828 lines
121 KiB
XML
828 lines
121 KiB
XML
<document id="27E577035DEA7B194BDF339FE42E47AA" ID-DOI="10.11646/zootaxa.3931.3.1" ID-GBIF-Dataset="3740223f-a8b0-4fd6-ae57-aec2e7eefde1" ID-ISSN="1175-5326" ID-Zenodo-Dep="244189" ID-ZooBank="5C5A381C-7EB6-4C1F-BAD2-30B45B019E1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461010551014" checkinUser="plazi" docAuthor="Moraza, María L. & Lindquist, Evert E." docDate="2015" docId="03E287ED1A7FCE45EAFAFDDCFDA4D320" docLanguage="en" docName="zt03931p351.pdf" docOrigin="Zootaxa 3931 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hispiniphis Moraza & Lindquist, 2015, new genus" docType="treatment" docVersion="8" lastPageNumber="330" masterDocId="FFDBFF951A66CE58EA6DFFB7FFD4D035" masterDocTitle="Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoidea)" masterLastPageNumber="351" masterPageNumber="301" pageNumber="326" updateTime="1698566786881" updateUser="plazi">
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<mods:titleInfo id="833EDEE7C0F20592C1A37F8ACCE62CD6">
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<mods:title id="2302D86DAC41F53FA3C2FC3A183314F9">Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoidea)</mods:title>
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<mods:name id="A5C59798182AC0C59C3A37872894A8F8" type="personal">
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<mods:roleTerm id="AF2FDA11A9B87CC8385A42FB219C1E49">Author</mods:roleTerm>
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<mods:namePart id="BC48450EE40C97D7B013F2EB2FD093F7">Moraza, María L.</mods:namePart>
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</mods:name>
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<mods:name id="1B3A50C43DBD30D1EE347C389E654A64" type="personal">
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<mods:roleTerm id="D9646F25AE58136D72CDA147780AC427">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7A502918BFD794D3D57FDDD8967D9E5B">Lindquist, Evert E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EF0BF48DC94666B27701425107ECE9CF">text</mods:typeOfResource>
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<mods:titleInfo id="29053B8C100F018259435EAE6903D85F">
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<mods:title id="81BE877834843658F412A2A218ACD4F1">Zootaxa</mods:title>
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<mods:part id="A1A9369823CBDEC4FF0AA1950A1A454A">
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<mods:date id="6ADAD9BD1757525202FAD381C0AE5CA4">2015</mods:date>
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<mods:detail id="20513B23E9783BAB2AF96F736DCA535B" type="volume">
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<mods:number id="E0C0474DF649E41F35EAA839222342BC">3931</mods:number>
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<mods:detail id="974EFC13B3EA0DFDE4B434E7355BBBEA" type="issue">
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<mods:number id="0D9AFACADA5B8A60D8FF6E46C5C81FB2">3</mods:number>
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<mods:classification id="A5D8D5F3FA26217EF11395E4F64DF1F3">journal article</mods:classification>
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<mods:identifier id="6766BEE5FC01881C64B3D8336316808E" type="DOI">10.11646/zootaxa.3931.3.1</mods:identifier>
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<mods:identifier id="A272BDA04D72A2F6BE741977870F9D74" type="GBIF-Dataset">3740223f-a8b0-4fd6-ae57-aec2e7eefde1</mods:identifier>
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<mods:identifier id="17A52BE4E1DF95F6FC3399FBCA5297E8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="ED30178E7CB93D03A3503DBF2956FB01" type="Zenodo-Dep">244189</mods:identifier>
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<treatment id="03E287ED1A7FCE45EAFAFDDCFDA4D320" ID-DOI="http://doi.org/10.5281/zenodo.6115414" ID-GBIF-Taxon="119600154" ID-Zenodo-Dep="6115414" LSID="urn:lsid:plazi:treatment:03E287ED1A7FCE45EAFAFDDCFDA4D320" httpUri="http://treatment.plazi.org/id/03E287ED1A7FCE45EAFAFDDCFDA4D320" lastPageId="29" lastPageNumber="330" pageId="25" pageNumber="326">
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<subSubSection id="C35165701A7FCE41EAFAFDDCFE43D292" pageId="25" pageNumber="326" type="nomenclature">
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<paragraph id="8BF436FB1A7FCE41EAFAFDDCFE78D2B3" blockId="25.[151,428,619,679]" box="[151,428,619,646]" pageId="25" pageNumber="326">
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<heading id="D0BC81971A7FCE41EAFAFDDCFE78D2B3" bold="true" box="[151,428,619,646]" fontSize="11" level="1" pageId="25" pageNumber="326" reason="1">
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<emphasis id="B93FEAE91A7FCE41EAFAFDDCFE78D2B3" bold="true" box="[151,428,619,646]" pageId="25" pageNumber="326">
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<taxonomicName id="4C4B4D781A7FCE41EAFAFDDCFEF7D2B0" box="[151,291,619,645]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B93FEAE91A7FCE41EAFAFDDCFEF7D2B0" bold="true" box="[151,291,619,645]" italics="true" pageId="25" pageNumber="326">Hispiniphis</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57921A7FCE41EB47FDDBFE78D2B3" box="[298,428,620,646]" pageId="25" pageNumber="326" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF436FB1A7FCE41EAFAFD39FE43D292" blockId="25.[151,428,619,679]" box="[151,407,654,679]" pageId="25" pageNumber="326">
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(
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<figureCitation id="13702A7E1A7FCE41EAF2FD39FEF5D292" box="[159,289,654,679]" captionStart-0="FIGURES 41 – 44" captionStart-1="FIGURES 45 – 49" captionStart-2="FIGURES 50 – 55" captionStart-3="FIGURES 56 – 59" captionStart-4="FIGURES 60 – 62" captionStart-5="FIGURES 63 – 74" captionStart-6="FIGURES 75 – 78" captionStart-7="FIGURES 79 – 87" captionStart-8="FIGURE 88" captionStart-9="FIGURES 89 – 92" captionStartId-0="31.[151,264,1905,1928]" captionStartId-1="32.[151,264,1577,1600]" captionStartId-2="33.[151,264,1480,1503]" captionStartId-3="34.[151,264,1557,1580]" captionStartId-4="35.[151,264,1331,1354]" captionStartId-5="36.[151,264,1680,1703]" captionStartId-6="37.[151,264,1869,1892]" captionStartId-7="38.[151,264,1700,1723]" captionStartId-8="39.[151,250,1491,1513]" captionStartId-9="40.[151,264,1884,1907]" captionTargetBox-0="[162,1426,449,1859]" captionTargetBox-1="[163,1431,199,1538]" captionTargetBox-2="[156,1427,349,1450]" captionTargetBox-3="[151,1424,400,1529]" captionTargetBox-4="[156,1429,206,1292]" captionTargetBox-5="[151,1433,305,1644]" captionTargetBox-6="[151,1436,375,1834]" captionTargetBox-7="[151,1436,197,1664]" captionTargetBox-8="[156,1429,204,1457]" captionTargetBox-9="[152,1436,301,1850]" captionTargetId-0="figure@31.[151,1436,445,1885]" captionTargetId-1="figure@32.[151,1436,193,1556]" captionTargetId-2="figure@33.[151,1436,337,1460]" captionTargetId-3="figure@34.[151,1436,391,1544]" captionTargetId-4="figure@35.[151,1436,193,1310]" captionTargetId-5="figure@36.[151,1436,301,1659]" captionTargetId-6="figure@37.[151,1436,373,1849]" captionTargetId-7="figure@38.[151,1436,193,1679]" captionTargetId-8="figure@39.[151,1436,193,1469]" captionTargetId-9="figure@40.[151,1436,301,1864]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="36" captionTargetPageId-6="37" captionTargetPageId-7="38" captionTargetPageId-8="39" captionTargetPageId-9="40" captionText-0="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." captionText-1="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." captionText-2="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." captionText-3="FIGURES 56 – 59. Hispiniphis parva n. sp., adult male: 56. Idiosoma, dorsal aspect; 57. Gnathotectum; 58. Chelicera, lateral view; 59. Detail of movable cheliceral digit and spermatodactyl." captionText-4="FIGURES 60 – 62. Hispiniphis spp., adult male. 60 – 61. Hispiniphis parva n. sp.: 60. Idiosoma, ventral aspect; 61. Subcapitulum. 62. Hispiniphis sp., detail of sternitigenital region." captionText-5="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." captionText-6="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." captionText-7="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." captionText-8="FIGURE 88. Hispiniphis spp., chaetotaxy of dorsal shield, female: 88 a. Hispiniphis parva; 88 b. Hispiniphis sp. 4; 88 c. Hispiniphis sp. 2; 88 d. Hispiniphis sp. 5; 88 e. Hispiniphis sp. 3." captionText-9="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri-0="https://zenodo.org/record/244198/files/figure.png" httpUri-1="https://zenodo.org/record/244199/files/figure.png" httpUri-2="https://zenodo.org/record/244200/files/figure.png" httpUri-3="https://zenodo.org/record/244202/files/figure.png" httpUri-4="https://zenodo.org/record/244203/files/figure.png" httpUri-5="https://zenodo.org/record/244204/files/figure.png" httpUri-6="https://zenodo.org/record/244205/files/figure.png" httpUri-7="https://zenodo.org/record/244206/files/figure.png" httpUri-8="https://zenodo.org/record/244207/files/figure.png" httpUri-9="https://zenodo.org/record/244208/files/figure.png" pageId="25" pageNumber="326">Figs 41–92</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7FCE41EB46FD39FE5AD292" box="[299,398,654,679]" captionStart-0="FIGURES 98 – 101" captionStart-1="FIGURE 102 - 105" captionStartId-0="42.[151,264,1744,1767]" captionStartId-1="43.[151,248,1903,1925]" captionTargetBox-0="[153,1431,185,1707]" captionTargetBox-1="[157,1425,411,1865]" captionTargetId-0="figure@42.[151,1436,180,1730]" captionTargetId-1="figure@43.[151,1436,411,1886]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 98 – 101. Adult idiosomas, ventral view. 98, 99. Makarovaia ornata n. sp.: 98. Female, idiosoma, ventral view; 99. Male, idiosoma, ventral view with attached fungi. 100, 101. Detail of sternal shield, female: 100. Makarovaia ornata n. sp; 101. Hispiniphis parva n. sp." captionText-1="FIGURE 102 - 105. Hispiniphis parva n. sp.: 102. Idiosoma, ventral view, adult female; 103. Detail of anal shield surrounded by soft striated opisthogastric cuticle flanked by thickened, pigmented marginal cuticle, adult female; 104. Detail of posterior ornamentation of dorsal shield and adjacent thickened, pigmented soft cuticle, adult female; 105. Detail of sternitigenital region, adult male" httpUri-0="https://zenodo.org/record/244210/files/figure.png" httpUri-1="https://zenodo.org/record/244211/files/figure.png" pageId="25" pageNumber="326">101–105</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7FCE41EAFAFD61FAAAD326" pageId="25" pageNumber="326" type="materials_examined">
|
||
<paragraph id="8BF436FB1A7FCE41EAFAFD61FAAAD326" blockId="25.[151,1437,726,2011]" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAFAFD61FEE3D2DA" bold="true" box="[151,311,726,751]" pageId="25" pageNumber="326">
|
||
<typeStatus id="54F088591A7FCE41EAFAFD61FF06D2DA" box="[151,210,726,751]" pageId="25" pageNumber="326">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C4B4D781A7FCE41EB2EFD61FDC7D2DB" box="[323,531,726,751]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
|
||
<emphasis id="B93FEAE91A7FCE41EB2EFD61FDC7D2DB" box="[323,531,726,751]" italics="true" pageId="25" pageNumber="326">Hispiniphis parva</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAE91A7FCE41E873FD61FD79D2DA" bold="true" box="[542,685,726,751]" pageId="25" pageNumber="326">
|
||
<taxonomicNameLabel id="A20C57921A7FCE41E873FD61FD79D2DA" box="[542,685,726,751]" pageId="25" pageNumber="326" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Genus based on adult female, adult male, nymphal and larval material representing six species, of which one is newly described here, the others to be described subsequently.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7FCE41EAAAFCA9FCF0D53F" pageId="25" pageNumber="326" type="diagnosis">
|
||
<paragraph id="8BF436FB1A7FCE41EAAAFCA9FCF0D53F" blockId="25.[151,1437,726,2011]" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAAAFCA9FEEED302" bold="true" box="[199,314,798,823]" pageId="25" pageNumber="326">Diagnosis</emphasis>
|
||
. Adult
|
||
<taxonomicName id="4C4B4D781A7FCE41EBFAFCA9FDCCD302" box="[407,536,798,823]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7FCE41EBFAFCA9FDCCD302" box="[407,536,798,823]" italics="true" pageId="25" pageNumber="326">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
are distinguished from those of
|
||
<taxonomicName id="4C4B4D781A7FCE41E9F1FCA9FBF0D302" box="[924,1060,798,823]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="311" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7FCE41E9F1FCA9FBF0D302" box="[924,1060,798,823]" italics="true" pageId="25" pageNumber="326">Makarovaia</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsal shield less extensive anterolaterally, leaving at least setae
|
||
<emphasis id="B93FEAE91A7FCE41E8C9FCF4FD69D36E" box="[676,701,835,859]" italics="true" pageId="25" pageNumber="326">z2</emphasis>
|
||
, all
|
||
<emphasis id="B93FEAE91A7FCE41E883FCF3FD2DD36E" box="[750,761,836,859]" italics="true" pageId="25" pageNumber="326">s</emphasis>
|
||
, and all
|
||
<emphasis id="B93FEAE91A7FCE41E937FCF3FCB1D36E" box="[858,869,836,859]" italics="true" pageId="25" pageNumber="326">r</emphasis>
|
||
setae on adjacent soft cuticle, which has a thicker, micropunctate appearance distinct from the normally striate ventral soft cuticle. Adults have less strongly sclerotised and ornamented shields on the venter, including a smaller, inversely subtriangular anal shield, slightly longer than wide. Male stenitigenital shield incompletely formed, deeply excavated posteriorly, leaving setae
|
||
<emphasis id="B93FEAE91A7FCE41EF34FC18FAAED3F3" box="[1369,1402,943,966]" italics="true" pageId="25" pageNumber="326">st5</emphasis>
|
||
on soft cuticle or on a separated platelet medially. Hypostome of nymphs and adults short, corniculi inserted at level only slightly behind insertions of hypostomatic setae
|
||
<emphasis id="B93FEAE91A7FCE41E972FC41FC9CD43A" box="[799,840,1014,1039]" italics="true" pageId="25" pageNumber="326">hp1</emphasis>
|
||
. Movable cheliceral digit of female and nymphs tridentate, and with pointed process discernible on midventral face. Male chelicerae lacking a digitiform process at level where movable digit hinged to fixed digit, and with spermatodactyl projecting freely anteriorly. In addition to leg chaetotactic reductions noted for genus-group description, femur I and tibia I lacking dorsal seta
|
||
<emphasis id="B93FEAE91A7FCE41EF65FBD5FAEFD44E" box="[1288,1339,1122,1147]" italics="true" pageId="25" pageNumber="326">ad -3</emphasis>
|
||
; genu II lacking lateral seta
|
||
<emphasis id="B93FEAE91A7FCE41EB15FB31FE71D4AA" box="[376,421,1158,1183]" italics="true" pageId="25" pageNumber="326">al -2</emphasis>
|
||
; genua II and III retaining ventral seta
|
||
<emphasis id="B93FEAE91A7FCE41E91FFB30FC5FD4AB" box="[882,907,1159,1182]" italics="true" pageId="25" pageNumber="326">av</emphasis>
|
||
. Tarsus I with ventrodistal acuminate process extending under pretarsus; tarsi II–IV with ventral projection under pretarsus seeming articulated, acuminate in immature instars and adults. Protonymph and to lesser extent deutonymph with opisthosoma reduced in form, size, sometimes flanked closely by posteriorly-directed legs IV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7FCE44EAAAFAA1FC08D63D" lastPageId="28" lastPageNumber="329" pageId="25" pageNumber="326" type="description">
|
||
<paragraph id="8BF436FB1A7FCE41EAAAFAA1FBDED6EA" blockId="25.[151,1437,726,2011]" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAAAFAA1FD99D51A" bold="true" box="[199,589,1302,1327]" pageId="25" pageNumber="326">
|
||
Description.
|
||
<emphasis id="B93FEAE91A7FCE41EB09FAA1FD99D51A" box="[356,589,1302,1327]" italics="true" pageId="25" pageNumber="326">Idiosomatic dorsum</emphasis>
|
||
</emphasis>
|
||
. Adult female (
|
||
<figureCitation id="13702A7E1A7FCE41E96BFAA1FC8BD51A" box="[774,863,1302,1327]" captionStart="FIGURES 41 – 44" captionStartId="31.[151,264,1905,1928]" captionTargetBox="[162,1426,449,1859]" captionTargetId="figure@31.[151,1436,445,1885]" captionTargetPageId="31" captionText="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." httpUri="https://zenodo.org/record/244198/files/figure.png" pageId="25" pageNumber="326">Fig. 42</figureCitation>
|
||
). Dorsal shield moderately well sclerotised, with reticulate ornamentation; shield relatively smaller in size, compared to overall dorsal surface area, not extensive enough to bear setae
|
||
<emphasis id="B93FEAE91A7FCE41EBE8FAE8FE4AD542" box="[389,414,1375,1399]" italics="true" pageId="25" pageNumber="326">s1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EBC1FAE8FE11D542" box="[428,453,1375,1399]" italics="true" pageId="25" pageNumber="326">z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EBB9FAE8FE39D542" box="[468,493,1375,1399]" italics="true" pageId="25" pageNumber="326">r2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EB96FAE8FDC0D542" box="[507,532,1375,1399]" italics="true" pageId="25" pageNumber="326">r3</emphasis>
|
||
which remain with
|
||
<emphasis id="B93FEAE91A7FCE41E89BFAE8FCDBD542" box="[758,783,1375,1399]" italics="true" pageId="25" pageNumber="326">s4</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41E970FAE8FCE2D543" box="[797,822,1375,1398]" italics="true" pageId="25" pageNumber="326">s5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41E928FAE9FC8AD542" box="[837,862,1374,1399]" italics="true" pageId="25" pageNumber="326">s6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41E901FAE8FC51D543" box="[876,901,1375,1398]" italics="true" pageId="25" pageNumber="326">r5</emphasis>
|
||
on surrounding soft cuticle; soft cuticle thickly striate, micropunctate, giving ornamented effect in contrast with opisthogastric soft cuticle (
|
||
<figureCitation id="13702A7E1A7FCE41EECFFA35FAEAD5AE" box="[1186,1342,1410,1435]" captionStart="FIGURE 102 - 105" captionStartId="43.[151,248,1903,1925]" captionTargetBox="[157,1425,411,1865]" captionTargetId="figure@43.[151,1436,411,1886]" captionTargetPageId="43" captionText="FIGURE 102 - 105. Hispiniphis parva n. sp.: 102. Idiosoma, ventral view, adult female; 103. Detail of anal shield surrounded by soft striated opisthogastric cuticle flanked by thickened, pigmented marginal cuticle, adult female; 104. Detail of posterior ornamentation of dorsal shield and adjacent thickened, pigmented soft cuticle, adult female; 105. Detail of sternitigenital region, adult male" httpUri="https://zenodo.org/record/244211/files/figure.png" pageId="25" pageNumber="326">Fig. 103, 104</figureCitation>
|
||
). Dorsal shield with a maximum complement of 13-20 pairs of setae, including seven to 12 podonotal (
|
||
<emphasis id="B93FEAE91A7FCE41EEADFA10FB02D58A" box="[1216,1238,1447,1471]" italics="true" pageId="25" pageNumber="326">j1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EE88FA10FACAD58A" box="[1253,1310,1446,1471]" italics="true" pageId="25" pageNumber="326">j3–j6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EF40FA10FA92D58A" box="[1325,1350,1447,1471]" italics="true" pageId="25" pageNumber="326">z4</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EF39FA10FABAD58B" box="[1364,1390,1447,1470]" italics="true" pageId="25" pageNumber="326">z5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EF11FA11FA41D58A" box="[1404,1429,1446,1471]" italics="true" pageId="25" pageNumber="326">z6</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7FCE41EB76FA7CFEE4D5D6" box="[283,304,1483,1507]" italics="true" pageId="25" pageNumber="326">j1</emphasis>
|
||
off shield, sometimes
|
||
<emphasis id="B93FEAE91A7FCE41E82EFA7CFD8DD5D6" box="[579,601,1483,1507]" italics="true" pageId="25" pageNumber="326">j4</emphasis>
|
||
duplicate) and six to 11 opisthonotal pairs (
|
||
<emphasis id="B93FEAE91A7FCE41EE02FA7CFB48D5D6" box="[1135,1180,1483,1507]" italics="true" pageId="25" pageNumber="326">J1+</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EEDDFA7CFB1FD5D6" box="[1200,1227,1483,1507]" italics="true" pageId="25" pageNumber="326">J3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EEB0FA7CFADED5D6" box="[1245,1290,1483,1507]" italics="true" pageId="25" pageNumber="326">J4+</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EF73FA7CFAECD5D7" box="[1310,1336,1483,1506]" italics="true" pageId="25" pageNumber="326">J5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EF21FA7CFA41D5D7" box="[1356,1429,1483,1507]" italics="true" pageId="25" pageNumber="326">Z2–Z5</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7FCE41EB7AFA58FEE0D632" box="[279,308,1519,1543]" italics="true" pageId="25" pageNumber="326">Z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41EB29FA58FEB5D632" box="[324,353,1519,1543]" italics="true" pageId="25" pageNumber="326">Z3</emphasis>
|
||
, and
|
||
<emphasis id="B93FEAE91A7FCE41EBCFFA58FE14D633" box="[418,448,1519,1542]" italics="true" pageId="25" pageNumber="326">Z5</emphasis>
|
||
duplicate) (
|
||
<figureCitation id="13702A7E1A7FCE41E825FA59FD49D632" box="[584,669,1518,1543]" captionStart="FIGURE 88" captionStartId="39.[151,250,1491,1513]" captionTargetBox="[156,1429,204,1457]" captionTargetId="figure@39.[151,1436,193,1469]" captionTargetPageId="39" captionText="FIGURE 88. Hispiniphis spp., chaetotaxy of dorsal shield, female: 88 a. Hispiniphis parva; 88 b. Hispiniphis sp. 4; 88 c. Hispiniphis sp. 2; 88 d. Hispiniphis sp. 5; 88 e. Hispiniphis sp. 3." httpUri="https://zenodo.org/record/244207/files/figure.png" pageId="25" pageNumber="326">Fig. 88</figureCitation>
|
||
); dorsal setae usually similar in form, somewhat variable in length, except for
|
||
<emphasis id="B93FEAE91A7FCE41EB78F9A4FEE4D61F" box="[277,304,1555,1578]" italics="true" pageId="25" pageNumber="326">J5</emphasis>
|
||
always minute. Dorsal shield with complement of 13–15 pairs of discernible pore-like structures (three to five podonotal, 10 opisthonotal), of which five to seven pairs (two to four podonotal, three or four opisthonotal) superficially appear secretory (gland pores) and seven to nine pairs (one or two podonotal, six or seven opisthonotal) non-secretory (poroids). Soft lateral cuticle with 20–45 pairs of lateral, marginal and submarginal setae, including
|
||
<emphasis id="B93FEAE91A7FCE41EB8FF914FE2FD68E" box="[482,507,1699,1723]" italics="true" pageId="25" pageNumber="326">z2</emphasis>
|
||
, all
|
||
<emphasis id="B93FEAE91A7FCE41E843F913FDEDD68E" box="[558,569,1700,1723]" italics="true" pageId="25" pageNumber="326">s</emphasis>
|
||
, all
|
||
<emphasis id="B93FEAE91A7FCE41E801F913FDA2D68E" box="[620,630,1700,1723]" italics="true" pageId="25" pageNumber="326">r</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7FCE41E968F914FCF7D68E" box="[773,803,1699,1723]" italics="true" pageId="25" pageNumber="326">Z2</emphasis>
|
||
, and all
|
||
<emphasis id="B93FEAE91A7FCE41E9EAF914FC41D68F" box="[903,917,1699,1722]" italics="true" pageId="25" pageNumber="326">S</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7FCE41E9C9F913FC61D68E" box="[932,949,1700,1723]" italics="true" pageId="25" pageNumber="326">R</emphasis>
|
||
, and
|
||
<emphasis id="B93FEAE91A7FCE41E998F913FBCED68E" box="[1013,1050,1700,1723]" italics="true" pageId="25" pageNumber="326">UR</emphasis>
|
||
, and at least six pairs of marginal poroids, including
|
||
<emphasis id="B93FEAE91A7FCE41EB07F971FE4BD6EA" box="[362,415,1734,1759]" italics="true" pageId="25" pageNumber="326">idRp</emphasis>
|
||
; few to many of setae on soft cuticle may be duplicate.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7FCE41EAAAF95DFF3FD77E" blockId="25.[151,1437,726,2011]" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAAAF95DFE8DD737" box="[199,345,1770,1795]" italics="true" pageId="25" pageNumber="326">Adult male (</emphasis>
|
||
<figureCitation id="13702A7E1A7FCE41EB37F95DFE65D736" box="[346,433,1770,1795]" captionStart="FIGURES 56 – 59" captionStartId="34.[151,264,1557,1580]" captionTargetBox="[151,1424,400,1529]" captionTargetId="figure@34.[151,1436,391,1544]" captionTargetPageId="34" captionText="FIGURES 56 – 59. Hispiniphis parva n. sp., adult male: 56. Idiosoma, dorsal aspect; 57. Gnathotectum; 58. Chelicera, lateral view; 59. Detail of movable cheliceral digit and spermatodactyl." httpUri="https://zenodo.org/record/244202/files/figure.png" pageId="25" pageNumber="326">Fig. 56</figureCitation>
