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<document id="AC92F48368EA8370A8C698AA803DFA86" ID-DOI="10.11646/zootaxa.3920.1.8" ID-GBIF-Dataset="895f5d96-6bbe-41d0-a95e-661022ee5e82" ID-ISSN="1175-5326" ID-Zenodo-Dep="240661" ID-ZooBank="7C444B23-6A42-4B63-93C3-9927D836C631" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994803421" checkinUser="plazi" docAuthor="Huang, Hao &amp; Chen, Chang-Chin" docDate="2015" docId="039B7E53B27BFFABFF09FCE8CECFC089" docLanguage="en" docName="zt03920p170.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoaesalus" docType="key" docVersion="9" lastPageNumber="170" masterDocId="FFA2062BB27DFFACFF9EFF9ACB53C321" masterDocTitle="Discovery of a second species of Aesalini from Taiwan, with description of the new species of the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae)" masterLastPageNumber="170" masterPageNumber="163" pageNumber="169" updateTime="1698562557755" updateUser="plazi">
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<mods:title id="8E75925EED8B6E1113B8EA6CB28B779A">Discovery of a second species of Aesalini from Taiwan, with description of the new species of the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae)</mods:title>
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<mods:namePart id="E5EFCA9F7531E08235C19E52E4EA20E2">Huang, Hao</mods:namePart>
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<mods:namePart id="7084A777B2406329461407C23BB42EC6">Chen, Chang-Chin</mods:namePart>
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<mods:date id="83D05DA2D0C71394D3966B1E76A195BB">2015</mods:date>
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<mods:number id="3E64BF523BFB232FD252F8B47F81379D">3920</mods:number>
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<mods:classification id="8CB447D24052EF0E8E1927F3E4840DE6">journal article</mods:classification>
<mods:identifier id="3C05475318A642636B602F5CDC5C1403" type="DOI">10.11646/zootaxa.3920.1.8</mods:identifier>
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<treatment id="039B7E53B27BFFABFF09FCE8CECFC089" ID-DOI="http://doi.org/10.5281/zenodo.6106049" ID-GBIF-Taxon="119597181" ID-Zenodo-Dep="6106049" LSID="urn:lsid:plazi:treatment:039B7E53B27BFFABFF09FCE8CECFC089" httpUri="http://treatment.plazi.org/id/039B7E53B27BFFABFF09FCE8CECFC089" lastPageId="7" lastPageNumber="170" pageId="6" pageNumber="169">
<subSubSection id="C3289CCEB27BFFAAFF09FCE8C9E9C719" pageId="6" pageNumber="169" type="nomenclature">
<paragraph id="8B8DCF45B27BFFAAFF09FCE8C9CFC0AD" blockId="6.[151,668,881,908]" box="[151,668,881,908]" pageId="6" pageNumber="169">
<heading id="D0C57829B27BFFAAFF09FCE8C9CFC0AD" bold="true" box="[151,668,881,908]" fontSize="11" level="1" pageId="6" pageNumber="169" reason="1">
<emphasis id="B9461357B27BFFAAFF09FCE8C9CFC0AD" bold="true" box="[151,668,881,908]" pageId="6" pageNumber="169">
Key to males of the
<taxonomicName id="4C32B4C6B27BFFAAFE13FCEBC96DC0AA" ID-CoL="632YD" box="[397,574,881,907]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B9461357B27BFFAAFE13FCEBC96DC0AA" bold="true" box="[397,574,881,907]" italics="true" pageId="6" pageNumber="169">Echinoaesalus</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09FC29C9E9C719" blockId="6.[151,1436,947,1080]" pageId="6" pageNumber="169">
The females of the most of the
<taxonomicName id="4C32B4C6B27BFFAAFE71FC29C9C1C0EA" box="[495,658,947,971]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B9461357B27BFFAAFE71FC29C9C1C0EA" box="[495,658,947,971]" italics="true" pageId="6" pageNumber="169">Echinoaesalus</emphasis>
</taxonomicName>
species have not been carefully studied, thus only the male characters are used in the following key. The characters of hindwing in veins 3dA1 and 3dA2 are probably also good key characters to separate the species groups, but these characters have not been checked for all species described by
<bibRefCitation id="EFA3B2B4B27BFFAAFF09FB85CA68C719" author="Zelenka" box="[151,315,1055,1080]" pageId="6" pageNumber="169" refString="Zelenka, W. (1993) Echinoaesalus gen. n. eine neue Lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau, 63, 235 - 237." type="journal article" year="1993">Zelenka (1993</bibRefCitation>
,
<bibRefCitation id="EFA3B2B4B27BFFAAFED9FBBACAD3C719" author="Zelenka" box="[327,384,1056,1080]" pageId="6" pageNumber="169" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">1994</bibRefCitation>
) and
<bibRefCitation id="EFA3B2B4B27BFFAAFE21FB85C936C719" author="Krikken" box="[447,613,1055,1080]" pageId="6" pageNumber="169" refString="Krikken, J. (1975) Aesalus timidus sp. nov., a remarkable addition to the Indonesian stag-beetle fauna. Entomologische Berichten (Amsterdam), 35, 49 - 51." type="journal article" year="1975">Krikken (1975</bibRefCitation>
,
<bibRefCitation id="EFA3B2B4B27BFFAAFDEFFBBAC9FDC719" author="Krikken" box="[625,686,1056,1080]" pageId="6" pageNumber="169" refString="Krikken, J. (2008) Zelenkaesalus subgen. n. for three Southeast Asian species of Echinoaesalus Zelenka, including E. javanus sp. n. (Coleoptera: Lucanidae: Aesalinae). Koleopterologische Rundschau, 78, 275 - 284." type="journal article" year="2008">2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3289CCEB27BFFABFF09FBFCCECFC089" lastPageId="7" lastPageNumber="170" pageId="6" pageNumber="169" type="key">
<keyStep id="30C6D7E0B27BFFAAFF09FBFCCEC8C7F1" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09FBFCCEC8C7B9" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FBFCCEC8C7B9" pageId="6" pageNumber="169">1 Dorsal surface of the body with only the stick-like bristles and tomenta, but devoid of the scale-like bristles. Cephalic lobe of the 9th abdominal segment of male obsolete................................................................ 8</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09FB05CEC8C7F1" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FB05CEC8C7F1" pageId="6" pageNumber="169">- Dorsal surface of the body more or less with clumps of the scale-like bristles. Cephalic lobe of the 9th abdominal segment of male well marked.................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27BFFAAFF09FB4DCECFC6B7" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09FB4DCECFC604" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FB4DCECFC604" pageId="6" pageNumber="169">
2 Abdominal ventrites without the sublongitudinal sulci. Semicircular punctures along the anterior margin of the abdominal ventrites 3–5 transverse and markedly broader than those of the abdominal ventrites 1–2. Dorsal surface of the body with the clumps of the scale-like bristles and the sparsely distributed stick-like bristles.......................3 (
<taxonomicName id="4C32B4C6B27BFFAAFB6DFA8ACE05C604" ID-CoL="7GQN4" box="[1267,1366,1296,1317]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="hidakai">
<emphasis id="B9461357B27BFFAAFB6DFA8ACE05C604" box="[1267,1366,1296,1317]" italics="true" pageId="6" pageNumber="169">E. hidakai</emphasis>
</taxonomicName>
group)
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09FAB6CECFC6B7" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FAB6CECFC6B7" pageId="6" pageNumber="169">
- Abdominal ventrites 1–3 on either side with the sharply defined sublongitudinal sulcus to receive metatibia and metatarsi. Semicircular punctures along the anterior margin of the abdominal ventrites 3–5 as broad as those of the abdominal ventrites 1–2. Dorsal surface of the body with the clumps of the scale-like bristles but devoid of the stick-like bristles................................................................. 5 (
<taxonomicName id="4C32B4C6B27BFFAAFCFAFA1BC896C6B7" box="[868,965,1409,1430]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="B9461357B27BFFAAFCFAFA1BC896C6B7" box="[868,965,1409,1430]" italics="true" pageId="6" pageNumber="169">E. timidus</emphasis>
</taxonomicName>
group = subgenus
<emphasis id="B9461357B27BFFAAFBE2FA1BCE50C6B7" box="[1148,1283,1409,1430]" italics="true" pageId="6" pageNumber="169">Zelenkaesalus</emphasis>
<bibRefCitation id="EFA3B2B4B27BFFAAFA97FA1BCEC6C6B7" author="Krikken" box="[1289,1429,1409,1430]" pageId="6" pageNumber="169" refString="Krikken, J. (2008) Zelenkaesalus subgen. n. for three Southeast Asian species of Echinoaesalus Zelenka, including E. javanus sp. n. (Coleoptera: Lucanidae: Aesalinae). Koleopterologische Rundschau, 78, 275 - 284." type="journal article" year="2008">Krikken, 2008</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27BFFAAFF09FA07CECFC6CA" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09FA07CEC8C693" blockId="6.[151,1437,1126,2024]" box="[151,1435,1437,1458]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FA07CEC8C693" box="[151,1435,1437,1458]" pageId="6" pageNumber="169">3 Anterior margin of the head projected medially. Scutellum about as long as wide.................................. 4</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09FA20CECFC6CA" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FA20CECFC6CA" pageId="6" pageNumber="169">
- Anterior margin of the head not projected medially. Scutellum markedly longer than wide. Malay Peninsula................................................................................................
<taxonomicName id="4C32B4C6B27BFFAAFB2DFA4CCECFC6CA" ID-CoL="38G54" authority="Zelenka, 1994" authorityName="Zelenka" authorityYear="1994" box="[1203,1436,1494,1515]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B9461357B27BFFAAFB2DFA4CCE5FC6CA" box="[1203,1292,1494,1515]" italics="true" pageId="6" pageNumber="169">E. schuhi</emphasis>
<bibRefCitation id="EFA3B2B4B27BFFAAFA8DFA4CCECFC6CA" author="Zelenka" box="[1299,1436,1494,1515]" pageId="6" pageNumber="169" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">Zelenka, 1994</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27BFFAAFF09FA68CEC8C57D" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09FA68CECFC502" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09FA68CECFC502" pageId="6" pageNumber="169">
4 Intercoxal process of the prosternum longer than wide. Paramere longer than half of the medial lobe of parameres. Malay Peninsula.........................................................
<taxonomicName id="4C32B4C6B27BFFAAFC53F994CFD1C502" ID-CoL="7GQN4" box="[973,1154,1550,1571]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="subSpecies" species="hidakai" subSpecies="hidakai">
<emphasis id="B9461357B27BFFAAFC53F994CFD1C502" box="[973,1154,1550,1571]" italics="true" pageId="6" pageNumber="169">E. hidakai hidakai</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA3B2B4B27BFFAAFB10F994CEC6C502" author="Araya" box="[1166,1429,1550,1571]" pageId="6" pageNumber="169" refString="Araya, K., Kon, M. &amp; Johki, Y. (1993) A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula. Elytra, 21, 93 - 107. [Tokyo]" type="journal article" year="1993">Araya, Kon, &amp; Johki, 1993</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09F9B1CEC8C57D" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F9B1CEC8C57D" pageId="6" pageNumber="169">
- Intercoxal process of the prosternum as long as wide. Paramere less than 1/3 times as long as medial lobe of parameres. Borneo................................................................
