308 lines
37 KiB
XML
308 lines
37 KiB
XML
<document id="2F51FBE4FA6E40D7CE50328D9552CC28" ID-CLB-Dataset="7420" ID-DOI="10.11646/zootaxa.4940.1.1" ID-GBIF-Dataset="a09df80c-5d8b-44da-8f25-723f107c6c49" ID-ISSN="1175-5326" ID-Zenodo-Dep="4580917" ID-ZooBank="5F073867-F6EA-4222-A190-77F68076BE9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614844133957" checkinUser="plazi" docAuthor="Booysen, Ruan & Haddad, Charles R." docDate="2021" docId="03AC33149C0BFFFC7CBDFC9BFBB0F868" docLanguage="en" docName="zootaxa.4940.1.1.pdf" docOrigin="Zootaxa 4940 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Micaria plana Booysen & Haddad 2021, sp. nov." docType="treatment" docVersion="10" lastPageNumber="59" masterDocId="FF954B6C9C32FFC67C2AFFFCFFCEFFEC" masterDocTitle="Revision and molecular phylogeny of the spider genus Micaria Westring, 1851 (Araneae: Gnaphosidae) in the Afrotropical Region" masterLastPageNumber="82" masterPageNumber="1" pageNumber="58" updateTime="1698932415381" updateUser="ExternalLinkService">
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<mods:mods id="DBF833721A3CEFFC299641D8870342B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C17ECF4A0CFAC2F80C7CC6F576B986DF">
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<mods:title id="5F9064AD328E6C6872EE24F12B896874">Revision and molecular phylogeny of the spider genus Micaria Westring, 1851 (Araneae: Gnaphosidae) in the Afrotropical Region</mods:title>
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<mods:name id="65CC94951617D390121B99BEEAC8333E" type="personal">
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<mods:roleTerm id="E77F16A467E3F5EBFE9C4B5A77D14F42">Author</mods:roleTerm>
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<mods:namePart id="883764CE1A97F143CA477978AB6660E4">Booysen, Ruan</mods:namePart>
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<mods:nameIdentifier id="85D8B035906E938F406CAF7085234BD1" type="email">booysenr@ufs.ac.za</mods:nameIdentifier>
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<mods:roleTerm id="A06340A91C63BA30E981267A738D76E6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="52E9D41C26B5D72D615A79C999FA1584">Haddad, Charles R.</mods:namePart>
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<mods:nameIdentifier id="7C8161F1B911DB1BC5FEF3FA67732B1C" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
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<mods:nameIdentifier id="D3B3D76A72919141E04B5260A1103496" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:typeOfResource id="2A90187F61ABEE428A047978D4BEE5C5">text</mods:typeOfResource>
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<mods:title id="29B7F13C34456001731FA83D1212D976">Zootaxa</mods:title>
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<mods:part id="6109201FD8C1C6B49EA8E39004E42048">
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<mods:date id="82A8934D1172D19CE080BA0D937AE36A">2021</mods:date>
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<mods:number id="9E40221781181A09EA7430A05AB79238">2021-03-04</mods:number>
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<mods:detail id="9B78EAC3F11AE107E916C6FBC4D4BDAE" type="volume">
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<mods:number id="40BE4F02F4C82F07D7D5181F3A272A90">4940</mods:number>
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<mods:detail id="123BCA316D51A3015EC2A64726BACD0A" type="issue">
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<mods:number id="958473FE2187FB968162B7FF20891364">1</mods:number>
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<mods:start id="9FED8462D0756F9393F403041788FA9D">1</mods:start>
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<mods:classification id="D75B1AD86FCCEB14FD1C52AE33D4CDD5">journal article</mods:classification>
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<mods:identifier id="C0E21E32450F0C902EA1C96035D950F7" type="CLB-Dataset">7420</mods:identifier>
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<mods:identifier id="237E358121367E8E073E0F0503D5900E" type="DOI">10.11646/zootaxa.4940.1.1</mods:identifier>
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</mods:mods>
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<treatment id="03AC33149C0BFFFC7CBDFC9BFBB0F868" ID-DOI="http://doi.org/10.5281/zenodo.4681018" ID-GBIF-Taxon="180991658" ID-Zenodo-Dep="4681018" LSID="urn:lsid:plazi:treatment:03AC33149C0BFFFC7CBDFC9BFBB0F868" httpUri="http://treatment.plazi.org/id/03AC33149C0BFFFC7CBDFC9BFBB0F868" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
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<subSubSection id="C31FD1899C0BFFFF7CBDFC9BFE66FC6D" box="[151,424,870,897]" pageId="57" pageNumber="58" type="nomenclature">
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<paragraph id="8BBA82029C0BFFFF7CBDFC9BFE66FC6D" blockId="57.[151,472,870,933]" box="[151,424,870,897]" pageId="57" pageNumber="58">
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<heading id="D0F2356E9C0BFFFF7CBDFC9BFE66FC6D" bold="true" box="[151,424,870,897]" fontSize="11" level="2" pageId="57" pageNumber="58" reason="4">
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<emphasis id="B9715E109C0BFFFF7CBDFC9BFE66FC6D" bold="true" box="[151,424,870,897]" pageId="57" pageNumber="58">
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<taxonomicName id="4C05F9819C0BFFFF7CBDFC9BFE8DFC6D" ID-CoL="98Z5G" authority="Booysen & Haddad, 2021" authorityName="Booysen & Haddad" authorityYear="2021" box="[151,323,870,897]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="plana" status="sp. nov.">
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<emphasis id="B9715E109C0BFFFF7CBDFC9BFE8DFC6D" bold="true" box="[151,323,870,897]" italics="true" pageId="57" pageNumber="58">Micariaplana</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A242E36B9C0BFFFF7D60FC9BFE66FC6D" box="[330,424,871,897]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31FD1899C0BFFFF7CBDFC77FE16FC49" box="[151,472,906,933]" pageId="57" pageNumber="58" type="description">
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||
<paragraph id="8BBA82029C0BFFFF7CBDFC77FE16FC49" blockId="57.[151,472,870,933]" box="[151,472,906,933]" pageId="57" pageNumber="58">
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<figureCitation id="133E9E879C0BFFFF7CBDFC77FED8FC49" box="[151,278,907,933]" captionStart="FIGURES 27–36" captionStartId="13.[151,264,1855,1879]" captionTargetBox="[151,1435,181,1827]" captionTargetId="figure-17@13.[151,1435,181,1830]" captionTargetPageId="13" captionText="FIGURES 27–36. Stereomicroscope photographs of the dorsal habitus of Afrotropical Micaria species: 27 M. parvotibialis sp. nov. male; 28 M. plana sp. nov. female and 29 male; 30 M. quadrata sp. nov. female; 31 M. quinquemaculosa sp. nov. female and 32 male; 33 M. rivonosy sp. nov. female; 34 M. sanipass sp. nov. male; 35 M. scutellata sp. nov. male; 36 M. tersissima Simon, 1910 male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580933" httpUri="https://zenodo.org/record/4580933/files/figure.png" pageId="57" pageNumber="58">Figs 28, 29</figureCitation>
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,
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<figureCitation id="133E9E879C0BFFFF7D08FC77FE8EFC49" box="[290,320,907,933]" captionStart="FIGURES 61–64" captionStartId="19.[151,264,1949,1973]" captionTargetBox="[151,1436,902,1933]" captionTargetId="figure-1089@19.[151,1436,902,1933]" captionTargetPageId="19" captionText="FIGURES 61–64. Scanning electron microscope images of the lyriform organs of Micaria: 61 M. koingnaas sp. nov. male, patella I; 62 M. scutellata sp. nov. male, patella IV; 63 M. plana sp. nov. male, patella I; 64 M. quadrata sp. nov. female, patella I." figureDoi="http://doi.org/10.5281/zenodo.4580943" httpUri="https://zenodo.org/record/4580943/files/figure.png" pageId="57" pageNumber="58">63</figureCitation>
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,
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<figureCitation id="133E9E879C0BFFFF7D66FC77FEA5FC49" box="[332,363,907,933]" captionStart="FIGURES 65–71" captionStartId="20.[151,264,1580,1604]" captionTargetBox="[151,1436,181,1563]" captionTargetId="figure-31@20.[151,1436,181,1563]" captionTargetPageId="20" captionText="FIGURES 65–71. Scanning electron microscope images of the lanceolate setae of Micaria: 65 M. basaliducta sp. nov. male, patella I; 66 M. beaufortia (Tucker, 1923) male, palpal tibia; 67 M. bispicula sp. nov. male, patella I; 68 M. koingnaas sp. nov. male, femur I; 69 M. parvotibialis sp. nov. male, tarsus IV; 70 M. plana sp. nov. male, patella I; 71 M. quadrata sp. nov. female, femur IV. Abbreviation: LS—lanceolate setae." figureDoi="http://doi.org/10.5281/zenodo.4580946" httpUri="https://zenodo.org/record/4580946/files/figure.png" pageId="57" pageNumber="58">70</figureCitation>
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,
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<figureCitation id="133E9E879C0BFFFF7D5CFC76FE16FC49" box="[374,472,906,933]" captionStart="FIGURES 151–154" captionStartId="58.[151,264,963,987]" captionTargetBox="[163,1430,243,931]" captionTargetId="figure-50@58.[151,1436,237,940]" captionTargetPageId="58" captionText="FIGURES 151–154. Genital morphology of Micaria plana sp. nov.: 151 female epigyne, ventral view; 152 same, dorsal view; 153 male palp, ventral view; 154 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580984" httpUri="https://zenodo.org/record/4580984/files/figure.png" pageId="57" pageNumber="58">
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151
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||
<emphasis id="B9715E109C0BFFFF7D8AFC77FE60FC48" box="[416,430,907,932]" italics="true" pageId="57" pageNumber="58">–</emphasis>
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||
154
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</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31FD1899C0BFFFF7CBDFC2FFBA0FBB5" pageId="57" pageNumber="58" type="materials_examined">
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<paragraph id="8BBA82029C0BFFFF7CBDFC2FFAF6FBFD" blockId="57.[151,1437,978,2013]" pageId="57" pageNumber="58">
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||
<emphasis id="B9715E109C0BFFFF7CBDFC2FFE8BFC00" bold="true" box="[151,325,978,1005]" pageId="57" pageNumber="58">Type material.</emphasis>
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||
<materialsCitation id="3B6D885F9C0BFFFF7D66FC2FFAF6FBFD" ID-GBIF-Occurrence="3071034301" collectingDate="1988-06-28" collectorName="A. Russell-Smith" country="Ethiopia" latitude="9.047133" location="Shewa Region" longLatPrecision="1" longitude="38.40753" municipality="Addis Alem" pageId="57" pageNumber="58" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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||
<emphasis id="B9715E109C0BFFFF7D66FC2FFE77FC01" bold="true" box="[332,441,979,1005]" pageId="57" pageNumber="58">
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||
<typeStatus id="54BE3CA09C0BFFFF7D66FC2FFE77FC01" box="[332,441,979,1005]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
<specimenCount id="9D03498B9C0BFFFF7DEAFC2FFE1BFC01" box="[448,469,979,1005]" pageId="57" pageNumber="58" type="female">♀</specimenCount>
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||
,
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||
<emphasis id="B9715E109C0BFFFF7DCEFC2EFCE1FC01" bold="true" box="[484,815,978,1005]" pageId="57" pageNumber="58">
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||
