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<document id="AB10D6D2E8354ABE1E0517AF5184B190" ID-DOI="10.11646/phytotaxa.597.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7929279" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683889203312" checkinUser="plazi" docAuthor="Wan, Xia, Knapp, Ralf, Chen, De-Kui, He, Hai &amp; Zhang, Li-Bing" docDate="2023" docId="0384467FFFBDFF873FC3028BFD072B43" docLanguage="en" docName="phytotaxa.597.2.3.pdf" docOrigin="Phytotaxa 597 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.597.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Polystichum pseudodeltodon Tagawa 1937" docType="treatment" docVersion="4" lastPageNumber="132" masterDocId="FFBD3E07FFB9FF833F4B026EFFD72907" masterDocTitle="Taxonomic notes on some East Asian pteridophytes" masterLastPageNumber="140" masterPageNumber="128" pageNumber="132" updateTime="1684155635051" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="95DEA3AE75C9F504B27DC15754CC7431">Taxonomic notes on some East Asian pteridophytes</mods:title>
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<mods:namePart id="4E6E275E917B254F7FA85A07D1C8CA31">Wan, Xia</mods:namePart>
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<mods:affiliation id="8CAAA335C04C0B8E95790DAA65D39BF4">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China; &amp; College of Life Sciences, Sichuan University, Chengdu 610065, China; &amp; University of Chinese Academy of Sciences, Beijing 100049, China; &amp; Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. &amp; xwan @ mobot. org; https: // orcid. org / 0000 - 0001 - 8312 - 6917</mods:affiliation>
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<mods:namePart id="16B17F0B45FD3C0F3D9EFCF721271E01">Knapp, Ralf</mods:namePart>
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<mods:affiliation id="186405B1F36165DC57C9A9D604C222BB">Correspondent, The Muséum national dHistoire naturelle (MNHN, Paris, France), Steigestrasse 78, 69412 Eberbach, Germany; &amp; ralf. knapp @ gmail. org; https: // orcid. org / 0000 - 0002 - 0147 - 5687</mods:affiliation>
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<mods:namePart id="EBC3BD3D8DA21CEB858B3BF3481E55FB">Chen, De-Kui</mods:namePart>
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<mods:affiliation id="C3B6FBEC5F6F514F19079BCF785841DC">College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 401331, China; &amp; No. 1 Middle School of Tongren, Tong Ren, Guizhou 554300, China &amp; cjxby 739847 @ 163. com; https: // orcid. org / 0000 - 0002 - 1224 - 7372</mods:affiliation>
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<mods:namePart id="A91E5C7588AFCF805EB38319365B0B6C">He, Hai</mods:namePart>
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<mods:affiliation id="CD75196651488FFA59FF5B6E5B8E35ED">College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 401331, China; &amp; hehai @ cqnu. edu. cn; https: // orcid. org / 0000 - 0001 - 9896 - 0971</mods:affiliation>
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<mods:namePart id="474BE53EBB73321E1FC2BB10096CF475">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="730D1BE7E059E204160A88125C999820">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China; &amp; Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<mods:title id="E3256F62639555A02C1D8F9965FB2F90">Phytotaxa</mods:title>
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<mods:date id="E206475A39E2CA529BE7DB6ECF7364B1">2023</mods:date>
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<treatment id="0384467FFFBDFF873FC3028BFD072B43" ID-DOI="http://doi.org/10.5281/zenodo.7929173" ID-Zenodo-Dep="7929173" LSID="urn:lsid:plazi:treatment:0384467FFFBDFF873FC3028BFD072B43" httpUri="http://treatment.plazi.org/id/0384467FFFBDFF873FC3028BFD072B43" lastPageNumber="132" pageId="4" pageNumber="132">
<subSubSection id="C337A4E2FFBDFF873FC3028BFCF92807" box="[136,814,229,257]" pageId="4" pageNumber="132" type="nomenclature">
<paragraph id="8B92F769FFBDFF873FC3028BFCF92807" blockId="4.[136,814,229,257]" box="[136,814,229,257]" pageId="4" pageNumber="132">
<heading id="D0DA4005FFBDFF873FC3028BFCF92807" box="[136,814,229,257]" fontSize="11" level="2" pageId="4" pageNumber="132" reason="3">
<treatmentCitationGroup id="AB3DD047FFBDFF873FC3028BFCF92807" box="[136,814,229,257]" pageId="4" pageNumber="132">
<treatmentCitation id="0A8CD178FFBDFF873FC3028BFD692806" author="Tagawa, M." box="[136,702,229,257]" page=")" pageId="4" pageNumber="132" year="1937">
<taxonomicName id="4C2D8CEAFFBDFF873FC3028BFD692806" ID-CoL="4LN6Q" ID-ENA="1814928" authority="Tagawa (1937: 159)" authorityName="Tagawa" authorityPageNumber="159" authorityYear="1937" box="[136,702,229,257]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="pseudodeltodon">
<emphasis id="B9592B7BFFBDFF873FC3028BFE052807" bold="true" box="[136,466,229,256]" italics="true" pageId="4" pageNumber="132">Polystichum pseudodeltodon</emphasis>
<bibRefCitation id="EFBC8A98FFBDFF873E920288FD692806" author="Tagawa, M." box="[473,702,230,257]" pageId="4" pageNumber="132" pagination="154 - 168" refId="ref5701" refString="Tagawa, M. (1937) Spicilegium Pteridographiae Asiae Orientalis 14. Acta Phytotaxonomica et Geobotanica 6: 154 - 168. https: // doi. org / 10.18942 / bunruichiri. KJ 00001078859" type="journal article" year="1937">Tagawa (1937: 159)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<figureCitation id="1316EBECFFBDFF873D830288FCFF2807" box="[712,808,230,256]" captionStart="FIGURE 3" captionStartId="5.[136,229,1928,1950]" captionTargetBox="[161,1426,199,1894]" captionTargetId="figure-21@5.[151,1436,190,1904]" captionTargetPageId="5" captionText="FIGURE 3. Polystichum pseudodeltodon.—A. Habit.—B. Adaxial view of middle portion of lamina.—C.Adaxial view of upper portion of lamina.—D.Abaxial view of middle portion of lamina.—F. Pinnae showing sori (photo credit: Ralf Knapp)." figureDoi="http://doi.org/10.5281/zenodo.7929289" httpUri="https://zenodo.org/record/7929289/files/figure.png" pageId="4" pageNumber="132">Figure 3</figureCitation>
.
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</heading>
</paragraph>
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<paragraph id="8B92F769FFBDFF873FC30363FB402825" blockId="4.[136,1175,269,291]" box="[136,1175,269,291]" pageId="4" pageNumber="132">
<materialsCitation id="3B45FD34FFBDFF873FC30363FC6B2824" box="[136,956,269,291]" collectingDate="1934-10-04" collectionCode="M" country="Taiwan Island" county="Tippon-zan" location="Taito" municipality="Kiriyama" pageId="4" pageNumber="132" specimenCount="1" typeStatus="holotype">
<typeStatus id="549649CBFFBDFF873FC30363FF6E2824" box="[136,185,269,291]" pageId="4" pageNumber="132">Type</typeStatus>
:—
<collectingCountry id="F33AB7F9FFBDFF873F9E0363FEB62824" box="[213,353,269,291]" name="Taiwan Island" pageId="4" pageNumber="132">Taiwan Island</collectingCountry>
:
<location id="8EF2A1B2FFBDFF873E210363FE4C2824" LSID="urn:lsid:plazi:treatment:0384467FFFBDFF873FC3028BFD072B43:8EF2A1B2FFBDFF873E210363FE4C2824" box="[362,411,269,291]" country="Taiwan Island" county="Tippon-zan" municipality="Kiriyama" name="Taito" pageId="4" pageNumber="132">Taito</location>
(Taitung), inter
<collectingMunicipality id="6BF66D13FFBDFF873D7C0363FD452824" box="[567,658,269,291]" pageId="4" pageNumber="132">Kiriyama</collectingMunicipality>
et
<collectingCounty id="62F38FE5FFBDFF873DE40363FCC82824" box="[687,799,269,291]" pageId="4" pageNumber="132">Tippon-zan</collectingCounty>
,
<date id="FF93D1A9FFBDFF873C620363FC4C2825" box="[809,923,269,290]" pageId="4" pageNumber="132" value="1934-10-04">
<collectingDate id="EFD72841FFBDFF873C620363FC4C2825" box="[809,923,269,290]" pageId="4" pageNumber="132" value="1934-10-04">4 Oct. 1934</collectingDate>
</date>
,
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<emphasis id="B9592B7BFFBDFF873CEE0360FC6B2824" box="[933,956,270,291]" italics="true" pageId="4" pageNumber="132">M</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="B9592B7BFFBDFF873CF60360FC162824" box="[957,961,270,291]" italics="true" pageId="4" pageNumber="132">.</emphasis>
<materialsCitation id="3B45FD34FFBDFF873C8E0360FB442825" box="[965,1171,269,291]" collectionCode="KYO" country="Taiwan Island" location="Tagawa" pageId="4" pageNumber="132" specimenCount="1" typeStatus="holotype">
<emphasis id="B9592B7BFFBDFF873C8E0360FB902825" box="[965,1095,269,291]" italics="true" pageId="4" pageNumber="132">
<location id="8EF2A1B2FFBDFF873C8E0360FBC72824" LSID="urn:lsid:plazi:treatment:0384467FFFBDFF873FC3028BFD072B43:8EF2A1B2FFBDFF873C8E0360FBC72824" box="[965,1040,270,291]" country="Taiwan Island" name="Tagawa" pageId="4" pageNumber="132">Tagawa</location>
1069
</emphasis>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337A4E2FFBDFF873FC3033CFCBD28FB" pageId="4" pageNumber="132" type="discussion">
<paragraph id="8B92F769FFBDFF873FC3033CFCBD28FB" blockId="4.[136,1452,338,580]" pageId="4" pageNumber="132">
<taxonomicName id="4C2D8CEAFFBDFF873FC3033CFE1C286B" authorityName="Tagawa" authorityYear="1937" box="[136,459,338,364]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="pseudodeltodon">
<emphasis id="B9592B7BFFBDFF873FC3033CFE1C286B" box="[136,459,338,364]" italics="true" pageId="4" pageNumber="132">Polystichum pseudodeltodon</emphasis>
</taxonomicName>
has long been identified as
<taxonomicName id="4C2D8CEAFFBDFF873C4C033DFAFE286B" authority="(Baker 1880: 494) Diels (1899: 191)" authorityName="Diels" authorityPageNumber="191" authorityYear="1899" baseAuthorityName="Baker" baseAuthorityPageNumber="494" baseAuthorityYear="1880" box="[775,1321,338,364]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="deltodon">
<emphasis id="B9592B7BFFBDFF873C4C033DFC52286B" box="[775,901,338,364]" italics="true" pageId="4" pageNumber="132">P. deltodon</emphasis>
(
<bibRefCitation id="EFBC8A98FFBDFF873CD8033CFB84286B" author="Baker, J. G." box="[915,1107,338,364]" pageId="4" pageNumber="132" pagination="494 - 495" refId="ref2905" refString="Baker, J. G. (1880) New ferns of China and Japan. The Gardeners' Chronicle, new series 14: 494 - 495." type="book chapter" year="1880">Baker 1880: 494</bibRefCitation>
)
<bibRefCitation id="EFBC8A98FFBDFF873B29033CFAFE286B" author="Diels, L." box="[1122,1321,338,364]" pageId="4" pageNumber="132" pagination="139 - 339" refId="ref3221" refString="Diels, L. (1899) Polypodiaceae. In: Engler, H. G. A. &amp; Prantl, K. A. E. (Eds.) Die Naturlichen Pflanzenfamilien, vol. 1 (4). W. Engelmann, Leipzig, pp. 139 - 339." type="book chapter" year="1899">Diels (1899: 191)</bibRefCitation>
</taxonomicName>
even in the recent works (e.g.,
<bibRefCitation id="EFBC8A98FFBDFF873E2B0318FE3B2897" author="Knapp, R." box="[352,492,374,400]" pageId="4" pageNumber="132" refId="ref4087" refString="Knapp, R. (2011) Ferns and fern allies of Taiwan. Taipei: KBCC Press &amp; Yuan-Liou Publishing." type="book" year="2011">Knapp 2011</bibRefCitation>
,
<bibRefCitation id="EFBC8A98FFBDFF873EB30318FD7E2897" author="Hsu, T. - C. &amp; Chen, C. - W. &amp; Knapp, R. &amp; Hung, H. - C." box="[504,681,374,400]" pageId="4" pageNumber="132" refId="ref4034" refString="Hsu, T. - C., Chen, C. - W., Knapp, R. &amp; Hung, H. - C. (2019) Illustrated Flora of Taiwan vol. 8: Ferns &amp; Lycophytes (I &amp; II). Maotouying Chuban, Taiwan." type="book" year="2019">
Hsu
<emphasis id="B9592B7BFFBDFF873D670319FDB22897" box="[556,613,374,400]" italics="true" pageId="4" pageNumber="132">et al.</emphasis>
2019
</bibRefCitation>
). However,
<taxonomicName id="4C2D8CEAFFBDFF873C7B0319FC792897" authorityName="Diels" authorityYear="1899" baseAuthorityName="Baker" baseAuthorityYear="1880" box="[816,942,374,400]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="deltodon">
<emphasis id="B9592B7BFFBDFF873C7B0319FC792897" box="[816,942,374,400]" italics="true" pageId="4" pageNumber="132">P. deltodon</emphasis>
</taxonomicName>
does not occur in
<collectingCountry id="F33AB7F9FFBDFF873B350318FB072897" box="[1150,1232,374,400]" name="Taiwan" pageId="4" pageNumber="132">Taiwan</collectingCountry>
but only in Central and southern
<collectingCountry id="F33AB7F9FFBDFF873E6903F4FEB228B3" box="[290,357,410,436]" name="China" pageId="4" pageNumber="132">China</collectingCountry>
(
<bibRefCitation id="EFBC8A98FFBDFF873E3E03F4FD7528B3" author="Zhang, L. - B. &amp; Barrington, D. S." box="[373,674,410,436]" pageId="4" pageNumber="132" pagination="629 - 713" refId="ref7470" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (eds.) Flora of China, vol. 2 - 3 (Lycopodiaceae- Polypodiaceae). Science Press and St. Louis, Missouri Botanical Garden Press, Beijing, pp. 629 - 713." type="book chapter" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
) and
<collectingCountry id="F33AB7F9FFBDFF873DA803F4FCF428B3" box="[739,803,410,436]" name="Japan" pageId="4" pageNumber="132">Japan</collectingCountry>
(
<bibRefCitation id="EFBC8A98FFBDFF873C7803F4FC0528B3" author="Ebihara, A." box="[819,978,410,436]" pageId="4" pageNumber="132" refId="ref3457" refString="Ebihara, A. (2016) The standard of ferns and lycophytes in Japan, vol. 1. Gakken Plus, Tokyo, 475 pp." type="book" year="2016">Ebihara 2016</bibRefCitation>
).