<taxonomicName id="4C32B4C6B27BFFAAFC78F9DDCEC8C57D" ID-CoL="6DVSB" authority="Huang &amp; Imura, 2011" authorityName="Huang &amp; Imura" authorityYear="2011" box="[998,1435,1607,1628]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="subSpecies" species="hidakai" subSpecies="borneoensis">
<emphasis id="B9461357B27BFFAAFC78F9DDCF90C57D" box="[998,1219,1607,1628]" italics="true" pageId="6" pageNumber="169">E. hidakai borneoensis</emphasis>
<bibRefCitation id="EFA3B2B4B27BFFAAFB57F9DDCEC8C57D" author="Huang" box="[1225,1435,1607,1628]" pageId="6" pageNumber="169" refString="Huang, H., Imura, Y. &amp; Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107. [Tokyo]" type="journal article" year="2011">Huang &amp; Imura, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27BFFAAFF09F9F9CEC8C5EC" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09F9F9CECFC5B4" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F9F9CECFC5B4" pageId="6" pageNumber="169">
5 Mentum with short (comma-shaped) pit-like groove on either side of mentum but without transversal groove in front. Dorsal line of the body in lateral view semicircular. Sumatra, Malay Peninsula,
<collectingCountry id="F3258FD5B27BFFAAFC08F91AC8BDC5B4" box="[918,1006,1664,1685]" name="Thailand" pageId="6" pageNumber="169">Thailand</collectingCountry>
, Borneo.......
<taxonomicName id="4C32B4C6B27BFFAAFB04F91ACECFC5B4" authority="Krikken, 1975" authorityName="Krikken" authorityYear="1975" box="[1178,1436,1664,1685]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="B9461357B27BFFAAFB04F91ACFA8C5B4" box="[1178,1275,1664,1685]" italics="true" pageId="6" pageNumber="169">E. timidus</emphasis>
(
<bibRefCitation id="EFA3B2B4B27BFFAAFA96F91ACEC6C5B4" author="Krikken" box="[1288,1429,1664,1685]" pageId="6" pageNumber="169" refString="Krikken, J. (1975) Aesalus timidus sp. nov., a remarkable addition to the Indonesian stag-beetle fauna. Entomologische Berichten (Amsterdam), 35, 49 - 51." type="journal article" year="1975">Krikken, 1975</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09F906CEC8C5EC" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F906CEC8C5EC" pageId="6" pageNumber="169">- Mentum with complete (variably deep) arcuate groove, transversely. Dorsal line of the body in lateral view produced at the summit.............................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27BFFAAFF09F94ECECFC47B" pageId="6" pageNumber="169">
<paragraph id="8B8DCF45B27BFFAAFF09F94ECECFC403" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F94ECECFC403" pageId="6" pageNumber="169">
6 Pronotum and elytra with yellowish brown bristle clumps but lacking black bristle clumps. Metasternum and abdominal ventrite shiny. Protibia slender. Anterolateral corner of metasternum shallowly impressed along mesocoxae. Anterolateral edge of mentum broadly rounded. Java......................................................
<taxonomicName id="4C32B4C6B27BFFAAFB38F897CECFC403" ID-CoL="38G4R" authority="Krikken, 2008" authorityName="Krikken" authorityYear="2008" box="[1190,1436,1805,1826]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B9461357B27BFFAAFB38F897CE5FC403" box="[1190,1292,1805,1826]" italics="true" pageId="6" pageNumber="169">E. javanus</emphasis>
<bibRefCitation id="EFA3B2B4B27BFFAAFA8FF897CECFC403" author="Krikken" box="[1297,1436,1805,1826]" pageId="6" pageNumber="169" refString="Krikken, J. (2008) Zelenkaesalus subgen. n. for three Southeast Asian species of Echinoaesalus Zelenka, including E. javanus sp. n. (Coleoptera: Lucanidae: Aesalinae). Koleopterologische Rundschau, 78, 275 - 284." type="journal article" year="2008">Krikken, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09F8B3CECFC47B" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F8B3CECFC47B" pageId="6" pageNumber="169">- Pronotum and elytra with black bristle clumps. Metasternum and abdominal ventrite matt. Protibia robust. Anterolateral corner of metasternum with deep furrow along mesocoxae. Anterolateral edge of mentum distinctly produced................ 7</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B8DCF45B27BFFAAFF09F8F8CEC8C48E" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F8F8CEC8C48E" pageId="6" pageNumber="169">
<keyStep id="30C6D7E0B27BFFAAFF09F8F8CEC8C48E" pageId="6" pageNumber="169">
7 Groove on mentum immediately behind anterior margin of mentum, interrupted in width at lateral sides, with the lateral ends broader and deeper. Parameres at most 1/4 times as long as the sclerotized part of medial lobe of parameres. West Java................................................................................
