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<document ID-DOI="10.5281/zenodo.184817" ID-GBIF-Dataset="e5c56f88-2105-49a1-99e0-39eb486f9c64" ID-ISSN="1175-5326" ID-Zenodo-Dep="184817" checkinTime="1460017920730" checkinUser="plazi" docAuthor="Shattuck, Steven O." docDate="2008" docId="038FD126FF8AC84FFF24F92D7DFDFAE7" docLanguage="en" docName="zt01926p019.pdf" docOrigin="Zootaxa 1926" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@b0d3e7" docStyleName="zootaxa.2007.monograph" docTitle="Aenictus ceylonicus Mayr" docType="treatment" docVersion="16" lastPageNumber="17" masterDocId="FFB6A95EFF85C85FFFB3FFA07D1DFFDD" masterDocTitle="Review of the ant genus Aenictus (Hymenoptera: Formicidae) in Australia with notes on A. ceylonicus (Mayr)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="16" updateTime="1661650143793" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the ant genus Aenictus (Hymenoptera: Formicidae) in Australia with notes on A. ceylonicus (Mayr)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shattuck, Steven O.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="volume">
<mods:number>1926</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>19</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.184817</mods:identifier>
<mods:identifier type="GBIF-Dataset">e5c56f88-2105-49a1-99e0-39eb486f9c64</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">184817</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5063924" ID-GBIF-Taxon="119381772" ID-Zenodo-Dep="5063924" LSID="urn:lsid:plazi:treatment:038FD126FF8AC84FFF24F92D7DFDFAE7" httpUri="http://treatment.plazi.org/id/038FD126FF8AC84FFF24F92D7DFDFAE7" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection box="[151,487,1677,1703]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph blockId="15.[151,487,1677,1703]" box="[151,487,1677,1703]" pageId="15" pageNumber="16">
<heading bold="true" box="[151,487,1677,1703]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName authority="Mayr" authorityName="Mayr" box="[151,487,1677,1703]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis bold="true" box="[151,487,1677,1703]" pageId="15" pageNumber="16">
<emphasis bold="true" box="[151,391,1677,1703]" italics="true" pageId="15" pageNumber="16">Aenictus ceylonicus</emphasis>
(Mayr)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="reference_group">
<paragraph blockId="15.[151,1339,1752,1839]" box="[151,1090,1752,1775]" pageId="15" pageNumber="16">
<treatmentCitationGroup box="[151,1090,1752,1775]" pageId="15" pageNumber="16">
<taxonomicName authority="Mayr, 1866: 505" authorityName="Mayr" authorityPageNumber="505" authorityYear="1866" box="[151,544,1752,1775]" class="Insecta" family="Formicidae" genus="Typhlatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonica">
<emphasis box="[151,360,1752,1775]" italics="true" pageId="15" pageNumber="16">Typhlatta ceylonica</emphasis>
<treatmentCitation ID-DOI="http://doi.org/10.5281/zenodo.6294516" ID-Zenodo-Dep="6294516" author="Mayr" box="[366,544,1752,1775]" httpUri="http://treatment.plazi.org/id/00F3BDAB04DA7004D800BE5245E601D4" page="505" pageId="15" pageNumber="16" year="1866">
<bibRefCitation author="Mayr" box="[366,494,1752,1775]" pageId="15" pageNumber="19" refString="Mayr, G. (1866) Myrmecologische Beitrage. Sitzungsberichte der k. Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 53, 484 - 517." type="journal article" year="1866">
Mayr,
<number box="[437,494,1752,1775]" pageId="15" pageNumber="16" value="1866.0">1866</number>
</bibRefCitation>
:
<number box="[504,544,1752,1775]" pageId="15" pageNumber="16" value="505.0">505</number>
</treatmentCitation>
</taxonomicName>
(combination in
<taxonomicName box="[727,818,1752,1775]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis box="[727,818,1752,1775]" italics="true" pageId="15" pageNumber="16">Aenictus</emphasis>
</taxonomicName>
by
<treatmentCitation author="Dalla" box="[859,1077,1752,1775]" page="7" pageId="15" pageNumber="16" year="1893">
<bibRefCitation author="Dalla" box="[859,1051,1752,1775]" pageId="15" pageNumber="19" refString="Dalla Torre, C. G. de. (1893) Catalogus Hymenopterorum, hucusque descriptorum systematicus et synonymicus, 7, 289 pp. Lipsiae." type="book" year="1893">
Dalla Torre,
<number box="[994,1051,1752,1775]" pageId="15" pageNumber="16" value="1893.0">1893</number>
</bibRefCitation>
:
<number box="[1061,1077,1752,1775]" pageId="15" pageNumber="16" value="7.0">7</number>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[151,1339,1752,1839]" pageId="15" pageNumber="16">
<treatmentCitationGroup pageId="15" pageNumber="16">
<taxonomicName box="[151,362,1784,1807]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[151,362,1784,1807]" italics="true" pageId="15" pageNumber="16">Aenictus ceylonicus</emphasis>
</taxonomicName>
<taxonomicName authority="Forel, 1901: 477" authorityName="Forel" authorityPageNumber="477" authorityYear="1901" box="[368,648,1784,1807]" pageId="15" pageNumber="16" rank="variety" variety="latro">
var.
<emphasis box="[414,465,1784,1807]" italics="true" pageId="15" pageNumber="16">latro</emphasis>
<treatmentCitation author="Forel" box="[471,648,1784,1807]" page="477" pageId="15" pageNumber="16" year="1901">
<bibRefCitation author="Forel" box="[471,598,1784,1807]" pageId="15" pageNumber="19" refString="Forel, A. (1901) Les formicides de l'Empire des Indes et de Ceylan. Part 8. Sous famille Dorylinae. Journal of the Bombay Natural History Society, 13, 462 - 477." type="journal article" year="1901">
Forel,
<number box="[541,598,1784,1807]" pageId="15" pageNumber="16" value="1901.0">1901</number>
</bibRefCitation>
:
<number box="[608,648,1784,1807]" pageId="15" pageNumber="16" value="477.0">477</number>
</treatmentCitation>
</taxonomicName>
(junior synonym of
<taxonomicName box="[868,1011,1784,1807]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[868,1011,1784,1807]" italics="true" pageId="15" pageNumber="16">A. ceylonicus</emphasis>
</taxonomicName>
by
<treatmentCitation author="Wilson" box="[1050,1251,1784,1807]" page="452" pageId="15" pageNumber="16" year="1964">
<bibRefCitation author="Wilson" box="[1050,1196,1784,1807]" pageId="15" pageNumber="19" refString="Wilson, E. O. (1964) The true army ants of the Indo-Australian area. Pacific Insects, 6, 427 - 483." type="journal article" year="1964">
Wilson,
<number box="[1139,1196,1784,1807]" pageId="15" pageNumber="16" value="1964.0">1964</number>
</bibRefCitation>
:
<number box="[1207,1251,1784,1807]" pageId="15" pageNumber="16" value="452.0">452</number>
</treatmentCitation>
).
<taxonomicName box="[151,362,1816,1839]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[151,362,1816,1839]" italics="true" pageId="15" pageNumber="16">Aenictus ceylonicus</emphasis>
</taxonomicName>
<taxonomicName authority="Forel, 1913: 188" authorityName="Forel" authorityPageNumber="188" authorityYear="1913" box="[368,725,1816,1839]" pageId="15" pageNumber="16" rank="variety" variety="formosensis">
var.
<emphasis box="[414,541,1816,1839]" italics="true" pageId="15" pageNumber="16">formosensis</emphasis>
<treatmentCitation author="Forel" box="[548,725,1816,1839]" page="188" pageId="15" pageNumber="16" year="1913">
<bibRefCitation author="Forel" box="[548,675,1816,1839]" pageId="15" pageNumber="19" refString="Forel, A. (1913) Fourmis de Tasmanie et d'Australie recoltees par MM. Lae, Froggatt etc. Bulletin de la Societe Vaudoise des Sciences Naturelles, (5) 49, 173 - 195." type="journal article" year="1913">
Forel,
<number box="[618,675,1816,1839]" pageId="15" pageNumber="16" value="1913.0">1913</number>
</bibRefCitation>
:
<number box="[685,725,1816,1839]" pageId="15" pageNumber="16" value="188.0">188</number>
</treatmentCitation>
</taxonomicName>
(junior synonym of
<taxonomicName box="[944,1087,1816,1839]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[944,1087,1816,1839]" italics="true" pageId="15" pageNumber="16">A. ceylonicus</emphasis>
</taxonomicName>
by
<treatmentCitation author="Wilson" box="[1126,1327,1816,1839]" page="452" pageId="15" pageNumber="16" year="1964">
<bibRefCitation author="Wilson" box="[1126,1273,1816,1839]" pageId="15" pageNumber="19" refString="Wilson, E. O. (1964) The true army ants of the Indo-Australian area. Pacific Insects, 6, 427 - 483." type="journal article" year="1964">
Wilson,
<number box="[1216,1273,1816,1839]" pageId="15" pageNumber="16" value="1964.0">1964</number>
</bibRefCitation>
:
<number box="[1283,1327,1816,1839]" pageId="15" pageNumber="16" value="452.0">452</number>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[151,1436,1887,2034]" pageId="15" pageNumber="16">
<emphasis bold="true" box="[151,229,1887,1913]" pageId="15" pageNumber="16">
<typeStatus box="[151,224,1887,1913]" pageId="15" pageNumber="16">Types</typeStatus>
.
