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<document id="9558BE8570912A5F5F4BC7C0864FB04D" ID-DOI="10.5281/zenodo.10667719" ID-ISSN="1407-8619" ID-Zenodo-Dep="10667719" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708024678478" checkinUser="felipe" docAuthor="Alekseev, Vitalii I. &amp; Bukejs, Andris" docDate="2021" docId="03A7878EFFDD0915FEBBFEADF9EEEAAF" docLanguage="en" docName="BalticJColeopterol.21.1.11-18.pdf" docOrigin="Baltic Journal of Coleopterology 21 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.1:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="1" docTitle="Petalium bruteno Alekseev &amp; Bukejs 2021, sp. nov." docType="treatment" docUuid="9423340F-8274-4F3C-A408-485E07EE5539" docUuidSource="ZooBank" docVersion="2" lastPageNumber="15" masterDocId="FF9EFFF6FFDF0911FF94FF92FD35E877" masterDocTitle="Two fossil species of Petaliini White (Coleoptera: Ptinidae: Dorcatominae) from Eocene Baltic amber" masterLastPageNumber="18" masterPageNumber="11" pageNumber="13" updateTime="1708024987982" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="77988DEB85C199D0F72FB260173232BB">Two fossil species of Petaliini White (Coleoptera: Ptinidae: Dorcatominae) from Eocene Baltic amber</mods:title>
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<mods:namePart id="CACC6C9E04B2E09F7B8610AF40232BFD">Alekseev, Vitalii I.</mods:namePart>
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<mods:namePart id="3B5D0431EC0274B084C8675B14842740">Bukejs, Andris</mods:namePart>
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<mods:title id="ABD2390A24D3C6091005BD8253B509D8">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="75C264746762B20C22CBC847BF03EACC">2021</mods:date>
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<treatment id="03A7878EFFDD0915FEBBFEADF9EEEAAF" LSID="urn:lsid:zoobank.org:act:9423340F-8274-4F3C-A408-485E07EE5539" httpUri="http://treatment.plazi.org/id/03A7878EFFDD0915FEBBFEADF9EEEAAF" lastPageId="4" lastPageNumber="15" pageId="2" pageNumber="13">
<subSubSection id="C3146513FFDD0913FEBBFEADFF70E920" box="[303,581,319,343]" pageId="2" pageNumber="13" type="nomenclature">
<paragraph id="8BB13698FFDD0913FEBBFEADFF70E920" blockId="2.[303,581,319,375]" box="[303,581,319,343]" pageId="2" pageNumber="13">
<heading id="D0F981F4FFDD0913FEBBFEADFF70E920" bold="true" box="[303,581,319,343]" fontSize="10" level="2" pageId="2" pageNumber="13" reason="4">
<emphasis id="B97AEA8AFFDD0913FEBBFEADFF70E920" bold="true" box="[303,581,319,343]" pageId="2" pageNumber="13">
<taxonomicName id="4C0E4D1BFFDD0913FEBBFEADFCD8E920" authority="Alekseev &amp; Bukejs, 2021" authorityName="Alekseev &amp; Bukejs" authorityYear="2021" box="[303,493,319,343]" class="Insecta" family="Anobiidae" genus="Petalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="bruteno" status="sp. nov.">
<emphasis id="B97AEA8AFFDD0913FEBBFEADFCD8E920" bold="true" box="[303,493,319,343]" italics="true" pageId="2" pageNumber="13">Petalium bruteno</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957F1FFDD0913FE67FEADFF70E920" box="[499,581,319,343]" pageId="2" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146513FFDD0913FEBBFECDFC95E900" box="[303,416,351,375]" pageId="2" pageNumber="13" type="description">
<paragraph id="8BB13698FFDD0913FEBBFECDFC95E900" blockId="2.[303,581,319,375]" box="[303,416,351,375]" pageId="2" pageNumber="13">
(
<figureCitation id="13352A1DFFDD0913FEA3FECDFCACE900" box="[311,409,351,375]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[303,342,1756,1780]" captionStartId-1="3.[227,266,886,910]" captionText-0="Fig. 1. Petalium bruteno sp. nov., holotype, 6781 [MAIG], habitus:A dorsal view; B ventral view" captionText-1="Fig. 2. Petalium bruteno sp. nov., paratype, 6782 [MAIG], habitus: A dorsal view; B ventral view" pageId="2" pageNumber="13">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146513FFDD0913FEBBFE0DF828EAC0" pageId="2" pageNumber="13" type="materials_examined">
<paragraph id="8BB13698FFDD0913FEBBFE0DFF30EAE0" blockId="2.[303,813,415,663]" pageId="2" pageNumber="13">
<emphasis id="B97AEA8AFFDD0913FEBBFE0DFCE1E9C0" bold="true" box="[303,468,415,439]" pageId="2" pageNumber="13">Type material</emphasis>
.
<materialsCitation id="3B663CC5FFDD0913FE72FE0DFF30EAE0" collectionCode="MAIG" collectorName="Jonas Damzen &amp; Syninclusions" pageId="2" pageNumber="13" specimenCount="1" typeStatus="holotype">
<emphasis id="B97AEA8AFFDD0913FE72FE0DFF6DE9C0" bold="true" box="[486,600,415,439]" pageId="2" pageNumber="13">
<typeStatus id="54B5883AFFDD0913FE72FE0DFF66E9C0" box="[486,595,415,439]" pageId="2" pageNumber="13" type="holotype">Holotype</typeStatus>
:
</emphasis>
collection number “6781” [
<collectionCode id="ED1FAE5DFFDD0913FE14FE2DFCF2E9A0" box="[384,455,447,471]" pageId="2" pageNumber="13">MAIG</collectionCode>
] (ex. coll.
<collectorName id="26FB534EFFDD0913FDBDFE52FF82E9A0" box="[553,695,448,471]" pageId="2" pageNumber="13">Jonas Damzen</collectorName>
JDC 7645); adult, sex unknown. A complete beetle included in a transparent, yellow amber piece with approximate dimensions of
<quantity id="4CF69B7DFFDD0913FE6BFD8DFF7CEA40" box="[511,585,543,567]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="13" unit="mm" value="24.0">24 mm</quantity>
×
<quantity id="4CF69B7DFFDD0913FDF7FD8DFF99EA40" box="[611,684,543,567]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="13" unit="mm" value="18.0">18 mm</quantity>
and a maximum thickness of
<quantity id="4CF69B7DFFDD0913FE64FDADFF04EA20" box="[496,561,575,599]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="13" unit="mm" value="5.0">5 mm</quantity>
; preserved without supplementary fixation.
<collectorName id="26FB534EFFDD0913FD9BFDCDFF9EEA00" box="[527,683,607,631]" pageId="2" pageNumber="13">Syninclusions</collectorName>
: few stellate fagacean trichomes.
</materialsCitation>
</paragraph>
<paragraph id="8BB13698FFDD0913FEBBFD2DFFE0EBA0" blockId="2.[303,814,703,984]" pageId="2" pageNumber="13">
<materialsCitation id="3B663CC5FFDD0913FEBBFD2DFFE4EBA0" collectionCode="MAIG" collectorName="Jonas Damzen &amp; Syninclusions &amp; Brachycera &amp; Diptera" pageId="2" pageNumber="13" specimenCount="1" typeStatus="paratype">
<emphasis id="B97AEA8AFFDD0913FEBBFD2DFCA8EAA0" bold="true" box="[303,413,703,727]" pageId="2" pageNumber="13">
<typeStatus id="54B5883AFFDD0913FEBBFD2DFCADEAA0" box="[303,408,703,727]" pageId="2" pageNumber="13" type="paratype">Paratype</typeStatus>
:
</emphasis>
collection number “6782” [
<collectionCode id="ED1FAE5DFFDD0913FD26FD2DFFCEEAA0" box="[690,763,703,727]" pageId="2" pageNumber="13">MAIG</collectionCode>
] (ex. coll.
<collectorName id="26FB534EFFDD0913FEF2FD72FF33EA80" box="[358,518,736,759]" pageId="2" pageNumber="13">Jonas Damzen</collectorName>
JDC 9805); adult, sex unknown. A complete beetle with partially exposed metathoracic wings included in a transparent, yellow amber piece with approximate dimensions of
<quantity id="4CF69B7DFFDD0913FEBBFCCDFC4CEB00" box="[303,377,863,887]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="13" unit="mm" value="51.0">51 mm</quantity>
×
<quantity id="4CF69B7DFFDD0913FE00FCCDFCD5EB00" box="[404,480,863,887]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="13" unit="mm" value="22.0">22 mm</quantity>
and a maximum thickness of
<quantity id="4CF69B7DFFDD0913FC8BFCCDFC68EBE0" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="13" unit="mm" value="9.0">9 mm</quantity>
; preserved without supplementary fixation.
<collectorName id="26FB534EFFDD0913FEBBFC0DFCD5EBC0" box="[303,480,927,951]" pageId="2" pageNumber="13">Syninclusions</collectorName>
: two
<collectorName id="26FB534EFFDD0913FDB3FC0DFF85EBC0" box="[551,688,927,951]" pageId="2" pageNumber="13">Brachycera</collectorName>
(
<collectorName id="26FB534EFFDD0913FD51FC0DFE11EBC0" box="[709,804,927,951]" pageId="2" pageNumber="13">Diptera</collectorName>
) apecomens, and few small gas vesicles
</materialsCitation>
.
</paragraph>
<paragraph id="8BB13698FFDD0913FCC7FF4DF940E9C0" blockId="2.[851,1361,223,439]" pageId="2" pageNumber="13">
<materialsCitation id="3B663CC5FFDD0913FCC7FF4DF945E9C0" collectionCode="JHAC" pageId="2" pageNumber="13" specimenCount="1" typeStatus="paratype">
<emphasis id="B97AEA8AFFDD0913FCC7FF4DFEF1E880" bold="true" box="[851,964,223,247]" pageId="2" pageNumber="13">
<typeStatus id="54B5883AFFDD0913FCC7FF4DFE8AE880" box="[851,959,223,247]" pageId="2" pageNumber="13" type="paratype">Paratype</typeStatus>
:
</emphasis>
collection number “PET01” [
<collectionCode id="ED1FAE5DFFDD0913FB68FF4DF876E880" box="[1276,1347,223,247]" pageId="2" pageNumber="13">JHAC</collectionCode>
]; adult, sex unknown. A complete beetle included in a transparent, yellow amber piece with approximate dimensions of
<quantity id="4CF69B7DFFDD0913FBB7FEADF958E920" box="[1059,1133,319,343]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="13" unit="mm" value="40.0">40 mm</quantity>
×
<quantity id="4CF69B7DFFDD0913FB12FEADF9E5E920" box="[1158,1232,319,343]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="13" unit="mm" value="18.0">18 mm</quantity>
and a maximum thickness of
<quantity id="4CF69B7DFFDD0913FB80FECDF961E900" box="[1044,1108,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="13" unit="mm" value="9.0">9 mm</quantity>
; preserved without supplementary fixation. Syninclusions: organic small residues of soil and plants
</materialsCitation>
.
</paragraph>
<paragraph id="8BB13698FFDD0913FCC7FE4DF9C0EA40" blockId="2.[851,1361,479,567]" pageId="2" pageNumber="13">
<emphasis id="B97AEA8AFFDD0913FCC7FE4DFEC8E980" bold="true" box="[851,1021,479,503]" pageId="2" pageNumber="13">Type stratum.</emphasis>
Mid-late Eocene, 4834 Ma (
<bibRefCitation id="EF9F4B69FFDD0913FCCEFE6DF905EA60" author="Sadowski E. - M. &amp; Seyfullah L. J. &amp; Schmidt A. R. &amp; Kunzmann L." box="[858,1072,511,535]" pageId="2" pageNumber="13" pagination="1 - 73" refId="ref4149" refString="Sadowski E. - M., Seyfullah L. J., Schmidt A. R., Kunzmann L. 2017. Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia 106: 1 - 73." type="journal article" year="2017">Sadowski et al. 2017</bibRefCitation>
,
<bibRefCitation id="EF9F4B69FFDD0913FBADFE6DF944EA60" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Denk T." box="[1081,1137,511,535]" pageId="2" pageNumber="13" pagination="405 - 517" refId="ref4192" refString="Sadowski E. - M., Schmidt A. R., Denk T. 2020. Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family). Willdenowia 50: 405 - 517. DOI: 10.3372 / wi. 50.50303" type="journal article" year="2020">2020</bibRefCitation>
;
<bibRefCitation id="EF9F4B69FFDD0913FBEDFE6DF879EA60" author="Seyfullah L. J. &amp; Beimforde C. &amp; Dal Corso J. &amp; Perrichot V. &amp; Rikkinen J. &amp; Schmidt A. R." box="[1145,1356,511,535]" pageId="2" pageNumber="13" pagination="1684 - 1714" refId="ref4244" refString="Seyfullah L. J., Beimforde C., Dal Corso J., Perrichot V., Rikkinen J., Schmidt A. R. 2018. Production and preservation of resins - past and present. Biological Reviews 93: 1684 - 1714." type="journal article" year="2018">Seyfullah et al. 2018</bibRefCitation>
;
<bibRefCitation id="EF9F4B69FFDD0913FCC7FD8DF93BEA40" author="Bukejs A. &amp; Alekseev V. I. &amp; Pollock D. A." box="[851,1038,543,567]" pageId="2" pageNumber="13" pagination="261 - 273" refId="ref3699" refString="Bukejs A., Alekseev V. I., Pollock D. A. 2019. Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa 4664: 261 - 273." type="journal article" year="2019">Bukejs et al. 2019</bibRefCitation>
;
<bibRefCitation id="EF9F4B69FFDD0913FB83FD8DF9DCEA40" author="Kasinski J. R. &amp; Kramarska R. &amp; Slodkowska B. &amp; Sivkov V. &amp; Piwocki, M." box="[1047,1257,543,567]" pageId="2" pageNumber="13" pagination="29 - 53" refId="ref4030" refString="Kasinski J. R., Kramarska R., Slodkowska B., Sivkov V., Piwocki, M. 2020. Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 bore hole, Kaliningrad region, Russia). Geological Quarterly 64: 29 - 53." type="journal article" year="2020">Kasiñski et al. 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8BB13698FFDD0913FCC7FDCDF828EAC0" blockId="2.[851,1361,607,695]" pageId="2" pageNumber="13">
<emphasis id="B97AEA8AFFDD0913FCC7FDCDFED9EA00" bold="true" box="[851,1004,607,631]" pageId="2" pageNumber="13">Type locality.</emphasis>
Baltic Sea coast, Yantarny settlement (formerly Palmnicken),
<collectingCountry id="F3197608FFDD0913FB1FFDEDF9EEEAE0" box="[1163,1243,639,663]" name="Zambia" pageId="2" pageNumber="13">Sambia</collectingCountry>
(Samland) Peninsula, the
<collectingRegion id="49CAF87AFFDD0913FC7AFD0DF9F3EAC0" box="[1006,1222,671,695]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="13">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F3197608FFDD0913FB5BFD0DF82FEAC0" box="[1231,1306,671,695]" name="Russia" pageId="2" pageNumber="13">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146513FFDD0915FCC7FD4DFCB6EEA0" lastPageId="4" lastPageNumber="15" pageId="2" pageNumber="13" type="description">
<paragraph id="8BB13698FFDD0912FCC7FD4DFF28EB8D" blockId="2.[851,1362,735,920]" lastBlockId="3.[227,736,962,1018]" lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
<emphasis id="B97AEA8AFFDD0913FCC7FD4DFEECEA80" bold="true" box="[851,985,735,759]" pageId="2" pageNumber="13">Description.</emphasis>
<typeStatus id="54B5883AFFDD0913FC49FD4DF90FEA80" box="[989,1082,735,759]" pageId="2" pageNumber="13" type="holotype">Holotype</typeStatus>
JDC 7645. Measurements: total body length about
<quantity id="4CF69B7DFFDD0913FBAAFD6DF9A9EB60" box="[1086,1180,767,791]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="13" unit="mm" value="1.46">1.46 mm</quantity>
; pronotum length
<quantity id="4CF69B7DFFDD0913FCC7FC8DFE9EEB40" box="[851,939,799,823]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="13" unit="mm" value="0.38">0.38 mm</quantity>
, pronotum maximum width
<quantity id="4CF69B7DFFDD0913FB2EFC8DF826EB40" box="[1210,1299,799,823]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="13" unit="mm" value="0.54">0.54 mm</quantity>
; elytra length
<quantity id="4CF69B7DFFDD0913FC0BFCADF931EB20" box="[927,1028,831,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="2" pageNumber="13" unit="mm" value="1.08">1.08 mm</quantity>
, combined maximum width of elytra
<quantity id="4CF69B7DFFDD0913FC06FCCDFEC4EB00" box="[914,1009,863,887]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="2" pageNumber="13" unit="mm" value="0.69">0.69 mm</quantity>
. Body elongate, subparallel-sided, convex; glabrous, elytra with very fine, inconspicuous recumbent pubescence; unicolorous brown-rufous (as preserved).
</paragraph>
<caption id="DF716610FFDD0913FEBBF94EF87AEE83" box="[303,1359,1756,1780]" pageId="2" pageNumber="13" startId="2.[303,342,1756,1780]" targetType="figure">
<paragraph id="8BB13698FFDD0913FEBBF94EF87AEE83" blockId="2.[303,1359,1756,1780]" box="[303,1359,1756,1780]" pageId="2" pageNumber="13">
Fig. 1.
<taxonomicName id="4C0E4D1BFFDD0913FEEDF94EFF07EE83" authority="Alekseev &amp; Bukejs, 2021" authorityName="Alekseev &amp; Bukejs" authorityYear="2021" box="[377,562,1756,1780]" class="Insecta" family="Anobiidae" genus="Petalium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="bruteno" status="sp. nov.">
<emphasis id="B97AEA8AFFDD0913FEEDF94EFF07EE83" box="[377,562,1756,1780]" italics="true" pageId="2" pageNumber="13">Petalium bruteno</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957F1FFDD0913FDA3F94FFFBEEE83" box="[567,651,1757,1780]" pageId="2" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, 6781 [MAIG], habitus: A dorsal view; B ventral view
</paragraph>
</caption>
<caption id="DF716610FFDC0912FF77FCE4F833EBF9" box="[227,1286,886,911]" pageId="3" pageNumber="14" startId="3.[227,266,886,910]" targetType="figure">
<paragraph id="8BB13698FFDC0912FF77FCE4F833EBF9" blockId="3.[227,1286,886,911]" box="[227,1286,886,911]" pageId="3" pageNumber="14">
Fig. 2.
<taxonomicName id="4C0E4D1BFFDC0912FEB9FCE4FCD2EBF9" authority="Alekseev &amp; Bukejs, 2021" authorityName="Alekseev &amp; Bukejs" authorityYear="2021" box="[301,487,886,910]" class="Insecta" family="Anobiidae" genus="Petalium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="14" phylum="Arthropoda" rank="species" species="bruteno" status="sp. nov.">
<emphasis id="B97AEA8AFFDC0912FEB9FCE4FCD2EBF9" box="[301,487,886,910]" italics="true" pageId="3" pageNumber="14">Petalium bruteno</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957F1FFDC0912FE78FCE5FF75EBF9" box="[492,576,887,910]" pageId="3" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, 6782 [MAIG], habitus: A dorsal view; B ventral view
</paragraph>
</caption>
<paragraph id="8BB13698FFDC0912FF77FBB0FF2AED0D" blockId="3.[227,737,1058,1403]" pageId="3" pageNumber="14">Head hypognathous, turned downwards and not visible in dorsal view; densely covered with coarse punctation, distance between punctures lesser than diameter of one puncture; frons wide, weakly convex, without carina or longitudinal impression. Compound eyes rather large, convex, oval, with inner margin slightly emarginate at antennal insertion, with distinct facets, without ommatidial setae, widely separated. Fronto-clypeal suture fine, slightly impressed. Labrum very small. Mandibulae large.</paragraph>
<paragraph id="8BB13698FFDC0912FF77FA31F830EC6D" blockId="3.[227,737,1442,1754]" lastBlockId="3.[774,1285,962,1051]" pageId="3" pageNumber="14">Antennae apparently with 11 antennomeres (antennomeres 14 not visible), clavate; antennomeres 5 and 7 widened, swollen, globose, nearly as long as wide; antennomeres 6 and 8 small, narrow, elongate, about 1.8× as long as wide, antennomere 8 about 0.5× as wide as antennomere 7; antennomeres 910 forming distinct club, that apparently longer than antennomeres 18 combined; antennomeres 9 and 10 elongate, conical, dilated apically, antennomere 9 about 2.4× as wide as and 2.4× as long as antennomere 8; antennomere 11 elongate oval with pointed apex, 2.5× as long as wide.</paragraph>
<paragraph id="8BB13698FFDC0912FC92FBD1F94CEE4D" blockId="3.[774,1286,1090,1594]" pageId="3" pageNumber="14">Pronotum convex, transverse, 1.4× as wide as long, with convex disc, margined, widest at base and narrowed anteriad, strongly constricted in anterior one-third, with transverse impression at anterior angles; covered with large and dense punctation, punctures smaller on disc and in anterior portion of pronotum, distance between punctures lesser than diameter of one puncture. Anterior pronotal angles subrectangular, slightly rounded, not visible in dorsal view; posterior pronotal angles widely rounded and barely marked. Lateral margins slightly rounded; posterior margin bisinuated with wide median lobe; anterior margin upturned, convex in dorsal view, arcuate in frontal view. Prohypomera impressed, with dense and coarse punctation.</paragraph>
<paragraph id="8BB13698FFDC0912FC92F9F0F914EECD" blockId="3.[774,1285,1634,1722]" pageId="3" pageNumber="14">Scutellar shield small, transverse, 1.3× as wide as long, pentagonal with widely rounded apex, covered with small punctures.</paragraph>
