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<mods:title id="DA27FABBF40FD4B60EAF28E502C47ED3">Studies on water mites of the family Hygrobatidae (Acari, Hydrachnidia) from Iran, I. The water mite genus Atractides Koch, with the description of five new species</mods:title>
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<treatment id="03FB7D20DD63FF8FFEAFF98848ED06FC" ID-DOI="http://doi.org/10.5281/zenodo.5689552" ID-GBIF-Taxon="119601532" ID-Zenodo-Dep="5689552" LSID="urn:lsid:plazi:treatment:03FB7D20DD63FF8FFEAFF98848ED06FC" httpUri="http://treatment.plazi.org/id/03FB7D20DD63FF8FFEAFF98848ED06FC" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24">
<subSubSection id="C3489FBDDD63FF83FEAFF9884BA102E3" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BEDCC36DD63FF83FEAFF98848D6023C" blockId="23.[264,722,1604,1665]" box="[264,722,1604,1630]" pageId="23" pageNumber="24">
<heading id="D0A57B5ADD63FF83FEAFF98848D6023C" bold="true" box="[264,722,1604,1630]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="4C52B7B5DD63FF83FEAFF98848D6023C" authority="Lundblad, 1956" authorityName="Lundblad" authorityYear="1956" box="[264,722,1604,1630]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD63FF83FEAFF98848D6023C" bold="true" box="[264,722,1604,1630]" pageId="23" pageNumber="24">
<emphasis id="B9261024DD63FF83FEAFF9884B86023C" bold="true" box="[264,386,1604,1630]" italics="true" pageId="23" pageNumber="24">Atractides</emphasis>
cf.
<emphasis id="B9261024DD63FF83FE08F9884803023C" bold="true" box="[431,519,1604,1630]" italics="true" pageId="23" pageNumber="24">lunipes</emphasis>
<bibRefCitation id="EFC3B1C7DD63FF83FDA9F98848D6023C" author="Lundblad" box="[526,722,1604,1630]" pageId="23" pageNumber="39" refString="Lundblad, O. (1956) Zur Kenntnis sud-und mitteleuropaischer Hydrachnellen. Arkiv for Zoology, 10 (1), 1 - 306." type="journal article" year="1956">Lundblad, 1956</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BEDCC36DD63FF83FEAFF9AB4BA102E3" blockId="23.[264,722,1604,1665]" box="[264,421,1639,1665]" pageId="23" pageNumber="24">
(
<figureCitation id="1369D0B3DD63FF83FEB6F9AB4B9902E3" box="[273,413,1639,1665]" captionStart-0="FIGURES 41 45" captionStart-1="FIGURES 46 49" captionStartId-0="24.[264,382,1555,1579]" captionStartId-1="26.[264,382,1410,1434]" captionTargetBox-0="[288,1269,315,1539]" captionTargetBox-1="[282,1280,318,1390]" captionTargetId-0="figure@24.[271,1305,306,1547]" captionTargetId-1="figure@26.[266,1321,303,1401]" captionTargetPageId-0="24" captionTargetPageId-1="26" captionText-0="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." captionText-1="FIGURES 46 49. Atractides cf. lunipes Lundblad, 1956, male: 46 = coxal field; 47 = I­L­ 5 / 6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm." httpUri-0="https://zenodo.org/record/157866/files/figure.png" httpUri-1="https://zenodo.org/record/157867/files/figure.png" pageId="23" pageNumber="24">Figs. 4149</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="B9261024DD63FF83FEAFF97D4BE302A8" box="[264,487,1713,1738]" italics="true" pageId="23" pageNumber="24">Material examined</emphasis>
:
<collectingCountry id="F3458CA6DD63FF83FE50F97D482F02A9" box="[503,555,1713,1739]" name="Iran" pageId="23" pageNumber="24">Iran</collectingCountry>
: IR30 Chahar Mahal and Bakhtiari Province, Omid Abad stream near Eisa Abad (After Farsan to Kuhrang) (ca. 32° 18'N, 50° 29'E),
<quantity id="4CAA61D3DD63FF83FB2CF9154EE90291" box="[1163,1261,1753,1779]" metricMagnitude="3" metricUnit="m" metricValue="2.077" pageId="23" pageNumber="24" unit="m" value="2077.0">2077 m</quantity>
asl., 0 1.08.2003, leg. Pesic (1/8/0); IR31 Chahar Mahal and Bakhtiari Province, Omid Abad stream near Eisa Abad (After Fill Abad to Kuhrang), 0 1.08.2003, leg. Pesic (0/9/0).
</paragraph>
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<caption id="DF2D9CBEDD6CFF8CFEAFF9DF498B022F" httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="24" pageNumber="25" targetBox="[288,1269,315,1539]" targetPageId="24">
<paragraph id="8BEDCC36DD6CFF8CFEAFF9DF498B022F" blockId="24.[264,1323,1555,1613]" pageId="24" pageNumber="25">
<emphasis id="B9261024DD6CFF8CFEAFF9DF4BCB0249" bold="true" box="[264,463,1555,1579]" pageId="24" pageNumber="25">FIGURES 4145.</emphasis>
<taxonomicName id="4C52B7B5DD6CFF8CFE71F9DF496A0248" authority="Lundblad, 1956" authorityName="Lundblad" authorityYear="1956" box="[470,878,1555,1578]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6CFF8CFE71F9DF48470248" box="[470,579,1555,1578]" italics="true" pageId="24" pageNumber="25">Atractides</emphasis>
cf.
<emphasis id="B9261024DD6CFF8CFDCAF9DF48BE0248" box="[621,698,1555,1578]" italics="true" pageId="24" pageNumber="25">lunipes</emphasis>
Lundblad, 1956
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, male: 41 = coxal field; 42 = palp; 43= I­ L­5/6; 44 = genital field; 45 = Vgl­1/2. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3489FBDDD6CFF8DFE9FF94D4B75032C" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
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<emphasis id="B9261024DD6CFF8CFE9FF94D4BC602F8" box="[312,450,1665,1690]" italics="true" pageId="24" pageNumber="25">Description</emphasis>
</paragraph>
<paragraph id="8BEDCC36DD6CFF8DFE9FF96548930614" blockId="24.[264,1323,1665,1891]" lastBlockId="25.[264,1324,284,1870]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B9261024DD6CFF8CFE9FF9654B7002A0" box="[312,372,1705,1730]" italics="true" pageId="24" pageNumber="25">Male</emphasis>
: Length of idiosoma 604, width 443. Dorsally the integument is lineated; muscle attachments: unsclerotized. Coxal field (
<figureCitation id="1369D0B3DD6CFF8CFD47F91D493C0289" box="[736,824,1745,1771]" captionStart="FIGURES 41 45" captionStartId="24.[264,382,1555,1579]" captionTargetBox="[288,1269,315,1539]" captionTargetId="figure@24.[271,1305,306,1547]" captionTargetPageId="24" captionText="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="24" pageNumber="25">Fig. 41</figureCitation>
): Cx­1+2 short and large, posterior margin Cx­1 medially straight, apodemes of Cx­
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an acute angle; length between anterior end of first coxae and posterior end of fourth coxae 354; Cx­3 width 438, Cx­1+2 width 350; Cx­1+2 medial suture
<quantity id="4CAA61D3DD6CFF8CFDB4F885485A0301" box="[531,606,1865,1891]" metricMagnitude="0" metricUnit="m" metricValue="2.667" pageId="24" pageNumber="25" unit="in" value="105.0">105 in</quantity>
length; distance from lateralmost tips of Cx­2 to the medioposterior edge of Cx­1+2 238. Measurements of mouthparts: chelicera length 200, claw 59.6; palp total length 312, dorsal length and relative length (% total length, in parentheses) of single segments: P­1 32 (10.3), P­2 73 (23.4), P­3 71 (22.8), P­4 103 (33), P­5 33 (10.6); ratio length P­2/P­4 0.70;palp (
<figureCitation id="1369D0B3DD6DFF8DFD43FE58494505CC" box="[740,833,404,430]" captionStart="FIGURES 41 45" captionStartId="24.[264,382,1555,1579]" captionTargetBox="[288,1269,315,1539]" captionTargetId="figure@24.[271,1305,306,1547]" captionTargetPageId="24" captionText="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="25" pageNumber="26">Fig. 42</figureCitation>
): ventral margin of P­2 with a strongly developed ventrodistal projection ending in a long, digitiform tip, P­3 slightly concave, P­ 4 ventral margin weakly extended near proximal hair insertion, sword seta curved and inserted between ventral hairs. Genital field (
<figureCitation id="1369D0B3DD6DFF8DFCBFFDC049750644" box="[792,881,524,550]" captionStart="FIGURES 41 45" captionStartId="24.[264,382,1555,1579]" captionTargetBox="[288,1269,315,1539]" captionTargetId="figure@24.[271,1305,306,1547]" captionTargetPageId="24" captionText="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="25" pageNumber="26">Fig. 44</figureCitation>
): anterior margin with shallow indentations;
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length,
<quantity id="4CAA61D3DD6DFF8DFD83FDF8486A062C" box="[548,622,564,590]" metricMagnitude="0" metricUnit="m" metricValue="3.1242" pageId="25" pageNumber="26" unit="in" value="123.0">123 in</quantity>
width, Ac­1­3 length 34­38.5­31.5; excretory pore smooth; Vgl­1: fused with Vgl­2 (
<figureCitation id="1369D0B3DD6DFF8DFD95FD90488D0614" box="[562,649,604,630]" captionStart="FIGURES 41 45" captionStartId="24.[264,382,1555,1579]" captionTargetBox="[288,1269,315,1539]" captionTargetId="figure@24.[271,1305,306,1547]" captionTargetPageId="24" captionText="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="25" pageNumber="26">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="8BEDCC36DD6DFF8DFE9FFD484E2807EC" blockId="25.[264,1324,284,1870]" pageId="25" pageNumber="26">
I­L (
<figureCitation id="1369D0B3DD6DFF8DFED5FD484BD406FC" box="[370,464,644,670]" captionStart="FIGURES 41 45" captionStartId="24.[264,382,1555,1579]" captionTargetBox="[288,1269,315,1539]" captionTargetId="figure@24.[271,1305,306,1547]" captionTargetPageId="24" captionText="FIGURES 41 45. Atractides cf. lunipes Lundblad, 1956, male: 41 = coxal field; 42 = palp; 43 = I­ L­ 5 / 6; 44 = genital field; 45 = Vgl­ 1 / 2. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157866/files/figure.png" pageId="25" pageNumber="26">Fig. 43</figureCitation>
): S­1/2 distanced, heteromorphic; I­L­6 slender and curved; I­L­3/4/5 forming distal sheaths each covering the base of the following segment; I­L­5 dorsal length 202, I­L­5 ventral length 124, ratio dorsal length I­L­5 /ventral length 1.63, I­L­5 central height 55.1, ratio dorsal length I­L­5/central height 3.67, S­1 length 102.5, ratio length S­1/width 11.4, S­2 length 77.6, ratio length S­2/ width 6.75, distance between sword setae of I­L­5 25.6; ratio length S­1/2 1.32; I­L­6 length 142.3, I­L­6 central height 19.2, ratio length I­L­6 /central height 7.4; ratio length I­L­5/6 1.42.
</paragraph>
<paragraph id="8BEDCC36DD6DFF8DFE9FFC504E8E026C" blockId="25.[264,1324,284,1870]" pageId="25" pageNumber="26">
<emphasis id="B9261024DD6DFF8DFE9FFC504B9407D7" box="[312,400,924,949]" italics="true" pageId="25" pageNumber="26">Female</emphasis>
(measurements based on two specimens): Length of idiosoma 907­1057, width 793­893. Dorsally the integument is lineated; muscle attachments: unsclerotized. Coxal field: Cx­1+2 short and large, posterior margin Cx­1 medially slightly concave, apodemes of Cx­
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an acute angle; coxal plates with distinct medial and posterior margins with secondary sclerotization (
<figureCitation id="1369D0B3DD6DFF8DFDCAFBF048C00034" box="[621,708,1084,1110]" captionStart="FIGURES 46 49" captionStartId="26.[264,382,1410,1434]" captionTargetBox="[282,1280,318,1390]" captionTargetId="figure@26.[266,1321,303,1401]" captionTargetPageId="26" captionText="FIGURES 46 49. Atractides cf. lunipes Lundblad, 1956, male: 46 = coxal field; 47 = I­L­ 5 / 6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/157867/files/figure.png" pageId="25" pageNumber="26">Fig. 46</figureCitation>
); length between anterior end of first coxae and posterior end of fourth coxae 416­508; Cx­3 width 585­696, Cx­1+2 width 473; Cx­1+2 medial suture
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length; distance from lateralmost tips of Cx­2 to the medioposterior edge of Cx­1+2 277­315. Measurements of mouthparts: chelicera 242­292, claw 73­ 82.7; palp total length 397.5­467.5, dorsal length and relative length (% total length, in parenthesis) of single segments: P­1 38.5­44 (9.4­9.7), P­2 85­106 (21.4­22.7), P­3 110­ 133 (27.7­28.5), P­4 122­138.5 (29.6­30.7), P­5 42­46 (9.8­10.6); ratio length P­2/P­4 0.7­ 0.77; palp (
<figureCitation id="1369D0B3DD6DFF8DFE37FA984BEE010C" box="[400,490,1364,1390]" captionStart="FIGURES 46 49" captionStartId="26.[264,382,1410,1434]" captionTargetBox="[282,1280,318,1390]" captionTargetId="figure@26.[266,1321,303,1401]" captionTargetPageId="26" captionText="FIGURES 46 49. Atractides cf. lunipes Lundblad, 1956, male: 46 = coxal field; 47 = I­L­ 5 / 6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/157867/files/figure.png" pageId="25" pageNumber="26">Fig. 48</figureCitation>
): P­2 ventral margin distally slightly protruding and convex, P­3 ventrally straight, P­4 more slender than in male, ventral margin weakly extended near proximal hair insertion, sword seta located halfway between ventral hairs. Genital field (
<figureCitation id="1369D0B3DD6DFF8DFB5EFA684B2E0184" captionStart="FIGURES 46 49" captionStartId="26.[264,382,1410,1434]" captionTargetBox="[282,1280,318,1390]" captionTargetId="figure@26.[266,1321,303,1401]" captionTargetPageId="26" captionText="FIGURES 46 49. Atractides cf. lunipes Lundblad, 1956, male: 46 = coxal field; 47 = I­L­ 5 / 6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/157867/files/figure.png" pageId="25" pageNumber="26">Fig. 49</figureCitation>
):
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length,
<quantity id="4CAA61D3DD6DFF8DFDA0FA00484D0184" box="[519,585,1484,1510]" metricMagnitude="0" metricUnit="m" metricValue="5.6134" pageId="25" pageNumber="26" unit="in" value="221.0">221in</quantity>
width, Ac­1­3 length 42­46­44; praegenital sclerite compact, thick, genital plates curved; excretory pore smooth; Vgl­1: fused with Vgl­2.
</paragraph>
<paragraph id="8BEDCC36DD6DFF8DFE9FF9D04B75032C" blockId="25.[264,1324,284,1870]" pageId="25" pageNumber="26">
I­L (
<figureCitation id="1369D0B3DD6DFF8DFECAF9D04BC20254" box="[365,454,1564,1590]" captionStart="FIGURES 46 49" captionStartId="26.[264,382,1410,1434]" captionTargetBox="[282,1280,318,1390]" captionTargetId="figure@26.[266,1321,303,1401]" captionTargetPageId="26" captionText="FIGURES 46 49. Atractides cf. lunipes Lundblad, 1956, male: 46 = coxal field; 47 = I­L­ 5 / 6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/157867/files/figure.png" pageId="25" pageNumber="26">Fig. 47</figureCitation>
): I­L­5 protruding between ventral seta and S­1 insertion, S­1/2 much distanced, heteromorphic; I­L­6 very slender and curved; I­L­5 dorsal length 271­298, I­L­5 ventral length 158­186.5, ratio dorsal length I­L­5/ventral length 1.6­1.71, I­L­5 central height 73­86, ratio dorsal length I­L­5/central height 3.47­3.7, S­1 length 122­131, ratio length S­1/width 11.4­12.7, S­2 length 99­102, ratio length S­2/ width 5.72­6.38, distance between sword setae of I­L­5 46­48; ratio length S­1/2 1.2­1.32; I­L­6 length 194­222, I­ L­6 central height 18­21.5, ratio length I­L­6 /central height 10.3­11.0; ratio length I­L­5/6 1.34­1.4.
</paragraph>
</subSubSection>
<caption id="DF2D9CBEDD6EFF8EFEAFFA4E48E401DE" httpUri="https://zenodo.org/record/157867/files/figure.png" pageId="26" pageNumber="27" targetBox="[282,1280,318,1390]" targetPageId="26">
<paragraph id="8BEDCC36DD6EFF8EFEAFFA4E48E401DE" blockId="26.[264,1322,1410,1468]" pageId="26" pageNumber="27">
<emphasis id="B9261024DD6EFF8EFEAFFA4E4BCB01F8" bold="true" box="[264,463,1410,1434]" pageId="26" pageNumber="27">FIGURES 4649.</emphasis>
<taxonomicName id="4C52B7B5DD6EFF8EFE72FA4E496F01FB" authority="Lundblad, 1956" authorityName="Lundblad" authorityYear="1956" box="[469,875,1410,1433]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6EFF8EFE72FA4E484601FB" box="[469,578,1410,1433]" italics="true" pageId="26" pageNumber="27">Atractides</emphasis>
cf.
<emphasis id="B9261024DD6EFF8EFDCBFA4E48BD01FB" box="[620,697,1410,1433]" italics="true" pageId="26" pageNumber="27">lunipes</emphasis>
Lundblad, 1956
</taxonomicName>
, male: 46 = coxal field; 47= I­L­5/6; 48 = palp; 49 = genital field. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3489FBDDD6EFF8FFE9FF9D44E48059C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BEDCC36DD6EFF8FFE9FF9D44E48059C" blockId="26.[264,1325,1560,1906]" lastBlockId="27.[264,1323,284,670]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B9261024DD6EFF8EFE9FF9D44B9A0253" box="[312,414,1560,1585]" italics="true" pageId="26" pageNumber="27">Remarks</emphasis>
: After the original description from the French Pyrenees (
<bibRefCitation id="EFC3B1C7DD6EFF8EFBC7F9D44F1F0250" author="Lundblad" box="[1120,1307,1560,1586]" pageId="26" pageNumber="39" refString="Lundblad, O. (1956) Zur Kenntnis sud-und mitteleuropaischer Hydrachnellen. Arkiv for Zoology, 10 (1), 1 - 306." type="journal article" year="1956">Lundblad 1956</bibRefCitation>
), the only other records of
<taxonomicName id="4C52B7B5DD6EFF8EFD9CF98C48B7023B" box="[571,691,1600,1625]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6EFF8EFD9CF98C48B7023B" box="[571,691,1600,1625]" italics="true" pageId="26" pageNumber="27">A. lunipes</emphasis>
</taxonomicName>
were published from
<collectingCountry id="F3458CA6DD6EFF8EFC1CF98C4E390238" box="[955,1085,1600,1626]" name="Macedonia" pageId="26" pageNumber="27">Macedonia</collectingCountry>
(Pesic 2003). Until now, only the female of
<taxonomicName id="4C52B7B5DD6EFF8EFD96F9A448AD02E3" box="[561,681,1640,1665]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6EFF8EFD96F9A448AD02E3" box="[561,681,1640,1665]" italics="true" pageId="26" pageNumber="27">A. lunipes</emphasis>
</taxonomicName>
was known. According to
<bibRefCitation id="EFC3B1C7DD6EFF8EFC48F9A44EAC02E0" author="Gerecke" box="[1007,1192,1640,1666]" pageId="26" pageNumber="39" refString="Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke (2003)</bibRefCitation>
<taxonomicName id="4C52B7B5DD6EFF8EFB14F9A44F2F02E3" box="[1203,1323,1640,1665]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6EFF8EFB14F9A44F2F02E3" box="[1203,1323,1640,1665]" italics="true" pageId="26" pageNumber="27">A. lunipes</emphasis>
</taxonomicName>
differs from all Western Palaearctic species in the very fine and slender seta S­1 (L/W ratio 20.4). The females from
<collectingCountry id="F3458CA6DD6EFF8EFD89F97448B402B0" box="[558,688,1720,1746]" name="Macedonia" pageId="26" pageNumber="27">Macedonia</collectingCountry>
(see Pesic 2003) and from
<collectingCountry id="F3458CA6DD6EFF8EFC51F9744E2C02B0" box="[1014,1064,1720,1746]" name="Iran" pageId="26" pageNumber="27">Iran</collectingCountry>
, differ from the original description (in parentheses the data of the
<typeStatus id="54E97294DD6EFF8EFC85F92C498C0298" box="[802,904,1760,1786]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
given by
<bibRefCitation id="EFC3B1C7DD6EFF8EFC5BF92C4EA60298" author="Gerecke" box="[1020,1186,1760,1786]" pageId="26" pageNumber="39" refString="Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke 2003</bibRefCitation>
) by the following measurements: S­1 length &lt;
<quantity id="4CAA61D3DD6EFF8EFD17F8C449090340" box="[688,781,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="26" pageNumber="27" unit="m" value="130.0">130 m</quantity>
(
<quantity id="4CAA61D3DD6EFF8EFCB8F8C449840340" box="[799,896,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="1.43" pageId="26" pageNumber="27" unit="m" value="143.0">143 m</quantity>
), ratio length S­1/width &lt;13 (20.4); ratio length S­2/width &lt;6.5 (7.7); ratio length I­L­5/6 1.31.45 (1.17); genital field L/W&gt; 180.0/200 (L/
<geoCoordinate id="EE66AAF1DD6EFF8EFE0CF89448130310" box="[427,535,1880,1906]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="-162.0">W 162.0</geoCoordinate>
/166.0). From the Iranian females, the Macedonian specimens (in parentheses data from Pesic 2003) differ in shorter segments I­L­5/6 (L I­L­5 235­239; L I­L­6 165­180), more slender I­L­5 (L/H 4.0­4.2), shorter S­2 (L&lt;85.0 m) and a more heteromorphic setae S­1/2 (ratio length S­1/2 1.51.6). For the time being they all are considered conspecific with
<taxonomicName id="4C52B7B5DD6FFF8FFD96FE5848AC05CF" box="[561,680,404,429]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6FFF8FFD96FE5848AC05CF" box="[561,680,404,429]" italics="true" pageId="27" pageNumber="28">A. lunipes</emphasis>
</taxonomicName>
, in this concept a very variable species. However, the assignement of the Macedonian and Iranian populations to this species should be verified by studying male specimens from
<collectingCountry id="F3458CA6DD6FFF8FFD3CFE284919059C" box="[667,797,484,510]" name="Macedonia" pageId="27" pageNumber="28">Macedonia</collectingCountry>
and the French Pyrenees.
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD6FFF8FFE9FFDC048ED06FC" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BEDCC36DD6FFF8FFE9FFDC048ED06FC" blockId="27.[264,1323,284,670]" pageId="27" pageNumber="28">
<emphasis id="B9261024DD6FFF8FFE9FFDC04BC30647" box="[312,455,524,549]" italics="true" pageId="27" pageNumber="28">Distribution</emphasis>
: Presently,
<taxonomicName id="4C52B7B5DD6FFF8FFDFDFDC0495A0647" box="[602,862,524,550]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lunipes">
<emphasis id="B9261024DD6FFF8FFDFDFDC048D60647" box="[602,722,524,549]" italics="true" pageId="27" pageNumber="28">Atractides</emphasis>
cf.
<emphasis id="B9261024DD6FFF8FFCAEFDC0495A0647" box="[777,862,524,549]" italics="true" pageId="27" pageNumber="28">lunipes</emphasis>
</taxonomicName>
is known from the French Pyrenees,
<collectingCountry id="F3458CA6DD6FFF8FFEAFFDF84B89062C" box="[264,397,564,590]" name="Macedonia" pageId="27" pageNumber="28">Macedonia</collectingCountry>
, and
<collectingCountry id="F3458CA6DD6FFF8FFE68FDF84806062C" box="[463,514,564,590]" name="Iran" pageId="27" pageNumber="28">Iran</collectingCountry>
. The only three findings in widely separate geographical regions is probably due its rareness as well as to our restricted knowledge of the genus
<taxonomicName id="4C52B7B5DD6FFF8FFB15FD904B4306FC" authority="Koch" authorityName="Koch" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9261024DD6FFF8FFB15FD904F2E0617" box="[1202,1322,604,629]" italics="true" pageId="27" pageNumber="28">Atractides</emphasis>
Koch
</taxonomicName>
in large parts of Palaearctic region.
</paragraph>
</subSubSection>
</treatment>
</document>