</emphasis>
<taxonomicName box="[241,473,1888,1913]" class="Insecta" family="Formicidae" genus="Typhlatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonica">
<emphasis box="[241,473,1888,1913]" italics="true" pageId="15" pageNumber="16">Typhlatta ceylonica</emphasis>
</taxonomicName>
: Worker
<typeStatus box="[587,690,1888,1914]" pageId="15" pageNumber="16">syntypes</typeStatus>
from
<collectingCountry box="[766,883,1888,1914]" name="Sri Lanka" pageId="15" pageNumber="16">Sri Lanka</collectingCountry>
(
<collectionCode box="[901,1000,1888,1914]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="16">NHMW</collectionCode>
, not examined).
<taxonomicName box="[1202,1436,1888,1913]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[1202,1436,1888,1913]" italics="true" pageId="15" pageNumber="16">Aenictus ceylonicus</emphasis>
</taxonomicName>
<taxonomicName authority="Three" authorityName="Three" box="[151,343,1928,1954]" pageId="15" pageNumber="16" rank="variety" variety="latro">
var.
<emphasis box="[202,258,1928,1953]" italics="true" pageId="15" pageNumber="16">latro</emphasis>
: Three
</taxonomicName>
worker
<typeStatus box="[443,546,1928,1954]" pageId="15" pageNumber="16">syntypes</typeStatus>
from Poona,
<collectingCountry box="[708,768,1928,1954]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
(
<collectionCode box="[784,872,1928,1954]" pageId="15" pageNumber="16">MCZC</collectionCode>
, examined).
<taxonomicName box="[1024,1257,1928,1953]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[1024,1257,1928,1953]" italics="true" pageId="15" pageNumber="16">Aenictus ceylonicus</emphasis>
</taxonomicName>
<taxonomicName pageId="15" pageNumber="16" rank="variety" variety="formosensis">
var.
<emphasis italics="true" pageId="15" pageNumber="16">formosensis</emphasis>
</taxonomicName>
: Worker
<typeStatus box="[291,394,1968,1994]" pageId="15" pageNumber="16">syntypes</typeStatus>
from
<collectingCountry box="[466,552,1968,1994]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
(not examined).
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph blockId="15.[151,1436,1887,2034]" lastBlockId="16.[151,1437,152,1338]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis bold="true" box="[198,341,2007,2033]" pageId="15" pageNumber="16">Comments.</emphasis>
<collectingCountry box="[348,380,2008,2034]" name="American Samoa" pageId="15" pageNumber="16">As</collectingCountry>
previously conceived (
<bibRefCitation author="Wilson" box="[652,872,2008,2034]" pageId="15" pageNumber="19" refString="Wilson, E. O. (1964) The true army ants of the Indo-Australian area. Pacific Insects, 6, 427 - 483." type="journal article" year="1964">
Wilson,
<number box="[752,815,2008,2034]" pageId="15" pageNumber="16" value="1964.0">1964</number>
:
<number box="[826,872,2008,2034]" pageId="15" pageNumber="16" value="452.0">452</number>
</bibRefCitation>
) this species extended from
<collectingCountry box="[1204,1264,2008,2034]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
and
<collectingCountry box="[1321,1436,2008,2034]" name="Sri Lanka" pageId="15" pageNumber="16">Sri Lanka</collectingCountry>
eastward to
<collectingCountry box="[300,386,152,178]" name="Taiwan" pageId="16" pageNumber="17">Taiwan</collectingCountry>
and south to
<collectingCountry box="[560,667,152,178]" name="Australia" pageId="16" pageNumber="17">Australia</collectingCountry>
and contained eight junior synonyms
<emphasis box="[1138,1288,152,177]" italics="true" pageId="16" pageNumber="17">
(
<taxonomicName authority="Forel" authorityName="Forel" box="[1146,1288,152,177]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosensis">formosensis</taxonomicName>
</emphasis>
Forel,
<taxonomicName authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="latro">
<emphasis box="[1380,1436,152,177]" italics="true" pageId="16" pageNumber="17">latro</emphasis>
Forel
</taxonomicName>
,
<taxonomicName authority="Karavaiev" authorityName="Karavaiev" box="[227,468,192,218]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis box="[227,340,192,217]" italics="true" pageId="16" pageNumber="17">orientalis</emphasis>
Karavaiev
</taxonomicName>
,
<taxonomicName authority="Donisthorpe" authorityName="Donisthorpe" box="[478,747,192,218]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis box="[478,592,192,217]" italics="true" pageId="16" pageNumber="17">papuanus</emphasis>
Donisthorpe
</taxonomicName>
,
<taxonomicName authority="Donisthorpe" authorityName="Donisthorpe" box="[758,989,192,218]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="similis">
<emphasis box="[758,835,192,217]" italics="true" pageId="16" pageNumber="17">similis</emphasis>
Donisthorpe
</taxonomicName>
, and
<taxonomicName authority="Forel" authorityName="Forel" box="[1049,1199,192,218]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="turneri">
<emphasis box="[1049,1131,192,217]" italics="true" pageId="16" pageNumber="17">turneri</emphasis>
Forel
</taxonomicName>
(with its junior synonyms
<taxonomicName authority="Crawley" authorityName="Crawley" box="[239,448,232,258]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="deuqueti">
<emphasis box="[239,340,232,257]" italics="true" pageId="16" pageNumber="17">deuqueti</emphasis>
Crawley
</taxonomicName>
and
<taxonomicName authority="Clark" authorityName="Clark" box="[510,680,232,258]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis box="[510,600,232,257]" italics="true" pageId="16" pageNumber="17">exiguus</emphasis>
Clark
</taxonomicName>
)). When discussing the specimens placed in
<taxonomicName authority="Wilson (1964)" authorityName="Wilson" authorityYear="1964" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[1219,1343,232,257]" italics="true" pageId="16" pageNumber="17">ceylonicus</emphasis>
<bibRefCitation author="Wilson" pageId="16" pageNumber="19" refString="Wilson, E. O. (1964) The true army ants of the Indo-Australian area. Pacific Insects, 6, 427 - 483." type="journal article" year="1964">
Wilson (
<number box="[159,220,272,298]" pageId="16" pageNumber="17" value="1964.0">1964</number>
)
</bibRefCitation>
</taxonomicName>
recognised at least some of the variation noted in this study (for example, see Wilsons figs.
<number box="[1346,1423,272,298]" pageId="16" pageNumber="17" value="40.5" valueMax="44.0" valueMin="37.0">3744</number>
), but interpreted this variation as intraspecific. For example he mentioned that the subpetiolar process varies considerably in its development, but did not appreciate that this variation occurs in discrete states and shows a strong geographic pattern suggesting that a series of species are involved. A careful re-examination of these characters, combined with considerably more material, has resulted in significantly different conclusions being drawn compared to
<bibRefCitation author="Wilson" box="[457,629,472,498]" pageId="16" pageNumber="19" refString="Wilson, E. O. (1964) The true army ants of the Indo-Australian area. Pacific Insects, 6, 427 - 483." type="journal article" year="1964">
Wilson (
<number box="[556,620,472,498]" pageId="16" pageNumber="17" value="1964.0">1964</number>
)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="16.[151,1437,152,1338]" pageId="16" pageNumber="17">
An examination of currently available material has found that the old “
<taxonomicName box="[1036,1160,512,537]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[1036,1160,512,537]" italics="true" pageId="16" pageNumber="17">ceylonicus</emphasis>
</taxonomicName>
” contains a large number of species, including
<taxonomicName box="[443,599,552,577]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[443,599,552,577]" italics="true" pageId="16" pageNumber="17">A. ceylonicus</emphasis>
</taxonomicName>
(strict sense),
<taxonomicName box="[769,894,552,577]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acerbus">
<emphasis box="[769,894,552,577]" italics="true" pageId="16" pageNumber="17">A. acerbus</emphasis>
</taxonomicName>
,
<taxonomicName box="[908,1053,552,577]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis box="[908,1053,552,577]" italics="true" pageId="16" pageNumber="17">A. orientalis</emphasis>
</taxonomicName>
,
<taxonomicName box="[1067,1213,552,577]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis box="[1067,1213,552,577]" italics="true" pageId="16" pageNumber="17">A. papuanus</emphasis>
</taxonomicName>
,
<taxonomicName box="[1227,1355,552,577]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis box="[1227,1355,552,577]" italics="true" pageId="16" pageNumber="17">A. prolixus</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="turneri">
<emphasis italics="true" pageId="16" pageNumber="17">A. turneri</emphasis>
</taxonomicName>
. To determine the identity of
<taxonomicName box="[601,761,592,617]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[601,761,592,617]" italics="true" pageId="16" pageNumber="17">A. ceylonicus</emphasis>
</taxonomicName>
itself will require considerable work and is beyond the scope of the present study. However, there are a wealth of morphological characters which allow the development of robust species hypotheses as has been demonstrated above for the Australian fauna. Having said that, morphological differences among species are often subtle and require considerable attention to detail to decipher. The following notes are provided as a starting point for a full revision of these ants.
</paragraph>
<paragraph blockId="16.[151,1437,152,1338]" pageId="16" pageNumber="17">
Most of the Indian specimens share the configuration of the subpetiolar process, which forms a rounded anterior lobe followed by a posterior flat to concave extension ending at the junction with the postpetiole. Others have an elongate rectangular subpetiolar process, including the
<typeStatus box="[947,1009,872,898]" pageId="16" pageNumber="17">types</typeStatus>
of
<taxonomicName box="[1050,1139,872,897]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="latro">
<emphasis box="[1050,1139,872,897]" italics="true" pageId="16" pageNumber="17">A. latro</emphasis>
</taxonomicName>
. Specimens with both of these morphologies can be found throughout Asia including in
<collectingCountry box="[895,984,912,938]" name="Taiwan" pageId="16" pageNumber="17">Taiwan</collectingCountry>
, the
<collectingCountry box="[1040,1175,912,938]" name="Philippines" pageId="16" pageNumber="17">Philippines</collectingCountry>
,
<collectingCountry box="[1187,1286,912,938]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
and
<collectingCountry name="Indonesia" pageId="16" pageNumber="17">Indonesia</collectingCountry>
. But while material from
<collectingCountry box="[503,602,952,978]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
has a rectangular subpetiolar process it has the dorsal surface of the mesosoma smooth and lacking any indication of the metanotal groove (most other species have at least a weak angle at the metanotal groove). Thus while the shape of the subpetiolar process is important it must be used in conjunction with other characters when determining species boundaries.
</paragraph>
<paragraph blockId="16.[151,1437,152,1338]" pageId="16" pageNumber="17">
While the work undertaken here is preliminary, it clearly shows that the situation surrounding this species, and close relatives, is much more complex than that recognised by earlier workers.
<collectingCountry box="[1124,1157,1152,1178]" name="American Samoa" pageId="16" pageNumber="17">As</collectingCountry>
a first step in clarifying this situation the names
<taxonomicName box="[433,578,1192,1217]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis box="[433,578,1192,1217]" italics="true" pageId="16" pageNumber="17">A. orientalis</emphasis>
</taxonomicName>
and
<taxonomicName box="[634,748,1192,1217]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="turneri">
<emphasis box="[634,748,1192,1217]" italics="true" pageId="16" pageNumber="17">A. turneri</emphasis>
</taxonomicName>
are treated as valid species,
<taxonomicName box="[1081,1228,1192,1217]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis box="[1081,1228,1192,1217]" italics="true" pageId="16" pageNumber="17">A. papuanus</emphasis>
</taxonomicName>
and
<taxonomicName box="[1284,1392,1192,1217]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="similis">
<emphasis box="[1284,1392,1192,1217]" italics="true" pageId="16" pageNumber="17">A. similis</emphasis>
</taxonomicName>
are transferred to synonymy with
<taxonomicName box="[506,652,1232,1257]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis box="[506,652,1232,1257]" italics="true" pageId="16" pageNumber="17">A. orientalis</emphasis>
</taxonomicName>
while
<taxonomicName box="[732,905,1232,1257]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis box="[732,905,1232,1257]" italics="true" pageId="16" pageNumber="17">A. formosensis</emphasis>
</taxonomicName>
and
<taxonomicName box="[964,1053,1232,1257]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="latro">
<emphasis box="[964,1053,1232,1257]" italics="true" pageId="16" pageNumber="17">A. latro</emphasis>
</taxonomicName>
are retained as junior synonyms of
<taxonomicName box="[185,342,1272,1297]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis box="[185,342,1272,1297]" italics="true" pageId="16" pageNumber="17">A. ceylonicus</emphasis>
</taxonomicName>
. However this should be treated as preliminary until all relevant material can be studied in detail.
</paragraph>
</subSubSection>
</treatment>
</document>