treatments-xml/data/03/A7/28/03A72877FF8EB353CDD2FBA5FCCEFED1.xml
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<document id="637D62D8AFB21E2560B019D8E2428FE6" ID-CLB-Dataset="8559" ID-DOI="10.11646/zootaxa.4852.1.5" ID-GBIF-Dataset="bd89417b-161a-4502-aa6b-17ac7e9ea2d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4408343" ID-ZooBank="5C7FCE64-497A-4D75-9779-6345A538CFAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609437762595" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2020" docId="03A72877FF8EB353CDD2FBA5FCCEFED1" docLanguage="en" docName="zootaxa.4852.1.5.pdf" docOrigin="Zootaxa 4852 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus deqenensis Mikhaljova 2020, new species" docType="treatment" docVersion="7" lastPageNumber="113" masterDocId="FF9E500FFF87B35FCD45FFBFFF98FF90" masterDocTitle="New data on the fauna of China, part II: the millipede genus Nepalmatoiulus Mauriès, 1983 (Diplopoda, Julidae) in continental China and Taiwan, with descriptions of seven new species" masterLastPageNumber="124" masterPageNumber="101" pageNumber="110" updateTime="1698850294185" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="89C6F6749D05C09CCAFC4613E8CF4E6E">New data on the fauna of China, part II: the millipede genus Nepalmatoiulus Mauriès, 1983 (Diplopoda, Julidae) in continental China and Taiwan, with descriptions of seven new species</mods:title>
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<mods:namePart id="D208A6038C471733BC67CEE95F8FA2A7">Mikhaljova, Elena V.</mods:namePart>
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<mods:date id="F0F7F1C40BF6213D77954122138D7FF5">2020</mods:date>
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<treatment id="03A72877FF8EB353CDD2FBA5FCCEFED1" ID-DOI="http://doi.org/10.5281/zenodo.4408337" ID-GBIF-Taxon="177200829" ID-Zenodo-Dep="4408337" LSID="urn:lsid:plazi:treatment:03A72877FF8EB353CDD2FBA5FCCEFED1" httpUri="http://treatment.plazi.org/id/03A72877FF8EB353CDD2FBA5FCCEFED1" lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110">
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<paragraph id="8BB19961FF8EB356CDD2FBA5FD19FBA4" blockId="9.[151,641,1050,1113]" box="[151,641,1050,1077]" pageId="9" pageNumber="110">
<heading id="D0F92E0DFF8EB356CDD2FBA5FD19FBA4" bold="true" box="[151,641,1050,1077]" fontSize="11" level="1" pageId="9" pageNumber="110" reason="1">
<emphasis id="B97A4573FF8EB356CDD2FBA5FD19FBA4" bold="true" box="[151,641,1050,1077]" pageId="9" pageNumber="110">
<taxonomicName id="4C0EE2E2FF8EB356CDD2FBA5FE7BFBA5" ID-CoL="246b9749-7781-4e4c-beb8-79e5a016cc50" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[151,483,1050,1077]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="deqenensis" status="sp. nov.">
<emphasis id="B97A4573FF8EB356CDD2FBA5FE7BFBA5" bold="true" box="[151,483,1050,1077]" italics="true" pageId="9" pageNumber="110">Nepalmatoiulus deqenensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A249F808FF8EB356CCB7FBA4FD19FBA4" box="[498,641,1050,1077]" pageId="9" pageNumber="110" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C314CAEAFF8EB356CDD2FB80FE8CFBC9" box="[151,276,1087,1113]" pageId="9" pageNumber="110" type="description">
<paragraph id="8BB19961FF8EB356CDD2FB80FE8CFBC9" blockId="9.[151,641,1050,1113]" box="[151,276,1087,1113]" pageId="9" pageNumber="110">
<figureCitation id="133585E4FF8EB356CDD2FB80FE8CFBC9" box="[151,276,1087,1113]" captionStart-0="FIGURES 3743" captionStart-1="FIGURES 4446" captionStartId-0="10.[151,264,1938,1962]" captionStartId-1="11.[151,264,1974,1998]" captionTargetBox-0="[151,1435,705,1914]" captionTargetBox-1="[195,1392,181,1950]" captionTargetId-0="figure-272@10.[151,1435,705,1914]" captionTargetId-1="figure-16@11.[194,1392,181,1950]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." captionText-1="FIGURES 4446. Nepalmatoiulus deqenensis sp. nov., male paratype. 44, opisthomere, mesal view; 45, solenomere; 46, apex of mesomeral process; d, outgrowth of opisthomere mesomeral process. Scales: 10 μm (Figs 4546), 100 μm (Fig. 44)." figureDoi-0="http://doi.org/10.5281/zenodo.4408363" figureDoi-1="http://doi.org/10.5281/zenodo.4408365" httpUri-0="https://zenodo.org/record/4408363/files/figure.png" httpUri-1="https://zenodo.org/record/4408365/files/figure.png" pageId="9" pageNumber="110">Figs 3746</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C314CAEAFF8EB356CDD2FB39FD02FA9D" pageId="9" pageNumber="110" type="materials_examined">
<paragraph id="8BB19961FF8EB356CDD2FB39FD02FA9D" blockId="9.[151,1437,1158,2013]" pageId="9" pageNumber="110">
<emphasis id="B97A4573FF8EB356CDD2FB39FE18FB30" bold="true" box="[151,384,1158,1184]" pageId="9" pageNumber="110">Material examined.</emphasis>
<materialsCitation id="3B66933CFF8EB356CCC3FB38FEBCFB79" ID-GBIF-Occurrence="3027951319" collectingDate="2013-06-14" collectionCode="ZMUM" collectorName="I. Belousov &amp; I. Kabak &amp; G. Davidian" country="China" county="Deqen" elevation="3950" latitude="27.650278" location="Dewei Line Mt. Range" longLatPrecision="20" longitude="99.18083" municipality="Weixi" pageId="9" pageNumber="110" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B97A4573FF8EB356CCC3FB38FE75FB31" box="[390,493,1159,1185]" italics="true" pageId="9" pageNumber="110">
<typeStatus id="54B527C3FF8EB356CCC3FB38FE75FB31" box="[390,493,1159,1185]" pageId="9" pageNumber="110" type="holotype">Holotype</typeStatus>
</emphasis>
: male (
<collectionCode id="ED1F01A4FF8EB356CF7AFB38FD04FB31" box="[575,668,1159,1185]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="110">ZMUM</collectionCode>
),
<collectingCountry id="F319D9F1FF8EB356CFEBFB38FD6CFB31" box="[686,756,1159,1185]" name="China" pageId="9" pageNumber="110">China</collectingCountry>
,
<collectingRegion id="49CA5783FF8EB356CFB8FB38FC26FB31" box="[765,958,1159,1185]" country="China" name="Yunnan" pageId="9" pageNumber="110">Yunnan Province</collectingRegion>
,
<collectingCounty id="62D0E1EDFF8EB356CE8CFB38FB8DFB31" box="[969,1045,1159,1185]" pageId="9" pageNumber="110">Deqen</collectingCounty>
,
<collectingMunicipality id="6BD5031BFF8EB356C95AFB38FBFCFB31" box="[1055,1124,1159,1185]" pageId="9" pageNumber="110">Weixi</collectingMunicipality>
,
<location id="8ED1CFBAFF8EB356C92BFB38FAF5FB31" LSID="urn:lsid:plazi:treatment:03A72877FF8EB353CDD2FBA5FCCEFED1:8ED1CFBAFF8EB356C92BFB38FAF5FB31" box="[1134,1389,1159,1185]" country="China" county="Deqen" latitude="27.650278" longLatPrecision="20" longitude="99.18083" municipality="Weixi" name="Dewei Line Mt. Range" pageId="9" pageNumber="110" stateProvince="Yunnan">Dewei Line Mt. Range</location>
btw
<collectorName id="26FBFCB7FF8EB356CDD2FB14FEFEFB55" box="[151,358,1195,1221]" pageId="9" pageNumber="110">Xiaruolisuzuxiang</collectorName>
&amp;
<collectorName id="26FBFCB7FF8EB356CCCDFB14FD98FB55" box="[392,512,1195,1221]" pageId="9" pageNumber="110">Yezhizhen</collectorName>
,
<geoCoordinate id="EE3AFFA6FF8EB356CF4EFB14FD0EFB55" box="[523,662,1195,1221]" degrees="27" direction="north" minutes="39" orientation="latitude" pageId="9" pageNumber="110" precision="15" seconds="1" value="27.650278">27°39´1´´ N</geoCoordinate>
,
<geoCoordinate id="EE3AFFA6FF8EB356CFE4FB15FCAEFB55" box="[673,822,1194,1221]" degrees="99" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="110" precision="15" seconds="51" value="99.18083">99°10´51´´ E</geoCoordinate>
, H =
<quantity id="4CF63484FF8EB356CE36FB15FC52FB55" box="[883,970,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="9" pageNumber="110" unit="m" value="3950.0">
<elevation id="00237E52FF8EB356CE36FB15FC52FB55" box="[883,970,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="9" pageNumber="110" unit="m" value="3950.0">3950 m</elevation>
</quantity>
,
<date id="FFB0BFA1FF8EB356CE93FB14FBF6FB55" box="[982,1134,1195,1221]" pageId="9" pageNumber="110" value="2013-06-14">
<collectingDate id="EFF44649FF8EB356CE93FB14FBF6FB55" box="[982,1134,1195,1221]" pageId="9" pageNumber="110" value="2013-06-14">14 June 2013</collectingDate>
</date>
, leg.
<collectorName id="26FBFCB7FF8EB356C9EDFB14FAB2FB55" box="[1192,1322,1195,1221]" pageId="9" pageNumber="110">I. Belousov</collectorName>
, I.
<collectorName id="26FBFCB7FF8EB356C809FB14FF36FB79" pageId="9" pageNumber="110">Kabak, G</collectorName>
. Davidian
</materialsCitation>
;
<materialsCitation id="3B66933CFF8EB356CC75FB70FCACFB79" ID-GBIF-Occurrence="3027951318" box="[304,820,1231,1257]" collectionCode="ZMUM" country="China" pageId="9" pageNumber="110" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B97A4573FF8EB356CC75FB70FE3AFB78" box="[304,418,1231,1256]" italics="true" pageId="9" pageNumber="110">
<typeStatus id="54B527C3FF8EB356CC75FB70FE3AFB78" box="[304,418,1231,1256]" pageId="9" pageNumber="110" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D0852E8FF8EB356CCF7FB70FD98FB79" box="[434,512,1231,1257]" pageId="9" pageNumber="110" type="male">1 male</specimenCount>
,
<specimenCount id="9D0852E8FF8EB356CF48FB70FD1AFB79" box="[525,642,1231,1257]" pageId="9" pageNumber="110" type="subadult">1 subadult</specimenCount>
male (
<collectionCode id="ED1F01A4FF8EB356CF8AFB70FCB4FB79" box="[719,812,1231,1257]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="110">ZMUM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B66933CFF8EB356CE05FB70FD02FA9D" ID-GBIF-Occurrence="3027951307" collectingDate="2013-06-14" collectionCode="FSCB" collectorName="I. Belousov &amp; Belousov, I &amp; Kabak, G." country="China" pageId="9" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D0852E8FF8EB356CE05FB70FC14FB79" box="[832,908,1231,1257]" pageId="9" pageNumber="110" type="male">1 male</specimenCount>
(
<collectionCode id="ED1F01A4FF8EB356CED9FB70FC7EFB79" box="[924,998,1231,1257]" pageId="9" pageNumber="110">FSCB</collectionCode>
), same locality as in
<typeStatus id="54B527C3FF8EB356C991FB70FAA1FB79" box="[1236,1337,1231,1257]" pageId="9" pageNumber="110" type="holotype">holotype</typeStatus>
,
<date id="FFB0BFA1FF8EB356C800FB70FF4AFA9D" pageId="9" pageNumber="110" value="2013-06-14">
<collectingDate id="EFF44649FF8EB356C800FB70FF4AFA9D" pageId="9" pageNumber="110" value="2013-06-14">14 June 2013</collectingDate>
</date>
, leg.
<collectorName id="26FBFCB7FF8EB356CC48FB4CFE08FA9D" box="[269,400,1267,1293]" pageId="9" pageNumber="110">I. Belousov</collectorName>
, I.
<collectorName id="26FBFCB7FF8EB356CCF7FB4CFDBCFA9D" box="[434,548,1267,1293]" pageId="9" pageNumber="110">Kabak, G.</collectorName>
Davidian.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C314CAEAFF8EB356CD82FAA9FAF6FAC5" pageId="9" pageNumber="110" type="diagnosis">
<paragraph id="8BB19961FF8EB356CD82FAA9FAF6FAC5" blockId="9.[151,1437,1158,2013]" pageId="9" pageNumber="110">
<emphasis id="B97A4573FF8EB356CD82FAA9FEDAFAA0" bold="true" box="[199,322,1302,1328]" pageId="9" pageNumber="110">Diagnosis.</emphasis>
Differs from congeners mainly by the hook-shaped mesomeral process of posterior gonopod with strong subapical mesal knob-like outgrowth and the absence of velar process coupled with a short solenomere.
</paragraph>
</subSubSection>
<subSubSection id="C314CAEAFF8EB353CD82FAE1FE09FE8D" lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110" type="description">
<paragraph id="8BB19961FF8EB355CD82FAE1FE71FD39" blockId="9.[151,1437,1158,2013]" lastBlockId="10.[151,1437,151,681]" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110">
<emphasis id="B97A4573FF8EB356CD82FAE1FEC1FAE8" bold="true" box="[199,345,1374,1400]" pageId="9" pageNumber="110">Description.</emphasis>
Male. Length
<quantity id="4CF63484FF8EB356CF43FAE1FD1CFAE9" box="[518,644,1374,1401]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="9" pageNumber="110" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">2325 mm</quantity>
, midbody vertical diameter about 2.0 mm, with 52 (1), 45 (2), 48 (2) segments, excluding telson. Coloration (in alcohol) marbled brown-gray with broad longitudinal dorsal stripe formed by lighter dorso-anterior metazonital portions; venter (including basal parts of legs) lighter. Distal portions of legs brown, antennae and telson dark brown. Occiput and forehead marbled brown, ventral parts of head light brown. Eyes black.
<emphasis id="B97A4573FF8EB356CC3CFA50FE21F999" bold="true" box="[377,441,1519,1545]" pageId="9" pageNumber="110">Head</emphasis>
smooth, covered with sparse setae. Epicranial setae not visible, 4 supralabral setae, not less than 20 labral setae. Eye patches almost oval, composed of more than 50 ocelli (in a male
<typeStatus id="54B527C3FF8EB356C9FEF9ACFAB9F9BD" box="[1211,1313,1555,1581]" pageId="9" pageNumber="110" type="paratype">paratype</typeStatus>
). Antennae medium-sized, rather slender and clavate. Gnathochilarium with not less than 10 nonapical stipital setae. Mandibular stipite with subtriangular rounded smooth lobe. More than 3 setae on each lamella linguales.
<emphasis id="B97A4573FF8EB356C9A5F9E5FAA1F9E4" bold="true" box="[1248,1337,1626,1652]" pageId="9" pageNumber="110">Collum</emphasis>
laterally with distinct lower striae of middle length at the posterior margin not reaching to anterior margin, dorsally with indistinct shorter striae at the posterior margin only. A transverse row of sparse setae at hind edge of collum. Body segments circular. Prozona smooth except for ventral part with obliquely located striae. The metazonite gradually a little narrower toward the venter. Metazona with dense, regular, longitudinal striae reaching hind margin (1213 striae in an approximate square with sides equal to metazonital length of a dorsal side of midbody ring). Limbus straight, smooth (of Type
<quantity id="4CF63484FF8EB356CCF0F88CFE78F8DD" box="[437,480,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="110" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EF9FE490FF8EB356CCA3F88CFD1BF8DD" author="Enghoff, H." box="[486,643,1843,1869]" pageId="9" pageNumber="110" pagination="241 - 331" refId="ref14079" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
). A transverse row of thin setae at hind edge of metazonites, setae gradually growing denser toward telson. Ozopores small, lying immediately behind suture dividing pro- and metazona. Caudal dorsal projection of telson straight and long, setose, tipped with a claw-shaped process curved dorsally. Preanal ring, anal valves, and subanal scale setose.
<emphasis id="B97A4573FF8EB356CF91F820FC94F829" bold="true" box="[724,780,1951,1977]" pageId="9" pageNumber="110">Legs</emphasis>
relatively short and slender. Very delicately serrate ventral pads present on postfemur and tibia, starting from legs 2; pads somewhat decreasing in size towards telson. The hind most legs without ventral pads on tibia. Claw of all legs at base with a long (longer than claw) setiform accessory claw ventrally but without accessory claw dorsally. Leg pair 1 forming hook, the distal segment not touching basal segments (“Open hook” type in
<bibRefCitation id="EF9FE490FF8DB355CF03FF60FD7DFF69" author="Enghoff, H." box="[582,741,223,249]" pageId="10" pageNumber="111" pagination="241 - 331" refId="ref14079" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
); postfemur with scaly-rugose ventral surface, tip indistinctly wrinkly; coxa with one seta, distal segment without seta and tarsal remnant (
<figureCitation id="133585E4FF8DB355CEBBFEBCFBCCFE8D" box="[1022,1108,259,285]" captionStart="FIGURES 3743" captionStartId="10.[151,264,1938,1962]" captionTargetBox="[151,1435,705,1914]" captionTargetId="figure-272@10.[151,1435,705,1914]" captionTargetPageId="10" captionText="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." figureDoi="http://doi.org/10.5281/zenodo.4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111">Fig. 37</figureCitation>
). Coxa 2 with one mesapical oral seta, a gland opening positioned in central and axial position according to
<bibRefCitation id="EF9FE490FF8DB355C95FFE98FB2DFED1" author="Enghoff, H." box="[1050,1205,295,321]" pageId="10" pageNumber="111" pagination="241 - 331" refId="ref14079" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
(
<figureCitation id="133585E4FF8DB355C980FE98FAD2FED1" box="[1221,1354,295,321]" captionStart="FIGURES 3743" captionStartId="10.[151,264,1938,1962]" captionTargetBox="[151,1435,705,1914]" captionTargetId="figure-272@10.[151,1435,705,1914]" captionTargetPageId="10" captionText="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." figureDoi="http://doi.org/10.5281/zenodo.4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111">Figs 38, 41</figureCitation>
). Penis relatively long, slender, about 1.8 longer than wide (
<figureCitation id="133585E4FF8DB355CF9CFEF4FCB4FEF5" box="[729,812,331,357]" captionStart="FIGURES 3743" captionStartId="10.[151,264,1938,1962]" captionTargetBox="[151,1435,705,1914]" captionTargetId="figure-272@10.[151,1435,705,1914]" captionTargetPageId="10" captionText="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." figureDoi="http://doi.org/10.5281/zenodo.4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111">Fig. 43</figureCitation>
). Ventral margin of body segment VII with lobes similar to figure
<quantity id="4CF63484FF8DB355CC47FED0FEA6FE19" box="[258,318,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="10" pageNumber="111" unit="in" value="32.0">32 in</quantity>
<bibRefCitation id="EF9FE490FF8DB355CC02FED0FE7EFE19" author="Enghoff, H." box="[327,486,367,393]" pageId="10" pageNumber="111" pagination="241 - 331" refId="ref14079" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
.
<emphasis id="B97A4573FF8DB355CCB1FED1FDF5FE18" bold="true" box="[500,621,366,392]" pageId="10" pageNumber="111">Gonopods</emphasis>
protruding. Anterior gonopod flattened, with parallel margins, apically rounded, in posterior view subapically excavated for accommodation of mesomeral process; distal margins of the excavation papillate. Subcentrally with knoblike flattened rudimentary telopodite (
<figureCitation id="133585E4FF8DB355C973FE08FB21FE41" box="[1078,1209,439,465]" captionStart="FIGURES 3743" captionStartId="10.[151,264,1938,1962]" captionTargetBox="[151,1435,705,1914]" captionTargetId="figure-272@10.[151,1435,705,1914]" captionTargetPageId="10" captionText="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." figureDoi="http://doi.org/10.5281/zenodo.4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111">Figs 3940</figureCitation>
). Flagellum slender, relatively short, covered with cuticular conical obtuse denticles in the distal part (
<figureCitation id="133585E4FF8DB355C968FE64FB1AFE65" box="[1069,1154,475,501]" captionStart="FIGURES 3743" captionStartId="10.[151,264,1938,1962]" captionTargetBox="[151,1435,705,1914]" captionTargetId="figure-272@10.[151,1435,705,1914]" captionTargetPageId="10" captionText="FIGURES 3743. Nepalmatoiulus deqenensis sp. nov., male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41)." figureDoi="http://doi.org/10.5281/zenodo.4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111">Fig. 42</figureCitation>
); basal part of the flagellum caudally covered with cuticular spikes. Opisthomere hook-shaped, smooth mesomeral process subapically with strong mesal knob-like outgrowth (
<emphasis id="B97A4573FF8DB355CF6BFD9CFDA6FDAD" bold="true" box="[558,574,547,573]" pageId="10" pageNumber="111">d</emphasis>
) (
<figureCitation id="133585E4FF8DB355CF1CFD9CFD79FDAE" box="[601,737,547,574]" captionStart="FIGURES 4446" captionStartId="11.[151,264,1974,1998]" captionTargetBox="[195,1392,181,1950]" captionTargetId="figure-16@11.[194,1392,181,1950]" captionTargetPageId="11" captionText="FIGURES 4446. Nepalmatoiulus deqenensis sp. nov., male paratype. 44, opisthomere, mesal view; 45, solenomere; 46, apex of mesomeral process; d, outgrowth of opisthomere mesomeral process. Scales: 10 μm (Figs 4546), 100 μm (Fig. 44)." figureDoi="http://doi.org/10.5281/zenodo.4408365" httpUri="https://zenodo.org/record/4408365/files/figure.png" pageId="10" pageNumber="111">Figs 44, 46</figureCitation>
). Membranous velum with sloped smooth margin, without a notch near the mesomeral process. Velum smooth. Solenomere short, spinose throughout (
<figureCitation id="133585E4FF8DB355C939FDF8FB57FDF1" box="[1148,1231,582,609]" captionStart="FIGURES 4446" captionStartId="11.[151,264,1974,1998]" captionTargetBox="[195,1392,181,1950]" captionTargetId="figure-16@11.[194,1392,181,1950]" captionTargetPageId="11" captionText="FIGURES 4446. Nepalmatoiulus deqenensis sp. nov., male paratype. 44, opisthomere, mesal view; 45, solenomere; 46, apex of mesomeral process; d, outgrowth of opisthomere mesomeral process. Scales: 10 μm (Figs 4546), 100 μm (Fig. 44)." figureDoi="http://doi.org/10.5281/zenodo.4408365" httpUri="https://zenodo.org/record/4408365/files/figure.png" pageId="10" pageNumber="111">Fig. 45</figureCitation>
). Margin of accessory membrane smooth. Acicular process of a flagellum-quiding flap arising from posterior margin of solenomeral furrow not seen in the figures.
</paragraph>
<caption id="DF71C9E9FF8DB355CDD2F82DFC85F862" ID-DOI="http://doi.org/10.5281/zenodo.4408363" ID-Zenodo-Dep="4408363" httpUri="https://zenodo.org/record/4408363/files/figure.png" pageId="10" pageNumber="111" startId="10.[151,264,1938,1962]" targetBox="[151,1435,705,1914]" targetPageId="10">
<paragraph id="8BB19961FF8DB355CDD2F82DFC85F862" blockId="10.[151,1437,1938,2035]" pageId="10" pageNumber="111">
<emphasis id="B97A4573FF8DB355CDD2F82DFECDF83A" bold="true" box="[151,341,1938,1962]" pageId="10" pageNumber="111">FIGURES 3743.</emphasis>
<taxonomicName id="4C0EE2E2FF8DB355CC19F82DFDEBF83A" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[348,627,1938,1962]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="deqenensis" status="sp. nov.">
<emphasis id="B97A4573FF8DB355CC19F82DFDEBF83A" box="[348,627,1938,1962]" italics="true" pageId="10" pageNumber="111">Nepalmatoiulus deqenensis</emphasis>
</taxonomicName>
<emphasis id="B97A4573FF8DB355CF3FF82DFD53F83A" bold="true" box="[634,715,1938,1962]" pageId="10" pageNumber="111">
<taxonomicNameLabel id="A249F808FF8DB355CF3FF82DFD53F83A" box="[634,715,1938,1962]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype. 37, leg pair 1; 38, leg pair 2, front view; 39, promere, caudal view; 40, distal part of promere; 41, coxae 2, front view; 42, distal part of flagellum; 43, penis. Scales: 10 μm (Fig. 42), 20 μm (Figs 37, 40), 30 μm (Fig. 43), 100 μm (Figs 3839, 41).