|
||
). Dorsal shield usually similar in size, ornamentation and setation to that of female; sometimes capturing one to three more pairs of setae than in female; form and relative lengths of shield setae as in female.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7FCE42EAAAF8E1FED4D29D" blockId="25.[151,1437,726,2011]" lastBlockId="26.[151,1437,151,2013]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAAAF8E1FD97D75A" box="[199,579,1878,1903]" italics="true" pageId="25" pageNumber="326">
|
||
<emphasis id="B93FEAE91A7FCE41EAAAF8E1FE71D75B" bold="true" box="[199,421,1878,1903]" italics="true" pageId="25" pageNumber="326">Idiosomatic venter.</emphasis>
|
||
Adult female
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13702A7E1A7FCE41E838F8E1FD79D75A" box="[597,685,1878,1903]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="25" pageNumber="326">Fig. 45</figureCitation>
|
||
). Tritosternum with laciniae fused for about half of their length, without basal elaborations; laciniae nearly devoid of pilosity (
|
||
<figureCitation id="13702A7E1A7FCE41E93FF8CDFC73D7A6" box="[850,935,1914,1939]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="25" pageNumber="326">Fig. 47</figureCitation>
|
||
). Ventral shields variably poorly sclerotised. Presternal region smooth or faintly lineated, without platelets (
|
||
<figureCitation id="13702A7E1A7FCE41E906F829FC17D782" box="[875,963,1950,1975]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="25" pageNumber="326">Fig. 45</figureCitation>
|
||
). Sternal shield fragmented, its form and setation as noted for genus-group description; sternal area with varying faint ornamentation; shield fragment sometimes with bases of setae
|
||
<emphasis id="B93FEAE91A7CCE42EB9CFF2FFDC6D085" box="[497,530,152,176]" italics="true" pageId="26" pageNumber="327">st1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42E84DFF2FFD95D085" box="[544,577,152,176]" italics="true" pageId="26" pageNumber="327">st2</emphasis>
|
||
circumscribed by sclerotised thickenings, while unsclerotised medial region sometimes faintly ornamented but not striated with soft cuticle. Epigynal shield lightly sclerotised, unornamented, narrow, with hyaline anterior margin hardly widened between legs III, and with posterior margin hardly widened but strongly rounded or bluntly pointed (
|
||
<figureCitation id="13702A7E1A7CCE42E81EFEB3FD1FD129" box="[627,715,260,285]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="26" pageNumber="327">Fig. 48</figureCitation>
|
||
). Opisthosomatic venter lacking metapodal platelets, with soft striated cuticle bearing four to six pairs of opisthogastric setae (
|
||
<emphasis id="B93FEAE91A7CCE42E90DFE9FFC5FD175" box="[864,907,296,320]" italics="true" pageId="26" pageNumber="327">JV1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42E9F4FE9FFC11D175" box="[921,965,296,320]" italics="true" pageId="26" pageNumber="327">JV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42E9BEFE9FFBD6D175" box="[979,1026,296,320]" italics="true" pageId="26" pageNumber="327">ZV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EE7DFE9FFBE8D10A" box="[1040,1084,296,319]" italics="true" pageId="26" pageNumber="327">JV5</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7CCE42EEA5FE9FFB27D175" box="[1224,1267,296,320]" italics="true" pageId="26" pageNumber="327">JV3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EF6FFE9FFAF9D175" box="[1282,1325,296,320]" italics="true" pageId="26" pageNumber="327">JV4</emphasis>
|
||
) and four pairs of poroids, flanked laterally and posteriorly by further thickened, ornate cuticle variably bearing several pairs of
|
||
<emphasis id="B93FEAE91A7CCE42EADBFEC6FF13D1BD" box="[182,199,369,392]" italics="true" pageId="26" pageNumber="327">R</emphasis>
|
||
- or
|
||
<emphasis id="B93FEAE91A7CCE42EA95FEC6FEC9D1BD" box="[248,285,369,392]" italics="true" pageId="26" pageNumber="327">UR</emphasis>
|
||
- setae. Anal shield small, well sclerotised, smooth or reticulated, pyriform or inversely subtriangular, slightly longer than wide; para-anal setae inserted at mid-level of anus, shorter than postanal seta; shield with pair of gland pores
|
||