<collectingCountry id="F312C2929C0BFFFF7DCEFC2EFDBEFC00" box="[484,624,978,1004]" name="Ethiopia" pageId="57" pageNumber="58">ETHIOPIA</collectingCountry>
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||
:
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||
<emphasis id="B9715E109C0BFFFF7EAAFC2EFCE8FC01" bold="true" box="[640,806,978,1005]" italics="true" pageId="57" pageNumber="58">
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||
<location id="8EDAD4D99C0BFFFF7EAAFC2EFCE8FC01" LSID="urn:lsid:plazi:treatment:03AC33149C0BFFFC7CBDFC9BFBB0F868:8EDAD4D99C0BFFFF7EAAFC2EFCE8FC01" box="[640,806,978,1005]" country="Ethiopia" latitude="9.047133" longLatPrecision="1" longitude="38.40753" municipality="Addis Alem" name="Shewa Region" pageId="57" pageNumber="58">Shewa Region</location>
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</emphasis>
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||
:
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</emphasis>
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<collectingMunicipality id="6BDE18789C0BFFFF7F1FFC2FFC71FC01" box="[821,959,979,1005]" pageId="57" pageNumber="58">Addis Alem</collectingMunicipality>
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,
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<locationDeviation id="9A089B209C0BFFFF7FE0FC2FFB2EFC01" box="[970,1248,979,1005]" location="Addis Alem" pageId="57" pageNumber="58">
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||
<quantity id="4CFD2FE79C0BFFFF7FE0FC2FFBDFFC01" box="[970,1041,979,1005]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="57" pageNumber="58" unit="km" value="10.0">10 km</quantity>
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Wof Addis Alem
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||
</locationDeviation>
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||
, under stones, in pasture, handcollecting,
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<geoCoordinate id="EE31E4C59C0BFFFF7D9AFC0BFD8AFBFD" box="[432,580,1015,1041]" degrees="09" direction="north" minutes="02.828" orientation="latitude" pageId="57" pageNumber="58" precision="1" value="9.047133">09°02.828’N</geoCoordinate>
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||
,
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<geoCoordinate id="EE31E4C59C0BFFFF7E7AFC0AFD2EFBFD" box="[592,736,1014,1041]" degrees="38" direction="east" minutes="24.452" orientation="longitude" pageId="57" pageNumber="58" precision="1" value="38.40753">38°24.452’E</geoCoordinate>
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, leg.
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<collectorName id="26F0E7D49C0BFFFF7F31FC0BFC11FBFD" box="[795,991,1015,1041]" pageId="57" pageNumber="58">A. Russell-Smith</collectorName>
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||
,
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||
<date id="FFBBA4C29C0BFFFF7FC0FC0BFBA4FBFD" box="[1002,1130,1015,1041]" pageId="57" pageNumber="58" value="1988-06-28">
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||
<collectingDate id="EFFF5D2A9C0BFFFF7FC0FC0BFBA4FBFD" box="[1002,1130,1015,1041]" pageId="57" pageNumber="58" value="1988-06-28">28.VI.1988</collectingDate>
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||
</date>
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(MRAC 225606).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BBA82029C0BFFFF7CEDFBE7FBA0FBB5" blockId="57.[151,1437,978,2013]" pageId="57" pageNumber="58">
|
||
<materialsCitation id="3B6D885F9C0BFFFF7CEDFBE7FBA4FBB5" ID-GBIF-Occurrence="3071034312" collectingDate="1983-04-27" collectorName="A. Russell-Smith" country="Ethiopia" latitude="9.683333" location="Shewa Region" longLatPrecision="1299" longitude="39.7" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Shewa Region" typeStatus="paratype">
|
||
<emphasis id="B9715E109C0BFFFF7CEDFBE7FD53FBD9" bold="true" box="[199,669,1050,1077]" pageId="57" pageNumber="58">
|
||
<typeStatus id="54BE3CA09C0BFFFF7CEDFBE7FE8DFBD9" box="[199,323,1051,1077]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<collectingCountry id="F312C2929C0BFFFF7D7AFBE6FE12FBD8" box="[336,476,1050,1076]" name="Ethiopia" pageId="57" pageNumber="58">ETHIOPIA</collectingCountry>
|
||
:
|
||
<emphasis id="B9715E109C0BFFFF7DC7FBE6FD5DFBD9" bold="true" box="[493,659,1050,1077]" italics="true" pageId="57" pageNumber="58">
|
||
<collectingRegion id="49C14CE09C0BFFFF7DC7FBE6FD5DFBD9" box="[493,659,1050,1077]" pageId="57" pageNumber="58">Shewa Region</collectingRegion>
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||
</emphasis>
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||
:
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||
</emphasis>
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||
<collectorName id="26F0E7D49C0BFFFF7E8FFBE7FC62FBD9" box="[677,940,1051,1077]" pageId="57" pageNumber="58">Between Debre Berhan</collectorName>
|
||
&
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||
<collectorName id="26F0E7D49C0BFFFF7FE5FBE7FBFDFBD9" box="[975,1075,1051,1077]" pageId="57" pageNumber="58">Ankober</collectorName>
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||
, under stones near stream, hand collecting,
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||
<geoCoordinate id="EE31E4C59C0BFFFF7D3EFBC3FEB8FBB5" box="[276,374,1087,1113]" degrees="09" direction="north" minutes="41" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="9.683333">09°41’N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE31E4C59C0BFFFF7DA8FBC3FE2CFBB5" box="[386,482,1087,1113]" degrees="39" direction="east" minutes="42" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="39.7">39°42’E</geoCoordinate>
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||
, leg.
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||
<collectorName id="26F0E7D49C0BFFFF7E37FBC3FD2FFBB5" box="[541,737,1087,1113]" pageId="57" pageNumber="58">A. Russell-Smith</collectorName>
|
||
,
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||
<date id="FFBBA4C29C0BFFFF7EC6FBC3FCA4FBB5" box="[748,874,1087,1113]" pageId="57" pageNumber="58" value="1983-04-27">
|
||
<collectingDate id="EFFF5D2A9C0BFFFF7EC6FBC3FCA4FBB5" box="[748,874,1087,1113]" pageId="57" pageNumber="58" value="1983-04-27">27.IV.1983</collectingDate>
|
||
</date>
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||
, 4 ♂ (MRAC 225570)
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31FD1899C0BFFFF7CEDFB9FFC84FB4D" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="8BBA82029C0BFFFF7CEDFB9FFC84FB4D" blockId="57.[151,1437,978,2013]" pageId="57" pageNumber="58">
|
||
<emphasis id="B9715E109C0BFFFF7CEDFB9FFE80FB91" bold="true" box="[199,334,1123,1149]" pageId="57" pageNumber="58">Etymology.</emphasis>
|
||
The species name is taken from the Latin adjective
|
||
<emphasis id="B9715E109C0BFFFF7F8EFB9FFC21FB91" box="[932,1007,1123,1149]" italics="true" pageId="57" pageNumber="58">planus</emphasis>
|
||
(flat), female
|
||
<emphasis id="B9715E109C0BFFFF78B9FB9FFB64FB90" box="[1171,1194,1123,1148]" italics="true" pageId="57" pageNumber="58">-a</emphasis>
|
||
, referring to the flattened and laterally extending copulatory ducts of the females.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C31FD1899C0BFFFF7CEDFB56FDFAFADD" pageId="57" pageNumber="58" type="diagnosis">
|
||
<paragraph id="8BBA82029C0BFFFF7CEDFB56FDFAFADD" blockId="57.[151,1437,978,2013]" pageId="57" pageNumber="58">
|
||
<emphasis id="B9715E109C0BFFFF7CEDFB56FE8CFB28" bold="true" box="[199,322,1194,1220]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
|
||
Females of this species can be recognised by the atypical structure of their copulatory ducts, in that they are laterally extending and tapered at the tip, opposed to the tubular structure of other
|
||
<taxonomicName id="4C05F9819C0BFFFF7884FB33FAC6FB04" authorityName="Westring" authorityYear="1851" box="[1198,1288,1231,1256]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9715E109C0BFFFF7884FB33FAC6FB04" box="[1198,1288,1231,1256]" italics="true" pageId="57" pageNumber="58">Micaria</emphasis>
|
||
</taxonomicName>
|
||
species. The males of this species typically have a short (free part approximately half the size of the MA) embolus that is thick. The MA is large and curves inwards.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31FD1899C0BFFFC7CEDFAC6FE01F88C" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
|
||
<paragraph id="8BBA82029C0BFFFF7CEDFAC6FE28F9E5" blockId="57.[151,1437,978,2013]" pageId="57" pageNumber="58">
|
||
<emphasis id="B9715E109C0BFFFF7CEDFAC6FAC4FAB9" bold="true" box="[199,1290,1338,1365]" pageId="57" pageNumber="58">
|
||
Description. Female (
|
||
<typeStatus id="54BE3CA09C0BFFFF7DECFAC6FDE0FAB8" box="[454,558,1338,1364]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
from Addis Alem,
|
||
<collectingCountry id="F312C2929C0BFFFF7F3AFAC6FCB6FAB8" box="[784,888,1338,1364]" name="Ethiopia" pageId="57" pageNumber="58">Ethiopia</collectingCountry>
|
||
, MRAC 225606): Measurements:
|
||
</emphasis>
|
||
TL 2.45, CL 1.03, CW 0.68, CL:CW 1.52, CLH 0.08, CLH:AME 1.65. Eyes: ALE 0.06, AME 0.05, PLE 0.05, PME 0.04, AME–AME 0.02, PME–PME 0.04, MOQAW 0.11, MOQPW 0.13, MOQL 0.16. SL 0.64, SW 0.49, AL 1.25, AW 0.80.
|
||
<emphasis id="B9715E109C0BFFFF7CFFFA5BFE73FA2D" bold="true" box="[213,445,1447,1473]" pageId="57" pageNumber="58">Leg measurements:</emphasis>
|
||
Palpal segment lengths: 0.29, 0.18, 0.16, 0.24. Leg I: 0.64, 0.30, 0.46, 0.37, 0.46. II: 0.59, 0.30, 0.41, 0.35, 0.43. III: 0.48, 0.27, 0.33, 0.38, 0.34. IV: 0.73, 0.33, 0.56, 0.59, 0.42. TLoflegs (I–IV): 2.23, 2.08, 1.80, 2.63; leg formula: 4123.
|
||
</paragraph>
|
||
<paragraph id="8BBA82029C0BFFFC7CEDF9EEFD80FF39" blockId="57.[151,1437,978,2013]" lastBlockId="58.[151,1436,151,213]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||
<emphasis id="B9715E109C0BFFFF7CEDF9EEFEAFF9C0" bold="true" box="[199,353,1554,1580]" pageId="57" pageNumber="58">Colouration:</emphasis>
|
||
carapace dark brown; abdomen dark brown or black; femora darkened; sternum, endites, labium, and chelicerae similar in colour to carapace (
|
||
<figureCitation id="133E9E879C0BFFFF7E8CF9CBFD30F9BD" box="[678,766,1591,1617]" captionStart="FIGURES 27–36" captionStartId="13.[151,264,1855,1879]" captionTargetBox="[151,1435,181,1827]" captionTargetId="figure-17@13.[151,1435,181,1830]" captionTargetPageId="13" captionText="FIGURES 27–36. Stereomicroscope photographs of the dorsal habitus of Afrotropical Micaria species: 27 M. parvotibialis sp. nov. male; 28 M. plana sp. nov. female and 29 male; 30 M. quadrata sp. nov. female; 31 M. quinquemaculosa sp. nov. female and 32 male; 33 M. rivonosy sp. nov. female; 34 M. sanipass sp. nov. male; 35 M. scutellata sp. nov. male; 36 M. tersissima Simon, 1910 male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580933" httpUri="https://zenodo.org/record/4580933/files/figure.png" pageId="57" pageNumber="58">Fig. 28</figureCitation>
|
||
).
|
||
<emphasis id="B9715E109C0BFFFF7F3FF9CAFC5CF9BC" bold="true" box="[789,914,1590,1616]" pageId="57" pageNumber="58">Carapace:</emphasis>
|
||
smooth in texture; decorated with squamose setae; dark pattern present medially, radiating; posterior margin indented medially; fovea absent; anterior margin narrowing slightly from coxa I; PER procurved in dorsal view, AER slightly recurved in anterior view.
|
||
<emphasis id="B9715E109C0BFFFF7939F982FA52F974" bold="true" box="[1299,1436,1662,1688]" pageId="57" pageNumber="58">Chelicerae:</emphasis>
|
||
decorated with plumose setae.
|
||
<emphasis id="B9715E109C0BFFFF7DC6F95EFD9BF950" bold="true" box="[492,597,1698,1724]" pageId="57" pageNumber="58">Labium:</emphasis>
|
||
triangular in shape, truncating where chelicerae meet.
|
||
<emphasis id="B9715E109C0BFFFF789EF95EFAD6F950" bold="true" box="[1204,1304,1698,1724]" pageId="57" pageNumber="58">Endites:</emphasis>
|
||
slanting inwards.
|
||
<emphasis id="B9715E109C0BFFFF7CCDF93AFE97F90C" bold="true" box="[231,345,1734,1760]" pageId="57" pageNumber="58">Sternum:</emphasis>
|
||
shield-like; broadest between coxa Iand II; posterior margin truncating behind coxa IV; decorated with aculeate setae.
|
||
<emphasis id="B9715E109C0BFFFF7D50F917FE75F8E9" bold="true" box="[378,443,1771,1797]" pageId="57" pageNumber="58">Legs:</emphasis>
|
||
femora laterally flattened, highest at base, leg Imore pronounced in this fashion; rest of legs normal, thin.
|
||
<collectingRegion id="49C14CE09C0BFFFF7D4DF8F3FE5DF8C5" box="[359,403,1807,1833]" country="South Sudan" name="Lakes" pageId="57" pageNumber="58">Leg</collectingRegion>
|
||
macrosetae: Palps: 1pl, 1do, 1pl, 1pl, 2plv, 1rlv, 3vt.
|
||
<collectingRegion id="49C14CE09C0BFFFF7FDDF8F3FBEDF8C5" box="[1015,1059,1807,1833]" country="South Sudan" name="Lakes" pageId="57" pageNumber="58">Leg</collectingRegion>
|
||
I: 1do, 1pl. II: 1d. III: 1do, 1plv, 2plv, 2rlv, 4vt. IV: 1do, 1plv, 2vt, 2plv, 3vt. Legs I–IV with two rows of scopulate setae on metatarsi and four rows on tarsi.
|
||
<emphasis id="B9715E109C0BFFFF7CD7F8AAFEB5F89C" bold="true" box="[253,379,1878,1904]" pageId="57" pageNumber="58">Abdomen:</emphasis>
|
||
cylindrical in shape; decorated with squamose setae; two white spots dorsally near anterior margin, white median transverse line broken into three smaller transverse stripes (
|
||
<figureCitation id="133E9E879C0BFFFF7807F887FB4CF879" box="[1069,1154,1915,1941]" captionStart="FIGURES 27–36" captionStartId="13.[151,264,1855,1879]" captionTargetBox="[151,1435,181,1827]" captionTargetId="figure-17@13.[151,1435,181,1830]" captionTargetPageId="13" captionText="FIGURES 27–36. Stereomicroscope photographs of the dorsal habitus of Afrotropical Micaria species: 27 M. parvotibialis sp. nov. male; 28 M. plana sp. nov. female and 29 male; 30 M. quadrata sp. nov. female; 31 M. quinquemaculosa sp. nov. female and 32 male; 33 M. rivonosy sp. nov. female; 34 M. sanipass sp. nov. male; 35 M. scutellata sp. nov. male; 36 M. tersissima Simon, 1910 male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580933" httpUri="https://zenodo.org/record/4580933/files/figure.png" pageId="57" pageNumber="58">Fig. 28</figureCitation>
|
||
); venter similar in colour to dorsum; sigilla not visible.
|
||
<emphasis id="B9715E109C0BFFFF7DC2F862FD9AF854" bold="true" box="[488,596,1950,1976]" pageId="57" pageNumber="58">Epigyne:</emphasis>
|
||
in ventral view, anterior hood divided, recurved; posterior pockets present medially, straight; copulatory openings at tipof copulatory ducts. In dorsal view, copulatory duct projecting laterally outwards from anterior margin of spermathecae, curved, tapered; fertilisation duct curved laterally outwards; spermathecae globular (
|
||
<figureCitation id="133E9E879C08FFFC7D89FF47FD8FFF39" box="[419,577,186,213]" captionStart="FIGURES 151–154" captionStartId="58.[151,264,963,987]" captionTargetBox="[163,1430,243,931]" captionTargetId="figure-50@58.[151,1436,237,940]" captionTargetPageId="58" captionText="FIGURES 151–154. Genital morphology of Micaria plana sp. nov.: 151 female epigyne, ventral view; 152 same, dorsal view; 153 male palp, ventral view; 154 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580984" httpUri="https://zenodo.org/record/4580984/files/figure.png" pageId="58" pageNumber="59">Figs 151, 152</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7AD28A9C08FFFC7CBDFC3FFC69FC13" ID-DOI="http://doi.org/10.5281/zenodo.4580984" ID-Zenodo-Dep="4580984" httpUri="https://zenodo.org/record/4580984/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,264,963,987]" targetBox="[163,1430,243,931]" targetPageId="58">
|
||
<paragraph id="8BBA82029C08FFFC7CBDFC3FFC69FC13" blockId="58.[151,1436,963,1023]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9715E109C08FFFC7CBDFC3FFEA3FC37" bold="true" box="[151,365,963,987]" pageId="58" pageNumber="59">FIGURES 151–154.</emphasis>
|
||
Genital morphology of
|
||
<taxonomicName id="4C05F9819C08FFFC7E4AFC38FD3CFC37" authority="Booysen & Haddad, 2021" authorityName="Booysen & Haddad" authorityYear="2021" box="[608,754,963,987]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="plana" status="sp. nov.">
|
||
<emphasis id="B9715E109C08FFFC7E4AFC38FD3CFC37" box="[608,754,963,987]" italics="true" pageId="58" pageNumber="59">Micaria plana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9715E109C08FFFC7ED2FC3FFC86FC37" bold="true" box="[760,840,963,987]" pageId="58" pageNumber="59">
|
||
<taxonomicNameLabel id="A242E36B9C08FFFC7ED2FC3FFC86FC37" box="[760,840,963,987]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 151 female epigyne, ventral view; 152 same, dorsal view; 153 male palp, ventral view; 154 same, retrolateral view. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBA82029C08FFFC7CEDFBD1FDEBFB17" blockId="58.[151,1437,1069,1924]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9715E109C08FFFC7CEDFBD1FB10FBAB" bold="true" box="[199,1246,1069,1095]" pageId="58" pageNumber="59">
|
||
<collectingRegion id="49C14CE09C08FFFC7CEDFBD1FECAFBAB" box="[199,260,1069,1095]" pageId="58" pageNumber="59">Male</collectingRegion>
|
||
(
|
||
<typeStatus id="54BE3CA09C08FFFC7D3BFBD1FEB0FBAB" box="[273,382,1069,1095]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
collected between
|
||
<collectingRegion id="49C14CE09C08FFFC7E72FBD1FC50FBAB" box="[600,926,1069,1095]" pageId="58" pageNumber="59">Debre Berhan and Ankober</collectingRegion>
|
||
,
|
||
<collectingCountry id="F312C2929C08FFFC7F82FBD1FBDEFBAB" box="[936,1040,1069,1095]" name="Ethiopia" pageId="58" pageNumber="59">Ethiopia</collectingCountry>
|
||
, MRAC 225570):
|
||
</emphasis>
|
||
<materialsCitation id="3B6D885F9C08FFFC78CEFBD1FF30FB5F" ID-GBIF-Occurrence="3071034350" collectionCode="MOQAW" country="Ethiopia" location="Eyes" pageId="58" pageNumber="59" specimenCount="1" stateProvince="Measurements">
|
||
<emphasis id="B9715E109C08FFFC78CEFBD1FA52FBAB" bold="true" box="[1252,1436,1069,1095]" pageId="58" pageNumber="59">
|
||
<collectingRegion id="49C14CE09C08FFFC78CEFBD1FA56FBAB" box="[1252,1432,1069,1095]" pageId="58" pageNumber="59">Measurements</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
TL 2.65, CL 1.10, CW 0.80, CL:CW 1.38, CLH 0.08, CLH:AME 1.41.
|
||
<location id="8EDAD4D99C08FFFC7F85FBAEFC24FB80" LSID="urn:lsid:plazi:treatment:03AC33149C0BFFFC7CBDFC9BFBB0F868:8EDAD4D99C08FFFC7F85FBAEFC24FB80" box="[943,1002,1106,1132]" country="Ethiopia" name="Eyes" pageId="58" pageNumber="59" stateProvince="Measurements">Eyes</location>
|
||
: ALE 0.06, AME 0.06, PLE 0.06, PME 0.05, AME–AME 0.02, PME–PME 0.04,
|
||
<collectionCode id="ED141AC79C08FFFC7E46FB89FD17FB63" box="[620,729,1141,1167]" pageId="58" pageNumber="59">MOQAW</collectionCode>
|
||
0.13, MOQPW 0.14, MOQL 0.16. SL 0.67, SW 0.49, AL 1.40, AW 0.26
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6D885F9C08FFFC7D22FB65FDEFFB17" ID-GBIF-Occurrence="3071034396" collectorName="Leg I" country="Ethiopia" location="Leg" pageId="58" pageNumber="59" specimenCount="1" stateProvince="Leg">
|
||
<emphasis id="B9715E109C08FFFC7D22FB65FE3FFB5F" bold="true" box="[264,497,1177,1203]" pageId="58" pageNumber="59">
|
||
<collectingRegion id="49C14CE09C08FFFC7D22FB65FEFBFB5F" box="[264,309,1177,1203]" pageId="58" pageNumber="59">Leg</collectingRegion>
|
||
measurements:
|
||
</emphasis>
|
||
Palpal segment lengths: 0.36, 0.19, 0.12, 0.30.
|
||
<collectorName id="26F0E7D49C08FFFC782AFB66FBF0FB58" box="[1024,1086,1178,1204]" pageId="58" pageNumber="59">Leg I</collectorName>
|
||
: 0.81, 0.40, 0.68, 0.52, 0.51. II: 0.64, 0.33, 0.49, 0.41, 0.46. III: 0.52, 0.27, 0.37, 0.41, 0.37. IV: 0.75, 0.35, 0.57, 0.65, 0.46. TLoflegs (I–IV): 2.92, 2.33, 1.94, 2.78; leg formula: 1423
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBA82029C08FFFC7CEDFAF9FA53FA67" blockId="58.[151,1437,1069,1924]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9715E109C08FFFC7CEDFAF9FEAFFAF3" bold="true" box="[199,353,1285,1311]" pageId="58" pageNumber="59">Colouration:</emphasis>
|
||
carapace dark brown; abdomen black; femora darkened; sternum, endites, labium and chelicerae similar in colour to carapace (
|
||
<figureCitation id="133E9E879C08FFFC7DCDFAD5FDF2FAAF" box="[487,572,1321,1347]" captionStart="FIGURES 27–36" captionStartId="13.[151,264,1855,1879]" captionTargetBox="[151,1435,181,1827]" captionTargetId="figure-17@13.[151,1435,181,1830]" captionTargetPageId="13" captionText="FIGURES 27–36. Stereomicroscope photographs of the dorsal habitus of Afrotropical Micaria species: 27 M. parvotibialis sp. nov. male; 28 M. plana sp. nov. female and 29 male; 30 M. quadrata sp. nov. female; 31 M. quinquemaculosa sp. nov. female and 32 male; 33 M. rivonosy sp. nov. female; 34 M. sanipass sp. nov. male; 35 M. scutellata sp. nov. male; 36 M. tersissima Simon, 1910 male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580933" httpUri="https://zenodo.org/record/4580933/files/figure.png" pageId="58" pageNumber="59">Fig. 29</figureCitation>
|
||
).
|
||
<emphasis id="B9715E109C08FFFC7E65FAD5FD02FAAF" bold="true" box="[591,716,1321,1347]" pageId="58" pageNumber="59">Carapace:</emphasis>
|
||
smooth in texture; decorated with squamose setae; broad somewhat rectangular dark marking medially, radiating dark lines project from it; posterior margin straight; fovea absent; anterior margin narrowing slightly from coxa I; PER procurved in dorsal view, AER recurved in anterior view.
|
||
</paragraph>
|
||
<paragraph id="8BBA82029C08FFFC7CBDFA69FE01F88C" blockId="58.[151,1437,1069,1924]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9715E109C08FFFC7CBDFA69FEEEFA43" bold="true" box="[151,288,1429,1455]" pageId="58" pageNumber="59">Chelicerae:</emphasis>
|
||
decorated with plumose setae.
|
||
<emphasis id="B9715E109C08FFFC7EADFA69FD21FA43" bold="true" box="[647,751,1429,1455]" pageId="58" pageNumber="59">Labium:</emphasis>
|
||
triangular in shape; cluster of setae present on tip.
|
||
<emphasis id="B9715E109C08FFFC7913FA69FA52FA43" bold="true" box="[1337,1436,1429,1455]" pageId="58" pageNumber="59">Endites:</emphasis>
|
||
oblique; constricted just above halfway; posterior margin rounded.
|
||
<emphasis id="B9715E109C08FFFC7FA7FA45FC30FA3F" bold="true" box="[909,1022,1465,1491]" pageId="58" pageNumber="59">Sternum:</emphasis>
|
||
shield-like; broadest between coxa I and II; posterior margin tapering behind coxa IV.
|
||
<emphasis id="B9715E109C08FFFC7EE2FA21FCC7FA1B" bold="true" box="[712,777,1501,1527]" pageId="58" pageNumber="59">Legs:</emphasis>
|
||
femora laterally flattened, highest at base, leg Imore pronounced in this fashion; rest of legs normal, thin; decorated with lanceolate setae (
|
||
<figureCitation id="133E9E879C08FFFC781BF9FDFB4BF9F7" box="[1073,1157,1537,1563]" captionStart="FIGURES 65–71" captionStartId="20.[151,264,1580,1604]" captionTargetBox="[151,1436,181,1563]" captionTargetId="figure-31@20.[151,1436,181,1563]" captionTargetPageId="20" captionText="FIGURES 65–71. Scanning electron microscope images of the lanceolate setae of Micaria: 65 M. basaliducta sp. nov. male, patella I; 66 M. beaufortia (Tucker, 1923) male, palpal tibia; 67 M. bispicula sp. nov. male, patella I; 68 M. koingnaas sp. nov. male, femur I; 69 M. parvotibialis sp. nov. male, tarsus IV; 70 M. plana sp. nov. male, patella I; 71 M. quadrata sp. nov. female, femur IV. Abbreviation: LS—lanceolate setae." figureDoi="http://doi.org/10.5281/zenodo.4580946" httpUri="https://zenodo.org/record/4580946/files/figure.png" pageId="58" pageNumber="59">Fig. 70</figureCitation>
|
||
).
|
||
<collectingRegion id="49C14CE09C08FFFC78B3F9FEFB0BF9F0" box="[1177,1221,1538,1564]" country="South Sudan" name="Lakes" pageId="58" pageNumber="59">Leg</collectingRegion>
|
||
macrosetae: Palps: 1do, 1pl, 1do, 1pl, 1pl, 1plv.
|
||
<collectingRegion id="49C14CE09C08FFFC7DF0F9DAFDC8F9AC" box="[474,518,1574,1600]" country="South Sudan" name="Lakes" pageId="58" pageNumber="59">Leg</collectingRegion>
|
||
I: 1do, 1pl. II: 1do, 1plv, 2vt. III: 1do, 1pl, 1plv, 2vt, 2plv, 2rlv, 4vt. IV: 1do, 1plv, 2vt, 2plv, 1rlv, 3vt. Legs I–IV with two rows of scopulate setae on metatarsi and four rows on tarsi.
|
||
<emphasis id="B9715E109C08FFFC78DEF9B5FABCF98F" bold="true" box="[1268,1394,1609,1635]" pageId="58" pageNumber="59">Abdomen:</emphasis>
|
||
cylindrical in shape; decorated with squamose setae; two white spots dorsally near anterior margin, broken transverse line medially (
|
||
<figureCitation id="133E9E879C08FFFC7D14F96DFE5BF947" box="[318,405,1681,1707]" captionStart="FIGURES 27–36" captionStartId="13.[151,264,1855,1879]" captionTargetBox="[151,1435,181,1827]" captionTargetId="figure-17@13.[151,1435,181,1830]" captionTargetPageId="13" captionText="FIGURES 27–36. Stereomicroscope photographs of the dorsal habitus of Afrotropical Micaria species: 27 M. parvotibialis sp. nov. male; 28 M. plana sp. nov. female and 29 male; 30 M. quadrata sp. nov. female; 31 M. quinquemaculosa sp. nov. female and 32 male; 33 M. rivonosy sp. nov. female; 34 M. sanipass sp. nov. male; 35 M. scutellata sp. nov. male; 36 M. tersissima Simon, 1910 male. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580933" httpUri="https://zenodo.org/record/4580933/files/figure.png" pageId="58" pageNumber="59">Fig. 29</figureCitation>
|
||
); venter similar in colour to dorsum; sigilla not visible.
|
||
<emphasis id="B9715E109C08FFFC7836F96DFB92F947" bold="true" box="[1052,1116,1681,1707]" pageId="58" pageNumber="59">Palp:</emphasis>
|
||
retrolateral tibial apophysis present apically on dorsal surface; median apophysis present, large, next to embolus, broad basally; sperm duct with slight bend posterior to embolus base, curved medially at an angle and continuing retrolaterally, straight, towards base of median apophysis; embolus base elongate, tip short, straight; cymbium elongate, pear-shaped and widest near basal third; two short spines present on tipof cymbium; three elongate ventral terminal spines present in front of embolus (
|
||
<figureCitation id="133E9E879C08FFFC7D09F8B9FE0CF88C" box="[291,450,1861,1888]" captionStart="FIGURES 151–154" captionStartId="58.[151,264,963,987]" captionTargetBox="[163,1430,243,931]" captionTargetId="figure-50@58.[151,1436,237,940]" captionTargetPageId="58" captionText="FIGURES 151–154. Genital morphology of Micaria plana sp. nov.: 151 female epigyne, ventral view; 152 same, dorsal view; 153 male palp, ventral view; 154 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580984" httpUri="https://zenodo.org/record/4580984/files/figure.png" pageId="58" pageNumber="59">Figs 153, 154</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31FD1899C08FFFC7CEDF895FBB0F868" box="[199,1150,1897,1924]" pageId="58" pageNumber="59" type="distribution">
|
||
<paragraph id="8BBA82029C08FFFC7CEDF895FBB0F868" blockId="58.[151,1437,1069,1924]" box="[199,1150,1897,1924]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9715E109C08FFFC7CEDF895FEAFF86F" bold="true" box="[199,353,1897,1923]" pageId="58" pageNumber="59">Distribution.</emphasis>
|
||
<taxonomicName id="4C05F9819C08FFFC7D4DF896FDC7F86F" authority="Booysen & Haddad, 2021" authorityName="Booysen & Haddad" authorityYear="2021" box="[359,521,1897,1923]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="plana" status="sp. nov.">
|
||
<emphasis id="B9715E109C08FFFC7D4DF896FDC7F86F" box="[359,521,1897,1923]" italics="true" pageId="58" pageNumber="59">Micaria plana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9715E109C08FFFC7E25F895FDA7F86F" bold="true" box="[527,617,1897,1923]" pageId="58" pageNumber="59">
|
||
<taxonomicNameLabel id="A242E36B9C08FFFC7E25F895FDA7F86F" box="[527,617,1897,1923]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is limited to two localities in
|
||
<collectingCountry id="F312C2929C08FFFC7F9DF895FBD8F86F" box="[951,1046,1897,1923]" name="Ethiopia" pageId="58" pageNumber="59">Ethiopia</collectingCountry>
|
||
(
|
||
<figureCitation id="133E9E879C08FFFC780EF895FBBEF868" box="[1060,1136,1897,1924]" captionStart="MAP 5" captionStartId="53.[151,209,1300,1324]" captionTargetBox="[151,1435,181,1276]" captionTargetId="figure-17@53.[151,1435,181,1276]" captionTargetPageId="53" captionText="MAP 5. The distribution of 13 Micaria species in the Afrotropical Region ." figureDoi="http://doi.org/10.5281/zenodo.4739030" httpUri="https://zenodo.org/record/4739030/files/figure.png" pageId="58" pageNumber="59">Map 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |