treatments-xml/data/03/AC/8C/03AC8C35FF95FFBDFCE8FC980EFAF8E6.xml

<document id="6683517036AD0C972DA25225BF8FF2A6" ID-CLB-Dataset="295750" ID-DOI="10.1590/1982-0224-2023-0051" ID-GBIF-Dataset="87d70df3-c16c-4043-b252-09b578b41a1d" ID-ISSN="1982-0224" ID-Zenodo-Dep="11125413" ID-ZooBank="3EE6F4B5-CA12-4372-87E6-6EC91627587A" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712846088107" checkinUser="felipe" docAuthor="Crispim-Rodrigues, Jefferson Luan, Silva, Gabriel de Souza da Costa e, Shibatta, Oscar Akio, Kuranaka, Mariana &amp; Oliveira, Claudio" docDate="2023" docId="03AC8C35FF95FFBDFCE8FC980EFAF8E6" docLanguage="en" docName="NeotropIchthyol.2023.21.3.e230051.pdf" docOrigin="Neotropical Ichthyology (e 230051) 21 (3)" docSource="http://dx.doi.org/10.1590/1982-0224-2023-0051" docStyle="DocumentStyle:FC0C3FD10650AF21F8DA93D98D275823.3:NeotropIchthyol.2020-.journal_article" docStyleId="FC0C3FD10650AF21F8DA93D98D275823" docStyleName="NeotropIchthyol.2020-.journal_article" docStyleVersion="3" docTitle="Rhyacoglanis beninei Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira 2023, new species" docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="FF95F44DFF96FFB4FFE8FFA00A6EFFA7" masterDocTitle="Description and phylogenetic position of a new species of Rhyacoglanis (Siluriformes: Pseudopimelodidae) from the Jamanxim River basin" masterLastPageNumber="15" masterPageNumber="1" pageNumber="4" updateTime="1715075245574" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F0936124A149A80AFE37981156157082">Description and phylogenetic position of a new species of Rhyacoglanis (Siluriformes: Pseudopimelodidae) from the Jamanxim River basin</mods:title>
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<mods:namePart id="B240AE5CEBBCA9112169C631D7CC999B">Crispim-Rodrigues, Jefferson Luan</mods:namePart>
<mods:affiliation id="F4BDF4571E14659821B2B98C4E2320BE">Laboratório de Biologia e Genética de Peixes, Setor Morfologia, Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências da Unesp de Botucatu, R. Celso Wagner Zanin, s / n, Rubião Júnior, 18618 - 970 Botucatu, SP, Brazil. (JLCR) jefferson. crispim @ unesp. br (corresponding author), (GSCS) gabriel _ biota @ hotmail. com, (MK) mariana. kuranaka @ unesp. br, (CO) claudio. oliveira @ unesp. br.</mods:affiliation>
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<mods:namePart id="D60EF8D9D97BB6769622AA6F4E58CCC3">Silva, Gabriel de Souza da Costa e</mods:namePart>
<mods:affiliation id="03902D845C1D770765C7ED0A9F940AB4">Laboratório de Biologia e Genética de Peixes, Setor Morfologia, Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências da Unesp de Botucatu, R. Celso Wagner Zanin, s / n, Rubião Júnior, 18618 - 970 Botucatu, SP, Brazil. (JLCR) jefferson. crispim @ unesp. br (corresponding author), (GSCS) gabriel _ biota @ hotmail. com, (MK) mariana. kuranaka @ unesp. br, (CO) claudio. oliveira @ unesp. br. &amp; Department of Ichthyology, The Academy of Natural Sciences of Drexel University, 1900, Benjamin Franklin Parkway, 19103 - 1195, Philadelphia, PA, USA.</mods:affiliation>
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<mods:namePart id="EA4BB3620A10EAA14C2ED5C31D07C62D">Shibatta, Oscar Akio</mods:namePart>
<mods:affiliation id="3FFF43F55F47FA894991078A89BB96F6">Museu de Zoologia, Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina, Centro de Ciências Biológicas, Rodovia Celso Garcia Cid, 86057 - 970 Londrina, PR, Brazil. (OAS) oscar. shibatta @ gmail. com.</mods:affiliation>
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<taxonomicName id="4C0546A0FF95FFB7FCE8FC980E67FCF7" authority="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira, 2023" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[768,1033,824,848]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="beninei" status="sp. nov.">
<emphasis id="B971E131FF95FFB7FCE8FC980E67FCF7" bold="true" box="[768,1033,824,848]" pageId="3" pageNumber="4">Rhyacoglanis beninei</emphasis>
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, 
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(
<figureCitation id="133E21A6FF95FFB7FC86FC6E09B9FC41" box="[878,983,974,998]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[488,561,1612,1632]" captionStartId-1="6.[136,209,1537,1557]" captionStartId-2="7.[488,561,529,549]" captionStartId-3="8.[488,561,1089,1109]" captionTargetBox-0="[488,1459,190,1590]" captionTargetBox-1="[136,1459,1046,1514]" captionTargetBox-2="[527,1431,207,486]" captionTargetBox-3="[488,1447,189,1067]" captionTargetId-0="figure-120@4.[488,1459,190,1590]" captionTargetId-1="figure-336@6.[136,1462,1046,1517]" captionTargetId-2="figure-332@7.[488,1459,183,507]" captionTargetId-3="figure-31@8.[488,1459,189,1067]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1 | Rhyacoglanis beninei, holotype, MZUSP 127014, 59.1 mm SL, from córrego Jussara, an affluent of Jamanxim River, Tapajós River basin. Scale bar = 10 mm." captionText-1="FIGURE 2 | Pigmentation of oblique dark bars in the predorsal region of Rhyacoglanis beninei. A. MZUEL 23049, 29.6 mm SL; B. LBP 32145, 32.9 mm SL; C. LBP 32145, 37.3 mm SL; D. MZUEL 23049, 42.6 mm SL; E. LBP 32145, 50.2 mm SL. Scale bars = 10 mm." captionText-2="FIGURE 3 | Rhyacoglanis beninei pectoral-fin spine, paratype, LBP 32145, 50.2 mm SL." captionText-3="FIGURE 4 | A. Habitat of Rhyacoglanis beninei in córrego Jussara, 07°21’08”S 55°17’45”W; B. A rock where specimens of R. beninei were associated; C. Paratype of R. beninei just after capture. Photo: Gabriel S. Costa e Silva." figureDoi-0="http://doi.org/10.5281/zenodo.11125415" figureDoi-1="http://doi.org/10.5281/zenodo.11125417" figureDoi-2="http://doi.org/10.5281/zenodo.11125421" figureDoi-3="http://doi.org/10.5281/zenodo.11125425" httpUri-0="https://zenodo.org/record/11125415/files/figure.png" httpUri-1="https://zenodo.org/record/11125417/files/figure.png" httpUri-2="https://zenodo.org/record/11125421/files/figure.png" httpUri-3="https://zenodo.org/record/11125425/files/figure.png" pageId="3" pageNumber="4">Figs. 1–4</figureCitation>
; 
<tableCitation id="C6870898FF95FFB7FC0BFC6E0E44FC41" box="[995,1066,974,998]" captionStart="TABLE 1" captionStartId="5.[488,550,417,437]" captionTargetBox="[495,1432,470,1876]" captionTargetId="graphics-39@5.[488,1459,543,1485]" captionTargetPageId="5" captionText="TABLE 1 | Morphometric data of Rhyacoglanis beninei. N = 21 specimens, including the holotype." httpUri="http://table.plazi.org/id/DF7A6DABFF93FFB1FE00FE010F09FE12" pageId="3" pageNumber="4" tableUuid="DF7A6DABFF93FFB1FE00FE010F09FE12">Tab. 1</tableCitation>
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<materialsCitation id="3B6D377EFF95FFB7FDEEFBB90E30FBDC" collectingDate="2022-08-20" collectionCode="T" collectorName="G. S. C. Silva &amp; C. Faria." country="Brazil" county="Jussara" latitude="-7.352222" location="Novo Progresso" longLatPrecision="21" longitude="-55.295834" municipality="Jamanxim River" pageId="3" pageNumber="4" specimenCode="MZUSP 127014" specimenCount="1" stateProvince="Para" typeStatus="holotype">
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<typeStatus id="54BE8381FF95FFB7FDEEFBB90817FB96" box="[518,633,1049,1073]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="DBA39558FF95FFB7FD6BFBB90952FB96" box="[643,828,1049,1073]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4">MZUSP 127014</specimenCode>
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SL, 
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<collectingRegion id="49C1F3C1FF95FFB7FBCDFBB90EFAFB96" box="[1061,1172,1049,1073]" country="Brazil" name="Para" pageId="3" pageNumber="4">Pará State</collectingRegion>
, córrego 
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, tributary of 
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, municipally of 
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basin, 
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<date id="FFBB1BE3FF95FFB7FD6DFBC30976FBDC" box="[645,792,1123,1147]" pageId="3" pageNumber="4" value="2022-08-20">
<collectingDate id="EFFFE20BFF95FFB7FD6DFBC30976FBDC" box="[645,792,1123,1147]" pageId="3" pageNumber="4" value="2022-08-20">20 Aug 2022</collectingDate>
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, 
<collectorName id="26F058F5FF95FFB7FCCCFBC309D7FBDC" box="[804,953,1123,1147]" pageId="3" pageNumber="4">G. S. C. Silva</collectorName>
&amp; 
<collectionCode id="ED14A5E6FF95FFB7FC33FBC4099FFBDC" box="[987,1009,1124,1147]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
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<collectorName id="26F058F5FF95FFB7FC15FBC30E30FBDC" box="[1021,1118,1123,1147]" pageId="3" pageNumber="4">C. Faria.</collectorName>
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<materialsCitation id="3B6D377EFF95FFB7FDEEFB0E08BEFAB6" collectingDate="2017-09-22" collectingDateMax="2022-08-22" collectingDateMin="2017-09-22" collectionCode="R, T" collectorName="A. C. Dias &amp; C. S. Souza &amp; C. Souto &amp; N. Flausino Jr. &amp; Devide" pageId="3" pageNumber="4" specimenCode="MZUEL 23049, 4" specimenCount="2" typeStatus="paratype">
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<typeStatus id="54BE8381FF95FFB7FDEEFB0E0815FB61" box="[518,635,1198,1222]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
.
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from the same locality of holotype. LBP 25081, 5, 
<quantity id="4CFD90C6FF95FFB7FB0DFB0E0FE9FB62" box="[1253,1415,1198,1222]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" metricValueMax="4.37" metricValueMin="2.67" pageId="3" pageNumber="4" unit="mm" value="35.2" valueMax="43.7" valueMin="26.7">26.7–43.7 mm</quantity>
SL, 
<date id="FFBB1BE3FF95FFB7FE00FB73081EFB4C" box="[488,624,1235,1259]" pageId="3" pageNumber="4" value="2017-09-22">
<collectingDate id="EFFFE20BFF95FFB7FE00FB73081EFB4C" box="[488,624,1235,1259]" pageId="3" pageNumber="4" value="2017-09-22">22 Sep 2017</collectingDate>
</date>
, 
<collectorName id="26F058F5FF95FFB7FD92FB730881FB4C" box="[634,751,1235,1259]" pageId="3" pageNumber="4">A. C. Dias</collectorName>
, 
<collectorName id="26F058F5FF95FFB7FD11FB730917FB4C" box="[761,889,1235,1259]" pageId="3" pageNumber="4">C. S. Souza</collectorName>
, 
<collectorName id="26F058F5FF95FFB7FC6BFB730986FB4C" box="[899,1000,1235,1259]" pageId="3" pageNumber="4">C. Souto</collectorName>
, 
<collectorName id="26F058F5FF95FFB7FC1AFB740EFAFB4C" box="[1010,1172,1235,1259]" pageId="3" pageNumber="4">N. Flausino Jr.</collectorName>
&amp; 
<collectionCode id="ED14A5E6FF95FFB7FB5BFB740EA7FB4C" box="[1203,1225,1236,1259]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="4">R</collectionCode>
. 
<collectorName id="26F058F5FF95FFB7FB3AFB730F45FB4C" box="[1234,1323,1235,1259]" pageId="3" pageNumber="4">Devidé.</collectorName>
LBP 32145, 8, 
<quantity id="4CFD90C6FF95FFB7FDEAFB5808CBFAB7" box="[514,677,1272,1296]" metricMagnitude="-2" metricUnit="m" metricValue="3.645" metricValueMax="5.02" metricValueMin="2.27" pageId="3" pageNumber="4" unit="mm" value="36.45" valueMax="50.2" valueMin="22.7">22.7–50.2 mm</quantity>
SL,
</materialsCitation>
<materialsCitation id="3B6D377EFF95FFB7FD30FB58090CFAFC" collectingDate="2017-09-22" collectingDateMax="2022-08-22" collectingDateMin="2017-09-22" collectionCode="R, T" collectorName="G. S. C. Silva &amp; C. Faria. &amp; C. Faria" pageId="3" pageNumber="4" specimenCode="MZUEL 23049, 4" specimenCount="2" typeStatus="paratype">
the largest specimen was C&amp;S, 
<specimenCode id="DBA39558FF95FFB7FBDFFB590E86FAB6" box="[1079,1256,1273,1297]" collectionCode="MZUEL" pageId="3" pageNumber="4">MZUEL 23049</specimenCode>
, 
<specimenCode id="DBA39558FF95FFB7FB1BFB590F6DFAB6" box="[1267,1283,1273,1297]" collectionCode="MZUEL" pageId="3" pageNumber="4">4</specimenCode>
, 
<quantity id="4CFD90C6FF95FFB7FAE7FB580FDCFAB7" box="[1295,1458,1272,1296]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.26" metricValueMin="2.94" pageId="3" pageNumber="4" unit="mm" value="36.0" valueMax="42.6" valueMin="29.4">29.4–42.6 mm</quantity>
SL, 
<date id="FFBB1BE3FF95FFB7FDFAFABE08CAFA91" box="[530,676,1310,1334]" pageId="3" pageNumber="4" value="2022-08-20">
<collectingDate id="EFFFE20BFF95FFB7FDFAFABE08CAFA91" box="[530,676,1310,1334]" pageId="3" pageNumber="4" value="2022-08-20">20 Aug 2022</collectingDate>
</date>
, 
<collectorName id="26F058F5FF95FFB7FD47FABE092CFA91" box="[687,834,1310,1334]" pageId="3" pageNumber="4">G. S. C. Silva</collectorName>
&amp; 
<collectionCode id="ED14A5E6FF95FFB7FC8AFABE0916FA92" box="[866,888,1310,1333]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
. 
<collectorName id="26F058F5FF95FFB7FC6BFABE098AFA91" box="[899,996,1310,1334]" pageId="3" pageNumber="4">C. Faria.</collectorName>
LBP 32163, 3, 27.0– 
<quantity id="4CFD90C6FF95FFB7FB24FABE0F5EFA92" box="[1228,1328,1310,1334]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" pageId="3" pageNumber="4" unit="mm" value="47.9">47.9 mm</quantity>
SL, 
<date id="FFBB1BE3FF95FFB7FA89FABE084EFAFC" pageId="3" pageNumber="4" value="2022-08-22">
<collectingDate id="EFFFE20BFF95FFB7FA89FABE084EFAFC" pageId="3" pageNumber="4" value="2022-08-22">22 Aug 2022</collectingDate>
</date>
, 
<collectorName id="26F058F5FF95FFB7FDC3FAE308AFFAFC" box="[555,705,1347,1371]" pageId="3" pageNumber="4">G. S. C. Silva</collectorName>
&amp; 
<collectionCode id="ED14A5E6FF95FFB7FD0BFAE40897FAFC" box="[739,761,1348,1371]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
. 
<collectorName id="26F058F5FF95FFB7FCECFAE3090CFAFC" box="[772,866,1347,1371]" pageId="3" pageNumber="4">C. Faria</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F6EA8FF95FFB0FDEEFA2E09A2F8E6" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BBA3D23FF95FFB0FDEEFA2E09A2F8E6" blockId="3.[486,1461,1422,1894]" lastBlockId="4.[487,1459,1720,1857]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B971E131FF95FFB7FDEEFA2E08E8FA01" bold="true" box="[518,646,1422,1446]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="4C0546A0FF95FFB7FD65FA2E0909FA01" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[653,871,1422,1446]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF95FFB7FD65FA2E0909FA01" box="[653,871,1422,1446]" italics="true" pageId="3" pageNumber="4">Rhyacoglanis beninei</emphasis>
</taxonomicName>
can be diagnosed from all congeners by two oblique dorsal dark brown bars on the predorsal region (
<figureCitation id="133E21A6FF95FFB7FBF2FA130E30FA6C" box="[1050,1118,1459,1483]" captionStart="FIGURE 2" captionStartId="6.[136,209,1537,1557]" captionTargetBox="[136,1459,1046,1514]" captionTargetId="figure-336@6.[136,1462,1046,1517]" captionTargetPageId="6" captionText="FIGURE 2 | Pigmentation of oblique dark bars in the predorsal region of Rhyacoglanis beninei. A. MZUEL 23049, 29.6 mm SL; B. LBP 32145, 32.9 mm SL; C. LBP 32145, 37.3 mm SL; D. MZUEL 23049, 42.6 mm SL; E. LBP 32145, 50.2 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125417" httpUri="https://zenodo.org/record/11125417/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
) (
<emphasis id="B971E131FF95FFB7FB91FA140EE0FA6C" box="[1145,1166,1460,1483]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. absent). Additionally, 
<taxonomicName id="4C0546A0FF95FFB7FA73FA14085FFA57" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF95FFB7FA73FA14085FFA57" italics="true" pageId="3" pageNumber="4">R. beninei</emphasis>
</taxonomicName>
is distinguished from some congeners by having a dorsal confluence between the dark subdorsal and subadipose bands in large juveniles and adults (&gt; 
<quantity id="4CFD90C6FF95FFB7FB30FA5E0F49F9B2" box="[1240,1319,1534,1558]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="4" unit="mm" value="28.0">28 mm</quantity>
SL) (
<emphasis id="B971E131FF95FFB7FA8CFA5F0F17F9B1" box="[1380,1401,1535,1558]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. lack dorsal confluence in 
<taxonomicName id="4C0546A0FF95FFB7FD3DF984090EF99C" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[725,864,1572,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="B971E131FF95FFB7FD3DF984090EF99C" box="[725,864,1572,1595]" italics="true" pageId="3" pageNumber="4">R. paranensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0546A0FF95FFB7FC85F984099AF99C" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[877,1012,1571,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B971E131FF95FFB7FC85F984099AF99C" box="[877,1012,1571,1595]" italics="true" pageId="3" pageNumber="4">R. annulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0546A0FF95FFB7FBEBF9840E3CF99C" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" box="[1027,1106,1572,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="varii">
<emphasis id="B971E131FF95FFB7FBEBF9840E3CF99C" box="[1027,1106,1572,1595]" italics="true" pageId="3" pageNumber="4">R. varii</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0546A0FF95FFB7FB7BF9840F21F99C" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[1171,1359,1571,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="B971E131FF95FFB7FB7BF9840F21F99C" box="[1171,1359,1571,1595]" italics="true" pageId="3" pageNumber="4">R. rapppydanielae</emphasis>
</taxonomicName>
); ventral confluence between the dark subadipose and caudal peduncle bands (
<emphasis id="B971E131FF95FFB7FAE1F9E90F70F9C7" box="[1289,1310,1609,1632]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. lack ventral confluence in 
<taxonomicName id="4C0546A0FF95FFB7FD6AF9CF096AF921" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[642,772,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B971E131FF95FFB7FD6AF9CF096AF921" box="[642,772,1646,1670]" italics="true" pageId="3" pageNumber="4">R. annulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0546A0FF95FFB7FCFAF9CF09FFF921" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[786,913,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="B971E131FF95FFB7FCFAF9CF09FFF921" box="[786,913,1646,1670]" italics="true" pageId="3" pageNumber="4">R. epiblepsis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0546A0FF95FFB7FC75F9CF0E4BF921" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[925,1061,1647,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="B971E131FF95FFB7FC75F9CF0E4BF921" box="[925,1061,1647,1670]" italics="true" pageId="3" pageNumber="4">R. paranensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0546A0FF95FFB7FBD8F9CF0EC1F921" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1072,1199,1647,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="seminiger">
<emphasis id="B971E131FF95FFB7FBD8F9CF0EC1F921" box="[1072,1199,1647,1670]" italics="true" pageId="3" pageNumber="4">R. seminiger</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0546A0FF95FFB7FB0FF9CF0FCAF921" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[1255,1444,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="B971E131FF95FFB7FB0FF9CF0E95F921" box="[1255,1275,1647,1670]" italics="true" pageId="3" pageNumber="4">R</emphasis>
. 
<emphasis id="B971E131FF95FFB7FAE1F9CE0FCAF921" box="[1289,1444,1646,1670]" italics="true" pageId="3" pageNumber="4">rapppydanielae</emphasis>
</taxonomicName>
); body without conspicuous dark brown spots (
<emphasis id="B971E131FF95FFB7FC1DF9340E64F90C" box="[1013,1034,1684,1707]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. conspicuous dark brown spots in 
<taxonomicName id="4C0546A0FF95FFB7FA75F9340827F977" authorityName="Shibatta &amp; Vari" authorityYear="2017" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="B971E131FF95FFB7FA75F9340827F977" italics="true" pageId="3" pageNumber="4">R. epiblepsis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0546A0FF95FFB7FD60F9190929F977" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[648,839,1720,1744]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="B971E131FF95FFB7FD60F9190929F977" box="[648,839,1720,1744]" italics="true" pageId="3" pageNumber="4">R. rapppydanielae</emphasis>
</taxonomicName>
); complete dark band on caudal peduncle (
<emphasis id="B971E131FF95FFB7FAAFF9190F0CF977" box="[1351,1378,1721,1744]" italics="true" pageId="3" pageNumber="4">vs.</emphasis>
caudal peduncle-band with a unpigmented central region in 
<taxonomicName id="4C0546A0FF95FFB7FBB0F97F0E8EF951" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1112,1248,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B971E131FF95FFB7FBB0F97F0E8EF951" box="[1112,1248,1758,1782]" italics="true" pageId="3" pageNumber="4">R. annulatus</emphasis>
</taxonomicName>
); body with three dark bands (
<emphasis id="B971E131FF95FFB7FD9BF8A408E6F8BC" box="[627,648,1796,1819]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. two dark bands in 
<taxonomicName id="4C0546A0FF95FFB7FC87F8A4099CF8BC" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[879,1010,1796,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="seminiger">
<emphasis id="B971E131FF95FFB7FC87F8A4099CF8BC" box="[879,1010,1796,1819]" italics="true" pageId="3" pageNumber="4">R. seminiger</emphasis>
</taxonomicName>
); a thin dark caudal-fin bands (
<emphasis id="B971E131FF95FFB7FAB0F8A40F03F8BC" box="[1368,1389,1796,1819]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. large caudal-fin bands in 
<taxonomicName id="4C0546A0FF95FFB7FD28F8890926F8E7" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[704,840,1833,1856]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="B971E131FF95FFB7FD28F8890926F8E7" box="[704,840,1833,1856]" italics="true" pageId="3" pageNumber="4">R. paranensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0546A0FF95FFB7FC94F8890995F8E7" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[892,1019,1832,1856]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="B971E131FF95FFB7FC94F8890995F8E7" box="[892,1019,1832,1856]" italics="true" pageId="3" pageNumber="4">R. epiblepsis</emphasis>
</taxonomicName>
); pectoral-fin spine with anterior serrae distributed along the entire margin (restricted to the proximal half in 
<taxonomicName id="4C0546A0FF95FFB7FB10F8EF0F0DF8C1" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[1272,1379,1870,1894]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="B971E131FF95FFB7FB10F8EF0F0DF8C1" box="[1272,1379,1870,1894]" italics="true" pageId="3" pageNumber="4">R. pulcher</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0546A0FF95FFB0FA73F8EF0827F977" authorityName="Shibatta &amp; Vari" authorityYear="2017" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="4" lastPageNumber="5" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="seminiger">
<emphasis id="B971E131FF95FFB0FA73F8EF0827F977" italics="true" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">R. seminiger</emphasis>
</taxonomicName>
); posterior tip of the post-cleithral process reaching vertical through the base of the dorsal-fin spine (
<emphasis id="B971E131FF92FFB0FD03F97F096EF951" box="[747,768,1759,1782]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. not reaching in 
<taxonomicName id="4C0546A0FF92FFB0FC29F97F0E2FF951" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[961,1089,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="B971E131FF92FFB0FC29F97F0E2FF951" box="[961,1089,1758,1782]" italics="true" pageId="4" pageNumber="5">R. epiblepsis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0546A0FF92FFB0FB90F97F0F5CF951" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[1144,1330,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="B971E131FF92FFB0FB90F97F0F5CF951" box="[1144,1330,1758,1782]" italics="true" pageId="4" pageNumber="5">R. rapppydanielae</emphasis>
</taxonomicName>
); hypural 5 free of hypural 3 and 4 (
<emphasis id="B971E131FF92FFB0FD1CF8A40967F8BC" box="[756,777,1796,1819]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. hypurals 4 and 5 fused in 
<taxonomicName id="4C0546A0FF92FFB0FBDBF8A40E83F8BC" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[1075,1261,1795,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="B971E131FF92FFB0FBDBF8A40E83F8BC" box="[1075,1261,1795,1819]" italics="true" pageId="4" pageNumber="5">R. rapppydanielae</emphasis>
</taxonomicName>
); pointed caudal-fin lobes (
<emphasis id="B971E131FF92FFB0FDBEF8890805F8E7" box="[598,619,1833,1856]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. rounded lobes in 
<taxonomicName id="4C0546A0FF92FFB0FCD3F88909D2F8E7" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[827,956,1832,1856]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="B971E131FF92FFB0FCD3F88909D2F8E7" box="[827,956,1832,1856]" italics="true" pageId="4" pageNumber="5">R. epiblepsis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF7A6DABFF92FFB0FE00F9EC0E7CF922" ID-DOI="http://doi.org/10.5281/zenodo.11125415" ID-Zenodo-Dep="11125415" httpUri="https://zenodo.org/record/11125415/files/figure.png" pageId="4" pageNumber="5" startId="4.[488,561,1612,1632]" targetBox="[488,1459,190,1590]" targetPageId="4" targetType="figure">
<paragraph id="8BBA3D23FF92FFB0FE00F9EC0E7CF922" blockId="4.[488,1459,1612,1669]" pageId="4" pageNumber="5">
<emphasis id="B971E131FF92FFB0FE00F9EC0961F9C7" bold="true" box="[488,783,1612,1632]" pageId="4" pageNumber="5">
FIGURE 1 | 
<taxonomicName id="4C0546A0FF92FFB0FDB2F9EC0961F9C7" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[602,783,1612,1632]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF92FFB0FDB2F9EC0961F9C7" bold="true" box="[602,783,1612,1632]" italics="true" pageId="4" pageNumber="5">Rhyacoglanis beninei</emphasis>
</taxonomicName>
</emphasis>
, holotype, MZUSP 127014, 59.1 mm SL, from córrego Jussara, an affluent of Jamanxim River, Tapajós River basin. Scale bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="C31F6EA8FF93FFB3FDEEFF1E0996FA56" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="8BBA3D23FF93FFB2FDEEFF1E0ED2FD81" blockId="5.[487,1461,190,363]" lastBlockId="6.[486,1462,190,961]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FDEEFF1E08F5FF71" bold="true" box="[518,667,190,214]" pageId="5" pageNumber="6">Description.</emphasis>
Morphometric data from 
<taxonomicName id="4C0546A0FF93FFB1FC2CFF1E0EF0FF71" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[964,1182,190,214]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF93FFB1FC2CFF1E0EF0FF71" box="[964,1182,190,214]" italics="true" pageId="5" pageNumber="6">Rhyacoglanis beninei</emphasis>
</taxonomicName>
is available in 
<tableCitation id="C6870898FF93FFB1FAAAFF1E0FE3FF71" box="[1346,1421,190,214]" captionStart="TABLE 1" captionStartId="5.[488,550,417,437]" captionTargetBox="[495,1432,470,1876]" captionTargetId="graphics-39@5.[488,1459,543,1485]" captionTargetPageId="5" captionText="TABLE 1 | Morphometric data of Rhyacoglanis beninei. N = 21 specimens, including the holotype." httpUri="http://table.plazi.org/id/DF7A6DABFF93FFB1FE00FE010F09FE12" pageId="5" pageNumber="6" tableUuid="DF7A6DABFF93FFB1FE00FE010F09FE12">Tab. 1</tableCitation>
. In lateral view, straight profile of body from snout tip to dorsal-fin origin followed by a straight profile from dorsal-fin base to caudal-fin insertion. Body ventral surface slightly convex from snout tip to opercular opening region, straight to slightly concave from head posterior end to anal-fin insertion, and straight-angled upward on caudal peduncle. Head depressed with numerous well-developed unculiferous tubercles. Head dorsal profile in a rounded trapezoidal shape, with most anterior region narrower than posterior region. Wide mouth, larger than distance between posterior nostrils. Posterior nostrils opening wider than anterior nostrils. Thick lips with lateral portion extended posteriorly. Dentigerous plates on premaxilla and dentary. Eyes small, covered by skin. Opercular opening covered by a well-developed membrane. Maxillary barbels reaching first branched pectoral-fin ray base. Inner mental barbel small, not reaching isthmus, and outer mental barbel reaching first pectoral-fin ray base. Head laterosensory system bearing conspicuous pores, including six infraorbital, five supraorbital, nine premaxillaries, one premaxillary-postorbital, and one postorbital.
</paragraph>
<caption id="DF7A6DABFF93FFB1FE00FE010F09FE12" ID-Table-UUID="DF7A6DABFF93FFB1FE00FE010F09FE12" box="[488,1383,417,437]" httpUri="http://table.plazi.org/id/DF7A6DABFF93FFB1FE00FE010F09FE12" pageId="5" pageNumber="6" startId="5.[488,550,417,437]" targetBox="[495,1432,470,1876]" targetIsTable="true" targetPageId="5">
<paragraph id="8BBA3D23FF93FFB1FE00FE010F09FE12" blockId="5.[488,1383,417,437]" box="[488,1383,417,437]" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FE00FE010829FE12" bold="true" box="[488,583,417,437]" pageId="5" pageNumber="6">TABLE 1 |</emphasis>
Morphometric data of 
<emphasis id="B971E131FF93FFB1FCC9FE0109B9FE12" bold="true" box="[801,983,417,437]" italics="true" pageId="5" pageNumber="6">Rhyacoglanis beninei</emphasis>
. N = 21 specimens, including the holotype.
</paragraph>
</caption>
<paragraph id="8BBA3D23FF93FFB1FC31FE770FFCF8F3" pageId="5" pageNumber="6">
<table id="F905CF83FF93004BFE07FE760FF6F8F3" box="[495,1432,470,1876]" gridcols="5" gridrows="32" pageId="5" pageNumber="6">
<tr id="35353F61FF93004BFE07FE760FF6FE4F" box="[495,1432,470,488]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1">
<th id="76E4561DFF93004BFC31FE760E41FE4F" box="[985,1071,470,488]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FC31FE770E41FE40" bold="true" box="[985,1071,471,487]" pageId="5" pageNumber="6">Holotype</emphasis>
</th>
<th id="76E4561DFF93004BFB93FE760EA0FE4F" box="[1147,1230,470,488]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FB60FE770EACFE40" bold="true" box="[1160,1218,471,487]" pageId="5" pageNumber="6">Range</emphasis>
</th>
<th id="76E4561DFF93004BFAE7FE760F2DFE4F" box="[1295,1347,470,488]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FAE7FE770F2DFE40" bold="true" box="[1295,1347,471,487]" pageId="5" pageNumber="6">Mean</emphasis>
</th>
<th id="76E4561DFF93004BFA9BFE760FF6FE4F" box="[1395,1432,470,488]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FA91FE760FFCFE4F" bold="true" box="[1401,1426,470,488]" pageId="5" pageNumber="6">SD</emphasis>
</th>
</tr>
<tr id="35353F61FF93004BFE07FDA30FF6FDB2" box="[495,1432,515,533]" gridrow="1" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FDA309CEFDB2" box="[495,928,515,533]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Standard length (mm)</th>
<td id="76E4561DFF93004BFC31FDA30E41FDB2" box="[985,1071,515,533]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">59.1</td>
<td id="76E4561DFF93004BFB93FDA30EA0FDB2" box="[1147,1230,515,533]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">22.7–59.1</td>
<td id="76E4561DFF93004BFAE7FDA30F2DFDB2" box="[1295,1347,515,533]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">36.6</td>
<td id="76E4561DFF93004BFA9BFDA30FF6FDB2" box="[1395,1432,515,533]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">–</td>
</tr>
<tr id="35353F61FF93004BFE07FD900FF6FDE5" box="[495,1432,560,578]" gridrow="2" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FD900FF6FDE5" box="[495,1432,560,578]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FE07FD910885FDE6" bold="true" box="[495,747,560,578]" pageId="5" pageNumber="6">Percent of standard length</emphasis>
</th>
</tr>
<tr id="35353F61FF93004BFE07FDFD0FF6FDC8" box="[495,1432,605,623]" gridrow="3" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FDFD09CEFDC8" box="[495,928,605,623]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Head length</th>
<td id="76E4561DFF93004BFC31FDFD0E41FDC8" box="[985,1071,605,623]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">30.7</td>
<td id="76E4561DFF93004BFB93FDFD0EA0FDC8" box="[1147,1230,605,623]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">27.5–31.9</td>
<td id="76E4561DFF93004BFAE7FDFD0F2DFDC8" box="[1295,1347,605,623]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">30.0</td>
<td id="76E4561DFF93004BFA9BFDFD0FF6FDC8" box="[1395,1432,605,623]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">1.3</td>
</tr>
<tr id="35353F61FF93004BFE07FD2A0FF6FD3B" box="[495,1432,650,668]" gridrow="4" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FD2A09CEFD3B" box="[495,928,650,668]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Pectoral-girdle width</th>
<td id="76E4561DFF93004BFC31FD2A0E41FD3B" box="[985,1071,650,668]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">34.1</td>
<td id="76E4561DFF93004BFB93FD2A0EA0FD3B" box="[1147,1230,650,668]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">26.7–34.1</td>
<td id="76E4561DFF93004BFAE7FD2A0F2DFD3B" box="[1295,1347,650,668]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">29.2</td>
<td id="76E4561DFF93004BFA9BFD2A0FF6FD3B" box="[1395,1432,650,668]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">1.7</td>
</tr>
<tr id="35353F61FF93004BFE07FD160FF6FD6F" box="[495,1432,694,712]" gridrow="5" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FD1609CEFD6F" box="[495,928,694,712]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Predorsal length</th>
<td id="76E4561DFF93004BFC31FD160E41FD6F" box="[985,1071,694,712]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">38.4</td>
<td id="76E4561DFF93004BFB93FD160EA0FD6F" box="[1147,1230,694,712]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">35.4–42.3</td>
<td id="76E4561DFF93004BFAE7FD160F2DFD6F" box="[1295,1347,694,712]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">39.0</td>
<td id="76E4561DFF93004BFA9BFD160FF6FD6F" box="[1395,1432,694,712]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">1.8</td>
</tr>
<tr id="35353F61FF93004BFE07FD430FF6FD52" box="[495,1432,739,757]" gridrow="6" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FD4309CEFD52" box="[495,928,739,757]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Dorsal-fin base length</th>
<td id="76E4561DFF93004BFC31FD430E41FD52" box="[985,1071,739,757]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">17.1</td>
<td id="76E4561DFF93004BFB93FD430EA0FD52" box="[1147,1230,739,757]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">13.8–18.4</td>
<td id="76E4561DFF93004BFAE7FD430F2DFD52" box="[1295,1347,739,757]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">16.3</td>
<td id="76E4561DFF93004BFA9BFD430FF6FD52" box="[1395,1432,739,757]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">1.2</td>
</tr>
<tr id="35353F61FF93004BFE07FCB00FF6FC85" box="[495,1432,784,802]" gridrow="7" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FCB009CEFC85" box="[495,928,784,802]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Adipose-fin base length</th>
<td id="76E4561DFF93004BFC31FCB00E41FC85" box="[985,1071,784,802]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">19.9</td>
<td id="76E4561DFF93004BFB93FCB00EA0FC85" box="[1147,1230,784,802]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">15.2–21.2</td>
<td id="76E4561DFF93004BFAE7FCB00F2DFC85" box="[1295,1347,784,802]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">18.0</td>
<td id="76E4561DFF93004BFA9BFCB00FF6FC85" box="[1395,1432,784,802]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">1.9</td>
</tr>
<tr id="35353F61FF93004BFE07FC9D0FF6FCE8" box="[495,1432,829,847]" gridrow="8" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FC9D09CEFCE8" box="[495,928,829,847]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">Prepelvic length</th>
<td id="76E4561DFF93004BFC31FC9D0E41FCE8" box="[985,1071,829,847]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">53.5</td>
<td id="76E4561DFF93004BFB93FC9D0EA0FCE8" box="[1147,1230,829,847]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">48.7–56.3</td>
<td id="76E4561DFF93004BFAE7FC9D0F2DFCE8" box="[1295,1347,829,847]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">51.8</td>
<td id="76E4561DFF93004BFA9BFC9D0FF6FCE8" box="[1395,1432,829,847]" gridcol="4" gridrow="8" pageId="5" pageNumber="6">1.6</td>
</tr>
<tr id="35353F61FF93004BFE07FCC90FF6FCDC" box="[495,1432,873,891]" gridrow="9" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FCC909CEFCDC" box="[495,928,873,891]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">Distance between pelvic and anal fins</th>
<td id="76E4561DFF93004BFC31FCC90E41FCDC" box="[985,1071,873,891]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">24.9</td>
<td id="76E4561DFF93004BFB93FCC90EA0FCDC" box="[1147,1230,873,891]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">22.0–27.3</td>
<td id="76E4561DFF93004BFAE7FCC90F2DFCDC" box="[1295,1347,873,891]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">25.6</td>
<td id="76E4561DFF93004BFA9BFCC90FF6FCDC" box="[1395,1432,873,891]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">1.3</td>
</tr>
<tr id="35353F61FF93004BFE07FC360FF6FC0F" box="[495,1432,918,936]" gridrow="10" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FC3609CEFC0F" box="[495,928,918,936]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Anal-fin base length</th>
<td id="76E4561DFF93004BFC31FC360E41FC0F" box="[985,1071,918,936]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">11.3</td>
<td id="76E4561DFF93004BFB93FC360EA0FC0F" box="[1147,1230,918,936]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">9.1–13.2</td>
<td id="76E4561DFF93004BFAE7FC360F2DFC0F" box="[1295,1347,918,936]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">10.8</td>
<td id="76E4561DFF93004BFA9BFC360FF6FC0F" box="[1395,1432,918,936]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">1.0</td>
</tr>
<tr id="35353F61FF93004BFE07FC630FF6FC72" box="[495,1432,963,981]" gridrow="11" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FC6309CEFC72" box="[495,928,963,981]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">Caudal-peduncle length</th>
<td id="76E4561DFF93004BFC31FC630E41FC72" box="[985,1071,963,981]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">16.2</td>
<td id="76E4561DFF93004BFB93FC630EA0FC72" box="[1147,1230,963,981]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">11.4–16.2</td>
<td id="76E4561DFF93004BFAE7FC630F2DFC72" box="[1295,1347,963,981]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">13.0</td>
<td id="76E4561DFF93004BFA9BFC630FF6FC72" box="[1395,1432,963,981]" gridcol="4" gridrow="11" pageId="5" pageNumber="6">1.4</td>
</tr>
<tr id="35353F61FF93004BFE07FC500FF6FBA5" box="[495,1432,1008,1026]" gridrow="12" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FC5009CEFBA5" box="[495,928,1008,1026]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">Body depth</th>
<td id="76E4561DFF93004BFC31FC500E41FBA5" box="[985,1071,1008,1026]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">19.2</td>
<td id="76E4561DFF93004BFB93FC500EA0FBA5" box="[1147,1230,1008,1026]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">16.8–25.1</td>
<td id="76E4561DFF93004BFAE7FC500F2DFBA5" box="[1295,1347,1008,1026]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">19.7</td>
<td id="76E4561DFF93004BFA9BFC500FF6FBA5" box="[1395,1432,1008,1026]" gridcol="4" gridrow="12" pageId="5" pageNumber="6">2.1</td>
</tr>
<tr id="35353F61FF93004BFE07FBBD0FF6FB88" box="[495,1432,1053,1071]" gridrow="13" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FBBD09CEFB88" box="[495,928,1053,1071]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">Caudal-peduncle depth</th>
<td id="76E4561DFF93004BFC31FBBD0E41FB88" box="[985,1071,1053,1071]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">10.3</td>
<td id="76E4561DFF93004BFB93FBBD0EA0FB88" box="[1147,1230,1053,1071]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">8.0–11.5</td>
<td id="76E4561DFF93004BFAE7FBBD0F2DFB88" box="[1295,1347,1053,1071]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">9.9</td>
<td id="76E4561DFF93004BFA9BFBBD0FF6FB88" box="[1395,1432,1053,1071]" gridcol="4" gridrow="13" pageId="5" pageNumber="6">0.9</td>
</tr>
<tr id="35353F61FF93004BFE07FBE90FF6FBFC" box="[495,1432,1097,1115]" gridrow="14" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FBE909CEFBFC" box="[495,928,1097,1115]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">Pectoral-fin spine length</th>
<td id="76E4561DFF93004BFC31FBE90E41FBFC" box="[985,1071,1097,1115]" gridcol="1" gridrow="14" pageId="5" pageNumber="6">15.4</td>
<td id="76E4561DFF93004BFB93FBE90EA0FBFC" box="[1147,1230,1097,1115]" gridcol="2" gridrow="14" pageId="5" pageNumber="6">14.8–20.0</td>
<td id="76E4561DFF93004BFAE7FBE90F2DFBFC" box="[1295,1347,1097,1115]" gridcol="3" gridrow="14" pageId="5" pageNumber="6">17.6</td>
<td id="76E4561DFF93004BFA9BFBE90FF6FBFC" box="[1395,1432,1097,1115]" gridcol="4" gridrow="14" pageId="5" pageNumber="6">1.4</td>
</tr>
<tr id="35353F61FF93004BFE07FBD60FF6FB2F" box="[495,1432,1142,1160]" gridrow="15" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FBD609CEFB2F" box="[495,928,1142,1160]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">Dorsal-fin spine length</th>
<td id="76E4561DFF93004BFC31FBD60E41FB2F" box="[985,1071,1142,1160]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">14.9</td>
<td id="76E4561DFF93004BFB93FBD60EA0FB2F" box="[1147,1230,1142,1160]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">11.1–20.2</td>
<td id="76E4561DFF93004BFAE7FBD60F2DFB2F" box="[1295,1347,1142,1160]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">17.2</td>
<td id="76E4561DFF93004BFA9BFBD60FF6FB2F" box="[1395,1432,1142,1160]" gridcol="4" gridrow="15" pageId="5" pageNumber="6">2.5</td>
</tr>
<tr id="35353F61FF93004BFE07FB030FF6FB12" box="[495,1432,1187,1205]" gridrow="16" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FB0309CEFB12" box="[495,928,1187,1205]" gridcol="0" gridrow="16" pageId="5" pageNumber="6">Pelvic-fin length</th>
<td id="76E4561DFF93004BFC31FB030E41FB12" box="[985,1071,1187,1205]" gridcol="1" gridrow="16" pageId="5" pageNumber="6">12.3</td>
<td id="76E4561DFF93004BFB93FB030EA0FB12" box="[1147,1230,1187,1205]" gridcol="2" gridrow="16" pageId="5" pageNumber="6">11.6–19.7</td>
<td id="76E4561DFF93004BFAE7FB030F2DFB12" box="[1295,1347,1187,1205]" gridcol="3" gridrow="16" pageId="5" pageNumber="6">14.6</td>
<td id="76E4561DFF93004BFA9BFB030FF6FB12" box="[1395,1432,1187,1205]" gridcol="4" gridrow="16" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr id="35353F61FF93004BFE07FB700FF6FB45" box="[495,1432,1232,1250]" gridrow="17" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FB7009CEFB45" box="[495,928,1232,1250]" gridcol="0" gridrow="17" pageId="5" pageNumber="6">Postcleithral-process length</th>
<td id="76E4561DFF93004BFC31FB700E41FB45" box="[985,1071,1232,1250]" gridcol="1" gridrow="17" pageId="5" pageNumber="6">15.2</td>
<td id="76E4561DFF93004BFB93FB700EA0FB45" box="[1147,1230,1232,1250]" gridcol="2" gridrow="17" pageId="5" pageNumber="6">9.8–15.7</td>
<td id="76E4561DFF93004BFAE7FB700F2DFB45" box="[1295,1347,1232,1250]" gridcol="3" gridrow="17" pageId="5" pageNumber="6">13.6</td>
<td id="76E4561DFF93004BFA9BFB700FF6FB45" box="[1395,1432,1232,1250]" gridcol="4" gridrow="17" pageId="5" pageNumber="6">1.7</td>
</tr>
<tr id="35353F61FF93004BFE07FB5C0FF6FAA9" box="[495,1432,1276,1294]" gridrow="18" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FB5C09CEFAA9" box="[495,928,1276,1294]" gridcol="0" gridrow="18" pageId="5" pageNumber="6">Distance between dorsal and pelvic fins</th>
<td id="76E4561DFF93004BFC31FB5C0E41FAA9" box="[985,1071,1276,1294]" gridcol="1" gridrow="18" pageId="5" pageNumber="6">24.9</td>
<td id="76E4561DFF93004BFB93FB5C0EA0FAA9" box="[1147,1230,1276,1294]" gridcol="2" gridrow="18" pageId="5" pageNumber="6">19.7–26.9</td>
<td id="76E4561DFF93004BFAE7FB5C0F2DFAA9" box="[1295,1347,1276,1294]" gridcol="3" gridrow="18" pageId="5" pageNumber="6">22.8</td>
<td id="76E4561DFF93004BFA9BFB5C0FF6FAA9" box="[1395,1432,1276,1294]" gridcol="4" gridrow="18" pageId="5" pageNumber="6">1.8</td>
</tr>
<tr id="35353F61FF93004BFE07FA890FF6FA9C" box="[495,1432,1321,1339]" gridrow="19" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FA8909CEFA9C" box="[495,928,1321,1339]" gridcol="0" gridrow="19" pageId="5" pageNumber="6">Distance between pelvic fins</th>
<td id="76E4561DFF93004BFC31FA890E41FA9C" box="[985,1071,1321,1339]" gridcol="1" gridrow="19" pageId="5" pageNumber="6">13.5</td>
<td id="76E4561DFF93004BFB93FA890EA0FA9C" box="[1147,1230,1321,1339]" gridcol="2" gridrow="19" pageId="5" pageNumber="6">10.4–14.4</td>
<td id="76E4561DFF93004BFAE7FA890F2DFA9C" box="[1295,1347,1321,1339]" gridcol="3" gridrow="19" pageId="5" pageNumber="6">11.6</td>
<td id="76E4561DFF93004BFA9BFA890FF6FA9C" box="[1395,1432,1321,1339]" gridcol="4" gridrow="19" pageId="5" pageNumber="6">2.8</td>
</tr>
<tr id="35353F61FF93004BFE07FAF60FF6FACF" box="[495,1432,1366,1384]" gridrow="20" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FAF609CEFACF" box="[495,928,1366,1384]" gridcol="0" gridrow="20" pageId="5" pageNumber="6">Distance between pelvic fin and anus</th>
<td id="76E4561DFF93004BFC31FAF60E41FACF" box="[985,1071,1366,1384]" gridcol="1" gridrow="20" pageId="5" pageNumber="6">13.9</td>
<td id="76E4561DFF93004BFB93FAF60EA0FACF" box="[1147,1230,1366,1384]" gridcol="2" gridrow="20" pageId="5" pageNumber="6">11.1–14.6</td>
<td id="76E4561DFF93004BFAE7FAF60F2DFACF" box="[1295,1347,1366,1384]" gridcol="3" gridrow="20" pageId="5" pageNumber="6">12.5</td>
<td id="76E4561DFF93004BFA9BFAF60FF6FACF" box="[1395,1432,1366,1384]" gridcol="4" gridrow="20" pageId="5" pageNumber="6">1.1</td>
</tr>
<tr id="35353F61FF93004BFE07FA230FF6FA32" box="[495,1432,1411,1429]" gridrow="21" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FA2309CEFA32" box="[495,928,1411,1429]" gridcol="0" gridrow="21" pageId="5" pageNumber="6">Distance between the anus and anal fin</th>
<td id="76E4561DFF93004BFC31FA230E41FA32" box="[985,1071,1411,1429]" gridcol="1" gridrow="21" pageId="5" pageNumber="6">11.2</td>
<td id="76E4561DFF93004BFB93FA230EA0FA32" box="[1147,1230,1411,1429]" gridcol="2" gridrow="21" pageId="5" pageNumber="6">11.1–17.7</td>
<td id="76E4561DFF93004BFAE7FA230F2DFA32" box="[1295,1347,1411,1429]" gridcol="3" gridrow="21" pageId="5" pageNumber="6">13.9</td>
<td id="76E4561DFF93004BFA9BFA230FF6FA32" box="[1395,1432,1411,1429]" gridcol="4" gridrow="21" pageId="5" pageNumber="6">1.7</td>
</tr>
<tr id="35353F61FF93004BFE07FA0F0FF6FA66" box="[495,1432,1455,1473]" gridrow="22" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FA0F0FF6FA66" box="[495,1432,1455,1473]" colspan="5" colspanRight="4" gridcol="0" gridrow="22" pageId="5" pageNumber="6">
<emphasis id="B971E131FF93FFB1FE07FA1008AAFA67" bold="true" box="[495,708,1455,1473]" pageId="5" pageNumber="6">Percent of head length</emphasis>
</th>
</tr>
<tr id="35353F61FF93004BFE07FA7C0FF6FA49" box="[495,1432,1500,1518]" gridrow="23" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07FA7C09CEFA49" box="[495,928,1500,1518]" gridcol="0" gridrow="23" pageId="5" pageNumber="6">Eye diameter</th>
<td id="76E4561DFF93004BFC31FA7C0E41FA49" box="[985,1071,1500,1518]" gridcol="1" gridrow="23" pageId="5" pageNumber="6">8.8</td>
<td id="76E4561DFF93004BFB93FA7C0EA0FA49" box="[1147,1230,1500,1518]" gridcol="2" gridrow="23" pageId="5" pageNumber="6">6.5–12.6</td>
<td id="76E4561DFF93004BFAE7FA7C0F2DFA49" box="[1295,1347,1500,1518]" gridcol="3" gridrow="23" pageId="5" pageNumber="6">9.5</td>
<td id="76E4561DFF93004BFA9BFA7C0FF6FA49" box="[1395,1432,1500,1518]" gridcol="4" gridrow="23" pageId="5" pageNumber="6">1.7</td>
</tr>
<tr id="35353F61FF93004BFE07F9A90FF6F9BC" box="[495,1432,1545,1563]" gridrow="24" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F9A909CEF9BC" box="[495,928,1545,1563]" gridcol="0" gridrow="24" pageId="5" pageNumber="6">Interorbital distance</th>
<td id="76E4561DFF93004BFC31F9A90E41F9BC" box="[985,1071,1545,1563]" gridcol="1" gridrow="24" pageId="5" pageNumber="6">34.1</td>
<td id="76E4561DFF93004BFB93F9A90EA0F9BC" box="[1147,1230,1545,1563]" gridcol="2" gridrow="24" pageId="5" pageNumber="6">26.2–44.5</td>
<td id="76E4561DFF93004BFAE7F9A90F2DF9BC" box="[1295,1347,1545,1563]" gridcol="3" gridrow="24" pageId="5" pageNumber="6">34.2</td>
<td id="76E4561DFF93004BFA9BF9A90FF6F9BC" box="[1395,1432,1545,1563]" gridcol="4" gridrow="24" pageId="5" pageNumber="6">4.1</td>
</tr>
<tr id="35353F61FF93004BFE07F9960FF6F9EF" box="[495,1432,1590,1608]" gridrow="25" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F99609CEF9EF" box="[495,928,1590,1608]" gridcol="0" gridrow="25" pageId="5" pageNumber="6">Snout length</th>
<td id="76E4561DFF93004BFC31F9960E41F9EF" box="[985,1071,1590,1608]" gridcol="1" gridrow="25" pageId="5" pageNumber="6">41.3</td>
<td id="76E4561DFF93004BFB93F9960EA0F9EF" box="[1147,1230,1590,1608]" gridcol="2" gridrow="25" pageId="5" pageNumber="6">32.6–41.3</td>
<td id="76E4561DFF93004BFAE7F9960F2DF9EF" box="[1295,1347,1590,1608]" gridcol="3" gridrow="25" pageId="5" pageNumber="6">37.1</td>
<td id="76E4561DFF93004BFA9BF9960FF6F9EF" box="[1395,1432,1590,1608]" gridcol="4" gridrow="25" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr id="35353F61FF93004BFE07F9C30FF6F9D2" box="[495,1432,1635,1653]" gridrow="26" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F9C309CEF9D2" box="[495,928,1635,1653]" gridcol="0" gridrow="26" pageId="5" pageNumber="6">Mouth width</th>
<td id="76E4561DFF93004BFC31F9C30E41F9D2" box="[985,1071,1635,1653]" gridcol="1" gridrow="26" pageId="5" pageNumber="6">65.2</td>
<td id="76E4561DFF93004BFB93F9C30EA0F9D2" box="[1147,1230,1635,1653]" gridcol="2" gridrow="26" pageId="5" pageNumber="6">44.8–65.2</td>
<td id="76E4561DFF93004BFAE7F9C30F2DF9D2" box="[1295,1347,1635,1653]" gridcol="3" gridrow="26" pageId="5" pageNumber="6">54.8</td>
<td id="76E4561DFF93004BFA9BF9C30FF6F9D2" box="[1395,1432,1635,1653]" gridcol="4" gridrow="26" pageId="5" pageNumber="6">6.2</td>
</tr>
<tr id="35353F61FF93004BFE07F92F0FF6F906" box="[495,1432,1679,1697]" gridrow="27" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F92F09CEF906" box="[495,928,1679,1697]" gridcol="0" gridrow="27" pageId="5" pageNumber="6">Head depth</th>
<td id="76E4561DFF93004BFC31F92F0E41F906" box="[985,1071,1679,1697]" gridcol="1" gridrow="27" pageId="5" pageNumber="6">53.2</td>
<td id="76E4561DFF93004BFB93F92F0EA0F906" box="[1147,1230,1679,1697]" gridcol="2" gridrow="27" pageId="5" pageNumber="6">30.7–53.2</td>
<td id="76E4561DFF93004BFAE7F92F0F2DF906" box="[1295,1347,1679,1697]" gridcol="3" gridrow="27" pageId="5" pageNumber="6">39.5</td>
<td id="76E4561DFF93004BFA9BF92F0FF6F906" box="[1395,1432,1679,1697]" gridcol="4" gridrow="27" pageId="5" pageNumber="6">5.4</td>
</tr>
<tr id="35353F61FF93004BFE07F91C0FF6F969" box="[495,1432,1724,1742]" gridrow="28" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F91C09CEF969" box="[495,928,1724,1742]" gridcol="0" gridrow="28" pageId="5" pageNumber="6">Maxillary-barbel length</th>
<td id="76E4561DFF93004BFC31F91C0E41F969" box="[985,1071,1724,1742]" gridcol="1" gridrow="28" pageId="5" pageNumber="6">90.7</td>
<td id="76E4561DFF93004BFB93F91C0EA0F969" box="[1147,1230,1724,1742]" gridcol="2" gridrow="28" pageId="5" pageNumber="6">47.5–94.0</td>
<td id="76E4561DFF93004BFAE7F91C0F2DF969" box="[1295,1347,1724,1742]" gridcol="3" gridrow="28" pageId="5" pageNumber="6">72.8</td>
<td id="76E4561DFF93004BFA9BF91C0FF6F969" box="[1395,1432,1724,1742]" gridcol="4" gridrow="28" pageId="5" pageNumber="6">12.2</td>
</tr>
<tr id="35353F61FF93004BFE07F9490FF6F95C" box="[495,1432,1769,1787]" gridrow="29" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F94909CEF95C" box="[495,928,1769,1787]" gridcol="0" gridrow="29" pageId="5" pageNumber="6">Distance between anterior and posterior nostrils</th>
<td id="76E4561DFF93004BFC31F9490E41F95C" box="[985,1071,1769,1787]" gridcol="1" gridrow="29" pageId="5" pageNumber="6">18.8</td>
<td id="76E4561DFF93004BFB93F9490EA0F95C" box="[1147,1230,1769,1787]" gridcol="2" gridrow="29" pageId="5" pageNumber="6">11.4–22.5</td>
<td id="76E4561DFF93004BFAE7F9490F2DF95C" box="[1295,1347,1769,1787]" gridcol="3" gridrow="29" pageId="5" pageNumber="6">16.4</td>
<td id="76E4561DFF93004BFA9BF9490FF6F95C" box="[1395,1432,1769,1787]" gridcol="4" gridrow="29" pageId="5" pageNumber="6">2.9</td>
</tr>
<tr id="35353F61FF93004BFE07F8B60FF6F88F" box="[495,1432,1814,1832]" gridrow="30" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F8B609CEF88F" box="[495,928,1814,1832]" gridcol="0" gridrow="30" pageId="5" pageNumber="6">Distance between the posterior nostril and eye</th>
<td id="76E4561DFF93004BFC31F8B60E41F88F" box="[985,1071,1814,1832]" gridcol="1" gridrow="30" pageId="5" pageNumber="6">7.3</td>
<td id="76E4561DFF93004BFB93F8B60EA0F88F" box="[1147,1230,1814,1832]" gridcol="2" gridrow="30" pageId="5" pageNumber="6">3.5–10.1</td>
<td id="76E4561DFF93004BFAE7F8B60F2DF88F" box="[1295,1347,1814,1832]" gridcol="3" gridrow="30" pageId="5" pageNumber="6">6.3</td>
<td id="76E4561DFF93004BFA9BF8B60FF6F88F" box="[1395,1432,1814,1832]" gridcol="4" gridrow="30" pageId="5" pageNumber="6">1.7</td>
</tr>
<tr id="35353F61FF93004BFE07F8E20FF6F8F3" box="[495,1432,1858,1876]" gridrow="31" pageId="5" pageNumber="6">
<th id="76E4561DFF93004BFE07F8E209CEF8F3" box="[495,928,1858,1876]" gridcol="0" gridrow="31" pageId="5" pageNumber="6">Distance between posterior nostrils</th>
<td id="76E4561DFF93004BFC31F8E20E41F8F3" box="[985,1071,1858,1876]" gridcol="1" gridrow="31" pageId="5" pageNumber="6">20.2</td>
<td id="76E4561DFF93004BFB93F8E20EA0F8F3" box="[1147,1230,1858,1876]" gridcol="2" gridrow="31" pageId="5" pageNumber="6">18.8–25.1</td>
<td id="76E4561DFF93004BFAE7F8E20F2DF8F3" box="[1295,1347,1858,1876]" gridcol="3" gridrow="31" pageId="5" pageNumber="6">21.5</td>
<td id="76E4561DFF93004BFA9BF8E20FF6F8F3" box="[1395,1432,1858,1876]" gridcol="4" gridrow="31" pageId="5" pageNumber="6">1.7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBA3D23FF90FFB2FDEEFD930E4FFC66" blockId="6.[486,1462,190,961]" pageId="6" pageNumber="7">
First dorsal-fin ray forming spine locking mechanism. Second dorsal-fin ray modified in a spine, smooth on anterior margin and serrated on posterior margin. Adiposefin base broad, generally larger than dorsal-fin base. Pectoral and pelvic fins roughly triangular, distal margin larger than base. First pectoral-fin ray as spine with retrorse serrations on anterior and posterior margin (
<figureCitation id="133E21A6FF90FFB2FC37FD680E4CFD46" box="[991,1058,712,737]" captionStart="FIGURE 3" captionStartId="7.[488,561,529,549]" captionTargetBox="[527,1431,207,486]" captionTargetId="figure-332@7.[488,1459,183,507]" captionTargetPageId="7" captionText="FIGURE 3 | Rhyacoglanis beninei pectoral-fin spine, paratype, LBP 32145, 50.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.11125421" httpUri="https://zenodo.org/record/11125421/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
). Pectoral, pelvic, anal, and adipose fins distal margins round. Caudal fin bifurcated with acute dorsal and ventral lobes. Dorsal-fin rays II,6* (1 C&amp;S, 11 eth); pectoral-fin rays I,6* (1 C&amp;S, 13 eth); pelvic-fin rays i,5* (1 C&amp;S, 13 eth); anal-fin rays iii,5 (2 eth), iv,6 (1 C&amp;S) or iii,6 (5), iii,7* (1); caudal-fin rays i,6,8,i* (1 C&amp;S, 11 eth) or i,5,7,i (1 eth). Dorsal procurrent caudal-fin rays 16 (1 C&amp;S), ventral procurrent caudal-fin rays 12 (1 C&amp;S). Branchiostegal rays 8 (1 C&amp;S). Total vertebrae 33 (1 C&amp;S). Ribs 9 (1 C&amp;S).
</paragraph>
<caption id="DF7A6DABFF90FFB2FF60F9A10EFBF99C" ID-DOI="http://doi.org/10.5281/zenodo.11125417" ID-Zenodo-Dep="11125417" httpUri="https://zenodo.org/record/11125417/files/figure.png" pageId="6" pageNumber="7" startId="6.[136,209,1537,1557]" targetBox="[136,1459,1046,1514]" targetPageId="6" targetType="figure">
<paragraph id="8BBA3D23FF90FFB2FF60F9A10EFBF99C" blockId="6.[136,1458,1537,1595]" pageId="6" pageNumber="7">
<emphasis id="B971E131FF90FFB2FF60F9A10A9CF9B2" bold="true" box="[136,242,1537,1557]" pageId="6" pageNumber="7">FIGURE 2 |</emphasis>
Pigmentation of oblique dark bars in the predorsal region of 
<taxonomicName id="4C0546A0FF90FFB2FCD9F9A10988F9B2" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[817,998,1537,1557]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF90FFB2FCD9F9A10988F9B2" bold="true" box="[817,998,1537,1557]" italics="true" pageId="6" pageNumber="7">Rhyacoglanis beninei</emphasis>
</taxonomicName>
. 
<emphasis id="B971E131FF90FFB2FC19F9A10E66F9B3" bold="true" box="[1009,1032,1537,1556]" pageId="6" pageNumber="7">A.</emphasis>
MZUEL 23049, 29.6 mm SL; 
<emphasis id="B971E131FF90FFB2FAE7F9A20F4AF9B2" bold="true" box="[1295,1316,1538,1557]" pageId="6" pageNumber="7">B.</emphasis>
LBP 32145, 32.9 mm SL; 
<emphasis id="B971E131FF90FFB2FF3CF9870A86F99D" bold="true" box="[212,232,1575,1594]" pageId="6" pageNumber="7">C.</emphasis>
LBP 32145, 37.3 mm SL; 
<emphasis id="B971E131FF90FFB2FE20F9870BB1F99D" bold="true" box="[456,479,1575,1594]" pageId="6" pageNumber="7">D.</emphasis>
MZUEL 23049, 42.6 mm SL; 
<emphasis id="B971E131FF90FFB2FD01F9870893F99D" bold="true" box="[745,765,1575,1594]" pageId="6" pageNumber="7">E.</emphasis>
LBP 32145, 50.2 mm SL. Scale bars = 10 mm.
</paragraph>
</caption>
<caption id="DF7A6DABFF91FFB3FE00FDB10E92FD82" ID-DOI="http://doi.org/10.5281/zenodo.11125421" ID-Zenodo-Dep="11125421" box="[488,1276,529,549]" httpUri="https://zenodo.org/record/11125421/files/figure.png" pageId="7" pageNumber="8" startId="7.[488,561,529,549]" targetBox="[527,1431,207,486]" targetPageId="7" targetType="figure">
<paragraph id="8BBA3D23FF91FFB3FE00FDB10E92FD82" blockId="7.[488,1276,529,549]" box="[488,1276,529,549]" pageId="7" pageNumber="8">
<emphasis id="B971E131FF91FFB3FE00FDB10961FD82" bold="true" box="[488,783,529,549]" pageId="7" pageNumber="8">
FIGURE 3 | 
<taxonomicName id="4C0546A0FF91FFB3FDB1FDB10961FD82" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[601,783,529,549]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF91FFB3FDB1FDB10961FD82" bold="true" box="[601,783,529,549]" italics="true" pageId="7" pageNumber="8">Rhyacoglanis beninei</emphasis>
</taxonomicName>
</emphasis>
pectoral-fin spine, paratype, LBP 32145, 50.2 mm SL.
</paragraph>
</caption>
<paragraph id="8BBA3D23FF91FFB3FDEEFDDE0899FB61" blockId="7.[485,1459,638,1222]" pageId="7" pageNumber="8">
<emphasis id="B971E131FF91FFB3FDEEFDDE0960FD31" bold="true" box="[518,782,638,662]" pageId="7" pageNumber="8">Coloration in alcohol.</emphasis>
Ground color yellowish to brownish, with three conspicuous dark brown bands on body (
<figureCitation id="133E21A6FF91FFB3FCF2FD03090DFD1C" box="[794,867,675,699]" captionStart="FIGURE 1" captionStartId="4.[488,561,1612,1632]" captionTargetBox="[488,1459,190,1590]" captionTargetId="figure-120@4.[488,1459,190,1590]" captionTargetPageId="4" captionText="FIGURE 1 | Rhyacoglanis beninei, holotype, MZUSP 127014, 59.1 mm SL, from córrego Jussara, an affluent of Jamanxim River, Tapajós River basin. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125415" httpUri="https://zenodo.org/record/11125415/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
, 
<figureCitation id="133E21A6FF91FFB3FC87FD030911FD1C" box="[879,895,675,699]" captionStart="FIGURE 2" captionStartId="6.[136,209,1537,1557]" captionTargetBox="[136,1459,1046,1514]" captionTargetId="figure-336@6.[136,1462,1046,1517]" captionTargetPageId="6" captionText="FIGURE 2 | Pigmentation of oblique dark bars in the predorsal region of Rhyacoglanis beninei. A. MZUEL 23049, 29.6 mm SL; B. LBP 32145, 32.9 mm SL; C. LBP 32145, 37.3 mm SL; D. MZUEL 23049, 42.6 mm SL; E. LBP 32145, 50.2 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125417" httpUri="https://zenodo.org/record/11125417/files/figure.png" pageId="7" pageNumber="8">2</figureCitation>
, 
<figureCitation id="133E21A6FF91FFB3FC60FD0309F9FD1C" box="[904,919,675,699]" captionStart="FIGURE 5" captionStartId="8.[136,209,1761,1781]" captionTargetBox="[136,1459,1235,1739]" captionTargetId="figure-70@8.[136,1459,1235,1739]" captionTargetPageId="8" captionText="FIGURE 5 | Variation pattern of dark body bands in Rhyacoglanis beninei. A. MZUEL 23049, 42.6 mm SL; B. LBP 32145, 29.8 mm SL;C. LBP 32163, 27.0 mm SL; D. LBP 32163, 42.9 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125429" httpUri="https://zenodo.org/record/11125429/files/figure.png" pageId="7" pageNumber="8">5</figureCitation>
). The first dark band under dorsal fin (subdorsal), the second under adipose fin (subadipose), and the third posterior adipose fin, extending to the caudal-fin base. Subadipose and caudal peduncle bands surrounding body; subdorsal band open in ventral region; all bands with variable connections pattern on lateral side (
<figureCitation id="133E21A6FF91FFB3FD72FC9808B1FCF7" box="[666,735,824,848]" captionStart="FIGURE 5" captionStartId="8.[136,209,1761,1781]" captionTargetBox="[136,1459,1235,1739]" captionTargetId="figure-70@8.[136,1459,1235,1739]" captionTargetPageId="8" captionText="FIGURE 5 | Variation pattern of dark body bands in Rhyacoglanis beninei. A. MZUEL 23049, 42.6 mm SL; B. LBP 32145, 29.8 mm SL;C. LBP 32163, 27.0 mm SL; D. LBP 32163, 42.9 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125429" httpUri="https://zenodo.org/record/11125429/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
) in individuals of different SL, sometimes not present in small juveniles (&lt;
<quantity id="4CFD90C6FF91FFB3FD83FCFE08D6FCD2" box="[619,696,862,886]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="8" unit="mm" value="28.0">28 mm</quantity>
SL – 
<figureCitation id="133E21A6FF91FFB3FD1CFCFE0925FCD1" box="[756,843,862,886]" captionStart="FIGURE 5" captionStartId="8.[136,209,1761,1781]" captionTargetBox="[136,1459,1235,1739]" captionTargetId="figure-70@8.[136,1459,1235,1739]" captionTargetPageId="8" captionText="FIGURE 5 | Variation pattern of dark body bands in Rhyacoglanis beninei. A. MZUEL 23049, 42.6 mm SL; B. LBP 32145, 29.8 mm SL;C. LBP 32163, 27.0 mm SL; D. LBP 32163, 42.9 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125429" httpUri="https://zenodo.org/record/11125429/files/figure.png" pageId="7" pageNumber="8">Fig. 5C</figureCitation>
). Two oblique dark brown bands on predorsal region in dorsal view, each beginning just laterally to parieto-supraoccipital process, continuing towards posterior cleithral process (
<figureCitation id="133E21A6FF91FFB3FC9AFC0809D8FC66" box="[882,950,936,961]" captionStart="FIGURE 2" captionStartId="6.[136,209,1537,1557]" captionTargetBox="[136,1459,1046,1514]" captionTargetId="figure-336@6.[136,1462,1046,1517]" captionTargetPageId="6" captionText="FIGURE 2 | Pigmentation of oblique dark bars in the predorsal region of Rhyacoglanis beninei. A. MZUEL 23049, 29.6 mm SL; B. LBP 32145, 32.9 mm SL; C. LBP 32145, 37.3 mm SL; D. MZUEL 23049, 42.6 mm SL; E. LBP 32145, 50.2 mm SL. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.11125417" httpUri="https://zenodo.org/record/11125417/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
). Pectoral and pelvic fins hyaline with middle dark brown stripe and numerous melanophores dispersed. Anal-fin hyaline with middle dark brown stripe and numerous melanophores spread; fin base with dark brown band and first unbranched ray base with clear dot. Caudal fin hyaline, dorsal, and ventral lobes with middle dark brown stripe and numerous melanophores dispersed. Dorsal-fin dark brown with numerous melanophores on hyaline margin. Adipose fin dark brown, clear dot on anterior base portion with numerous melanophores. Head grayish with lighter region on cheek.