<paragraph id="8BB13698FFDB0915FEBBFF2DFF82EA00" blockId="4.[303,814,191,631]" pageId="4" pageNumber="15">Elytra elongate, 1.6× as long as wide, subparallel-sided, widest in posterior one-third, slightly tapered posteriad, with transverse impression in anterior one-fourth; humeri distinctly prominent. Elytral punctures fine, round, regular, each elytron apparently with 10 rows; distance between punctures in rows equal to 12× diameter of puncture, distance between rows equal to 23× diameter of puncture; intervals wide, flat, covered with micropunctation. Metaventrite with convex disc; medial groove distinct in posterior one-fourth; densely covered with small punctation; anterior median lobe large, with oval excavation posteriolaterally for reception of mesotarsus.</paragraph>
<paragraph id="8BB13698FFDB0915FEBBFD32FCC8EC60" blockId="4.[303,814,671,1047]" pageId="4" pageNumber="15">Legs short, slender. Metacoxae narrow, strongly transverse, separated. Trochanters small, slightly elongate. Femora flattened, slightly dilated medially, carinate dorsally, with deep ventral groove for tibia reception; metafemora not extending beyond elytral lateral margins. Tibiae almost straight, narrow, slightly longer than femora. Tarsi pentamerous, short, shorter than tibia, tarsomeres subcylindrical; protarsi distinctly dilated; relative length ratios of protarsomeres 15 equal to 5-3-2-2-4. Pretarsal claws thin, simple, free, strongly divergent.</paragraph>
<paragraph id="8BB13698FFDB0915FEBBFBADFC54ED00" blockId="4.[303,814,1087,1399]" pageId="4" pageNumber="15">Abdomen with five freely articulated, visible ventrites, covered with small and dense punctation; ventrite 1 with short, triangular intercoxal process, and with transverse excavation in anterior half for reception of metathoracic legs; ventrite 2 largest; ventrite 5 with widely rounded apical margin; sutures slightly concave. Relative length ratios of ventrites 15 equal to 13-16-9-7-9 (medially; including intercoxal process).</paragraph>
<paragraph id="8BB13698FFDB0915FEBBFA32FCBAEE40" blockId="4.[303,814,1439,1591]" pageId="4" pageNumber="15">
<materialsCitation id="3B663CC5FFDB0915FEBBFA32FCB6EE40" collectionCode="MAIG" pageId="4" pageNumber="15" specimenCount="2" typeStatus="paratype">
<typeStatus id="54B5883AFFDB0915FEBBFA32FCBEEDCF" box="[303,395,1440,1464]" pageId="4" pageNumber="15" type="paratype">Paratype</typeStatus>
6782 [
<collectionCode id="ED1FAE5DFFDB0915FE44FA0DFF29EDC0" box="[464,540,1439,1463]" pageId="4" pageNumber="15">MAIG</collectionCode>
] Body length is
<quantity id="4CF69B7DFFDB0915FD5CFA0DFE1FEDC0" box="[712,810,1439,1463]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="15" unit="mm" value="1.42">1.42 mm</quantity>
. Sutures between abdominal ventrites almost straight. Otherwise, this specimen is similar in all visible morphological characters to
<typeStatus id="54B5883AFFDB0915FD0FFA6DFFC1EE60" box="[667,756,1535,1559]" pageId="4" pageNumber="15" type="holotype">holotype</typeStatus>
6781 [
<collectionCode id="ED1FAE5DFFDB0915FEA1F98DFC4BEE40" box="[309,382,1567,1591]" pageId="4" pageNumber="15">MAIG</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BB13698FFDB0915FEBBF9F2FCB6EEA0" blockId="4.[303,814,1631,1751]" pageId="4" pageNumber="15">
<materialsCitation id="3B663CC5FFDB0915FEBBF9F2FCB6EEA0" collectionCode="JHAC, MAIG" pageId="4" pageNumber="15" specimenCount="2" typeStatus="paratype">
<typeStatus id="54B5883AFFDB0915FEBBF9F2FCBDEE0F" box="[303,392,1632,1656]" pageId="4" pageNumber="15" type="paratype">Paratype</typeStatus>
PET01 [
<collectionCode id="ED1FAE5DFFDB0915FE74F9CDFF1DEE00" box="[480,552,1631,1655]" pageId="4" pageNumber="15">JHAC</collectionCode>
]. Body length is
<quantity id="4CF69B7DFFDB0915FD5FF9CDFE1FEE00" box="[715,810,1631,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="15" unit="mm" value="1.45">1.45 mm</quantity>
. Otherwise, this specimen is similar in all visible morphological characters to
<typeStatus id="54B5883AFFDB0915FD11F90DFFDDEEC0" box="[645,744,1695,1719]" pageId="4" pageNumber="15" type="holotype">holotype</typeStatus>
6781 [
<collectionCode id="ED1FAE5DFFDB0915FEA1F92DFC48EEA0" box="[309,381,1727,1751]" pageId="4" pageNumber="15">MAIG</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146513FFDB0915FCC7FF2DF93EEA60" pageId="4" pageNumber="15" type="diagnosis">
<paragraph id="8BB13698FFDB0915FCC7FF2DF93EEA60" blockId="4.[851,1362,191,535]" pageId="4" pageNumber="15">
<emphasis id="B97AEA8AFFDB0915FCC7FF2DF96DE8A0" bold="true" box="[851,1112,191,215]" pageId="4" pageNumber="15">Differential diagnosis.</emphasis>
<taxonomicName id="4C0E4D1BFFDB0915FBF7FF2DF813E8A0" authority="Alekseev &amp; Bukejs, 2021" authorityName="Alekseev &amp; Bukejs" authorityYear="2021" box="[1123,1318,191,215]" class="Insecta" family="Anobiidae" genus="Petalium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="bruteno" status="sp. nov.">
<emphasis id="B97AEA8AFFDB0915FBF7FF2DF813E8A0" box="[1123,1318,191,215]" italics="true" pageId="4" pageNumber="15">Petalium bruteno</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957F1FFDB0915FAA5FF52FEB2E880" pageId="4" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
can easily distinguished from two dorcatomine ptinids described from Baltic amber,
<taxonomicName id="4C0E4D1BFFDB0915FCEBFEB2F811E940" authority="Hava &amp; Zahradnik, 2020" authorityName="Hava &amp; Zahradnik" authorityYear="2020" box="[895,1316,287,311]" class="Insecta" family="Anobiidae" genus="Stagetus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="arturi">
<emphasis id="B97AEA8AFFDB0915FCEBFEB2F92EE940" box="[895,1051,288,311]" italics="true" pageId="4" pageNumber="15">Stagetus arturi</emphasis>
Háva &amp; Zahradník, 2020
</taxonomicName>
and
<taxonomicName id="4C0E4D1BFFDB0915FCC7FEADFEB9E900" authority="Hava &amp; Zahradnik, 2020" authorityName="Hava &amp; Zahradnik" authorityYear="2020" class="Insecta" family="Anobiidae" genus="Calymmaderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="ambericus">
<emphasis id="B97AEA8AFFDB0915FCC7FEADF945E920" box="[851,1136,319,343]" italics="true" pageId="4" pageNumber="15">Calymmaderus ambericus</emphasis>
Háva &amp; Zahradník, 2020
</taxonomicName>
, in the metathoracic ventrite with broad rounded lobe, in another body form and elytral striation (
<taxonomicName id="4C0E4D1BFFDB0915FC56FE0DF9D0E9C0" authorityName="Hava &amp; Zahradnik" authorityYear="2020" box="[962,1253,415,439]" class="Insecta" family="Anobiidae" genus="Calymmaderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="ambericus">
<emphasis id="B97AEA8AFFDB0915FC56FE0DF9D0E9C0" box="[962,1253,415,439]" italics="true" pageId="4" pageNumber="15">Calymmaderus ambericus</emphasis>
</taxonomicName>
has only three distinct striae on apex of each elytron only), distinctly longer ventrite 2 (as long as ventrites 3 and 4 combined).
</paragraph>
</subSubSection>
<subSubSection id="C3146513FFDB0915FCC7FDADF9EEEAAF" pageId="4" pageNumber="15" type="etymology">
<paragraph id="8BB13698FFDB0915FCC7FDADF9EEEAAF" blockId="4.[851,1361,575,728]" pageId="4" pageNumber="15">
<emphasis id="B97AEA8AFFDB0915FCC7FDADF917EA20" bold="true" box="[851,1058,575,599]" pageId="4" pageNumber="15">Derivatio nominis.</emphasis>
Noun in apposition. Bruteno (also Pruteno or Prutanis) is the mythologized Prussian cultural hero, elder brother of Widewuto, the first high priest (Kriwe-Kriwajto) of the pagan Prussians during the VI century.
</paragraph>
</subSubSection>
</treatment>
</document>