148" phylum="Arthropoda" rank="species" species="torosa" status="gen. et sp. nov.">
|
||
<emphasis id="B915EAECC45D7C3ADBE9F841B9A6F812" box="[465,699,2009,2033]" italics="true" pageId="2" pageNumber="148">Rhomeocalpsua torosa</emphasis>
|
||
Li, Tomaszewska & Cai
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B915EAECC45D7C3AD9FFF842BF75F811" bold="true" box="[967,1128,2010,2034]" pageId="2" pageNumber="148">
|
||
<taxonomicNameLabel id="A2265797C45D7C3AD9FFF842BF75F811" box="[967,1128,2010,2034]" pageId="2" pageNumber="148" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, NIGP180054, under widefield fluorescence.
|
||
<emphasis id="B915EAECC45D7C3ADB43F865BA90F7F6" bold="true" box="[379,397,2045,2069]" pageId="2" pageNumber="148">A</emphasis>
|
||
, Dorsal view.
|
||
<emphasis id="B915EAECC45D7C3AD818F866B92CF7F5" bold="true" box="[544,561,2046,2070]" pageId="2" pageNumber="148">B</emphasis>
|
||
, Ventral view. Scale bars = 500 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1E6676C45C7C3BDAB0FB1DB891FB21" ID-DOI="http://doi.org/10.5281/zenodo.6530511" ID-Zenodo-Dep="6530511" httpUri="https://zenodo.org/record/6530511/files/figure.png" pageId="3" pageNumber="149" startId="3.[136,235,1157,1181]" targetBox="[215,1372,189,1134]" targetPageId="3">
|
||
<paragraph id="8BDE36FEC45C7C3BDAB0FB1DB891FB21" blockId="3.[136,1452,1157,1218]" pageId="3" pageNumber="149">
|
||
<emphasis id="B915EAECC45C7C3BDAB0FB1DBA18FB7E" bold="true" box="[136,261,1157,1181]" pageId="3" pageNumber="149">FIGURE 3.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="4C614D7DC45C7C3BDBE9FB1EB8A1FB7D" authority="Li, Tomaszewska & Cai" authorityName="Li, Tomaszewska & Cai" authorityYear="2022" box="[465,956,1158,1182]" class="Insecta" family="Endomychidae" genus="Rhomeocalpsua" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="149" phylum="Arthropoda" rank="species" species="torosa" status="gen. et sp. nov.">
|
||
<emphasis id="B915EAECC45C7C3BDBE9FB1EB9A6FB7E" box="[465,699,1158,1182]" italics="true" pageId="3" pageNumber="149">Rhomeocalpsua torosa</emphasis>
|
||
Li, Tomaszewska & Cai
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B915EAECC45C7C3BD9FFFB1EBF75FB7D" bold="true" box="[967,1128,1158,1182]" pageId="3" pageNumber="149">
|
||
<taxonomicNameLabel id="A2265797C45C7C3BD9FFFB1EBF75FB7D" box="[967,1128,1158,1182]" pageId="3" pageNumber="149" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, NIGP180054, under confocal microscopy.
|
||
<emphasis id="B915EAECC45C7C3BDB5FFB31BA64FB22" bold="true" box="[359,377,1193,1217]" pageId="3" pageNumber="149">A</emphasis>
|
||
, Dorsal view.
|
||
<emphasis id="B915EAECC45C7C3BD834FB32B900FB21" bold="true" box="[524,541,1194,1218]" pageId="3" pageNumber="149">B</emphasis>
|
||
, Ventral view. Scale bars = 500 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDE36FEC45C7C3BDA85F9B1BAB6F91D" blockId="3.[136,778,1277,1902]" pageId="3" pageNumber="149">
|
||
Legs relatively long and slender. Trochanterofemoral joint oblique. Femora with deep and long groove (nearly throughout their length) for receiving tibiae (
|
||
<figureCitation id="135A2A7BC45C7C3BD8BEF9EBB9E0F96D" box="[646,765,1651,1678]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4E, F</figureCitation>
|
||
). Tibiae without apical spurs, with short spines surrounding apex (
|
||
<figureCitation id="135A2A7BC45C7C3BDAF6F926BA36F93B" box="[206,299,1726,1752]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4H</figureCitation>
|
||
). Tarsi 3-3-3; tarsomeres simple (
|
||
<figureCitation id="135A2A7BC45C7C3BD892F926BB8EF91D" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4H, I</figureCitation>
|
||
). Pretarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="8BDE36FEC45C7C3BDA85F891BA2AF88D" blockId="3.[136,778,1277,1902]" pageId="3" pageNumber="149">
|
||
Abdomen (
|
||
<figureCitation id="135A2A7BC45C7C3BDB79F891BAA2F8C0" box="[321,447,1801,1827]" captionStart="FIGURE 4" captionStartId="4.[136,235,1533,1557]" captionTargetBox="[145,1443,198,1509]" captionTargetId="figure-132@4.[145,1443,198,1509]" captionTargetPageId="4" captionText="FIGURE 4. Details of Rhomeocalpsua torosa Li, Tomaszewska & Cai, gen. et sp. nov., holotype, NIGP180054, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Antenna, dorsal view. D, Antenna, ventral view. E, Mesothorax, ventral view. F, Abdominal base, ventral view. G, Abdominal apex, ventral view. H, Mesotarsus, ventral view. I, Metatarsus, dorsal view. Abbreviations: a1–9, antennomeres 1–9; cl, clypeus; lbp, labial palp; msc, mesocoxa; msf, mesofemur; mst1–3, mesotarsomeres 1–3; msv, mesoventrite; mtf, metafemur; mtt1–3, metatarsomeres 1–3; mtv, metaventrite; mxp, maxillary palp; v1–6, ventrites 1–6. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6530513" httpUri="https://zenodo.org/record/6530513/files/figure.png" pageId="3" pageNumber="149">Fig. 4F, G</figureCitation>
|
||
) with six ventrites; ventrite 1 slightly longer than ventrite 2–4 combined, without postcoxal lines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |