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<document id="E775AE3172D0B633A563E90CDA3D2EAC" ID-DOI="10.11646/zootaxa.5418.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10718140" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1709045721138" checkinUser="plazi" docAuthor="Tiunova, Tatiana M." docDate="2024" docId="03FFE945FFB81F5655CFFC06D2E91BF4" docLanguage="en" docName="zootaxa.5418.1.6.pdf" docOrigin="Zootaxa 5418 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torleya mikhaili Tiunova 1995" docType="treatment" docVersion="1" lastPageNumber="97" masterDocId="FFC6913DFFB81F5E5558FFCAD0641C0E" masterDocTitle="Redescription of the Mayfly Torleya mikhaili Tiunova, 1995 (Ephemeroptera, Ephemerellidae)" masterLastPageNumber="100" masterPageNumber="89" pageNumber="89" updateTime="1709247786406" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="E979E37D1B47DD262A3FFD4EA5B29B11">Redescription of the Mayfly Torleya mikhaili Tiunova, 1995 (Ephemeroptera, Ephemerellidae)</mods:title>
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<treatment id="03FFE945FFB81F5655CFFC06D2E91BF4" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4" httpUri="http://treatment.plazi.org/id/03FFE945FFB81F5655CFFC06D2E91BF4" lastPageId="8" lastPageNumber="97" pageId="0" pageNumber="89">
<subSubSection id="C34C0BD8FFB81F5E55CFFC06D12C1FEA" box="[151,328,972,996]" pageId="0" pageNumber="89" type="nomenclature">
<paragraph id="8BE95853FFB81F5E55CFFC06D12C1FEA" blockId="0.[151,1437,972,2040]" box="[151,328,972,996]" pageId="0" pageNumber="89">
<taxonomicName id="4C5623D0FFB81F5E55CFFC06D12C1FEA" ID-CoL="57FS4" authorityName="Tiunova" authorityYear="1995" box="[151,328,972,996]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB81F5E55CFFC06D12C1FEA" box="[151,328,972,996]" italics="true" pageId="0" pageNumber="89">Torleya mikhaili</emphasis>
</taxonomicName>
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<subSubSection id="C34C0BD8FFB81F5E540AFC06D33E1ABA" pageId="0" pageNumber="89" type="description">
<paragraph id="8BE95853FFB81F5E540AFC06D54119D2" blockId="0.[151,1437,972,2040]" pageId="0" pageNumber="89">
was first described by the author (1995) from the south of the Russian Far East, based on the male adult stage, reared from a larva (
<bibRefCitation id="EFC725A2FFB81F5E54ADFC3AD2EB1806" author="Tiunova, T. M." box="[501,655,1008,1032]" pageId="0" pageNumber="89" pagination="51 - 56" refId="ref4850" refString="Tiunova, T. M. (1995) A new species of Torleya Lestage, 1917 (Ephemeroptera: Ephemerellidae) from the Far East of Russia. Aquatic Insects, 17, 51 - 56. https: // doi. org / 10.1080 / 01650429509361569" type="journal article" year="1995">Tiunova 1995</bibRefCitation>
). At that time, only four other species of the genus were known:
<taxonomicName id="4C5623D0FFB81F5E5014FC3AD2701822" authority="(Allen &amp; Edmunds, 1963)" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="nepalica">
<emphasis id="B9228441FFB81F5E5014FC3AD0971822" italics="true" pageId="0" pageNumber="89">Torleya nepalica</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E545AFBDED2681822" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[258,524,1044,1068]" pageId="0" pageNumber="89" pagination="583 - 600" refId="ref3973" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) A revision of the genus Ephemerella (Ephemeroptera: Ephemerellidae). VI. The subgenus Serratella in North America. Annals of the Entomological Society of America, 56 (5), 583 - 600. https: // doi. org / 10.1093 / aesa / 56.5.583" type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F34118C3FFB81F5E570EFBDED2FC1822" box="[598,664,1044,1068]" name="Nepal" pageId="0" pageNumber="89">Nepal</collectingCountry>
,
<taxonomicName id="4C5623D0FFB81F5E57FCFBDED3D81822" authority="(Klapalek, 1905)" baseAuthorityName="Klapalek" baseAuthorityYear="1905" box="[676,956,1044,1068]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B9228441FFB81F5E57FCFBDED29B1825" box="[676,767,1044,1067]" italics="true" pageId="0" pageNumber="89">T. major</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E5655FBDED3D01822" author="Klapalek, F." box="[781,948,1044,1068]" pageId="0" pageNumber="89" pagination="75 - 79" refId="ref4605" refString="Klapalek, F. (1905) Ephemeridarum species quator novae. Casopis Ceske Spolecnosti Entomologicke, 2, 75 - 79." type="journal article" year="1905">Klapalék, 1905</bibRefCitation>
)
</taxonomicName>
from Europe,
<taxonomicName id="4C5623D0FFB81F5E5102FBDED5F81822" authority="Kazlauskas, 1963" authorityName="Kazlauskas" authorityYear="1963" box="[1114,1436,1044,1068]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="padunica">
<emphasis id="B9228441FFB81F5E5102FBDED4BC1822" box="[1114,1240,1044,1068]" italics="true" pageId="0" pageNumber="89">T. padunica</emphasis>
<bibRefCitation id="EFC725A2FFB81F5E5187FBDED5F81822" author="Kazlauskas, R." box="[1247,1436,1044,1068]" pageId="0" pageNumber="89" pagination="582 - 592" refId="ref4575" refString="Kazlauskas, R. (1963) New and little known mayfly (Ephemeroptera) species of the USSR fauna. Entomologicheskoe Obozrenie, 42, 582 - 592." type="journal article" year="1963">Kazlauskas, 1963</bibRefCitation>
</taxonomicName>
from eastern Siberia and
<taxonomicName id="4C5623D0FFB81F5E54F8FBF2D2F1185E" authority="Gose, 1980" authorityName="Gose" authorityYear="1980" box="[416,661,1080,1104]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B9228441FFB81F5E54F8FBF2D2731841" box="[416,535,1080,1103]" italics="true" pageId="0" pageNumber="89">T. japonica</emphasis>
<bibRefCitation id="EFC725A2FFB81F5E5745FBF2D2F1185E" author="Gose, K." box="[541,661,1080,1104]" pageId="0" pageNumber="89" pagination="286 - 288" refId="ref4041" refString="Gose, K. (1980) The mayflies of Japanese. 8. Key to families, genera, and species. Aquabiology, 9, 286 - 288." type="journal article" year="1980">Gose, 1980</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F34118C3FFB81F5E578DFBF2D3711841" box="[725,789,1080,1103]" name="Japan" pageId="0" pageNumber="89">Japan</collectingCountry>
. In 2004, a review of
<taxonomicName id="4C5623D0FFB81F5E56A1FBF2D491185E" box="[1017,1269,1080,1104]" pageId="0" pageNumber="89">
<emphasis id="B9228441FFB81F5E56A1FBF2D42E185E" box="[1017,1098,1080,1104]" italics="true" pageId="0" pageNumber="89">Torleya</emphasis>
Lestage species
</taxonomicName>
was carried out that included eight valid
<taxonomicName id="4C5623D0FFB81F5E54FDFB96D22E187A" box="[421,586,1116,1140]" pageId="0" pageNumber="89">
<emphasis id="B9228441FFB81F5E54FDFB96D192187A" box="[421,502,1116,1140]" italics="true" pageId="0" pageNumber="89">Torleya</emphasis>
species
</taxonomicName>
from the Palaearctic and Oriental Regions (Jacobus
<emphasis id="B9228441FFB81F5E51DDFB96D4D1187A" box="[1157,1205,1116,1140]" italics="true" pageId="0" pageNumber="89">et al</emphasis>
. 2004). The previous five species and three others were included:
<taxonomicName id="4C5623D0FFB81F5E572EFB4AD4321896" authority="Zhou, Su &amp; Gui, 2000" authorityName="Zhou, Su &amp; Gui" authorityYear="2000" box="[630,1110,1152,1176]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="grandipennis">
<emphasis id="B9228441FFB81F5E572EFB4AD33F1896" box="[630,859,1152,1176]" italics="true" pageId="0" pageNumber="89">Torleya grandipennis</emphasis>
<bibRefCitation id="EFC725A2FFB81F5E563AFB4AD4321896" author="Zhou, C. - F. &amp; Su, C. - R. &amp; Gui, H." box="[866,1110,1152,1176]" pageId="0" pageNumber="89" pagination="312 - 314" refId="ref5025" refString="Zhou, C. - F., Su, C. - R. &amp; Gui, H. (2000) The first record of the genus Torleya in China with description of a new species (Ephemeroptera: Ephemerellidae). Acta Zootaxonomica Sinica, 25 (3), 312 - 314." type="journal article" year="2000">Zhou, Su &amp; Gui, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5623D0FFB81F5E513AFB4AD5FC1896" authority="Kang &amp; Yang, 1995" authorityName="Kang &amp; Yang" authorityYear="1995" box="[1122,1432,1152,1176]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="lutosa">
<emphasis id="B9228441FFB81F5E513AFB4AD4D91896" box="[1122,1213,1152,1176]" italics="true" pageId="0" pageNumber="89">T. lutosa</emphasis>
<bibRefCitation id="EFC725A2FFB81F5E519DFB4AD5FC1896" author="Kang, S. - C. &amp; Yang, C. - T." box="[1221,1432,1152,1176]" pageId="0" pageNumber="89" pagination="95 - 116" refId="ref4534" refString="Kang, S. - C. &amp; Yang, C. - T. (1995) Ephemerellidae of Taiwan (Insecta, Ephemeroptera). Bulletin of National Museum of Natural Science, 5, 95 - 116." type="journal article" year="1995">Kang &amp;Yang, 1995</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5623D0FFB81F5E559DFB6ED26F18B2" authority="Jacobus &amp; McCafferty" authorityName="Jacobus &amp; McCafferty" authorityYear="2004" box="[197,523,1188,1212]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="naga">
<emphasis id="B9228441FFB81F5E559DFB6ED17218B5" box="[197,278,1188,1211]" italics="true" pageId="0" pageNumber="89">T. naga</emphasis>
Jacobus &amp;McCafferty
</taxonomicName>
(in Jacobus
<emphasis id="B9228441FFB81F5E57CBFB6ED2A718B2" box="[659,707,1188,1212]" italics="true" pageId="0" pageNumber="89">et al</emphasis>
.), 2004. Later, in a work devoted to the revision of
<taxonomicName id="4C5623D0FFB81F5E51ADFB6ED5F818B2" authorityName="Klapalek" authorityYear="1909" box="[1269,1436,1188,1212]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
genera, three more species were added.
<taxonomicName id="4C5623D0FFB81F5E571AFB02D39818EE" authority="(Allen &amp; Edmunds, 1963)" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" box="[578,1020,1224,1248]" class="Insecta" family="Ephemerellidae" genus="Torlyea" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="coheri">
<emphasis id="B9228441FFB81F5E571AFB02D2B918EE" box="[578,733,1224,1248]" italics="true" pageId="0" pageNumber="89">Torlyea coheri</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E57B3FB02D39018EE" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[747,1012,1224,1248]" pageId="0" pageNumber="89" pagination="583 - 600" refId="ref3973" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) A revision of the genus Ephemerella (Ephemeroptera: Ephemerellidae). VI. The subgenus Serratella in North America. Annals of the Entomological Society of America, 56 (5), 583 - 600. https: // doi. org / 10.1093 / aesa / 56.5.583" type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C5623D0FFB81F5E5168FB02D173190A" authority="(Jacobus, McCafferty &amp; Sites, 2007)" baseAuthorityName="Jacobus, McCafferty &amp; Sites" baseAuthorityYear="2007" class="Insecta" family="Ephemerellidae" genus="Torlyea" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="lacuna">
<emphasis id="B9228441FFB81F5E5168FB02D4F718EE" box="[1072,1171,1224,1248]" italics="true" pageId="0" pageNumber="89">T. lacuna</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E51F9FB02D175190A" author="Jacobus, L. M. &amp; McCafferty, W. P. &amp; Sites, R. W." pageId="0" pageNumber="89" pagination="45 - 53" refId="ref4414" refString="Jacobus, L. M., McCafferty, W. P. &amp; Sites, R. W. (2007) New species and stage associations in the genus Crinitella (Ephemeroptera: Ephemerellidae). Zootaxa, 1611 (1), 45 - 53." type="journal article" year="2007">Jacobus, McCafferty &amp; Sites, 2007</bibRefCitation>
)
</taxonomicName>
were moved from the genus
<taxonomicName id="4C5623D0FFB81F5E570EFB26D3E8190A" authority="Allen &amp; Edmunds" authorityName="Allen &amp; Edmunds" authorityYear="1963" box="[598,908,1260,1284]" class="Insecta" family="Ephemerellidae" genus="Crinitella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E570EFB26D2DA190A" box="[598,702,1260,1284]" italics="true" pageId="0" pageNumber="89">Crinitella</emphasis>
Allen &amp; Edmunds
</taxonomicName>
, and
<taxonomicName id="4C5623D0FFB81F5E569EFB26D50F190A" authority="(Gui, Zhou &amp; Su, 1999)" baseAuthorityName="Gui, Zhou &amp; Su" baseAuthorityYear="1999" box="[966,1387,1260,1284]" class="Insecta" family="Ephemerellidae" genus="Torlyea" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="longforceps">
<emphasis id="B9228441FFB81F5E569EFB26D43A190A" box="[966,1118,1260,1284]" italics="true" pageId="0" pageNumber="89">T. longforceps</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E5136FB26D500190A" author="Gui, H. &amp; Zhou, C. - F. &amp; Su, C. - R." box="[1134,1380,1260,1284]" pageId="0" pageNumber="89" refId="ref4072" refString="Gui, H., Zhou, C. - F. &amp; Su, C. - R. (1999) Ephemeroptera. In: Huang, B. - K. (Ed.), Fujian Insect Fauna. Vol. 1. Fujian Science &amp; Technology Press, Fuzhou, pp. 324 - 346." type="book" year="1999">Gui, Zhou &amp; Su, 1999</bibRefCitation>
)
</taxonomicName>
was transferred from the genus
<taxonomicName id="4C5623D0FFB81F5E54E6FADAD2FA1926" authority="Edmunds." authorityName="Edmunds." box="[446,670,1296,1320]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E54E6FADAD24D1926" box="[446,553,1296,1320]" italics="true" pageId="0" pageNumber="89">Serratella</emphasis>
Edmunds.
</taxonomicName>
<taxonomicName id="4C5623D0FFB81F5E57FDFADAD31F1926" baseAuthorityName="Gui, Zhou &amp; Su" baseAuthorityYear="1999" box="[677,891,1296,1320]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="longforceps">
<emphasis id="B9228441FFB81F5E57FDFADAD31F1926" box="[677,891,1296,1320]" italics="true" pageId="0" pageNumber="89">Torleya longforceps</emphasis>
</taxonomicName>
was pleased in this genus on the basis of its male genitalia being similar to those of
<taxonomicName id="4C5623D0FFB81F5E575FFAFED2021942" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" box="[519,614,1332,1356]" class="Insecta" family="Ephemerellidae" genus="Torlyea" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="coheri">
<emphasis id="B9228441FFB81F5E575FFAFED2021942" box="[519,614,1332,1356]" italics="true" pageId="0" pageNumber="89">T. coheri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5623D0FFB81F5E572CFAFED2BC1942" baseAuthorityName="Jacobus, McCafferty &amp; Sites" baseAuthorityYear="2007" box="[628,728,1332,1356]" class="Insecta" family="Ephemerellidae" genus="Torlyea" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="lacuna">
<emphasis id="B9228441FFB81F5E572CFAFED2BC1942" box="[628,728,1332,1356]" italics="true" pageId="0" pageNumber="89">T. lacuna</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5623D0FFB81F5E564CFAFED3E21942" authorityName="Tiunova" authorityYear="1995" box="[788,902,1332,1356]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB81F5E564CFAFED3E21942" box="[788,902,1332,1356]" italics="true" pageId="0" pageNumber="89">T. mikhaili</emphasis>
</taxonomicName>
. At the same time,
<taxonomicName id="4C5623D0FFB81F5E5100FAFED5641942" authorityName="Zhou, Su &amp; Gui" authorityYear="2000" box="[1112,1280,1332,1356]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="grandipennis">
<emphasis id="B9228441FFB81F5E5100FAFED5641942" box="[1112,1280,1332,1356]" italics="true" pageId="0" pageNumber="89">T. grandipennis</emphasis>
</taxonomicName>
was moved to the genus
<taxonomicName id="4C5623D0FFB81F5E545BFA92D3ED197E" authority="Allen &amp; Edmunds (Jacobus &amp; McCafferty 2008)" authorityName="Allen &amp; Edmunds (Jacobus &amp; McCafferty" authorityYear="2008" box="[259,905,1368,1392]" class="Insecta" family="Ephemerellidae" genus="Hyrtanella" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E545BFA92D113197E" box="[259,375,1368,1392]" italics="true" pageId="0" pageNumber="89">Hyrtanella</emphasis>
Allen &amp; Edmunds (
<bibRefCitation id="EFC725A2FFB81F5E5714FA92D3E5197E" author="Jacobus, L. M. &amp; McCafferty, W. P." box="[588,897,1368,1392]" pageId="0" pageNumber="89" pagination="185 - 274" refId="ref4222" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2008) Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society, 134 (1 - 2), 185 - 274. https: // doi. org / 10.3157 / 0002 - 8320 (2008) 134 [185: ROEGE] 2.0. CO; 2" type="journal article" year="2008">Jacobus &amp; McCafferty 2008</bibRefCitation>
)
</taxonomicName>
. Currently, the genus
<taxonomicName id="4C5623D0FFB81F5E5128FA92D4A5197E" authorityName="Lestage" authorityYear="1917" box="[1136,1217,1368,1392]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E5128FA92D4A5197E" box="[1136,1217,1368,1392]" italics="true" pageId="0" pageNumber="89">Torleya</emphasis>
</taxonomicName>
includes 13 species. This includes two species based on larvae from
<collectingCountry id="F34118C3FFB81F5E57D4FAB6D2A6199A" box="[652,706,1404,1428]" name="India" pageId="0" pageNumber="89">India</collectingCountry>
—
<taxonomicName id="4C5623D0FFB81F5E5785FAB6D5F8199A" authority="Selvakumar, Subramanian, Martynov &amp; Jacobus" authorityName="Selvakumar, Subramanian, Martynov &amp; Jacobus" authorityYear="2021" box="[733,1436,1404,1428]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="simbalbarensis">
<emphasis id="B9228441FFB81F5E5785FAB6D3F3199A" box="[733,919,1404,1428]" italics="true" pageId="0" pageNumber="89">T. simbalbarensis</emphasis>
Selvakumar, Subramanian, Martynov &amp; Jacobus
</taxonomicName>
(in Martynov
<emphasis id="B9228441FFB81F5E5468FA6AD10519B6" box="[304,353,1440,1464]" italics="true" pageId="0" pageNumber="89">et al</emphasis>
.), 2021 and
<taxonomicName id="4C5623D0FFB81F5E54B6FA6AD49319B6" authority="Selvakumar, Martynov &amp; Jacobus (Martynov et al.) 2021" box="[494,1271,1440,1464]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="dibruensis">
<emphasis id="B9228441FFB81F5E54B6FA6AD21E19B6" box="[494,634,1440,1464]" italics="true" pageId="0" pageNumber="89">T. dibruensis</emphasis>
Selvakumar, Martynov &amp; Jacobus (Martynov
<emphasis id="B9228441FFB81F5E5122FA6AD4D619B6" box="[1146,1202,1440,1464]" italics="true" pageId="0" pageNumber="89">et al.</emphasis>
) 2021
</taxonomicName>
—and
<taxonomicName id="4C5623D0FFB81F5E501BFA6AD27919D2" authority="(Jacobus, Zhou &amp; McCafferty 2009)" baseAuthorityName="Jacobus, Zhou &amp; McCafferty" baseAuthorityYear="2009" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="B9228441FFB81F5E501BFA6AD5F819B6" box="[1347,1436,1440,1464]" italics="true" pageId="0" pageNumber="89">T. elissa</emphasis>
(
<bibRefCitation id="EFC725A2FFB81F5E55C7FA0ED27219D2" author="Jacobus, L. M. &amp; Zhou, C. - F. &amp; McCafferty, W. P." box="[159,534,1476,1500]" pageId="0" pageNumber="89" pagination="52 - 58" refId="ref4475" refString="Jacobus, L. M., Zhou, C. - F. &amp; McCafferty, W. P. (2009) Two new species of Asian Serratella Edmunds (Ephemeroptera: Ephemerellidae). Zootaxa, 2268, 1, 52 - 58. https: // doi. org / 10.11646 / zootaxa. 2268.1.4" type="journal article" year="2009">Jacobus, Zhou &amp; McCafferty 2009</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F34118C3FFB81F5E5706FA0ED2E819D5" box="[606,652,1476,1499]" name="Iran" pageId="0" pageNumber="89">Iran</collectingCountry>
, which was moved from
<taxonomicName id="4C5623D0FFB81F5E56C0FA0ED46719D2" box="[920,1027,1476,1500]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E56C0FA0ED46719D2" box="[920,1027,1476,1500]" italics="true" pageId="0" pageNumber="89">Serratella</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC725A2FFB81F5E5173FA0ED54519D2" author="Martynov, A. V. &amp; Palatov, D. M. &amp; Godunko, R. J." box="[1067,1313,1476,1500]" pageId="0" pageNumber="89" pagination="1 - 30" refId="ref4783" refString="Martynov, A. V., Palatov, D. M. &amp; Godunko, R. J. (2023) The tribe Hyrtanellini Allen, 1980 (Ephemeroptera: Ephemerellidae) of Western and Central Asia with description of a new species. Insects, 14, 87, 1 - 30. https: // doi. org / 10.3390 / insects 14010087" type="journal article" year="2023">
Martynov
<emphasis id="B9228441FFB81F5E51C3FA0ED4AE19D2" box="[1179,1226,1476,1500]" italics="true" pageId="0" pageNumber="89">et al</emphasis>
. (2023)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BE95853FFB81F5E559FFA22D33E1ABA" blockId="0.[151,1437,972,2040]" pageId="0" pageNumber="89">
The increased number of species in the genus
<taxonomicName id="4C5623D0FFB81F5E57FDFA22D2921A0E" authorityName="Lestage" authorityYear="1917" box="[677,758,1512,1536]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9228441FFB81F5E57FDFA22D2921A0E" box="[677,758,1512,1536]" italics="true" pageId="0" pageNumber="89">Torleya</emphasis>
</taxonomicName>
over the last thirty years has been associated with the description of new species and the movement of some species from other genera. Thus, a number of morphological characters that are necessary for a diagnosis of species are often absent from earlier descriptions (Jacobus
<emphasis id="B9228441FFB81F5E513BF9FAD4F61A46" box="[1123,1170,1584,1608]" italics="true" pageId="0" pageNumber="89">et al</emphasis>
. 2004). Considering that since the description of
<taxonomicName id="4C5623D0FFB81F5E54CEF99ED2631A62" authorityName="Tiunova" authorityYear="1995" box="[406,519,1620,1644]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB81F5E54CEF99ED2631A62" box="[406,519,1620,1644]" italics="true" pageId="0" pageNumber="89">T. mikhaili</emphasis>
</taxonomicName>
the author has collected new material of both larvae and adults from different regions of the Russian Far East, it was decided to present additional characteristics and original illustrations for larvae and adults, as well as to describe for the first time the female imago and eggs.
</paragraph>
</subSubSection>
<subSubSection id="C34C0BD8FFB81F5F559FF975D2CB1D15" lastPageId="1" lastPageNumber="90" pageId="0" pageNumber="89" type="materials_examined">
<paragraph id="8BE95853FFB81F5E559FF975D29A1AF2" blockId="0.[151,1437,972,2040]" pageId="0" pageNumber="89">
<emphasis id="B9228441FFB81F5E559FF975D1C21AD9" box="[199,422,1727,1751]" inLineHeading="true" pageId="0" pageNumber="89" reason="1">
<emphasis id="B9228441FFB81F5E559FF975D1FB1AD9" bold="true" box="[199,415,1727,1751]" pageId="0" pageNumber="89">Material examined</emphasis>
.
</emphasis>
<materialsCitation id="3B3E520EFFB81F5E54F4F90AD29A1AF2" collectingDate="1992-08-02" collectorName="T. Tiunova." country="Russia" location="Primorye Territory" municipality="Ussuri River" pageId="0" pageNumber="89" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F34118C3FFB81F5E54F4F90AD2191AD6" box="[428,637,1728,1752]" name="Russia" pageId="0" pageNumber="89">Russian Federation</collectingCountry>
:
<emphasis id="B9228441FFB81F5E57DFF90AD3351AD9" box="[647,849,1728,1751]" italics="true" pageId="0" pageNumber="89">
<location id="8E890E88FFB81F5E57DFF90AD3351AD9" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E57DFF90AD3351AD9" box="[647,849,1728,1751]" country="Russia" municipality="Ussuri River" name="Primorye Territory" pageId="0" pageNumber="89">Primorye Territory</location>
</emphasis>
:
<emphasis id="B9228441FFB81F5E5607F90AD3AA1AD6" bold="true" box="[863,974,1728,1752]" pageId="0" pageNumber="89">
<typeStatus id="54EDE6F1FFB81F5E5607F90AD3AD1AD6" box="[863,969,1728,1752]" pageId="0" pageNumber="89" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D5093DAFFB81F5E568CF90AD38C1AD9" box="[980,1000,1728,1751]" count="1" pageId="0" pageNumber="89" type="male">♂</specimenCount>
imago (reared from larva).
<collectingMunicipality id="6B8DC229FFB81F5E5057F90AD5FC1AD6" box="[1295,1432,1728,1752]" pageId="0" pageNumber="89">Ussuri River</collectingMunicipality>
,
<locationDeviation id="9A5B4171FFB81F5E55CFF92ED1841AF2" box="[151,480,1764,1788]" location="Utyos Mountain" pageId="0" pageNumber="89">
<quantity id="4CAEF5B6FFB81F5E55CFF92ED0851AF2" box="[151,225,1764,1788]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="89" unit="km" value="1.5">1.5 km</quantity>
below Utyos Mountain
</locationDeviation>
,
<date id="FFE87E93FFB81F5E54B3F92ED21D1AF2" box="[491,633,1764,1788]" pageId="0" pageNumber="89" value="1992-08-02">
<collectingDate id="EFAC877BFFB81F5E54B3F92ED21D1AF2" box="[491,633,1764,1788]" pageId="0" pageNumber="89" value="1992-08-02">02.VIII.1992</collectingDate>
</date>
,
<collectorName id="26A33D85FFB81F5E57DBF92ED29A1AF2" box="[643,766,1764,1788]" pageId="0" pageNumber="89">T. Tiunova.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BE95853FFB81F5F559FF8C2D2CB1D15" blockId="0.[151,1437,972,2040]" lastBlockId="1.[151,1437,151,572]" lastPageId="1" lastPageNumber="90" pageId="0" pageNumber="89">
<emphasis id="B9228441FFB81F5E559FF8C2D1241B2E" bold="true" box="[199,320,1800,1824]" inLineHeading="true" pageId="0" pageNumber="89" reason="1">
<typeStatus id="54EDE6F1FFB81F5E559FF8C2D15F1B2E" box="[199,315,1800,1824]" pageId="0" pageNumber="89" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D5093DAFFB81F5E5411F8C2D1151B2E" box="[329,369,1800,1824]" count="5" pageId="0" pageNumber="89" type="male">5♂</specimenCount>
<specimenCount id="9D5093DAFFB81F5E5429F8C2D1E81B2E" box="[369,396,1800,1824]" count="1" pageId="0" pageNumber="89" type="female">1♀</specimenCount>
imagoes,
<specimenCount id="9D5093DAFFB81F5E54A7F8C2D27A1B2E" box="[511,542,1800,1824]" count="5" pageId="0" pageNumber="89" type="male">5♂</specimenCount>
<specimenCount id="9D5093DAFFB81F5E5746F8C2D2501B2E" box="[542,564,1800,1824]" count="1" pageId="0" pageNumber="89" type="female">1♀</specimenCount>
subimagoes, all reared from larvae,
<materialsCitation id="3B3E520EFFB81F5E5696F8C2D1FB1B4A" collectingDate="1994-08-04" location="Ussuri River" pageId="0" pageNumber="89" specimenCount="4" specimenCount-larva="4" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E5696F8C2D44D1B2E" box="[974,1065,1800,1824]" count="4" pageId="0" pageNumber="89" type="larva">4 larvae</specimenCount>
,
<location id="8E890E88FFB81F5E516EF8C2D4A61B2E" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E516EF8C2D4A61B2E" box="[1078,1218,1800,1824]" name="Ussuri River" pageId="0" pageNumber="89">Ussuri River</location>
,
<locationDeviation id="9A5B4171FFB81F5E5188F8C2D1651B4A" location="Utyos Mountain" pageId="0" pageNumber="89">
<quantity id="4CAEF5B6FFB81F5E5188F8C2D56C1B2E" box="[1232,1288,1800,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="89" unit="km" value="1.0">1 km</quantity>
below Utyos Mountain
</locationDeviation>
,
<date id="FFE87E93FFB81F5E5448F8E6D1FB1B4A" box="[272,415,1836,1860]" pageId="0" pageNumber="89" value="1994-08-04">
<collectingDate id="EFAC877BFFB81F5E5448F8E6D1FB1B4A" box="[272,415,1836,1860]" pageId="0" pageNumber="89" value="1994-08-04">04.VIII.1994</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5093DAFFB81F5E54F5F8E6D1AA1B4A" box="[429,462,1836,1860]" count="5" pageId="0" pageNumber="89" type="male">5♂</specimenCount>
imagoes,
<specimenCount id="9D5093DAFFB81F5E571BF8E6D2001B4A" box="[579,612,1836,1860]" count="2" pageId="0" pageNumber="89" type="male">2♂</specimenCount>
subimagoes, all reared from larvae,
<materialsCitation id="3B3E520EFFB81F5E5158F8E6D3B31B66" collectingDate="1996-07-18" collectorName="Vladivostok-Khabarovsk" location="Gornye Kluchi Village" municipality="Ussuri River" pageId="0" pageNumber="89" specimenCount="1" specimenCount-larva="1" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E5158F8E6D4341B4A" box="[1024,1104,1836,1860]" count="1" pageId="0" pageNumber="89" type="larva">1 larva</specimenCount>
,
<collectingMunicipality id="6B8DC229FFB81F5E5106F8E6D48F1B4A" box="[1118,1259,1836,1860]" pageId="0" pageNumber="89">Ussuri River</collectingMunicipality>
,
<location id="8E890E88FFB81F5E51A1F8E6D0831B66" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E51A1F8E6D0831B66" municipality="Ussuri River" name="Gornye Kluchi Village" pageId="0" pageNumber="89">Gornye Kluchi Village</location>
,
<quantity id="4CAEF5B6FFB81F5E55ADF89AD14A1B66" box="[245,302,1872,1896]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="89" unit="km" value="2.0">2 km</quantity>
below bridge, highway
<collectorName id="26A33D85FFB81F5E5764F89AD32C1B66" box="[572,840,1872,1896]" pageId="0" pageNumber="89">Vladivostok-Khabarovsk</collectorName>
<date id="FFE87E93FFB81F5E560AF89AD3B31B66" box="[850,983,1872,1896]" pageId="0" pageNumber="89" value="1996-07-18">
<collectingDate id="EFAC877BFFB81F5E560AF89AD3B31B66" box="[850,983,1872,1896]" pageId="0" pageNumber="89" value="1996-07-18">18.VII.1996</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5093DAFFB81F5E56BDF89AD4631B66" box="[997,1031,1872,1896]" count="1" pageId="0" pageNumber="89" type="male">1♂</specimenCount>
imago,1 larval skin
<specimenCount id="9D5093DAFFB81F5E51B6F89AD5661B69" box="[1262,1282,1872,1895]" count="1" pageId="0" pageNumber="89" type="female">♀</specimenCount>
, 1 larval skin
<materialsCitation id="3B3E520EFFB81F5E55CFF8BED3B61B82" box="[151,978,1908,1932]" collectingDate="1996-07-20" location="Bolshaya Ussurka River" pageId="0" pageNumber="89" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E55CFF8BED0C91B85" box="[151,173,1908,1931]" count="1" pageId="0" pageNumber="89" type="male">♂</specimenCount>
,
<location id="8E890E88FFB81F5E55E2F8BED1A51B82" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E55E2F8BED1A51B82" box="[186,449,1908,1932]" name="Bolshaya Ussurka River" pageId="0" pageNumber="89">Bolshaya Ussurka River</location>
,
<locationDeviation id="9A5B4171FFB81F5E5494F8BED3251B82" box="[460,833,1908,1932]" location="Zvenigorogka Village" pageId="0" pageNumber="89">
<quantity id="4CAEF5B6FFB81F5E5494F8BED2601B82" box="[460,516,1908,1932]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="89" unit="km" value="1.0">1 km</quantity>
below Zvenigorogka Village
</locationDeviation>
,
<date id="FFE87E93FFB81F5E5614F8BED3B61B82" box="[844,978,1908,1932]" pageId="0" pageNumber="89" value="1996-07-20">
<collectingDate id="EFAC877BFFB81F5E5614F8BED3B61B82" box="[844,978,1908,1932]" pageId="0" pageNumber="89" value="1996-07-20">20.VII.1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB81F5E5686F8BED4341BBE" collectingDate="1996-07-20" collectorName="Vladivostok-Khabarovsk" elevation="50" location="Malinovka River" municipality="Bolshaya Ussurka River" pageId="0" pageNumber="89" specimenCount="13" specimenCount-larva="13" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E5686F8BED4201B82" box="[990,1092,1908,1932]" count="13" pageId="0" pageNumber="89" type="larva">13 larvae</specimenCount>
,
<collectingMunicipality id="6B8DC229FFB81F5E5108F8BED5311B82" box="[1104,1365,1908,1932]" pageId="0" pageNumber="89">Bolshaya Ussurka River</collectingMunicipality>
basin,
<location id="8E890E88FFB81F5E55CFF852D1371BBE" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E55CFF852D1371BBE" box="[151,339,1944,1968]" municipality="Bolshaya Ussurka River" name="Malinovka River" pageId="0" pageNumber="89">Malinovka River</location>
,
<quantity id="4CAEF5B6FFB81F5E5439F852D1F81BA1" box="[353,412,1944,1968]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="89" unit="m" value="50.0">
<elevation id="007BBF60FFB81F5E5439F852D1F81BA1" box="[353,412,1944,1968]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="89" unit="m" value="50.0">50 m</elevation>
</quantity>
above bridge, highway
<collectorName id="26A33D85FFB81F5E57F5F852D3D81BBE" box="[685,956,1944,1968]" pageId="0" pageNumber="89">Vladivostok-Khabarovsk</collectorName>
,
<date id="FFE87E93FFB81F5E5692F852D4341BBE" box="[970,1104,1944,1968]" pageId="0" pageNumber="89" value="1996-07-20">
<collectingDate id="EFAC877BFFB81F5E5692F852D4341BBE" box="[970,1104,1944,1968]" pageId="0" pageNumber="89" value="1996-07-20">20.VII.1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB81F5E5107F852D3D91BDA" collectingDate="2007-08-05" collectorName="Vladivostok-Khabarovsk" elevation="700" location="Ilistaya River" pageId="0" pageNumber="89" specimenCount="8" specimenCount-male="8" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E5107F852D4EF1BA1" box="[1119,1163,1944,1968]" count="8" pageId="0" pageNumber="89" type="male">8 ♂</specimenCount>
imagoes (light),
<location id="8E890E88FFB81F5E5016F852D0B01BDA" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E5016F852D0B01BDA" name="Ilistaya River" pageId="0" pageNumber="89">Ilistaya River</location>
,
<quantity id="4CAEF5B6FFB81F5E5586F876D1461BDD" box="[222,290,1980,2004]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="89" unit="m" value="700.0">
<elevation id="007BBF60FFB81F5E5586F876D1461BDD" box="[222,290,1980,2004]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="89" unit="m" value="700.0">700 m</elevation>
</quantity>
below bridge, highway
<collectorName id="26A33D85FFB81F5E577BF876D3561BDA" box="[547,818,1980,2004]" pageId="0" pageNumber="89">Vladivostok-Khabarovsk</collectorName>
,
<date id="FFE87E93FFB81F5E5664F876D3D91BDA" box="[828,957,1980,2004]" pageId="0" pageNumber="89" value="2007-08-05">
<collectingDate id="EFAC877BFFB81F5E5664F876D3D91BDA" box="[828,957,1980,2004]" pageId="0" pageNumber="89" value="2007-08-05">5.VIII.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB81F5E5690F876D1961BF6" collectingDate="2007-08-04" location="Zarechnoe Village" municipality="Razdol'naya River" pageId="0" pageNumber="89" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E5690F876D38D1BDA" box="[968,1001,1980,2004]" count="1" pageId="0" pageNumber="89" type="male">1♂</specimenCount>
imago (light),
<collectingMunicipality id="6B8DC229FFB81F5E51D0F876D5361BDA" box="[1160,1362,1980,2004]" pageId="0" pageNumber="89">Razdol’naya River</collectingMunicipality>
, above
<location id="8E890E88FFB81F5E55CFF82AD1071BF6" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB81F5E55CFF82AD1071BF6" box="[151,355,2016,2040]" municipality="Razdol'naya River" name="Zarechnoe Village" pageId="0" pageNumber="89">Zarechnoe Village</location>
,
<date id="FFE87E93FFB81F5E5429F82AD1961BF6" box="[369,498,2016,2040]" pageId="0" pageNumber="89" value="2007-08-04">
<collectingDate id="EFAC877BFFB81F5E5429F82AD1961BF6" box="[369,498,2016,2040]" pageId="0" pageNumber="89" value="2007-08-04">4.VIII.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB81F5F575AF82AD21E1CA1" collectingDate="2002-08-14" collectingDateMax="2002-08-15" collectingDateMin="2002-08-14" county="Razdol'naya River" lastPageId="1" lastPageNumber="90" location="Kraunovka River" municipality="Medveditsa River" pageId="0" pageNumber="89" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D5093DAFFB81F5E575AF82AD2471BF6" box="[514,547,2016,2040]" count="2" pageId="0" pageNumber="89" type="male">2♂</specimenCount>
imagoes (light),
<collectingCounty id="628820DFFFB81F5E57BFF82AD3D01BF6" box="[743,948,2016,2040]" pageId="0" pageNumber="89">Razdol’naya River</collectingCounty>
Basin,
<collectingMunicipality id="6B8DC229FFB81F5E5157F82AD4B51BF6" box="[1039,1233,2016,2040]" pageId="0" pageNumber="89">Medveditsa River</collectingMunicipality>
at the confluence with the
<location id="8E890E88FFB91F5F55A1FF5DD1D21CA1" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB91F5F55A1FF5DD1D21CA1" box="[249,438,151,175]" county="Razdol'naya River" municipality="Medveditsa River" name="Kraunovka River" pageId="1" pageNumber="90">Kraunovka River</location>
,
<date id="FFE87E93FFB91F5F549BFF5DD21E1CA1" box="[451,634,151,175]" pageId="1" pageNumber="90" value="2002-08-14" valueMax="2002-08-15" valueMin="2002-08-14">
<collectingDate id="EFAC877BFFB91F5F549BFF5DD21E1CA1" box="[451,634,151,175]" pageId="1" pageNumber="90" value="2002-08-14" valueMax="2002-08-15" valueMin="2002-08-14">14–15.VIII.2002</collectingDate>
</date>
</materialsCitation>
;
<emphasis id="B9228441FFB91F5F57DFFF5DD30B1CA1" box="[647,879,151,175]" italics="true" pageId="1" pageNumber="90">
<collectingRegion id="499296B1FFB91F5F57DFFF5DD3631CA1" box="[647,775,151,175]" country="Russia" name="Khabarovsk" pageId="1" pageNumber="90">Khabarovsk</collectingRegion>
Territory
</emphasis>
:
<materialsCitation id="3B3E520EFFB91F5F56D8FF52D2571CDD" collectingDate="1996-07-22" county="River Basin" location="Zvenevoy Village" municipality="Bikin River" pageId="1" pageNumber="90" specimenCount="1" specimenCount-male="1" stateProvince="Amur" typeStatus="paratype">
<specimenCount id="9D5093DAFFB91F5F56D8FF52D3C51CBE" box="[896,929,152,176]" count="1" pageId="1" pageNumber="90" type="male">1♂</specimenCount>
(reared from larva),
<collectingRegion id="499296B1FFB91F5F51D1FF52D4A31CBE" box="[1161,1223,152,176]" country="Russia" name="Amur" pageId="1" pageNumber="90">Amur</collectingRegion>
<collectingCounty id="628820DFFFB91F5F5188FF5DD5371CA1" box="[1232,1363,151,175]" pageId="1" pageNumber="90">River Basin</collectingCounty>
,
<collectingMunicipality id="6B8DC229FFB91F5F5039FF5DD0B01CDD" pageId="1" pageNumber="90">Bikin River</collectingMunicipality>
,
<location id="8E890E88FFB91F5F55B8FF76D1C61CDD" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB91F5F55B8FF76D1C61CDD" box="[224,418,187,212]" county="River Basin" municipality="Bikin River" name="Zvenevoy Village" pageId="1" pageNumber="90" stateProvince="Amur">Zvenevoy Village</location>
,
<date id="FFE87E93FFB91F5F54F6FF71D2571CDD" box="[430,563,187,211]" pageId="1" pageNumber="90" value="1996-07-22">
<collectingDate id="EFAC877BFFB91F5F54F6FF71D2571CDD" box="[430,563,187,211]" pageId="1" pageNumber="90" value="1996-07-22">22.VII.1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB91F5F5767FF76D1791CF6" collectingDate="2000-07-13" collectorName="Vladivostok-Khabarovsk" elevation="300" location="Bikin River" pageId="1" pageNumber="90" specimenCount="13" specimenCount-larva="13" stateProvince="Amur" typeStatus="paratype">
<specimenCount id="9D5093DAFFB91F5F5767FF76D2C21CDD" box="[575,678,187,212]" count="13" pageId="1" pageNumber="90" type="larva">13 larvae</specimenCount>
,
<location id="8E890E88FFB91F5F57EAFF71D3561CDD" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB91F5F57EAFF71D3561CDD" box="[690,818,187,211]" name="Bikin River" pageId="1" pageNumber="90" stateProvince="Amur">Bikin River</location>
,
<quantity id="4CAEF5B6FFB91F5F5666FF76D3E71CDD" box="[830,899,188,212]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="90" unit="m" value="300.0">
<elevation id="007BBF60FFB91F5F5666FF76D3E71CDD" box="[830,899,188,212]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="90" unit="m" value="300.0">300 m</elevation>
</quantity>
below bridge, highway
<collectorName id="26A33D85FFB91F5F51D1FF71D5FC1CDD" box="[1161,1432,187,211]" pageId="1" pageNumber="90">Vladivostok-Khabarovsk</collectorName>
,
<date id="FFE87E93FFB91F5F55CFFF2AD1791CF6" box="[151,285,224,248]" pageId="1" pageNumber="90" value="2000-07-13">
<collectingDate id="EFAC877BFFB91F5F55CFFF2AD1791CF6" box="[151,285,224,248]" pageId="1" pageNumber="90" value="2000-07-13">13.VII.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E520EFFB91F5F5477FF2AD1791D15" collectingDate="1996-07-22" collectorName="Khabarovsk-Komsomolsk" location="Gur River" municipality="River Basin" pageId="1" pageNumber="90" specimenCount="1" specimenCount-male="1" stateProvince="Amur" typeStatus="paratype">
<specimenCount id="9D5093DAFFB91F5F5477FF2AD1351CF6" box="[303,337,224,248]" count="1" pageId="1" pageNumber="90" type="male">1♂</specimenCount>
(reared from larva),
<collectingRegion id="499296B1FFB91F5F5714FF2AD2EE1CF6" box="[588,650,224,248]" country="Russia" name="Amur" pageId="1" pageNumber="90">Amur</collectingRegion>
<collectingMunicipality id="6B8DC229FFB91F5F57C1FF15D3451CF9" box="[665,801,223,247]" pageId="1" pageNumber="90">River Basin</collectingMunicipality>
,
<location id="8E890E88FFB91F5F566CFF2AD3CC1CF9" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB91F5F566CFF2AD3CC1CF9" box="[820,936,223,248]" municipality="River Basin" name="Gur River" pageId="1" pageNumber="90" stateProvince="Amur">Gur River</location>
, bridge, highway
<collectorName id="26A33D85FFB91F5F5126FF15D5FD1CF9" box="[1150,1433,223,247]" pageId="1" pageNumber="90">Khabarovsk-Komsomolsk</collectorName>
,
<date id="FFE87E93FFB91F5F55CFFEC9D1791D15" box="[151,285,259,283]" pageId="1" pageNumber="90" value="1996-07-22">
<collectingDate id="EFAC877BFFB91F5F55CFFEC9D1791D15" box="[151,285,259,283]" pageId="1" pageNumber="90" value="1996-07-22">22.VII.1996</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B3E520EFFB91F5F547DFEC9D2CB1D15" box="[293,687,259,283]" collectorName="T. Tiunova." location="All" pageId="1" pageNumber="90" specimenCount="1" stateProvince="Amur" typeStatus="paratype">
<location id="8E890E88FFB91F5F547DFEC9D1231D15" LSID="urn:lsid:plazi:treatment:03FFE945FFB81F5655CFFC06D2E91BF4:8E890E88FFB91F5F547DFEC9D1231D15" box="[293,327,259,283]" name="All" pageId="1" pageNumber="90" stateProvince="Amur">All</location>
material collected by
<collectorName id="26A33D85FFB91F5F576CFECED2CB1D15" box="[564,687,259,283]" pageId="1" pageNumber="90">T. Tiunova.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C0BD8FFB91F5F559FFEEDD4DF1E35" pageId="1" pageNumber="90" type="diagnosis">
<paragraph id="8BE95853FFB91F5F559FFEEDD4DF1E35" blockId="1.[151,1437,151,572]" pageId="1" pageNumber="90">
<emphasis id="B9228441FFB91F5F559FFEEDD1581D31" bold="true" box="[199,316,295,319]" inLineHeading="true" pageId="1" pageNumber="90" reason="1">Diagnosis.</emphasis>
<taxonomicName id="4C5623D0FFB91F5F5419FEEDD18B1D31" authorityName="Tiunova" authorityYear="1995" box="[321,495,295,319]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB91F5F5419FEEDD18B1D31" box="[321,495,295,319]" italics="true" pageId="1" pageNumber="90">Torleya mikhaili</emphasis>
</taxonomicName>
can be separated from the other species of the genus by the combination of the following characteristics.
<emphasis id="B9228441FFB91F5F5463FE81D1DD1D6D" box="[315,441,331,355]" italics="true" pageId="1" pageNumber="90">Male imago</emphasis>
: (1) forceps segment II which has a slight distal expansion; (2) penis with elongated apical lobes rounded at the apex; (3) U-shape a median notch; (4) genital forceps segment III relatively short, length approximately 1.5–1.7 times the width.
<emphasis id="B9228441FFB91F5F54FDFE5ED1951DA5" box="[421,497,404,427]" italics="true" pageId="1" pageNumber="90">Larvae</emphasis>
: (5) posterior margins of abdominal terga without paired tubercles; (6) forefemur with distal transverse row of long, stout and pointed setae; length of these setae ½ of width of femur; (7) claw of fore leg with two groups of denticles; apical group with four acute and long denticles; subdistal setae absent; (8) posterolateral projection on terga IV–IX well developed; (9) posterior margins of terga IV–IX with paired submedian short, clavate setae.
<emphasis id="B9228441FFB91F5F5582FDEED16B1E35" box="[218,271,548,571]" italics="true" pageId="1" pageNumber="90">Eggs</emphasis>
: (10) the ridges about 2–3 µm in width; (11) the surface of the ridges relatively smooth.
</paragraph>
</subSubSection>
<subSubSection id="C34C0BD8FFB91F5655CFFD47D2E91BF4" lastPageId="8" lastPageNumber="97" pageId="1" pageNumber="90" type="description">
<paragraph id="8BE95853FFB91F5F55CFFD47D1211EA9" blockId="1.[151,1437,653,1940]" box="[151,325,653,679]" pageId="1" pageNumber="90">
<heading id="D0A1EF3FFFB91F5F55CFFD47D1211EA9" bold="true" box="[151,325,653,679]" fontSize="11" level="1" pageId="1" pageNumber="90" reason="1">
<emphasis id="B9228441FFB91F5F55CFFD47D1211EA9" bold="true" box="[151,325,653,679]" pageId="1" pageNumber="90">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BE95853FFB91F5F55CFFD79D4C01FAA" blockId="1.[151,1437,653,1940]" pageId="1" pageNumber="90">
<emphasis id="B9228441FFB91F5F55CFFD79D17C1EC5" box="[151,280,691,715]" italics="true" pageId="1" pageNumber="90">Male imago</emphasis>
(in alcohol) (
<figureCitation id="136D44D6FFB91F5F54F1FD79D18C1EC2" box="[425,488,691,716]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[167,1421,181,1919]" captionTargetId="figure-19@2.[167,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Torleya mikhaili, paratypes (material from type locality), lateral view: 1, imago male; 2, imago female; 2A, subanal plate female." figureDoi="http://doi.org/10.5281/zenodo.10718142" httpUri="https://zenodo.org/record/10718142/files/figure.png" pageId="1" pageNumber="90">Fig. 1</figureCitation>
). Length (mm): body, 4.7–5.6; fore wings, 4.6–5.6; caudal filaments, 4.5–5.3. Length of segments in foreleg (mm): femora 0.95–1.0, tibia 2.3–2.6, tarsal segments 0.45–0.5, 0.35–0.4, 0.2–0.25, 0.15–0.2, 0.1. Abdominal terga I–IV or I–VII dirty brownish, translucent; tergum V–IX or VII–IX light brown, opaque (
<figureCitation id="136D44D6FFB91F5F505AFD31D5251F1A" box="[1282,1345,763,788]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[167,1421,181,1919]" captionTargetId="figure-19@2.[167,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Torleya mikhaili, paratypes (material from type locality), lateral view: 1, imago male; 2, imago female; 2A, subanal plate female." figureDoi="http://doi.org/10.5281/zenodo.10718142" httpUri="https://zenodo.org/record/10718142/files/figure.png" pageId="1" pageNumber="90">Fig. 1</figureCitation>
). Wings hyaline with translucent milky tinge along costal margin; longitudinal veins, intercalaries, and crossveins hyaline (
<figureCitation id="136D44D6FFB91F5F5035FCD5D0A71F55" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1675,1699]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-15@3.[151,1436,181,1651]" captionTargetPageId="3" captionText="FIGURES 3–7. Imago male of Torleya mikhaili, paratypes (material from type locality): 3, genitalia, ventral; 4, genitalia, dorsal; 5, penis lobes, dorsal; 6, hind wing; 7, fore wing." figureDoi="http://doi.org/10.5281/zenodo.10718144" httpUri="https://zenodo.org/record/10718144/files/figure.png" pageId="1" pageNumber="90">Figs 6–7</figureCitation>
). Second segment of genital forceps with slight distal expansion (
<figureCitation id="136D44D6FFB91F5F5621FC89D3BE1F52" box="[889,986,835,860]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1675,1699]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-15@3.[151,1436,181,1651]" captionTargetPageId="3" captionText="FIGURES 3–7. Imago male of Torleya mikhaili, paratypes (material from type locality): 3, genitalia, ventral; 4, genitalia, dorsal; 5, penis lobes, dorsal; 6, hind wing; 7, fore wing." figureDoi="http://doi.org/10.5281/zenodo.10718144" httpUri="https://zenodo.org/record/10718144/files/figure.png" pageId="1" pageNumber="90">Figs 3–4</figureCitation>
); third segment length approximately 1.7x width. Penis lobes slightly elongated and rounded at apex and with U-shaped, shallow and relatively wide median notch (
<figureCitation id="136D44D6FFB91F5F55C7FC41D1641FAA" box="[159,256,907,932]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1675,1699]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-15@3.[151,1436,181,1651]" captionTargetPageId="3" captionText="FIGURES 3–7. Imago male of Torleya mikhaili, paratypes (material from type locality): 3, genitalia, ventral; 4, genitalia, dorsal; 5, penis lobes, dorsal; 6, hind wing; 7, fore wing." figureDoi="http://doi.org/10.5281/zenodo.10718144" httpUri="https://zenodo.org/record/10718144/files/figure.png" pageId="1" pageNumber="90">Figs 3–4</figureCitation>
); penis lobes with small subapical projection (
<figureCitation id="136D44D6FFB91F5F57B6FC41D3281FAD" box="[750,844,907,931]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1675,1699]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-15@3.[151,1436,181,1651]" captionTargetPageId="3" captionText="FIGURES 3–7. Imago male of Torleya mikhaili, paratypes (material from type locality): 3, genitalia, ventral; 4, genitalia, dorsal; 5, penis lobes, dorsal; 6, hind wing; 7, fore wing." figureDoi="http://doi.org/10.5281/zenodo.10718144" httpUri="https://zenodo.org/record/10718144/files/figure.png" pageId="1" pageNumber="90">Fig. 4–5</figureCitation>
). Caudal filaments pale (
<figureCitation id="136D44D6FFB91F5F510DFC41D4F01FAA" box="[1109,1172,907,932]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[167,1421,181,1919]" captionTargetId="figure-19@2.[167,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Torleya mikhaili, paratypes (material from type locality), lateral view: 1, imago male; 2, imago female; 2A, subanal plate female." figureDoi="http://doi.org/10.5281/zenodo.10718142" httpUri="https://zenodo.org/record/10718142/files/figure.png" pageId="1" pageNumber="90">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BE95853FFB91F5F559FFC65D1E3183D" blockId="1.[151,1437,653,1940]" pageId="1" pageNumber="90">
<emphasis id="B9228441FFB91F5F559FFC65D1041FC9" box="[199,352,943,967]" italics="true" pageId="1" pageNumber="90">Female imago</emphasis>
(in alcohol, reared from larva) (
<figureCitation id="136D44D6FFB91F5F57ECFC65D2911FC6" box="[692,757,943,968]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[167,1421,181,1919]" captionTargetId="figure-19@2.[167,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Torleya mikhaili, paratypes (material from type locality), lateral view: 1, imago male; 2, imago female; 2A, subanal plate female." figureDoi="http://doi.org/10.5281/zenodo.10718142" httpUri="https://zenodo.org/record/10718142/files/figure.png" pageId="1" pageNumber="90">Fig. 2</figureCitation>
). Length (mm): body 4.7; fore wings, 5.0; caudal filaments, 4.5. Head, thorax and abdomen yellowish brown; thorax somewhat darker than abdomen. Wings hyaline, all veins pale. Legs pale. Abdomen with eggs yellowish or brownish; abdominal sterna yellowish. Subanal plate triangular with rounded apex. Caudal filaments pale.
</paragraph>
<paragraph id="8BE95853FFB91F5F559FFBF5D5E21859" blockId="1.[151,1437,653,1940]" box="[199,1414,1087,1112]" pageId="1" pageNumber="90">
<emphasis id="B9228441FFB91F5F559FFBF5D1091859" box="[199,365,1087,1111]" italics="true" pageId="1" pageNumber="90">Male subimago</emphasis>
(in alcohol) (
<figureCitation id="136D44D6FFB91F5F54A5FBF5D25A1856" box="[509,574,1087,1112]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 8</figureCitation>
). Similar to male imago, but color of abdomen uniformly yellowish-brownish.
</paragraph>
<paragraph id="8BE95853FFB91F5F559FFBA9D30A1B9D" blockId="1.[151,1437,653,1940]" pageId="1" pageNumber="90">
<emphasis id="B9228441FFB91F5F559FFBA9D1E81875" box="[199,396,1123,1147]" italics="true" pageId="1" pageNumber="90">Final larval instar</emphasis>
(in alcohol) (
<figureCitation id="136D44D6FFB91F5F5745FBA9D23B1872" box="[541,607,1123,1148]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Length (mm): body, 3.8–5.5; caudal filaments: 1.4–2.3. General colouration light brown with brown markings (
<figureCitation id="136D44D6FFB91F5F574DFB4DD23C18AE" box="[533,600,1159,1184]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Head with pair of long hairlike setae on occiput (
<figureCitation id="136D44D6FFB91F5F51D0FB4DD4BD18AE" box="[1160,1241,1159,1184]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Fig. 10</figureCitation>
). Mouthparts (see also Tiunova 1955: figs 3–5): clypeus with anterior row of long, stout, hairlike setae (
<figureCitation id="136D44D6FFB91F5F5169FB61D4E518CA" box="[1073,1153,1195,1220]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Fig. 10</figureCitation>
); dorsal surface of labrum densely covered with long, fine setae (
<figureCitation id="136D44D6FFB91F5F576FFB05D2EC18E6" box="[567,648,1231,1256]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Fig. 10</figureCitation>
). Maxilla without palp (see Tiunova 1955: fig. 4). Pronotum without any protuberances (
<figureCitation id="136D44D6FFB91F5F5466FB39D1E71902" box="[318,387,1267,1292]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Forefemur with distal transverse row of long, stout and pointed setae (
<figureCitation id="136D44D6FFB91F5F51FDFB39D5471902" box="[1189,1315,1267,1292]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Figs 11–13</figureCitation>
); length of these setae ½ of width of femur (
<figureCitation id="136D44D6FFB91F5F54ACFADDD20B193E" box="[500,623,1303,1328]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Figs 11–12</figureCitation>
); mid and hind femora each with longitudinal, sparse row of long thick setae and long thin hairlike setae along dorsal and ventral margins (
<figureCitation id="136D44D6FFB91F5F5677FAF1D316195A" box="[815,882,1339,1364]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Fore tibia with row of stout, brown setae and pale hairlike setae along ventral margin; mid and hind tibia and tarsi with long, thin hairs along dorsal and ventral margins (
<figureCitation id="136D44D6FFB91F5F55C7FA49D0861992" box="[159,226,1411,1436]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Claw with two groups of denticles; apical group with four acute and long denticles, gradually longer apically; basal group with four or five blunt denticles, middle denticles largest; subdistal setae absent (
<figureCitation id="136D44D6FFB91F5F51DAFA6DD4B719CE" box="[1154,1235,1447,1472]" captionStart="FIGURES 10–14" captionStartId="5.[151,264,1865,1889]" captionTargetBox="[151,1435,181,1841]" captionTargetId="figure-15@5.[151,1435,181,1841]" captionTargetPageId="5" captionText="FIGURES 10–14. Larvae of Torleya mikhaili, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput." figureDoi="http://doi.org/10.5281/zenodo.10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="1" pageNumber="90">Fig. 14</figureCitation>
). Lengths (mm) of leg segments. Fore leg: femur 0.7–0.75; tibia 0.5–0.55; and tarsus 0.45–0.5. Middle leg: femur 0.75–0.85; tibia 0.5–0.6; and tarsus 0.45–0.5. Hind leg: femur 0.85–0.9; tibia 0.6–0.75; and tarsus 0.5–0.6. Posterior margins of terga IV–IX with paired submedian short, clavate setae, which may be single on terga IV and IX, tuft of two or three setae on terga V–VII, and row of three or four setae on tergum VIII which continues to lateral margin (
<figureCitation id="136D44D6FFB91F5F513CF9FDD4D11A41" box="[1124,1205,1591,1615]" captionStart="FIGURES 15–18" captionStartId="6.[151,264,1683,1707]" captionTargetBox="[151,1436,181,1659]" captionTargetId="figure-19@6.[151,1436,181,1659]" captionTargetPageId="6" captionText="FIGURES 15–18. Larvae of Torleya mikhaili: 15, abdomen, dorsal; 16, submedian setae on posterior margins of terga; 17, abdominal sternum IX and caudal filaments; 18, setae on caudal filaments." figureDoi="http://doi.org/10.5281/zenodo.10718156" httpUri="https://zenodo.org/record/10718156/files/figure.png" pageId="1" pageNumber="90">Fig. 15</figureCitation>
); terga II–III without posterolateral projections; posterolateral projections on terga IV–IX well developed (
<figureCitation id="136D44D6FFB91F5F5162F991D4E81A7D" box="[1082,1164,1627,1651]" captionStart="FIGURES 15–18" captionStartId="6.[151,264,1683,1707]" captionTargetBox="[151,1436,181,1659]" captionTargetId="figure-19@6.[151,1436,181,1659]" captionTargetPageId="6" captionText="FIGURES 15–18. Larvae of Torleya mikhaili: 15, abdomen, dorsal; 16, submedian setae on posterior margins of terga; 17, abdominal sternum IX and caudal filaments; 18, setae on caudal filaments." figureDoi="http://doi.org/10.5281/zenodo.10718156" httpUri="https://zenodo.org/record/10718156/files/figure.png" pageId="1" pageNumber="90">Fig. 15</figureCitation>
); anterolateral margin of tergum II with tuft of long, hairlike setae; lateral margins of terga IV–IX each with row of long, stout setae of various sizes; lateral setae on tergum IX shorter than those on other terga (
<figureCitation id="136D44D6FFB91F5F563CF969D3C21AB2" box="[868,934,1699,1724]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[206,1382,181,1919]" captionTargetId="figure-19@4.[206,1382,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–9. Habitus of Torleya mikhaili, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae." figureDoi="http://doi.org/10.5281/zenodo.10718149" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="1" pageNumber="90">Fig. 9</figureCitation>
). Caudal filaments about one fourth as long as body; setae at apex of each segment; setae of different shapes: long and hairlike, short stout, oval (
<figureCitation id="136D44D6FFB91F5F5192F90DD52C1AEE" box="[1226,1352,1735,1760]" captionStart="FIGURES 15–18" captionStartId="6.[151,264,1683,1707]" captionTargetBox="[151,1436,181,1659]" captionTargetId="figure-19@6.[151,1436,181,1659]" captionTargetPageId="6" captionText="FIGURES 15–18. Larvae of Torleya mikhaili: 15, abdomen, dorsal; 16, submedian setae on posterior margins of terga; 17, abdominal sternum IX and caudal filaments; 18, setae on caudal filaments." figureDoi="http://doi.org/10.5281/zenodo.10718156" httpUri="https://zenodo.org/record/10718156/files/figure.png" pageId="1" pageNumber="90">Figs 17–18</figureCitation>
). Distal margins of lamellate portion of gills III–V with shallow notch in middle part and rounded protrusion on outer side (
<figureCitation id="136D44D6FFB91F5F5035F921D0BA1B26" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[211,1366,187,1913]" captionTargetId="figure-15@7.[205,1382,181,1919]" captionTargetPageId="7" captionText="FIGURES 19–22. Larvae of Torleya mikhaili: gills of I (18), II (19), III (21), and IV (22) pairs." figureDoi="http://doi.org/10.5281/zenodo.10718158" httpUri="https://zenodo.org/record/10718158/files/figure.png" pageId="1" pageNumber="90">Figs 19–22</figureCitation>
); gill III lamella largest, slightly tapering distally without any weak transverse band (
<figureCitation id="136D44D6FFB91F5F5132F8C5D4DD1B26" box="[1130,1209,1807,1832]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[211,1366,187,1913]" captionTargetId="figure-15@7.[205,1382,181,1919]" captionTargetPageId="7" captionText="FIGURES 19–22. Larvae of Torleya mikhaili: gills of I (18), II (19), III (21), and IV (22) pairs." figureDoi="http://doi.org/10.5281/zenodo.10718158" httpUri="https://zenodo.org/record/10718158/files/figure.png" pageId="1" pageNumber="90">Fig. 19</figureCitation>
); lamellae of gills IV and V almost same shape, with strongly attenuated, rounded anterior outer margin (
<figureCitation id="136D44D6FFB91F5F5152F8F9D4E21B42" box="[1034,1158,1843,1868]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[211,1366,187,1913]" captionTargetId="figure-15@7.[205,1382,181,1919]" captionTargetPageId="7" captionText="FIGURES 19–22. Larvae of Torleya mikhaili: gills of I (18), II (19), III (21), and IV (22) pairs." figureDoi="http://doi.org/10.5281/zenodo.10718158" httpUri="https://zenodo.org/record/10718158/files/figure.png" pageId="1" pageNumber="90">Figs 20–21</figureCitation>
); lamella of gill VI almost symmetrical in shape, with rounded distal margin, and well-defined rounded projection on anterior edge of outer margin (
<figureCitation id="136D44D6FFB91F5F55C7F8B1D08B1B9A" box="[159,239,1915,1940]" captionStart="FIGURES 19–22" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[211,1366,187,1913]" captionTargetId="figure-15@7.[205,1382,181,1919]" captionTargetPageId="7" captionText="FIGURES 19–22. Larvae of Torleya mikhaili: gills of I (18), II (19), III (21), and IV (22) pairs." figureDoi="http://doi.org/10.5281/zenodo.10718158" httpUri="https://zenodo.org/record/10718158/files/figure.png" pageId="1" pageNumber="90">Fig. 22</figureCitation>
); gill VII transparent, tiny, with weakly defined attachment.
</paragraph>
<caption id="DF2908DBFFBA1F5C55CFF85DD1891BDA" ID-DOI="http://doi.org/10.5281/zenodo.10718142" ID-Zenodo-Dep="10718142" httpUri="https://zenodo.org/record/10718142/files/figure.png" pageId="2" pageNumber="91" startId="2.[151,264,1943,1967]" targetBox="[167,1421,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BE95853FFBA1F5C55CFF85DD1891BDA" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="91">
<emphasis id="B9228441FFBA1F5C55CFF85DD1531BA1" bold="true" box="[151,311,1943,1967]" pageId="2" pageNumber="91">FIGURES 1–2</emphasis>
. Habitus of
<taxonomicName id="4C5623D0FFBA1F5C54E3F852D2001BBE" authorityName="Tiunova" authorityYear="1995" box="[443,612,1944,1968]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBA1F5C54E3F852D2001BBE" box="[443,612,1944,1968]" italics="true" pageId="2" pageNumber="91">Torleya mikhaili</emphasis>
</taxonomicName>
, paratypes (material from type locality), lateral view: 1, imago male; 2, imago female; 2A, subanal plate female.
</paragraph>
</caption>
<caption id="DF2908DBFFBB1F5D55CFF941D2B01AC9" ID-DOI="http://doi.org/10.5281/zenodo.10718144" ID-Zenodo-Dep="10718144" httpUri="https://zenodo.org/record/10718144/files/figure.png" pageId="3" pageNumber="92" startId="3.[151,264,1675,1699]" targetBox="[151,1436,181,1651]" targetPageId="3" targetType="figure">
<paragraph id="8BE95853FFBB1F5D55CFF941D2B01AC9" blockId="3.[151,1437,1675,1735]" pageId="3" pageNumber="92">
<emphasis id="B9228441FFBB1F5D55CFF941D1531AAD" bold="true" box="[151,311,1675,1699]" pageId="3" pageNumber="92">FIGURES 3–7</emphasis>
. Imago male of
<taxonomicName id="4C5623D0FFBB1F5D54B3F941D2F01AAD" authorityName="Tiunova" authorityYear="1995" box="[491,660,1675,1699]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBB1F5D54B3F941D2F01AAD" box="[491,660,1675,1699]" italics="true" pageId="3" pageNumber="92">Torleya mikhaili</emphasis>
</taxonomicName>
, paratypes (material from type locality): 3, genitalia, ventral; 4, genitalia, dorsal; 5, penis lobes, dorsal; 6, hind wing; 7, fore wing.
</paragraph>
</caption>
<paragraph id="8BE95853FFBB1F5D559FF8D6D24D1BCD" blockId="3.[151,1436,1819,1987]" pageId="3" pageNumber="92">
<emphasis id="B9228441FFBB1F5D559FF8D6D1601B3D" box="[199,260,1820,1843]" italics="true" pageId="3" pageNumber="92">Eggs.</emphasis>
General form of egg ovoid with length of 129–145 µm and width of 75–85 µm (
<figureCitation id="136D44D6FFBB1F5D510BF8D1D4B41B3D" box="[1107,1232,1819,1843]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1672,1696]" captionTargetBox="[151,1435,181,1648]" captionTargetId="figure-19@8.[151,1435,181,1648]" captionTargetPageId="8" captionText="FIGURES 23–26. Eggs of Torleya mikhaili: 23–24, general view; 25, structure of chorion micropyles; 26, polar cap. The arrow shows the micropyles (mr)." figureDoi="http://doi.org/10.5281/zenodo.10718160" httpUri="https://zenodo.org/record/10718160/files/figure.png" pageId="3" pageNumber="92">Figs 23–24</figureCitation>
); polar cap present, composed of dense threads (
<figureCitation id="136D44D6FFBB1F5D549FF8F5D2731B59" box="[455,535,1855,1879]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1672,1696]" captionTargetBox="[151,1435,181,1648]" captionTargetId="figure-19@8.[151,1435,181,1648]" captionTargetPageId="8" captionText="FIGURES 23–26. Eggs of Torleya mikhaili: 23–24, general view; 25, structure of chorion micropyles; 26, polar cap. The arrow shows the micropyles (mr)." figureDoi="http://doi.org/10.5281/zenodo.10718160" httpUri="https://zenodo.org/record/10718160/files/figure.png" pageId="3" pageNumber="92">Fig. 26</figureCitation>
). Chorion with geometric macrorelief constituted of large-meshed reticulations with relatively deeply mesh units; each mesh unit with flat, smooth bottom; ridges about 2–3 µm in width; surface of ridges relatively smooth; inner margin of ridges with numerous papillae (
<figureCitation id="136D44D6FFBB1F5D560AF84DD3C51B91" box="[850,929,1927,1951]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1672,1696]" captionTargetBox="[151,1435,181,1648]" captionTargetId="figure-19@8.[151,1435,181,1648]" captionTargetPageId="8" captionText="FIGURES 23–26. Eggs of Torleya mikhaili: 23–24, general view; 25, structure of chorion micropyles; 26, polar cap. The arrow shows the micropyles (mr)." figureDoi="http://doi.org/10.5281/zenodo.10718160" httpUri="https://zenodo.org/record/10718160/files/figure.png" pageId="3" pageNumber="92">Fig. 25</figureCitation>
). Micropyles in subequatorial area; sperm guide rounded or teardrop-shaped (
<figureCitation id="136D44D6FFBB1F5D5495F861D2781BCD" box="[461,540,1963,1987]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1672,1696]" captionTargetBox="[151,1435,181,1648]" captionTargetId="figure-19@8.[151,1435,181,1648]" captionTargetPageId="8" captionText="FIGURES 23–26. Eggs of Torleya mikhaili: 23–24, general view; 25, structure of chorion micropyles; 26, polar cap. The arrow shows the micropyles (mr)." figureDoi="http://doi.org/10.5281/zenodo.10718160" httpUri="https://zenodo.org/record/10718160/files/figure.png" pageId="3" pageNumber="92">Fig. 25</figureCitation>
).
</paragraph>
<caption id="DF2908DBFFBC1F5A55CFF85DD5FF1BBE" ID-DOI="http://doi.org/10.5281/zenodo.10718149" ID-Zenodo-Dep="10718149" box="[151,1435,1943,1968]" httpUri="https://zenodo.org/record/10718149/files/figure.png" pageId="4" pageNumber="93" startId="4.[151,264,1943,1967]" targetBox="[206,1382,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="8BE95853FFBC1F5A55CFF85DD5FF1BBE" blockId="4.[151,1435,1943,1968]" box="[151,1435,1943,1968]" pageId="4" pageNumber="93">
<emphasis id="B9228441FFBC1F5A55CFF85DD1501BA1" bold="true" box="[151,308,1943,1967]" pageId="4" pageNumber="93">FIGURES 8–9</emphasis>
. Habitus of
<taxonomicName id="4C5623D0FFBC1F5A54E9F852D2331BBE" authorityName="Tiunova" authorityYear="1995" box="[433,599,1944,1968]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBC1F5A54E9F852D2331BBE" box="[433,599,1944,1968]" italics="true" pageId="4" pageNumber="93">Torleya mikhaili</emphasis>
</taxonomicName>
, paratypes (material from type locality): dorsal view: 8, subimago male; 9, larvae.
</paragraph>
</caption>
<caption id="DF2908DBFFBD1F5B55CFF883D08D1BA7" ID-DOI="http://doi.org/10.5281/zenodo.10718154" ID-Zenodo-Dep="10718154" httpUri="https://zenodo.org/record/10718154/files/figure.png" pageId="5" pageNumber="94" startId="5.[151,264,1865,1889]" targetBox="[151,1435,181,1841]" targetPageId="5" targetType="figure">
<paragraph id="8BE95853FFBD1F5B55CFF883D08D1BA7" blockId="5.[151,1436,1865,1961]" pageId="5" pageNumber="94">
<emphasis id="B9228441FFBD1F5B55CFF883D1351B6F" bold="true" box="[151,337,1865,1889]" pageId="5" pageNumber="94">FIGURES 10–14</emphasis>
. Larvae of
<taxonomicName id="4C5623D0FFBD1F5B5496F883D2131B6F" authorityName="Tiunova" authorityYear="1995" box="[462,631,1865,1889]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBD1F5B5496F883D2131B6F" box="[462,631,1865,1889]" italics="true" pageId="5" pageNumber="94">Torleya mikhaili</emphasis>
</taxonomicName>
, paratype (material from type locality): 10, head, dorsal; 11, foreleg, dorsal; 12, fore femur and tibia, dorsal; 13, fore femur, latero-dorsal; 14, fore claw. The arrow shows the long hairlike setae on the occiput.
</paragraph>
</caption>
<caption id="DF2908DBFFBE1F5855CFF959D3E91AC1" ID-DOI="http://doi.org/10.5281/zenodo.10718156" ID-Zenodo-Dep="10718156" httpUri="https://zenodo.org/record/10718156/files/figure.png" pageId="6" pageNumber="95" startId="6.[151,264,1683,1707]" targetBox="[151,1436,181,1659]" targetPageId="6" targetType="figure">
<paragraph id="8BE95853FFBE1F5855CFF959D3E91AC1" blockId="6.[151,1437,1683,1743]" pageId="6" pageNumber="95">
<emphasis id="B9228441FFBE1F5855CFF959D1341AA5" bold="true" box="[151,336,1683,1707]" pageId="6" pageNumber="95">FIGURES 15–18</emphasis>
. Larvae of
<taxonomicName id="4C5623D0FFBE1F585493F959D2171AA5" authorityName="Tiunova" authorityYear="1995" box="[459,627,1683,1707]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBE1F585493F959D2171AA5" box="[459,627,1683,1707]" italics="true" pageId="6" pageNumber="95">Torleya mikhaili</emphasis>
</taxonomicName>
: 15, abdomen, dorsal; 16, submedian setae on posterior margins of terga; 17, abdominal sternum IX and caudal filaments; 18, setae on caudal filaments.
</paragraph>
</caption>
<paragraph id="8BE95853FFBE1F58559FF935D3661BA8" blockId="6.[151,1437,1791,2031]" pageId="6" pageNumber="95">
<emphasis id="B9228441FFBE1F58559FF935D17C1B19" box="[199,280,1791,1815]" italics="true" pageId="6" pageNumber="95">Habitat</emphasis>
. Larvae and adults of
<taxonomicName id="4C5623D0FFBE1F585745F8CAD2F61B19" authorityName="Tiunova" authorityYear="1995" box="[541,658,1791,1815]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBE1F585745F8CAD2F61B19" box="[541,658,1791,1815]" italics="true" pageId="6" pageNumber="95">T. mikhaili</emphasis>
</taxonomicName>
were collected in the south of the Russian Far East (
<collectingRegion id="499296B1FFBE1F5851AAF935D50E1B19" box="[1266,1386,1791,1815]" country="Russia" name="Primorskiy" pageId="6" pageNumber="95">Primorskiy</collectingRegion>
and Khabarovskiy Territories) at the beginning July–middle of August. Larvae were collected in rivers (
<figureCitation id="136D44D6FFBE1F5851E8F8E9D54E1B35" box="[1200,1322,1827,1851]" captionStart="FIGURES 27–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[205,1383,181,1919]" captionTargetId="figure-15@9.[205,1383,181,1919]" captionTargetPageId="9" captionText="FIGURES 27–28. Localities of Torleya mikhaili, Primorskiy Kray: 27, Ussuri River; 28, Ilistaya River." figureDoi="http://doi.org/10.5281/zenodo.10718162" httpUri="https://zenodo.org/record/10718162/files/figure.png" pageId="6" pageNumber="95">Figs 27–28</figureCitation>
) at a depth of
<quantity id="4CAEF5B6FFBE1F5855EDF88DD12C1B50" box="[181,328,1863,1887]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="6" pageNumber="95" unit="cm" value="65.0" valueMax="100.0" valueMin="30.0">30 to 100 cm</quantity>
. The width of the rivers in the places where larvae were collected exceeded
<quantity id="4CAEF5B6FFBE1F5851D9F88DD4AD1B50" box="[1153,1225,1863,1887]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="95" unit="m" value="100.0">100 m</quantity>
; the length of these rivers is more than
<quantity id="4CAEF5B6FFBE1F585435F8A1D1A71B8D" box="[365,451,1899,1923]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="95" unit="km" value="300.0">300 km</quantity>
. The water temperature in this period exceeds 20–25°C. The larvae inhabited stone-gravel substrates with sand and detritus, mainly in slow currents.
</paragraph>
<paragraph id="8BE95853FFBE1F58559FF879D17D1BE1" blockId="6.[151,1437,1791,2031]" pageId="6" pageNumber="95">
<emphasis id="B9228441FFBE1F58559FF879D12D1BC5" box="[199,329,1971,1995]" italics="true" pageId="6" pageNumber="95">Distribution</emphasis>
. East Palaearctic: the south portion of the Russian Far East, northeastern
<collectingCountry id="F34118C3FFBE1F585102F879D4FE1BC5" box="[1114,1178,1971,1995]" name="China" pageId="6" pageNumber="95">China</collectingCountry>
(Jacobus
<emphasis id="B9228441FFBE1F58505DF87ED55F1BC5" box="[1285,1339,1971,1995]" italics="true" pageId="6" pageNumber="95">et al.</emphasis>
2004;
<bibRefCitation id="EFC725A2FFBE1F5850DDF879D1691BE1" author="Li, R. &amp; Zhang, W. &amp; Ma, Z. &amp; Zhou, C." pageId="6" pageNumber="95" pagination="3106 - 3114" refId="ref4628" refString="Li, R., Zhang, W., Ma, Z. &amp; Zhou, C. (2020) Novel gene rearrangement pattern in the mitochondrial genomes of Torleya mikhaili and Cincticostella fusca (Ephemeroptera: Ephemerellidae). International Journal of Biological Macromolecules, 165, 3106 - 3114. https: // doi. org / 10.1016 / j. ijbiomac. 2020.10.124" type="journal article" year="2020">
Li
<emphasis id="B9228441FFBE1F5855CFF812D0A21BE1" box="[151,198,2007,2031]" italics="true" pageId="6" pageNumber="95">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
<caption id="DF2908DBFFBF1F5955CFF85DD4061BBE" ID-DOI="http://doi.org/10.5281/zenodo.10718158" ID-Zenodo-Dep="10718158" box="[151,1122,1943,1968]" httpUri="https://zenodo.org/record/10718158/files/figure.png" pageId="7" pageNumber="96" startId="7.[151,264,1943,1967]" targetBox="[211,1366,187,1913]" targetPageId="7" targetType="figure">
<paragraph id="8BE95853FFBF1F5955CFF85DD4061BBE" blockId="7.[151,1122,1943,1968]" box="[151,1122,1943,1968]" pageId="7" pageNumber="96">
<emphasis id="B9228441FFBF1F5955CFF85DD12A1BA1" bold="true" box="[151,334,1943,1967]" pageId="7" pageNumber="96">FIGURES 19–22</emphasis>
. Larvae of
<taxonomicName id="4C5623D0FFBF1F59549AF852D20C1BBE" authorityName="Tiunova" authorityYear="1995" box="[450,616,1944,1968]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFBF1F59549AF852D20C1BBE" box="[450,616,1944,1968]" italics="true" pageId="7" pageNumber="96">Torleya mikhaili</emphasis>
</taxonomicName>
: gills of I (18), II (19), III (21), and IV (22) pairs.
</paragraph>
</caption>
<caption id="DF2908DBFFB01F5655CFF942D1C91ACA" ID-DOI="http://doi.org/10.5281/zenodo.10718160" ID-Zenodo-Dep="10718160" httpUri="https://zenodo.org/record/10718160/files/figure.png" pageId="8" pageNumber="97" startId="8.[151,264,1672,1696]" targetBox="[151,1435,181,1648]" targetPageId="8" targetType="figure">
<paragraph id="8BE95853FFB01F5655CFF942D1C91ACA" blockId="8.[151,1436,1672,1732]" pageId="8" pageNumber="97">
<emphasis id="B9228441FFB01F5655CFF942D1291AAE" bold="true" box="[151,333,1672,1696]" pageId="8" pageNumber="97">FIGURES 23–26</emphasis>
. Eggs of
<taxonomicName id="4C5623D0FFB01F5654F4F942D2351AAE" authorityName="Tiunova" authorityYear="1995" box="[428,593,1672,1696]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB01F5654F4F942D2351AAE" box="[428,593,1672,1696]" italics="true" pageId="8" pageNumber="97">Torleya mikhaili</emphasis>
</taxonomicName>
: 23–24, general view; 25, structure of chorion micropyles; 26, polar cap. The arrow shows the micropyles (mr).
</paragraph>
</caption>
<paragraph id="8BE95853FFB01F56559FF8C1D2E91BF4" blockId="8.[151,1437,1803,2043]" pageId="8" pageNumber="97">
<emphasis id="B9228441FFB01F56559FF8C1D1401B2D" box="[199,292,1803,1827]" italics="true" pageId="8" pageNumber="97">Remarks</emphasis>
. In an article devoted to the revision of
<taxonomicName id="4C5623D0FFB01F565780F8C1D3E41B2D" box="[728,896,1803,1827]" pageId="8" pageNumber="97">
<emphasis id="B9228441FFB01F565780F8C1D34D1B2D" box="[728,809,1803,1827]" italics="true" pageId="8" pageNumber="97">Torleya</emphasis>
species
</taxonomicName>
, the authors were the first to indicate
<collectingCountry id="F34118C3FFB01F565040F8C1D53C1B2D" box="[1304,1368,1803,1827]" name="China" pageId="8" pageNumber="97">China</collectingCountry>
in the distribution of
<taxonomicName id="4C5623D0FFB01F565462F8E5D2C31B49" authority="(Jacobus et al., 2004)" baseAuthorityName="Jacobus" baseAuthorityYear="2004" box="[314,679,1839,1863]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB01F565462F8E5D1CA1B49" box="[314,430,1839,1863]" italics="true" pageId="8" pageNumber="97">T. mikhaili</emphasis>
(Jacobus
<emphasis id="B9228441FFB01F565745F8E5D22B1B49" box="[541,591,1839,1863]" italics="true" pageId="8" pageNumber="97">et al</emphasis>
., 2004)
</taxonomicName>
. At the same time, when comparing larvae from
<collectingCountry id="F34118C3FFB01F56519AF8E5D56D1B49" box="[1218,1289,1839,1863]" name="Russia" pageId="8" pageNumber="97">Russia</collectingCountry>
and
<collectingCountry id="F34118C3FFB01F565019F8E5D5E71B49" box="[1345,1411,1839,1863]" name="China" pageId="8" pageNumber="97">China</collectingCountry>
, a number of differences were noted that arose due to the absence of some morphological characteristics in the description of
<taxonomicName id="4C5623D0FFB01F5655ECF8BDD1421B81" box="[180,294,1911,1935]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB01F5655ECF8BDD1421B81" box="[180,294,1911,1935]" italics="true" pageId="8" pageNumber="97">T. mikhaili</emphasis>
</taxonomicName>
. Additional data presented in this work confirm the presence of this species in
<collectingCountry id="F34118C3FFB01F56512CF8BDD4D31B81" box="[1140,1207,1911,1935]" name="China" pageId="8" pageNumber="97">China</collectingCountry>
. Note that in 2020 an article was published that provided molecular data for
<taxonomicName id="4C5623D0FFB01F5657B8F851D3351BBD" box="[736,849,1947,1971]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB01F5657B8F851D3351BBD" box="[736,849,1947,1971]" italics="true" pageId="8" pageNumber="97">T. mikhaili</emphasis>
</taxonomicName>
larvae collected in
<collectingCountry id="F34118C3FFB01F56517BF851D4071BBD" box="[1059,1123,1947,1971]" name="China" pageId="8" pageNumber="97">China</collectingCountry>
(
<bibRefCitation id="EFC725A2FFB01F56512AF851D56C1BBD" author="Li, R. &amp; Zhang, W. &amp; Ma, Z. &amp; Zhou, C." box="[1138,1288,1947,1971]" pageId="8" pageNumber="97" pagination="3106 - 3114" refId="ref4628" refString="Li, R., Zhang, W., Ma, Z. &amp; Zhou, C. (2020) Novel gene rearrangement pattern in the mitochondrial genomes of Torleya mikhaili and Cincticostella fusca (Ephemeroptera: Ephemerellidae). International Journal of Biological Macromolecules, 165, 3106 - 3114. https: // doi. org / 10.1016 / j. ijbiomac. 2020.10.124" type="journal article" year="2020">
Li
<emphasis id="B9228441FFB01F5651CAF851D4AC1BBD" box="[1170,1224,1947,1971]" italics="true" pageId="8" pageNumber="97">et al.</emphasis>
2020
</bibRefCitation>
). Therefore, a goal of further research will be to collect material for the molecular study of
<taxonomicName id="4C5623D0FFB01F56569CF875D4511BD9" box="[964,1077,1983,2007]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="mikhaili">
<emphasis id="B9228441FFB01F56569CF875D4511BD9" box="[964,1077,1983,2007]" italics="true" pageId="8" pageNumber="97">T. mikhaili</emphasis>
</taxonomicName>
from
<collectingCountry id="F34118C3FFB01F56512EF875D4D91BD9" box="[1142,1213,1983,2007]" name="Russia" pageId="8" pageNumber="97">Russia</collectingCountry>
so those populations can be compared to their Chinese counterparts.
</paragraph>
</subSubSection>
</treatment>
</document>
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