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<document id="861BF944E24133A31A94E376D78329CB" ID-CLB-Dataset="8114" ID-DOI="10.37520/aemnp.2020.24" ID-GBIF-Dataset="9cf87cc9-79ad-4a31-83ca-b635d495a0b9" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489252" ID-ZooBank="AC387BAF-E7A8-40B2-9486-E5642074587D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612221266920" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2020" docId="039A87F1FFC4FFB1FF09B2F5FBDEF9FF" docLanguage="en" docName="ActaEntMusNatPrag.60.2.397-409.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Gnathoncus nannetensis" docType="treatment" docVersion="8" lastPageNumber="403" masterDocId="FFA3FF89FFC2FFB7FFEBB448FFB2FFA3" masterDocTitle="A review of Gnathoncus of Southeast Asia (Coleoptera: Histeridae: Saprininae)" masterLastPageNumber="409" masterPageNumber="397" pageNumber="403" updateTime="1698926172273" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="3A76517E5A480E1293BC26129B4B786C">A review of Gnathoncus of Southeast Asia (Coleoptera: Histeridae: Saprininae)</mods:title>
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<treatment id="039A87F1FFC4FFB1FF09B2F5FBDEF9FF" ID-DOI="http://doi.org/10.5281/zenodo.4549635" ID-GBIF-Taxon="177425808" ID-Zenodo-Dep="4549635" LSID="urn:lsid:plazi:treatment:039A87F1FFC4FFB1FF09B2F5FBDEF9FF" httpUri="http://treatment.plazi.org/id/039A87F1FFC4FFB1FF09B2F5FBDEF9FF" lastPageNumber="403" pageId="6" pageNumber="403">
<subSubSection id="C329656CFFC4FFB1FF09B2F5FD1FF976" box="[226,685,1725,1749]" pageId="6" pageNumber="403" type="nomenclature">
<paragraph id="8B8C36E7FFC4FFB1FF09B2F5FD1FF976" blockId="6.[226,685,1725,1749]" box="[226,685,1725,1749]" pageId="6" pageNumber="403">
<heading id="D0C4818BFFC4FFB1FF09B2F5FD1FF976" box="[226,685,1725,1749]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="403" reason="3">
<taxonomicName id="4C334D64FFC4FFB1FF09B2F5FD1FF976" ID-CoL="3GLRX" authority="(Marseul, 1862)" baseAuthorityName="Marseul" baseAuthorityYear="1862" box="[226,685,1725,1749]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="nannetensis">
<emphasis id="B947EAF5FFC4FFB1FF09B2F5FD1FF976" bold="true" box="[226,685,1725,1749]" pageId="6" pageNumber="403">
<emphasis id="B947EAF5FFC4FFB1FF09B2F5FE41F976" bold="true" box="[226,499,1725,1749]" italics="true" pageId="6" pageNumber="403">Gnathoncus nannetensis</emphasis>
(Marseul, 1862)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FE9DB294FDABF94C" box="[374,537,1756,1775]" pageId="6" pageNumber="403" type="description">
<paragraph id="8B8C36E7FFC4FFB1FE9DB294FDABF94C" blockId="6.[151,765,1756,2076]" box="[374,537,1756,1775]" pageId="6" pageNumber="403">
(
<figureCitation id="13082A62FFC4FFB1FE97B294FE01F94C" box="[380,435,1756,1775]" captionStart="Figs 15" captionStartId="2.[151,188,2037,2056]" captionTargetBox="[156,1415,199,1995]" captionTargetId="figure-13@2.[151,1435,189,2006]" captionTargetPageId="2" captionText="Figs 15. Dorsal (14) and ventral (5) habitus of SE Asian Gnathoncus species. 1 G. brevisternus Lewis, 1907; 2 G. nannetensis (Marseul, 1862); 3 G. rotundatus (Kugelann, 1792); 45 G. sechuanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4489256" httpUri="https://zenodo.org/record/4489256/files/figure.png" pageId="6" pageNumber="403">Figs 2</figureCitation>
,
<figureCitation id="13082A62FFC4FFB1FE57B294FE46F94C" box="[444,500,1756,1775]" captionStart="Figs 1651" captionStartId="5.[151,188,1822,1841]" captionTargetBox="[154,1434,196,1684]" captionTargetId="figure-13@5.[148,1439,189,1684]" captionTargetPageId="5" captionText="Figs 1651. Male genitalia of SE Asian Gnathoncus species. 1624 G. brevisternus Lewis, 1907: 1618 VIII sternite and tergite (16 ventral view; 17 dorsal view; 18 lateral view). 1920 IX and X tergites (19 dorsal view; 20 lateral view). 2122 IX sternite (spiculum gastrale) (21 ventral view; 22 lateral view). 2324 aedeagus (23 dorsal view; 24 lateral view). 2534 G. nannetensis (Marseul, 1862) (after ÔΗΑRΑ 1994). 2527 VIII sternite and tergite (25 apical half in ventral view; 26 dorsal view; 27 lateral view).2830 aedeagus (28 apex, apical view; 29 dorsal view; 30 lateral view). 3132 IX and X tergite (31 dorsal view; 32 lateral view). 3334 IX sternite (spiculum gastrale) (33 ventral view; 34 lateral view). 3542 G. rotundatus (Kugelann, 1792) (after ÔΗΑRΑ 1994). 3537 VIII sternite and tergite (35 apical half in ventral view; 36 dorsal view; 37 lateral view). 3839 IX and X tergite (38 dorsal view; 39 lateral view). 40 IX sternite (spiculum gastrale), ventral view. 4142 aedeagus (41 dorsal view; 42 lateral view). 4351 G. sechuanus sp. nov. 4345 VIII sternite and tergite (43 ventral view; 44 dorsal view; 45 lateral view). 4647 IX and X tergites (46 dorsal view; 47 lateral view). 4849 IX sternite (spiculum gastrale) (48 ventral view; 49 lateral view). 5051 aedeagus (51 dorsal view; 51 lateral view)" figureDoi="http://doi.org/10.5281/zenodo.4489268" httpUri="https://zenodo.org/record/4489268/files/figure.png" pageId="6" pageNumber="403">2534</figureCitation>
,
<figureCitation id="13082A62FFC4FFB1FE17B294FDA0F94C" box="[508,530,1756,1775]" captionStart="Figs 5253" captionStartId="6.[828,865,774,793]" captionTargetBox="[830,1434,192,745]" captionTargetId="figure-794@6.[825,1439,186,756]" captionTargetPageId="6" captionText="Figs 5253. Pygidial microsculptures. 52 Gnathoncus nannetensis (Marseul, 1862); 53 G. rotundatus (Kugelann, 1792) (from SτοΚ- ΜΑΝΝ 1957)." figureDoi="http://doi.org/10.5281/zenodo.4489272" httpUri="https://zenodo.org/record/4489272/files/figure.png" pageId="6" pageNumber="403">52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FF7CB2B6FEF3F7BF" pageId="6" pageNumber="403" type="reference_group">
<paragraph id="8B8C36E7FFC4FFB1FF7CB2B6FD08F8B2" blockId="6.[151,765,1756,2076]" box="[151,698,1790,1809]" pageId="6" pageNumber="403">
<taxonomicName id="4C334D64FFC4FFB1FF7CB2B6FE4BF8B2" ID-CoL="9N39H" authority="Marseul, 1862: 499" authorityName="Marseul" authorityPageNumber="499" authorityYear="1862" box="[151,505,1790,1809]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="nannetensis">
<emphasis id="B947EAF5FFC4FFB1FF7CB2B6FEFFF8B2" box="[151,333,1790,1809]" italics="true" pageId="6" pageNumber="403">Saprinus nannetensis</emphasis>
Marseul, 1862: 499
</taxonomicName>
(original description).
</paragraph>
<paragraph id="8B8C36E7FFC4FFB1FF7CB351FE43F8E4" blockId="6.[151,765,1756,2076]" pageId="6" pageNumber="403">
<treatmentCitationGroup id="AB2311C9FFC4FFB1FF7CB351FE43F8E4" pageId="6" pageNumber="403">
<treatmentCitation id="0A9210F6FFC4FFB1FF7CB351FE4EF88F" author="REITTER E." box="[151,508,1817,1836]" page="307" pageId="6" pageNumber="403" year="1896">
<taxonomicName id="4C334D64FFC4FFB1FF7CB351FE4EF88F" ID-CoL="9MPBS" authority="Reitter, 1896: 307" authorityName="Reitter" authorityPageNumber="307" authorityYear="1896" box="[151,508,1817,1836]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="urganensis">
<emphasis id="B947EAF5FFC4FFB1FF7CB351FED2F88F" box="[151,352,1817,1836]" italics="true" pageId="6" pageNumber="403">Gnathoncus urganensis</emphasis>
<bibRefCitation id="EFA24B16FFC4FFB1FE8FB351FE4EF88F" author="REITTER E." box="[356,508,1817,1836]" pageId="6" pageNumber="403" pagination="306 - 308" refId="ref9599" refString="REITTER E. 1896: Uebersicht der mir bekannten Gnathoncus Arten der palearktischen Fauna. Entomologische Nachrichten 22: 306 - 308." type="journal article" year="1896">Reitter, 1896: 307</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description) synonymized by RΕιΗΑRƉτ (1941): 162.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8C36E7FFC4FFB1FF7CB306FE45F8DF" blockId="6.[151,765,1756,2076]" pageId="6" pageNumber="403">
<taxonomicName id="4C334D64FFC4FFB1FF7CB306FD96F8C2" ID-CoL="9TLM2" authority="Ganglbauer, 1899: 380" authorityName="Ganglbauer" authorityPageNumber="380" authorityYear="1899" box="[151,548,1870,1889]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B947EAF5FFC4FFB1FF7CB306FEE0F8C2" box="[151,338,1870,1889]" italics="true" pageId="6" pageNumber="403">Gnathoncus suturalis</emphasis>
Ganglbauer, 1899: 380
</taxonomicName>
(original description) synonymized by MΑƵUR (1984): 104.
</paragraph>
<paragraph id="8B8C36E7FFC4FFB1FF7CB3CCFEF3F7BF" blockId="6.[151,765,1756,2076]" pageId="6" pageNumber="403">
<taxonomicName id="4C334D64FFC4FFB1FF7CB3CCFED5F834" ID-CoL="3GLRX" baseAuthorityName="Marseul" baseAuthorityYear="1862" box="[151,359,1924,1943]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="nannetensis">
<emphasis id="B947EAF5FFC4FFB1FF7CB3CCFED5F834" box="[151,359,1924,1943]" italics="true" pageId="6" pageNumber="403">Gnathoncus nannetensis</emphasis>
</taxonomicName>
: RΕιττΕR (1896): 308 (keyed); KRΥƵΗΑΝοඏඌΚιΙ +|RΕιΗΑRƉτ (1976):114, 117 (keyed, redescription); MΑƵUR (1984): 105 (catalogue); ÔΗΑRΑ (1994): 215, figs 130, 131, 132, 135 (key, redescription); MΑƵUR (1997): 214 (catalogue); MΑƵUR (2004): 92 (catalogue); MΑƵUR (2011): 176 (catalogue); LΑΚΝΕR et al. (2015): 114 (catalogue).
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FCD7B7EDFBB6FBDF" pageId="6" pageNumber="403" type="description">
<paragraph id="8B8C36E7FFC4FFB1FCD7B7EDFBB6FBDF" blockId="6.[828,1437,933,1628]" pageId="6" pageNumber="403">
<emphasis id="B947EAF5FFC4FFB1FCD7B7EDFCCAFC1F" bold="true" box="[828,888,933,956]" pageId="6" pageNumber="403">Note.</emphasis>
This species was very thoroughly redescribed and figured in detail by ÔΗΑRΑ (1994: 215), and the reader is referred to the redescription and detailed figures there. For the sake of better recognition of this species among others distributed in Southeast Asia, the habitus (
<figureCitation id="13082A62FFC4FFB1FAEAB06DFAF3FB9F" box="[1281,1345,1061,1084]" captionStart="Figs 15" captionStartId="2.[151,188,2037,2056]" captionTargetBox="[156,1415,199,1995]" captionTargetId="figure-13@2.[151,1435,189,2006]" captionTargetPageId="2" captionText="Figs 15. Dorsal (14) and ventral (5) habitus of SE Asian Gnathoncus species. 1 G. brevisternus Lewis, 1907; 2 G. nannetensis (Marseul, 1862); 3 G. rotundatus (Kugelann, 1792); 45 G. sechuanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4489256" httpUri="https://zenodo.org/record/4489256/files/figure.png" pageId="6" pageNumber="403">Fig. 2</figureCitation>
) as well as pygidium (
<figureCitation id="13082A62FFC4FFB1FC27B00DFBABFBFF" box="[972,1049,1093,1116]" captionStart="Figs 5253" captionStartId="6.[828,865,774,793]" captionTargetBox="[830,1434,192,745]" captionTargetId="figure-794@6.[825,1439,186,756]" captionTargetPageId="6" captionText="Figs 5253. Pygidial microsculptures. 52 Gnathoncus nannetensis (Marseul, 1862); 53 G. rotundatus (Kugelann, 1792) (from SτοΚ- ΜΑΝΝ 1957)." figureDoi="http://doi.org/10.5281/zenodo.4489272" httpUri="https://zenodo.org/record/4489272/files/figure.png" pageId="6" pageNumber="403">Fig. 52</figureCitation>
) and male genitalia (
<figureCitation id="13082A62FFC4FFB1FB1EB00DFADCFBFF" box="[1269,1390,1093,1116]" captionStart="Figs 1651" captionStartId="5.[151,188,1822,1841]" captionTargetBox="[154,1434,196,1684]" captionTargetId="figure-13@5.[148,1439,189,1684]" captionTargetPageId="5" captionText="Figs 1651. Male genitalia of SE Asian Gnathoncus species. 1624 G. brevisternus Lewis, 1907: 1618 VIII sternite and tergite (16 ventral view; 17 dorsal view; 18 lateral view). 1920 IX and X tergites (19 dorsal view; 20 lateral view). 2122 IX sternite (spiculum gastrale) (21 ventral view; 22 lateral view). 2324 aedeagus (23 dorsal view; 24 lateral view). 2534 G. nannetensis (Marseul, 1862) (after ÔΗΑRΑ 1994). 2527 VIII sternite and tergite (25 apical half in ventral view; 26 dorsal view; 27 lateral view).2830 aedeagus (28 apex, apical view; 29 dorsal view; 30 lateral view). 3132 IX and X tergite (31 dorsal view; 32 lateral view). 3334 IX sternite (spiculum gastrale) (33 ventral view; 34 lateral view). 3542 G. rotundatus (Kugelann, 1792) (after ÔΗΑRΑ 1994). 3537 VIII sternite and tergite (35 apical half in ventral view; 36 dorsal view; 37 lateral view). 3839 IX and X tergite (38 dorsal view; 39 lateral view). 40 IX sternite (spiculum gastrale), ventral view. 4142 aedeagus (41 dorsal view; 42 lateral view). 4351 G. sechuanus sp. nov. 4345 VIII sternite and tergite (43 ventral view; 44 dorsal view; 45 lateral view). 4647 IX and X tergites (46 dorsal view; 47 lateral view). 4849 IX sternite (spiculum gastrale) (48 ventral view; 49 lateral view). 5051 aedeagus (51 dorsal view; 51 lateral view)" figureDoi="http://doi.org/10.5281/zenodo.4489268" httpUri="https://zenodo.org/record/4489268/files/figure.png" pageId="6" pageNumber="403">Figs 2534</figureCitation>
) are reproduced herein.
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FCD7B0CDFAFAFADF" pageId="6" pageNumber="403" type="diagnosis">
<paragraph id="8B8C36E7FFC4FFB1FCD7B0CDFAFAFADF" blockId="6.[828,1437,933,1628]" pageId="6" pageNumber="403">
<emphasis id="B947EAF5FFC4FFB1FCD7B0CDFB83FB3F" bold="true" box="[828,1073,1157,1180]" pageId="6" pageNumber="403">Differential diagnosis.</emphasis>
In the rather shiny elytral disc, this species resembles both
<taxonomicName id="4C334D64FFC4FFB1FBD9B0EDFB76FB1F" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[1074,1220,1189,1212]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B947EAF5FFC4FFB1FBD9B0EDFB76FB1F" box="[1074,1220,1189,1212]" italics="true" pageId="6" pageNumber="403">G. rotundatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C334D64FFC4FFB1FB1FB0EDFA24FB1F" authorityName="Lewis" authorityYear="1907" box="[1268,1430,1189,1212]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="brevisternus">
<emphasis id="B947EAF5FFC4FFB1FB1FB0EDFA24FB1F" box="[1268,1430,1189,1212]" italics="true" pageId="6" pageNumber="403">G. brevisternus</emphasis>
</taxonomicName>
, but differs from them in round pygidial punctures (
<figureCitation id="13082A62FFC4FFB1FAA2B08DFA27FB7F" box="[1353,1429,1221,1244]" captionStart="Figs 5253" captionStartId="6.[828,865,774,793]" captionTargetBox="[830,1434,192,745]" captionTargetId="figure-794@6.[825,1439,186,756]" captionTargetPageId="6" captionText="Figs 5253. Pygidial microsculptures. 52 Gnathoncus nannetensis (Marseul, 1862); 53 G. rotundatus (Kugelann, 1792) (from SτοΚ- ΜΑΝΝ 1957)." figureDoi="http://doi.org/10.5281/zenodo.4489272" httpUri="https://zenodo.org/record/4489272/files/figure.png" pageId="6" pageNumber="403">Fig. 52</figureCitation>
) and dark-brown legs (
<figureCitation id="13082A62FFC4FFB1FBCDB0ADFBDAFB5F" box="[1062,1128,1253,1276]" captionStart="Figs 15" captionStartId="2.[151,188,2037,2056]" captionTargetBox="[156,1415,199,1995]" captionTargetId="figure-13@2.[151,1435,189,2006]" captionTargetPageId="2" captionText="Figs 15. Dorsal (14) and ventral (5) habitus of SE Asian Gnathoncus species. 1 G. brevisternus Lewis, 1907; 2 G. nannetensis (Marseul, 1862); 3 G. rotundatus (Kugelann, 1792); 45 G. sechuanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4489256" httpUri="https://zenodo.org/record/4489256/files/figure.png" pageId="6" pageNumber="403">Fig. 2</figureCitation>
). From
<taxonomicName id="4C334D64FFC4FFB1FB57B0ADFA34FB5F" authorityName="Bickhardt" authorityYear="1920" box="[1212,1414,1253,1276]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="semimarginatus">
<emphasis id="B947EAF5FFC4FFB1FB57B0ADFA34FB5F" box="[1212,1414,1253,1276]" italics="true" pageId="6" pageNumber="403">G. semimarginatus</emphasis>
</taxonomicName>
it differs in the complete marginal pronotal stria; from
<taxonomicName id="4C334D64FFC4FFB1FA69B14DFC08FA9F" authorityName="Kryzhanovskij" authorityYear="1972" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B947EAF5FFC4FFB1FA69B14DFC08FA9F" italics="true" pageId="6" pageNumber="403">G. vietnamicus</emphasis>
</taxonomicName>
in the absence of longer apical fragment of the sutural elytral stria, and from
<taxonomicName id="4C334D64FFC4FFB1FB98B10DFAB0FAFF" authorityName="Lackner" authorityYear="2020" box="[1139,1282,1349,1372]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="sechuanus">
<emphasis id="B947EAF5FFC4FFB1FB98B10DFAB0FAFF" box="[1139,1282,1349,1372]" italics="true" pageId="6" pageNumber="403">G. sechuanus</emphasis>
</taxonomicName>
in the absence of strong alutaceous microsculpture of the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FCD7B1CDFAB4FA5F" pageId="6" pageNumber="403" type="biology_ecology">
<paragraph id="8B8C36E7FFC4FFB1FCD7B1CDFAB4FA5F" blockId="6.[828,1437,933,1628]" pageId="6" pageNumber="403">
<emphasis id="B947EAF5FFC4FFB1FCD7B1CDFC25FA3F" bold="true" box="[828,919,1413,1436]" pageId="6" pageNumber="403">Biology.</emphasis>
This species is predominantly found in bird nests, but has also been recorded in burrows of small mammals, on (mostly bird) carrion, decaying fish, animal droppings, etc. (KRΥƵΗΑΝοඏඌΚιΙ +| RΕιΗΑRƉτ 1976).
</paragraph>
</subSubSection>
<subSubSection id="C329656CFFC4FFB1FCD7B24DFBDEF9FF" pageId="6" pageNumber="403" type="distribution">
<paragraph id="8B8C36E7FFC4FFB1FCD7B24DFBDEF9FF" blockId="6.[828,1437,933,1628]" pageId="6" pageNumber="403">
<emphasis id="B947EAF5FFC4FFB1FCD7B24DFC7CF9BF" bold="true" box="[828,974,1541,1564]" pageId="6" pageNumber="403">Distribution.</emphasis>
Widespread in almost the entire Palaearctic Region; from Southeast Asia recorded from
<collectingCountry id="F3247677FFC4FFB1FAE9B26DFAF7F99F" box="[1282,1349,1573,1596]" name="China" pageId="6" pageNumber="403">China</collectingCountry>
:
<collectingRegion id="49F7F805FFC4FFB1FAA6B26DFCC3F9FF" country="China" name="Guangdong" pageId="6" pageNumber="403">Guangdong</collectingRegion>
(LΑΚΝΕR et al. 2015).
</paragraph>
</subSubSection>
</treatment>
</document>