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<document id="807A17CFF72C189BF2FD367F927B9F6F" ID-DOI="10.11646/zootaxa.4999.6.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5119668" ID-ZooBank="2355BCA5-4260-4CDF-8545-8FD854D3E6A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626879889455" checkinUser="plazi" docAuthor="Benzel, Joseph &amp; Bright, Donald E." docDate="2021" docId="039F9144FFFAF86E40AAFAB6FA741809" docLanguage="en" docName="zootaxa.4999.6.1.pdf" docOrigin="Zootaxa 4999 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyrhinus Schonherr 1823" docType="key" docVersion="3" lastPageNumber="531" masterDocId="FFA6E93CFFE4F870403DFF8DFFE91E76" masterDocTitle="A revision of the genus Pachyrhinus Schӧnherr 1823 (Coleoptera: Curculionidae Entiminae) in the Nearctic Region" masterLastPageNumber="533" masterPageNumber="501" pageNumber="531" updateTime="1699012250246" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DC6CAE9859304D8D10D4C7F8F4E278F1">A revision of the genus Pachyrhinus Schӧnherr 1823 (Coleoptera: Curculionidae Entiminae) in the Nearctic Region</mods:title>
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<mods:namePart id="BE9FA3307875821EF989E97378556592">Benzel, Joseph</mods:namePart>
<mods:affiliation id="BE59E27F1699130D7E2FFA4534677BD6">C. P. Gillete Museum of Arthropod Diversity, Department of Biological Sciences and Pest Management. Colorado State University, Fort Collins, CO 80525 &amp; Current address: 2265 Wolfensberger Road, Castle Rock, CO 80109</mods:affiliation>
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<mods:namePart id="6010CCA1F752D81EE4F8C836714343E2">Bright, Donald E.</mods:namePart>
<mods:affiliation id="70CE1413A06598E63154D30AA7CA924C">C. P. Gillete Museum of Arthropod Diversity, Department of Biological Sciences and Pest Management. Colorado State University, Fort Collins, CO 80525</mods:affiliation>
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<mods:title id="9173F9431E7F236242155DF8AAC441F0">Zootaxa</mods:title>
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<mods:date id="BC60B4A5C28E44BB13F8035B16590007">2021</mods:date>
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<mods:number id="9EBDE0FA00F911D0B0EC43E3DBF39EA7">2021-07-15</mods:number>
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<mods:number id="B9070B461184276E3938990E52F7E13C">4999</mods:number>
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<mods:identifier id="27D4D9F945153F4FFDC7A7F7279709CB" type="DOI">10.11646/zootaxa.4999.6.1</mods:identifier>
<mods:identifier id="934827F15678CBEEB75747BED662D49D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039F9144FFFAF86E40AAFAB6FA741809" LSID="urn:lsid:plazi:treatment:039F9144FFFAF86E40AAFAB6FA741809" httpUri="http://treatment.plazi.org/id/039F9144FFFAF86E40AAFAB6FA741809" lastPageNumber="531" pageId="30" pageNumber="531">
<subSubSection id="C32C73D9FFFAF86E40AAFAB6FD411B23" box="[151,680,1338,1365]" pageId="30" pageNumber="531" type="nomenclature">
<paragraph id="8B892052FFFAF86E40AAFAB6FD411B23" blockId="30.[151,680,1338,1365]" box="[151,680,1338,1365]" pageId="30" pageNumber="531">
<heading id="D0C1973EFFFAF86E40AAFAB6FD411B23" bold="true" box="[151,680,1338,1365]" fontSize="11" level="1" pageId="30" pageNumber="531" reason="1">
<emphasis id="B942FC40FFFAF86E40AAFAB6FD411B23" bold="true" box="[151,680,1338,1365]" pageId="30" pageNumber="531">
Key to the Nearctic species of
<taxonomicName id="4C365BD1FFFAF86E4230FAB7FD411B23" ID-CoL="8HMFJ" authorityName="Schonherr" authorityYear="1823" box="[525,680,1338,1365]" class="Insecta" family="Curculionidae" genus="Pachyrhinus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="B942FC40FFFAF86E4230FAB7FD411B23" bold="true" box="[525,680,1338,1365]" italics="true" pageId="30" pageNumber="531">Pachyrhinus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C73D9FFFAF86E40AAFA0DFA741809" pageId="30" pageNumber="531" type="key">
<keyStep id="30C238F7FFFAF86E40AAFA0DFA751B9B" pageId="30" pageNumber="531">
<paragraph id="8B892052FFFAF86E40AAFA0DFA751BC4" blockId="30.[151,1437,1407,1664]" pageId="30" pageNumber="531">
<keyLead id="30C78367FFFAF86E40AAFA0DFA751BC4" pageId="30" pageNumber="531">
1 Scales in mottled color pattern on elytral disc (
<figureCitation id="130D3CD7FFFAF86E429DFA0DFCFD1BE0" box="[672,788,1408,1430]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1745,1769]" captionTargetBox="[165,1422,885,1717]" captionTargetId="figure-48@4.[165,1422,885,1720]" captionTargetPageId="4" captionText="FIGURES 10–15. Elytral Vestiture. 10. Pachyrhinus elegans. 11. Pachyrhinus californicus. 12. Pachyrhinus cinereus. 13. Pachyrhinus squamulosus. 14. Pachyrhinus lethierryi. 15. Polydrusus sericeus." figureDoi="http://doi.org/10.5281/zenodo.5119674" httpUri="https://zenodo.org/record/5119674/files/figure.png" pageId="30" pageNumber="531">Figs. 11–12</figureCitation>
). Aedeagus highly arcuate with a tapered tip (
<figureCitation id="130D3CD7FFFAF86E44F4FA0DFAD71BE3" box="[1225,1342,1407,1430]" captionStart="FIGURES 73–78" captionStartId="17.[151,264,1859,1883]" captionTargetBox="[174,1415,417,1836]" captionTargetId="figure-138@17.[173,1415,417,1836]" captionTargetPageId="17" captionText="FIGURES 73–78. 73. Pachyrhinus californicus left elytron. 74. Stria. 75. Aedeagus, lateral view. 76. Dorsal view. 77. Ventral view. 78. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119700" httpUri="https://zenodo.org/record/5119700/files/figure.png" pageId="30" pageNumber="531">Figs. 75–77</figureCitation>
), spermatheca variable........................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="8B892052FFFAF86E40AAFA36FA751B9B" blockId="30.[151,1437,1407,1664]" pageId="30" pageNumber="531">
<keyLead id="30C78367FFFAF86E40AAFA36FA751B9B" pageId="30" pageNumber="531">
- Scales not in mottled pattern (
<figureCitation id="130D3CD7FFFAF86E41C3FA37FDAF1BA6" box="[510,582,1466,1488]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1745,1769]" captionTargetBox="[165,1422,885,1717]" captionTargetId="figure-48@4.[165,1422,885,1720]" captionTargetPageId="4" captionText="FIGURES 10–15. Elytral Vestiture. 10. Pachyrhinus elegans. 11. Pachyrhinus californicus. 12. Pachyrhinus cinereus. 13. Pachyrhinus squamulosus. 14. Pachyrhinus lethierryi. 15. Polydrusus sericeus." figureDoi="http://doi.org/10.5281/zenodo.5119674" httpUri="https://zenodo.org/record/5119674/files/figure.png" pageId="30" pageNumber="531">Fig. 10</figureCitation>
). Aedeagus feebly arcuate with a wide blunt tip (
<figureCitation id="130D3CD7FFFAF86E4428FA37FB691BA6" box="[1045,1152,1466,1488]" captionStart="FIGURES 39–44" captionStartId="10.[151,264,1955,1979]" captionTargetBox="[165,1422,489,1930]" captionTargetId="figure-135@10.[162,1426,489,1931]" captionTargetPageId="10" captionText="FIGURES 39–44. Pachyrhinus elegans 39. Left elytron, dorsal view. 40. Stria and interstria. 41. Aedeagus, lateral view. 42. Aedeagus, ventral view. 43. Aedeagus, dorsal view. 44. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119688" httpUri="https://zenodo.org/record/5119688/files/figure.png" pageId="30" pageNumber="531">Fig. 41–43</figureCitation>
), spermatheca C-shaped (
<figureCitation id="130D3CD7FFFAF86E4548FA37FF141B98" captionStart="FIGURES 39–44" captionStartId="10.[151,264,1955,1979]" captionTargetBox="[165,1422,489,1930]" captionTargetId="figure-135@10.[162,1426,489,1931]" captionTargetPageId="10" captionText="FIGURES 39–44. Pachyrhinus elegans 39. Left elytron, dorsal view. 40. Stria and interstria. 41. Aedeagus, lateral view. 42. Aedeagus, ventral view. 43. Aedeagus, dorsal view. 44. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119688" httpUri="https://zenodo.org/record/5119688/files/figure.png" pageId="30" pageNumber="531">Fig. 44</figureCitation>
). Temperate North America...................................................................