</paragraph>
<paragraph id="8BBA3D23FF91FFB3FDEEFB580E33FA91" blockId="7.[488,1459,1272,1334]" pageId="7" pageNumber="8">
<emphasis id="B971E131FF91FFB3FDEEFB580884FAB7" bold="true" box="[518,746,1272,1296]" pageId="7" pageNumber="8">Coloration in life.</emphasis>
Same as alcohol-preserved specimens, but the light regions brighter, ranging from yellowish to brownish (
<figureCitation id="133E21A6FF91FFB3FC10FABE0E3FFA91" box="[1016,1105,1310,1334]" captionStart="FIGURE 4" captionStartId="8.[488,561,1089,1109]" captionTargetBox="[488,1447,189,1067]" captionTargetId="figure-31@8.[488,1459,189,1067]" captionTargetPageId="8" captionText="FIGURE 4 | A. Habitat of Rhyacoglanis beninei in córrego Jussara, 07°21’08”S 55°17’45”W; B. A rock where specimens of R. beninei were associated; C. Paratype of R. beninei just after capture. Photo: Gabriel S. Costa e Silva." figureDoi="http://doi.org/10.5281/zenodo.11125425" httpUri="https://zenodo.org/record/11125425/files/figure.png" pageId="7" pageNumber="8">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BBA3D23FF91FFB3FDEEFAC80996FA56" blockId="7.[488,1461,1384,1521]" pageId="7" pageNumber="8">
<emphasis id="B971E131FF91FFB3FDEEFAC80920FA27" bold="true" box="[518,846,1384,1408]" pageId="7" pageNumber="8">Geographical distribution.</emphasis>
<taxonomicName id="4C0546A0FF91FFB3FCB4FAC80E50FA27" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[860,1086,1384,1408]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF91FFB3FCB4FAC80E50FA27" box="[860,1086,1384,1408]" italics="true" pageId="7" pageNumber="8">Rhyacoglanis beninei</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="54BE8381FF91FFB3FA6AFAC90FDDFA26" box="[1410,1459,1385,1409]" pageId="7" pageNumber="8">type</typeStatus>
locality in the 
<location id="8EDA6BF8FF91FFB3FD67FA2E0927FA01" LSID="urn:lsid:plazi:treatment:03AC8C35FF95FFBDFCE8FC980EFAF8E6:8EDA6BF8FF91FFB3FD67FA2E0927FA01" box="[655,841,1422,1446]" country="Brazil" county="Corrego Jussara" municipality="The" name="Jamanxim River" pageId="7" pageNumber="8">Jamanxim River</location>
, 
<location id="8EDA6BF8FF91FFB3FCBEFA2E099AFA01" LSID="urn:lsid:plazi:treatment:03AC8C35FF95FFBDFCE8FC980EFAF8E6:8EDA6BF8FF91FFB3FCBEFA2E099AFA01" box="[854,1012,1422,1446]" country="Brazil" county="Corrego Jussara" municipality="The" name="Tapajos River" pageId="7" pageNumber="8">Tapajós River</location>
basin, 
<collectingCountry id="F3127DB3FF91FFB3FBAFFA2E0EE7FA01" box="[1095,1161,1422,1446]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
(
<figureCitation id="133E21A6FF91FFB3FB72FA2E0E8EFA01" box="[1178,1248,1422,1446]" captionStart="FIGURE 6" captionStartId="9.[136,209,1201,1221]" captionTargetBox="[136,1442,175,1173]" captionTargetId="figure-40@9.[136,1459,175,1179]" captionTargetPageId="9" captionText="FIGURE 6 | The type locality of Rhyacoglanis beninei in córrego Jussara, 07°21’08”S 55°17’45”W (yellow star), Tapajós River basin." figureDoi="http://doi.org/10.5281/zenodo.11125431" httpUri="https://zenodo.org/record/11125431/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
). 
<collectingMunicipality id="6BDEA759FF91FFB3FB1EFA2E0F4AFA01" box="[1270,1316,1422,1446]" pageId="7" pageNumber="8">The</collectingMunicipality>
new species was collected in fast-flowing currents of 
<collectingCounty id="62DB45AFFF91FFB3FC50FA130E03FA6C" box="[952,1133,1459,1483]" pageId="7" pageNumber="8">Córrego Jussara</collectingCounty>
, characterized by clear water and the bottom with rocks and gravels (
<figureCitation id="133E21A6FF91FFB3FC4FFA780985FA56" box="[935,1003,1496,1521]" captionStart="FIGURE 4" captionStartId="8.[488,561,1089,1109]" captionTargetBox="[488,1447,189,1067]" captionTargetId="figure-31@8.[488,1459,189,1067]" captionTargetPageId="8" captionText="FIGURE 4 | A. Habitat of Rhyacoglanis beninei in córrego Jussara, 07°21’08”S 55°17’45”W; B. A rock where specimens of R. beninei were associated; C. Paratype of R. beninei just after capture. Photo: Gabriel S. Costa e Silva." figureDoi="http://doi.org/10.5281/zenodo.11125425" httpUri="https://zenodo.org/record/11125425/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F6EA8FF91FFBDFDEEF9830EFAF8E6" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BBA3D23FF91FFB3FDEEF98308C9F90C" blockId="7.[488,1459,1571,1707]" pageId="7" pageNumber="8">
<emphasis id="B971E131FF91FFB3FDEEF98308FFF99C" bold="true" box="[518,657,1571,1595]" pageId="7" pageNumber="8">Etymology.</emphasis>
<taxonomicName id="4C0546A0FF91FFB3FD74F9830912F99C" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[668,892,1571,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF91FFB3FD74F9830912F99C" box="[668,892,1571,1595]" italics="true" pageId="7" pageNumber="8">Rhyacoglanis beninei</emphasis>
</taxonomicName>
is named in honor of Ricardo Cardoso Benine, Professor at Universidade Estadual Paulista “Júlio de Mesquita Filho”, in recognition of his dedication and remarkable contributions to the knowledge of Neotropical freshwater fishes.
</paragraph>
<caption id="DF7A6DABFF9EFFBCFE00FBE1080BFB07" ID-DOI="http://doi.org/10.5281/zenodo.11125425" ID-Zenodo-Dep="11125425" httpUri="https://zenodo.org/record/11125425/files/figure.png" pageId="8" pageNumber="9" startId="8.[488,561,1089,1109]" targetBox="[488,1447,189,1067]" targetPageId="8" targetType="figure">
<paragraph id="8BBA3D23FF9EFFBCFE00FBE1080BFB07" blockId="8.[487,1460,1089,1184]" pageId="8" pageNumber="9">
<emphasis id="B971E131FF9EFFBCFE00FBE1081EFBF3" bold="true" box="[488,624,1089,1109]" pageId="8" pageNumber="9">FIGURE 4 | A.</emphasis>
Habitat of 
<taxonomicName id="4C0546A0FF9EFFBCFD31FBE109E1FBF2" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[729,911,1089,1109]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9EFFBCFD31FBE109E1FBF2" bold="true" box="[729,911,1089,1109]" italics="true" pageId="8" pageNumber="9">Rhyacoglanis beninei</emphasis>
</taxonomicName>
in córrego Jussara, 07°21’08”S 55°17’45”W; 
<emphasis id="B971E131FF9EFFBCFAF3FBE20F5FFBF2" bold="true" box="[1307,1329,1090,1109]" pageId="8" pageNumber="9">B.</emphasis>
A rock where specimens of 
<taxonomicName id="4C0546A0FF9EFFBCFD84FBC708ADFBDC" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[620,707,1127,1147]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9EFFBCFD84FBC708ADFBDC" bold="true" box="[620,707,1127,1147]" italics="true" pageId="8" pageNumber="9">R. beninei</emphasis>
</taxonomicName>
were associated; 
<emphasis id="B971E131FF9EFFBCFC86FBC709EDFBDD" bold="true" box="[878,899,1127,1146]" pageId="8" pageNumber="9">C.</emphasis>
Paratype of 
<taxonomicName id="4C0546A0FF9EFFBCFC17FBC70E38FBDC" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[1023,1110,1127,1147]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9EFFBCFC17FBC70E38FBDC" bold="true" box="[1023,1110,1127,1147]" italics="true" pageId="8" pageNumber="9">R. beninei</emphasis>
</taxonomicName>
just after capture. Photo: Gabriel S. Costa e Silva.
</paragraph>
</caption>
<caption id="DF7A6DABFF9EFFBCFF60F94108C2F8BC" ID-DOI="http://doi.org/10.5281/zenodo.11125429" ID-Zenodo-Dep="11125429" httpUri="https://zenodo.org/record/11125429/files/figure.png" pageId="8" pageNumber="9" startId="8.[136,209,1761,1781]" targetBox="[136,1459,1235,1739]" targetPageId="8" targetType="figure">
<paragraph id="8BBA3D23FF9EFFBCFF60F94108C2F8BC" blockId="8.[135,1460,1761,1819]" pageId="8" pageNumber="9">
<emphasis id="B971E131FF9EFFBCFF60F9410A9FF952" bold="true" box="[136,241,1761,1781]" pageId="8" pageNumber="9">FIGURE 5 |</emphasis>
Variation pattern of dark body bands in 
<taxonomicName id="4C0546A0FF9EFFBCFD84F941094FF952" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[620,801,1761,1781]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9EFFBCFD84F941094FF952" bold="true" box="[620,801,1761,1781]" italics="true" pageId="8" pageNumber="9">Rhyacoglanis beninei</emphasis>
</taxonomicName>
. 
<emphasis id="B971E131FF9EFFBCFCC4F941092DF953" bold="true" box="[812,835,1761,1780]" pageId="8" pageNumber="9">A.</emphasis>
MZUEL 23049, 42.6 mm SL; 
<emphasis id="B971E131FF9EFFBCFBAFF9420E33F952" bold="true" box="[1095,1117,1762,1781]" pageId="8" pageNumber="9">B.</emphasis>
LBP 32145, 29.8 mm SL; 
<emphasis id="B971E131FF9EFFBCFAD1F9410F20F953" bold="true" box="[1337,1358,1761,1780]" pageId="8" pageNumber="9">C.</emphasis>
LBP 32163, 27.0 mm SL; 
<emphasis id="B971E131FF9EFFBCFF15F8A70B7BF8BD" bold="true" box="[253,277,1799,1818]" pageId="8" pageNumber="9">D.</emphasis>
LBP 32163, 42.9 mm SL. Scale bars = 10 mm.