<taxonomicName id="4C32B4C6B27BFFAAFBFDF800CEC8C48E" ID-CoL="38G4Y" authority="Huang &amp; Wu, 2011" authorityName="Huang &amp; Wu" authorityYear="2011" box="[1123,1435,1946,1967]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="169" phylum="Arthropoda" rank="species" species="gedeensis">
<emphasis id="B9461357B27BFFAAFBFDF800CF89C48E" box="[1123,1242,1946,1967]" italics="true" pageId="6" pageNumber="169">E. gedeensis</emphasis>
<bibRefCitation id="EFA3B2B4B27BFFAAFB7EF800CEC8C48E" author="Huang" box="[1248,1435,1946,1967]" pageId="6" pageNumber="169" refString="Huang, H., Imura, Y. &amp; Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107. [Tokyo]" type="journal article" year="2011">Huang &amp; Wu, 2011</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27BFFAAFF09F82CCEC4C4C9" blockId="6.[151,1437,1126,2024]" pageId="6" pageNumber="169">
<keyLead id="30C36C70B27BFFAAFF09F82CCEC4C4C9" pageId="6" pageNumber="169">- Groove on mentum remoter from anterior margin of mentum, not interrupted in width at lateral sides, with the lateral ends not broader. Parameres about half as long as the sclerotized part of medial lobe of parameres, Borneo.......................</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF7DFF0DCECFC3C4" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
..............................................................................
<taxonomicName id="4C32B4C6B27AFFABFB16FF0DCECFC38D" ID-CoL="38G53" authority="Zelenka, 1994" authorityName="Zelenka" authorityYear="1994" box="[1160,1436,151,172]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="B9461357B27AFFABFB16FF0DCE5FC38D" box="[1160,1292,151,172]" italics="true" pageId="7" pageNumber="170">E. sabahensis</emphasis>
<bibRefCitation id="EFA3B2B4B27AFFABFA8DFF0DCECFC38D" author="Zelenka" box="[1299,1436,151,172]" pageId="7" pageNumber="170" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">Zelenka, 1994</bibRefCitation>
</taxonomicName>
<keyLead id="30C36C70B27AFFABFF09FF2ECECFC3C4" pageId="7" pageNumber="170">
<keyStep id="30C6D7E0B27AFFABFF09FF2ECECFC3C4" pageId="7" pageNumber="170">
8 All punctures on the elytra placed serially in longitudinal lines. Intercoxal process of the prosternum with anterior margin produced medially, Sulawesi.............................................................
<taxonomicName id="4C32B4C6B27AFFABFB28FF4ACECFC3C4" ID-CoL="38G4Z" authority="Zelenka, 1993" authorityName="Zelenka" authorityYear="1993" box="[1206,1436,208,229]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B9461357B27AFFABFB28FF4ACE5EC3C4" box="[1206,1293,208,229]" italics="true" pageId="7" pageNumber="170">E. jaechi</emphasis>
<bibRefCitation id="EFA3B2B4B27AFFABFA8DFF4ACECFC3C4" author="Zelenka" box="[1299,1436,208,229]" pageId="7" pageNumber="170" refString="Zelenka, W. (1993) Echinoaesalus gen. n. eine neue Lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau, 63, 235 - 237." type="journal article" year="1993">Zelenka, 1993</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FF76CECFC23C" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FF76CECFC23C" pageId="7" pageNumber="170">
- Some punctures on the elytra placed irregularly, not serially in longitudinal lines. Intercoxal process of the prosternum with anterior margin rather flat............................................................... 9 (
<taxonomicName id="4C32B4C6B27AFFABFB6AFE92CE05C23C" ID-CoL="38G52" box="[1268,1366,264,285]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="matsuii">
<emphasis id="B9461357B27AFFABFB6AFE92CE05C23C" box="[1268,1366,264,285]" italics="true" pageId="7" pageNumber="170">E. matsuii</emphasis>
</taxonomicName>
group)
</keyLead>
</paragraph>
<keyStep id="30C6D7E0B27AFFABFF09FEBFCECFC253" pageId="7" pageNumber="170">
<paragraph id="8B8DCF45B27AFFABFF09FEBFCECFC277" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FEBFCECFC277" pageId="7" pageNumber="170">
9 Outline of elytra in lateral view unevenly rounded or obtusely angled, with anterior half more arched than posterior half. Sumatra.........................................................................
<taxonomicName id="4C32B4C6B27AFFABFB38FEDBCECFC277" ID-CoL="38G4T" authority="Zelenka, 1993" authorityName="Zelenka" authorityYear="1993" box="[1190,1436,321,342]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="barriesi">
<emphasis id="B9461357B27AFFABFB38FEDBCE5FC277" box="[1190,1292,321,342]" italics="true" pageId="7" pageNumber="170">E. barriesi</emphasis>
<bibRefCitation id="EFA3B2B4B27AFFABFA8DFEDBCECFC277" author="Zelenka" box="[1299,1436,321,342]" pageId="7" pageNumber="170" refString="Zelenka, W. (1993) Echinoaesalus gen. n. eine neue Lucaniden-Gattung aus Sudostasien (Coleoptera, Lucanidae). Koleopterologische Rundschau, 63, 235 - 237." type="journal article" year="1993">Zelenka, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FEC7CECFC253" blockId="7.[151,1437,151,936]" box="[151,1436,349,370]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FEC7CECFC253" box="[151,1436,349,370]" pageId="7" pageNumber="170">- Outline of elytra in lateral view rather evenly rounded, not more arched at anterior half than at posterior half........... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27AFFABFF09FEE0CEC8C13D" pageId="7" pageNumber="170">
<paragraph id="8B8DCF45B27AFFABFF09FEE0CECFC2E6" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FEE0CECFC2E6" pageId="7" pageNumber="170">
10 Pair of cavities on the metasternum just behind the mesocoxae oblique. Sulcus on mentum indistinct. Parameres very long, almost reaching the distal end of the pigmented part of medial lobe of parameres (not counting the distal sclerite of medial lobe of parameres). Malay Peninsula.......................................................
<taxonomicName id="4C32B4C6B27AFFABFB30FE28CECFC2E6" ID-CoL="38G52" authority="Araya, 1993" authorityName="Araya" authorityYear="1993" box="[1198,1436,434,455]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="matsuii">
<emphasis id="B9461357B27AFFABFB30FE28CE43C2E6" box="[1198,1296,434,455]" italics="true" pageId="7" pageNumber="170">E. matsuii</emphasis>
(Araya, 1993)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FE54CEC8C13D" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FE54CEC8C13D" pageId="7" pageNumber="170">- Pair of cavities on the metasternum just behind the mesocoxae almost longitudinal. Sulcus on mentum clearly marked. Parameres short and remote from the distal end of the pigmented part of medial lobe of parameres (not counting the distal sclerite of medial lobe of parameres).................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27AFFABFF09FDB9CEC8C1AC" pageId="7" pageNumber="170">
<paragraph id="8B8DCF45B27AFFABFF09FDB9CEC8C175" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FDB9CEC8C175" pageId="7" pageNumber="170">11 Parameres markedly shorter than the width of the medial lobe of parameres in dorsal or ventral view. Medial lobe of parameres wider and globe-like in basal portion and contracted in distal portion........................................... 12</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FDC6CEC8C1AC" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FDC6CEC8C1AC" pageId="7" pageNumber="170">- Parameres markedly longer than the width of the medial lobe of parameres in dorsal or ventral view. Medial lobe of parameres nearly even in width in dorsal view..................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27AFFABFF09FD0ECECFC1DF" pageId="7" pageNumber="170">
<paragraph id="8B8DCF45B27AFFABFF09FD0EC8FBC1E4" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FD0EC8FBC1E4" pageId="7" pageNumber="170">
12 Elytra longer than wide. 9th abdominal segment of male more elongate. Well pigmented part of medial lobe of parameres longer.
<collectingCountry id="F3258FD5B27AFFABFE94FD2ACA0CC1E4" box="[266,351,688,709]" name="Thailand" pageId="7" pageNumber="170">Thailand</collectingCountry>
...
<taxonomicName id="4C32B4C6B27AFFABFEE3FD2ACAB0C1E4" ID-CoL="38G4X" box="[381,483,688,709]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="dharma">
<emphasis id="B9461357B27AFFABFEE3FD2ACAB0C1E4" box="[381,483,688,709]" italics="true" pageId="7" pageNumber="170">E. dharma</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA3B2B4B27AFFABFE71FD2AC8F2C1E4" author="Araya" box="[495,929,688,709]" pageId="7" pageNumber="170" refString="Araya, K., Kon, M., Matsui, M., Nabhitabhata, J. &amp; Panha, S. (1995) A new bristly Aesalus (Coleoptera, Lucanidae) from Peninsular Thailand. Giornale Italiano di Entomologia, 7, 73 - 77." type="journal article" year="1995">Araya, Matsui, Nabhitabhata, &amp; Panha, 1995</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FD57CECFC1DF" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FD57CECFC1DF" pageId="7" pageNumber="170">
- Elytra nearly as long as wide. 9th abdominal segment of male stouter. Well pigmented part of medial lobe of parameres shorter. Malay Peninsula, Borneo.............................................................
<taxonomicName id="4C32B4C6B27AFFABFB20FD73CECFC1DF" ID-CoL="38G55" authority="Araya, 1993" authorityName="Araya" authorityYear="1993" box="[1214,1436,745,766]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="yongi">
<emphasis id="B9461357B27AFFABFB20FD73CE43C1DF" box="[1214,1296,745,766]" italics="true" pageId="7" pageNumber="170">E. yongi</emphasis>
(Araya, 1993)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C6D7E0B27AFFABFF09FC9FCECFC089" pageId="7" pageNumber="170">
<paragraph id="8B8DCF45B27AFFABFF09FC9FCEC8C072" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FC9FCEC8C072" pageId="7" pageNumber="170">
13 Size larger. Elytra nearly as long as wide. Insertion of the bristle placed centrally in the puncture on dorsal surface of the body. Bristles on elytra not placed serially. Medial lobe of parameres with most part of dorsal surface less pigmented, and with distal sclerite continuous. Borneo.....................................................
<taxonomicName id="4C32B4C6B27AFFABFBF4FCA4CEC8C072" ID-CoL="38G4S" authority="Huang &amp; Imura, 2011" authorityName="Huang &amp; Imura" authorityYear="2011" box="[1130,1435,830,851]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="arayai">
<emphasis id="B9461357B27AFFABFBF4FCA4CF91C072" box="[1130,1218,830,851]" italics="true" pageId="7" pageNumber="170">E. arayai</emphasis>
<bibRefCitation id="EFA3B2B4B27AFFABFB57FCA4CEC8C072" author="Huang" box="[1225,1435,830,851]" pageId="7" pageNumber="170" refString="Huang, H., Imura, Y. &amp; Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107. [Tokyo]" type="journal article" year="2011">Huang &amp; Imura, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8DCF45B27AFFABFF09FCC0CECFC089" blockId="7.[151,1437,151,936]" pageId="7" pageNumber="170">
<keyLead id="30C36C70B27AFFABFF09FCC0CECFC089" pageId="7" pageNumber="170">
- Size smaller. Elytra longer than wide. Insertion of the bristle placed anteriorly in the puncture on dorsal surface of the body. Bristles on elytra placed serially. Medial lobe of parameres with only distal half of dorsal surface less pigmented, and with distal sclerite interrupted.
<collectingCountry id="F3258FD5B27AFFABFE21FC09C956C089" box="[447,517,915,936]" name="Taiwan" pageId="7" pageNumber="170">Taiwan</collectingCountry>
.......................................................
<taxonomicName id="4C32B4C6B27AFFABFB37FC09CECFC089" ID-CoL="38G4W" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" box="[1193,1436,915,936]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="chungi">
<emphasis id="B9461357B27AFFABFB37FC09CE57C089" box="[1193,1284,915,936]" italics="true" pageId="7" pageNumber="170">E. chungi</emphasis>
Huang &amp; Chen
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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