<taxonomicName id="4C365BD1FFFAF86E4506FA55FA751B9B" ID-CoL="75HPF" authorityName=": O'Brien &amp; Wibmer and Wibmer" authorityYear="1982" box="[1339,1436,1496,1517]" class="Insecta" family="Curculionidae" genus="Pachyrhinus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="531" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B942FC40FFFAF86E4506FA55FA751B9B" box="[1339,1436,1496,1517]" italics="true" pageId="30" pageNumber="531">P. elegans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C238F7FFFAF86E40AAFA78FA741809" pageId="30" pageNumber="531">
<paragraph id="8B892052FFFAF86E40AAFA78FA75185E" blockId="30.[151,1437,1407,1664]" pageId="30" pageNumber="531">
<keyLead id="30C78367FFFAF86E40AAFA78FA75185E" pageId="30" pageNumber="531">
2 Scales 1.5X as long as wide, with blunt tips, teardrop shaped (
<figureCitation id="130D3CD7FFFAF86E4309FA78FC84187D" box="[820,877,1525,1547]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,801,825]" captionTargetBox="[164,1424,181,777]" captionTargetId="figure-20@4.[163,1424,181,777]" captionTargetPageId="4" captionText="FIGURES 6–9. Elytral Scales. 6. Pachyrhinus elegans. 7. Pachyrhinus californicus. 8. Pachyrhinus cinereus scale. 9. Polydrusus impressifrons Gyllenhall 1834." figureDoi="http://doi.org/10.5281/zenodo.5119672" httpUri="https://zenodo.org/record/5119672/files/figure.png" pageId="30" pageNumber="531">Fig. 8</figureCitation>
). Elongate interspersed scales on elytra bluntly tipped and recumbent to slightly appressed (
<figureCitation id="130D3CD7FFFAF86E421DF99FFD8E185E" box="[544,615,1554,1576]" captionStart="FIGURES 96–101" captionStartId="24.[151,264,1661,1685]" captionTargetBox="[157,1429,181,1637]" captionTargetId="figure-20@24.[157,1430,181,1637]" captionTargetPageId="24" captionText="FIGURES 96–101. 96. Pachyrhinus cinereus left elytron. 97. P. cinereus stria. 98. P. cinereus aedeagus lateral view. 99. Ventral view. 100. Dorsal view. 101. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119710" httpUri="https://zenodo.org/record/5119710/files/figure.png" pageId="30" pageNumber="531">Fig. 97</figureCitation>
). Spermatheca comma-shaped (
<figureCitation id="130D3CD7FFFAF86E43A4F99FFC04185E" box="[921,1005,1554,1576]" captionStart="FIGURES 96–101" captionStartId="24.[151,264,1661,1685]" captionTargetBox="[157,1429,181,1637]" captionTargetId="figure-20@24.[157,1430,181,1637]" captionTargetPageId="24" captionText="FIGURES 96–101. 96. Pachyrhinus cinereus left elytron. 97. P. cinereus stria. 98. P. cinereus aedeagus lateral view. 99. Ventral view. 100. Dorsal view. 101. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119710" httpUri="https://zenodo.org/record/5119710/files/figure.png" pageId="30" pageNumber="531">Fig. 101</figureCitation>
). Western North America......
<taxonomicName id="4C365BD1FFFAF86E4509F99EFA75185E" ID-CoL="BFVBK" authorityName=": O'Brien &amp; Wibmer" authorityYear="1982" box="[1332,1436,1555,1576]" class="Insecta" family="Curculionidae" genus="Pachyrhinus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="531" phylum="Arthropoda" rank="species" species="cinereus">
<emphasis id="B942FC40FFFAF86E4509F99EFA75185E" box="[1332,1436,1555,1576]" italics="true" pageId="30" pageNumber="531">P. cinereus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B892052FFFAF86E40AAF9BDFA741809" blockId="30.[151,1437,1407,1664]" pageId="30" pageNumber="531">
<keyLead id="30C78367FFFAF86E40AAF9BDFA741809" pageId="30" pageNumber="531">
- Scales 2.0 to 4.0X long as wide with tapered ends sometimes setose in form (
<figureCitation id="130D3CD7FFFAF86E4387F9BDFC1D1833" box="[954,1012,1584,1606]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,801,825]" captionTargetBox="[164,1424,181,777]" captionTargetId="figure-20@4.[163,1424,181,777]" captionTargetPageId="4" captionText="FIGURES 6–9. Elytral Scales. 6. Pachyrhinus elegans. 7. Pachyrhinus californicus. 8. Pachyrhinus cinereus scale. 9. Polydrusus impressifrons Gyllenhall 1834." figureDoi="http://doi.org/10.5281/zenodo.5119672" httpUri="https://zenodo.org/record/5119672/files/figure.png" pageId="30" pageNumber="531">Fig. 7</figureCitation>
). Interspersed scales long, tightly appressed, elytral surface often setose (
<figureCitation id="130D3CD7FFFAF86E41CDF9C0FDDE1815" box="[496,567,1613,1635]" captionStart="FIGURES 73–78" captionStartId="17.[151,264,1859,1883]" captionTargetBox="[174,1415,417,1836]" captionTargetId="figure-138@17.[173,1415,417,1836]" captionTargetPageId="17" captionText="FIGURES 73–78. 73. Pachyrhinus californicus left elytron. 74. Stria. 75. Aedeagus, lateral view. 76. Dorsal view. 77. Ventral view. 78. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119700" httpUri="https://zenodo.org/record/5119700/files/figure.png" pageId="30" pageNumber="531">Fig. 74</figureCitation>
), spermatheca C-shaped (
<figureCitation id="130D3CD7FFFAF86E4313F9C0FC9F1815" box="[814,886,1613,1635]" captionStart="FIGURES 73–78" captionStartId="17.[151,264,1859,1883]" captionTargetBox="[174,1415,417,1836]" captionTargetId="figure-138@17.[173,1415,417,1836]" captionTargetPageId="17" captionText="FIGURES 73–78. 73. Pachyrhinus californicus left elytron. 74. Stria. 75. Aedeagus, lateral view. 76. Dorsal view. 77. Ventral view. 78. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5119700" httpUri="https://zenodo.org/record/5119700/files/figure.png" pageId="30" pageNumber="531">Fig. 78</figureCitation>
).
<collectingRegion id="49F2EEB0FFFAF86E43BAF9C0FA661815" box="[903,1423,1613,1635]" isEnumeration="true" pageId="30" pageNumber="531">California, Oregon, Washington and British Columbia</collectingRegion>
..........................................................................................
<taxonomicName id="4C365BD1FFFAF86E452FF9E6FA741809" ID-CoL="B6MTF" authorityName=": O'Brien &amp; Wibmer and Wibmer" authorityYear="1982" box="[1298,1437,1642,1664]" class="Insecta" family="Curculionidae" genus="Pachyrhinus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="531" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B942FC40FFFAF86E452FF9E6FA741809" box="[1298,1437,1642,1664]" italics="true" pageId="30" pageNumber="531">P. californicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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