</paragraph>
</caption>
<caption id="DF7A6DABFF9FFFBDFF60FB110F43FB62" ID-DOI="http://doi.org/10.5281/zenodo.11125431" ID-Zenodo-Dep="11125431" box="[136,1325,1201,1221]" httpUri="https://zenodo.org/record/11125431/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,209,1201,1221]" targetBox="[136,1442,175,1173]" targetPageId="9" targetType="figure">
<paragraph id="8BBA3D23FF9FFFBDFF60FB110F43FB62" blockId="9.[136,1325,1201,1221]" box="[136,1325,1201,1221]" pageId="9" pageNumber="10">
<emphasis id="B971E131FF9FFFBDFF60FB110A9DFB62" bold="true" box="[136,243,1201,1221]" pageId="9" pageNumber="10">FIGURE 6 |</emphasis>
The type locality of 
<taxonomicName id="4C0546A0FF9FFFBDFE5BFB110806FB62" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[435,616,1201,1221]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9FFFBDFE5BFB110806FB62" bold="true" box="[435,616,1201,1221]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis beninei</emphasis>
</taxonomicName>
in córrego Jussara, 07°21’08”S 55°17’45”W (yellow star), Tapajós River basin.
</paragraph>
</caption>
<paragraph id="8BBA3D23FF9FFFBDFDEEFB5809CDFA6C" blockId="9.[486,1459,1272,1484]" pageId="9" pageNumber="10">
<emphasis id="B971E131FF9FFFBDFDEEFB580892FAB6" bold="true" box="[518,764,1272,1297]" pageId="9" pageNumber="10">Conservation status.</emphasis>
All specimens of 
<taxonomicName id="4C0546A0FF9FFFBDFC24FB580EC4FAB7" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[972,1194,1272,1296]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9FFFBDFC24FB580EC4FAB7" box="[972,1194,1272,1296]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis beninei</emphasis>
</taxonomicName>
were collected at only one sample site. However, several specimens were collected at the 
<typeStatus id="54BE8381FF9FFFBDFAF3FABE0F22FA91" box="[1307,1356,1310,1334]" pageId="9" pageNumber="10">type</typeStatus>
locality, indicating a common occurrence and no apparent threat of extinction. 
<location id="8EDA6BF8FF9FFFBDFB1EFAE30F03FAFC" LSID="urn:lsid:plazi:treatment:03AC8C35FF95FFBDFCE8FC980EFAF8E6:8EDA6BF8FF9FFFBDFB1EFAE30F03FAFC" box="[1270,1389,1347,1371]" county="Conservation of Nature" municipality="International Union" name="According" pageId="9" pageNumber="10">According</location>
to the 
<collectingMunicipality id="6BDEA759FF9FFFBDFE0EFAC808A9FA27" box="[486,711,1384,1408]" pageId="9" pageNumber="10">International Union</collectingMunicipality>
for 
<collectingCounty id="62DB45AFFF9FFFBDFD1CFAC80993FA27" box="[756,1021,1384,1408]" pageId="9" pageNumber="10">Conservation of Nature</collectingCounty>
(
<collectionCode id="ED14A5E6FF9FFFBDFBE4FAC90E39FA26" box="[1036,1111,1385,1409]" pageId="9" pageNumber="10">IUCN</collectionCode>
) categories and criteria (
<collectionCode id="ED14A5E6FF9FFFBDFA8FFAC90FDCFA26" box="[1383,1458,1385,1409]" pageId="9" pageNumber="10">IUCN</collectionCode>
Standards and Petitions Subcommittee, 2023), we propose classifying 
<taxonomicName id="4C0546A0FF9FFFBDFAC1FA2E085FFA6C" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9FFFBDFAC1FA2E085FFA6C" italics="true" pageId="9" pageNumber="10">Rhyacoglanis beninei</emphasis>
</taxonomicName>
as category Least Concern (
<collectionCode id="ED14A5E6FF9FFFBDFC86FA1309F8FA6C" box="[878,918,1459,1483]" pageId="9" pageNumber="10">LC</collectionCode>
).
</paragraph>
<paragraph id="8BBA3D23FF9FFFBDFDEEFA5E0EFAF8E6" blockId="9.[486,1465,1534,1857]" pageId="9" pageNumber="10">
<emphasis id="B971E131FF9FFFBDFDEEFA5E08E2F9B1" bold="true" box="[518,652,1534,1558]" pageId="9" pageNumber="10">Phylogeny.</emphasis>
Sequencing and data filtering yielded a 70% complete matrix with 1082 loci and 385,841 bp. Results support the monophyly of 
<taxonomicName id="4C0546A0FF9FFFBDFB32F9830FDDF99C" box="[1242,1459,1571,1595]" family="Pseudopimelodidae" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="family">Pseudopimelodidae</taxonomicName>
and the monophyly of Pseudopimelodinae and Batrochoglaninae subfamilies. Inside Pseudopimelodinae, 
<taxonomicName id="4C0546A0FF9FFFBDFD30F9CE0921F921" authorityName="Ortega-Lara &amp; Lehmann" authorityYear="2006" box="[728,847,1646,1670]" family="Pseudopimelodidae" genus="Cruciglanis" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B971E131FF9FFFBDFD30F9CE0921F921" box="[728,847,1646,1670]" italics="true" pageId="9" pageNumber="10">Cruciglanis</emphasis>
</taxonomicName>
is the sister to 
<taxonomicName id="4C0546A0FF9FFFBDFBF6F9CE0EBEF921" authorityName="Bleeker" authorityYear="1858" box="[1054,1232,1646,1670]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B971E131FF9FFFBDFBF6F9CE0EBEF921" box="[1054,1232,1646,1670]" italics="true" pageId="9" pageNumber="10">Pseudopimelodus</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C0546A0FF9FFFBDFB16F9CE0FE6F921" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1278,1416,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B971E131FF9FFFBDFB16F9CE0FE6F921" box="[1278,1416,1646,1670]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis</emphasis>
</taxonomicName>
, as supported by 
<bibRefCitation id="EF9440D2FF9FFFBDFD61F9330939F90C" author="In Silva" box="[649,855,1683,1707]" pageId="9" pageNumber="10" pagination="2021" refId="ref4507" refString="We also re-build a phylogeny of Pseudopimelodus, using the UCEs matrix dataset published by Silva et al. (2021), adding Pseudopimelodus charus and P. bufonius. In Silva et al. (2021), the monophyly of Pseudopimelodus was impossible to test since only one species (P. mangurus) was included in that phylogeny. Rangel-Mendrano et al. (2021), in a genetic analysis using a multi-locus approach, supported the monophyly of Pseudopimelodus, with P. charus (from the Sao Francisco basin) sister to P. bufonius (Amazon basin) + P. mangurus (Parana basin). Our result also supports the monophyly of Pseudopimelodus but places P. bufonius sister to P. charus plus P. mangurus. This arrangement is congruent with other molecular phylogenetic studies, under which monophyletic components from the upper Parana and Sao Francisco river basins have been recognized as sister group (Montoya-Burgos, 2003; Roxo et al., 2014; Ochoa et al., 2020)." type="book chapter" year="2021">
Silva 
<emphasis id="B971E131FF9FFFBDFD21F9340894F90C" box="[713,762,1683,1707]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (2021)
</bibRefCitation>
. Internally to 
<taxonomicName id="4C0546A0FF9FFFBDFBE8F9330EDCF90C" authorityName="Bleeker" authorityYear="1858" box="[1024,1202,1683,1707]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B971E131FF9FFFBDFBE8F9330EDCF90C" box="[1024,1202,1683,1707]" italics="true" pageId="9" pageNumber="10">Pseudopimelodus</emphasis>
</taxonomicName>
, our analysis found 
<taxonomicName id="4C0546A0FF9FFFBDFA4BF9340851F977" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bufonius">
<emphasis id="B971E131FF9FFFBDFA4BF9340851F977" italics="true" pageId="9" pageNumber="10">P. bufonius</emphasis>
</taxonomicName>
sister to 
<taxonomicName id="4C0546A0FF9FFFBDFD75F919089AF977" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[669,756,1720,1744]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="charus">
<emphasis id="B971E131FF9FFFBDFD75F919089AF977" box="[669,756,1720,1744]" italics="true" pageId="9" pageNumber="10">P. charus</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C0546A0FF9FFFBDFCE7F9190E14F977" authority="(Valenciennes, 1835)" baseAuthorityName="Valenciennes" baseAuthorityYear="1835" box="[783,1146,1720,1744]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mangurus">
<emphasis id="B971E131FF9FFFBDFCE7F91909E5F977" box="[783,907,1721,1744]" italics="true" pageId="9" pageNumber="10">P. mangurus</emphasis>
(Valenciennes, 1835)
</taxonomicName>
. 
<taxonomicName id="4C0546A0FF9FFFBDFB6CF9180F60F977" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1156,1294,1720,1744]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B971E131FF9FFFBDFB6CF9180F60F977" box="[1156,1294,1720,1744]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis</emphasis>
</taxonomicName>
also recovered as a monophyletic group, where 
<taxonomicName id="4C0546A0FF9FFFBDFCB4F97F09A8F951" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[860,966,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="B971E131FF9FFFBDFCB4F97F09A8F951" box="[860,966,1758,1782]" italics="true" pageId="9" pageNumber="10">R. pulcher</emphasis>
</taxonomicName>
was recovered as sister to a clade composed of two subclades, the first composed of 
<taxonomicName id="4C0546A0FF9FFFBDFC50F8A40E24F8BC" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[952,1098,1796,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="semininger">
<emphasis id="B971E131FF9FFFBDFC50F8A40E24F8BC" box="[952,1098,1796,1819]" italics="true" pageId="9" pageNumber="10">R. semininger</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C0546A0FF9FFFBDFB85F8A40E94F8BC" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1133,1274,1796,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="B971E131FF9FFFBDFB85F8A40E94F8BC" box="[1133,1274,1796,1819]" italics="true" pageId="9" pageNumber="10">R. paranensis</emphasis>
</taxonomicName>
and the second composed of 
<taxonomicName id="4C0546A0FF9FFFBDFD95F8890888F8E7" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[637,742,1832,1856]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="beninei">
<emphasis id="B971E131FF9FFFBDFD95F8890888F8E7" box="[637,742,1832,1856]" italics="true" pageId="9" pageNumber="10">R. beninei</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C0546A0FF9FFFBDFCEAF88809E2F8E7" authority="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira, 2023" authorityName="Crispim-Rodrigues &amp; Silva &amp; Shibatta &amp; Kuranaka &amp; Oliveira" authorityYear="2023" box="[770,908,1832,1856]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B971E131FF9FFFBDFCEAF88809E2F8E7" box="[770,908,1832,1856]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2425C4AFF9FFFBDFC7BF88909A1F8E7" box="[915,975,1833,1856]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
“Xingu” (
<figureCitation id="133E21A6FF9FFFBDFBABF8880EE9F8E6" box="[1091,1159,1832,1857]" captionStart="FIGURE 7" captionStartId="10.[136,209,1089,1109]" captionTargetBox="[160,1443,213,1040]" captionTargetId="figure-76@10.[136,1459,184,1066]" captionTargetPageId="10" captionText="FIGURE 7 | Maximum likelihood tree of Pseudopimelodidae on the 70% complete matrix (1082 loci, 385,841 bp). Nodes without symbols represent 100% support from 1000 bootstrap pseudo-replicates; nodal support between 99% and 75% denote by black circles; gray nodes indicate support between 75% and 50%." figureDoi="http://doi.org/10.5281/zenodo.11125435" httpUri="https://zenodo.org/record/11125435/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>