</paragraph>
</caption>
<caption id="DF71C9E9FF8CB354CDD2F809FAC8F862" ID-DOI="http://doi.org/10.5281/zenodo.4408365" ID-Zenodo-Dep="4408365" httpUri="https://zenodo.org/record/4408365/files/figure.png" pageId="11" pageNumber="112" startId="11.[151,264,1974,1998]" targetBox="[195,1392,181,1950]" targetPageId="11">
<paragraph id="8BB19961FF8CB354CDD2F809FAC8F862" blockId="11.[151,1436,1974,2035]" pageId="11" pageNumber="112">
<emphasis id="B97A4573FF8CB354CDD2F809FECCF85E" bold="true" box="[151,340,1974,1998]" pageId="11" pageNumber="112">FIGURES 4446.</emphasis>
<taxonomicName id="4C0EE2E2FF8CB354CC1FF809FDE8F85E" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[346,624,1974,1998]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="deqenensis" status="sp. nov.">
<emphasis id="B97A4573FF8CB354CC1FF809FDE8F85E" box="[346,624,1974,1998]" italics="true" pageId="11" pageNumber="112">Nepalmatoiulus deqenensis</emphasis>
</taxonomicName>
<emphasis id="B97A4573FF8CB354CF33F809FD5FF85E" bold="true" box="[630,711,1974,1998]" pageId="11" pageNumber="112">
<taxonomicNameLabel id="A249F808FF8CB354CF33F809FD5FF85E" box="[630,711,1974,1998]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype. 44, opisthomere, mesal view; 45, solenomere; 46, apex of mesomeral process;
<emphasis id="B97A4573FF8CB354CCC4F865FE17F862" bold="true" box="[385,399,2010,2034]" pageId="11" pageNumber="112">d</emphasis>
, outgrowth of opisthomere mesomeral process. Scales: 10 μm (Figs 4546), 100 μm (Fig. 44).
</paragraph>
</caption>
<paragraph id="8BB19961FF8BB353CD82FF28FE4FFF69" blockId="12.[151,1437,150,321]" pageId="12" pageNumber="113">
Subadult male. Length and midbody vertical diameter as in
<typeStatus id="54B527C3FF8BB353CE26FF28FC50FF21" box="[867,968,151,177]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
, i.e. about
<quantity id="4CF63484FF8BB353C905FF29FB09FF21" box="[1088,1169,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="113" unit="mm" value="25.0">25 mm</quantity>
, with 47 (2) segments, excluding telson. Leg pair 1 normal (not modificated). Gonopods very slightly protruding, probably undeveloped. The gonopods not dissected.
</paragraph>
<paragraph id="8BB19961FF8BB353CD82FEBCFE09FE8D" blockId="12.[151,1437,150,321]" box="[199,401,259,285]" pageId="12" pageNumber="113">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C314CAEAFF8BB353CD82FE98FCCEFED1" box="[199,854,295,321]" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8BB19961FF8BB353CD82FE98FCCEFED1" blockId="12.[151,1437,150,321]" box="[199,854,295,321]" pageId="12" pageNumber="113">
<emphasis id="B97A4573FF8BB353CD82FE98FED6FED1" bold="true" box="[199,334,295,321]" pageId="12" pageNumber="113">Etymology.</emphasis>
The specific epithet refers to the
<typeStatus id="54B527C3FF8BB353CF80FE98FD6DFED1" box="[709,757,295,321]" pageId="12" pageNumber="113">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>