<taxonomicName id="4C2D8CEAFFBDFF873CAD03F4FAFD28B3" authorityName="Tagawa" authorityYear="1937" box="[998,1322,410,436]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="pseudodeltodon">
<emphasis id="B9592B7BFFBDFF873CAD03F4FAFD28B3" box="[998,1322,410,436]" italics="true" pageId="4" pageNumber="132">Polystichum pseudodeltodon</emphasis>
</taxonomicName>
has pinnae with rounded auricles and hardly toothed pinna margins, whereas
<taxonomicName id="4C2D8CEAFFBDFF873CC603D1FBD928DF" authorityName="Diels" authorityYear="1899" baseAuthorityName="Baker" baseAuthorityYear="1880" box="[909,1038,446,472]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="deltodon">
<emphasis id="B9592B7BFFBDFF873CC603D1FBD928DF" box="[909,1038,446,472]" italics="true" pageId="4" pageNumber="132">P. deltodon</emphasis>
</taxonomicName>
has pinnae with acute auricles and toothed pinna margins (
<bibRefCitation id="EFBC8A98FFBDFF873EDB038CFDFC28FB" author="Tagawa, M." box="[400,555,482,508]" pageId="4" pageNumber="132" pagination="154 - 168" refId="ref5701" refString="Tagawa, M. (1937) Spicilegium Pteridographiae Asiae Orientalis 14. Acta Phytotaxonomica et Geobotanica 6: 154 - 168. https: // doi. org / 10.18942 / bunruichiri. KJ 00001078859" type="journal article" year="1937">Tagawa 1937</bibRefCitation>
,
<bibRefCitation id="EFBC8A98FFBDFF873D7D038CFC8828FB" author="Zhang, L. - B. &amp; Barrington, D. S." box="[566,863,482,508]" pageId="4" pageNumber="132" pagination="629 - 713" refId="ref7470" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (eds.) Flora of China, vol. 2 - 3 (Lycopodiaceae- Polypodiaceae). Science Press and St. Louis, Missouri Botanical Garden Press, Beijing, pp. 629 - 713." type="book chapter" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337A4E2FFBDFF873FF60068FD072B43" pageId="4" pageNumber="132" type="distribution">
<paragraph id="8B92F769FFBDFF873FF60068FD072B43" blockId="4.[136,1452,338,580]" pageId="4" pageNumber="132">
<taxonomicName id="4C2D8CEAFFBDFF873FF60068FDD22B27" authorityName="Tagawa" authorityYear="1937" box="[189,517,518,544]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="pseudodeltodon">
<emphasis id="B9592B7BFFBDFF873FF60068FDD22B27" box="[189,517,518,544]" italics="true" pageId="4" pageNumber="132">Polystichum pseudodeltodon</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F33AB7F9FFBDFF873DF00068FCDA2B27" box="[699,781,518,544]" name="Taiwan" pageId="4" pageNumber="132">Taiwan</collectingCountry>
(
<bibRefCitation id="EFBC8A98FFBDFF873C6A0068FB8D2B27" author="Zhang, L. - B. &amp; Barrington, D. S." box="[801,1114,518,544]" pageId="4" pageNumber="132" pagination="629 - 713" refId="ref7470" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (eds.) Flora of China, vol. 2 - 3 (Lycopodiaceae- Polypodiaceae). Science Press and St. Louis, Missouri Botanical Garden Press, Beijing, pp. 629 - 713." type="book chapter" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
). Morphologically it is quite variable (
<bibRefCitation id="EFBC8A98FFBDFF873FBA0044FEAA2B43" author="Knapp, R." box="[241,381,554,580]" pageId="4" pageNumber="132" refId="ref4087" refString="Knapp, R. (2011) Ferns and fern allies of Taiwan. Taipei: KBCC Press &amp; Yuan-Liou Publishing." type="book" year="2011">Knapp 2011</bibRefCitation>
) and more studies are needed.
</paragraph>
</subSubSection>
</treatment>
</document>