<emphasis id="B93FEAE91A7CCE42EB20FE0FFEA1D1E5" box="[333,373,440,464]" italics="true" pageId="26" pageNumber="327">gv3</emphasis>
|
||
on lateral edges at level of insertions of para-anal setae, and with cribrum along tapered posterior margin (
|
||
<figureCitation id="13702A7E1A7CCE42EB0BFE6BFE69D1C1" box="[358,445,476,501]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="26" pageNumber="327">Fig. 49</figureCitation>
|
||
). Peritrematal shield weakly sclerotised, extending without interruption to about midlevel of coxae I; shield with gland pore
|
||
<emphasis id="B93FEAE91A7CCE42E839FDB7FDAAD22D" box="[596,638,512,536]" italics="true" pageId="26" pageNumber="327">gp1</emphasis>
|
||
located at anterior extension at level of coxa I (
|
||
<figureCitation id="13702A7E1A7CCE42EEF9FE48FB3CD22D" box="[1172,1256,511,536]" captionStart="FIGURES 41 – 44" captionStartId="31.[151,264,1905,1928]" captionTargetBox="[162,1426,449,1859]" captionTargetId="figure@31.[151,1436,445,1885]" captionTargetPageId="31" captionText="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." httpUri="https://zenodo.org/record/244198/files/figure.png" pageId="26" pageNumber="327">Fig. 43</figureCitation>
|
||
); exopodal strip continuous alongside peritrematal shield only to level of coxae III, extensions between bases of coxae II–III rudimentary or absent. Peritremes thick, reaching at most to level of anterior margin of coxae II (
|
||
<figureCitation id="13702A7E1A7CCE42EF6AFDF0FAB5D255" box="[1287,1377,583,608]" captionStart="FIGURES 41 – 44" captionStartId="31.[151,264,1905,1928]" captionTargetBox="[162,1426,449,1859]" captionTargetId="figure@31.[151,1436,445,1885]" captionTargetPageId="31" captionText="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." httpUri="https://zenodo.org/record/244198/files/figure.png" pageId="26" pageNumber="327">Figs 43</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7CCE42EF1DFDF0FA44D255" box="[1392,1424,583,608]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="26" pageNumber="327">45</figureCitation>
|
||
). Spermathecal apparatus sometimes with small section of tubular piece discernibly sclerotised, slightly widened (
|
||
<figureCitation id="13702A7E1A7CCE42EAF2FD38FF27D29D" box="[159,243,655,680]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Fig. 55</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7CCE42EAAAFD03FDB9D3FD" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="327">
|
||
<emphasis id="B93FEAE91A7CCE42EAAAFD03FE90D2F8" box="[199,324,692,717]" italics="true" pageId="26" pageNumber="327">Adult male</emphasis>
|
||
(
|
||
<figureCitation id="13702A7E1A7CCE42EB35FD03FE65D2F8" box="[344,433,692,717]" captionStart="FIGURES 60 – 62" captionStartId="35.[151,264,1331,1354]" captionTargetBox="[156,1429,206,1292]" captionTargetId="figure@35.[151,1436,193,1310]" captionTargetPageId="35" captionText="FIGURES 60 – 62. Hispiniphis spp., adult male. 60 – 61. Hispiniphis parva n. sp.: 60. Idiosoma, ventral aspect; 61. Subcapitulum. 62. Hispiniphis sp., detail of sternitigenital region." httpUri="https://zenodo.org/record/244203/files/figure.png" pageId="26" pageNumber="327">Fig. 60</figureCitation>
|
||
). Form of tritosternum and presternal area as in female. Sternitigenital shield anterior margin well defined at level of genital opening; shield lightly sclerotised, united with endopodal strips between coxae I–II, II–III, III–IV; shield with four pairs of setae,
|
||
<emphasis id="B93FEAE91A7CCE42E95CFD4BFCAFD321" box="[817,891,764,788]" italics="true" pageId="26" pageNumber="327">st1-st4</emphasis>
|
||
, and two pairs of poroids; shield incompletely formed, deeply excavated posteriorly, leaving setae
|
||
<emphasis id="B93FEAE91A7CCE42E8B3FC97FD2BD302" box="[734,767,800,823]" italics="true" pageId="26" pageNumber="327">st5</emphasis>
|
||
on soft cuticle or on a separated platelet medially (
|
||
<figureCitation id="13702A7E1A7CCE42EF2DFCA8FF63D368" captionStart="FIGURES 60 – 62" captionStartId="35.[151,264,1331,1354]" captionTargetBox="[156,1429,206,1292]" captionTargetId="figure@35.[151,1436,193,1310]" captionTargetPageId="35" captionText="FIGURES 60 – 62. Hispiniphis spp., adult male. 60 – 61. Hispiniphis parva n. sp.: 60. Idiosoma, ventral aspect; 61. Subcapitulum. 62. Hispiniphis sp., detail of sternitigenital region." httpUri="https://zenodo.org/record/244203/files/figure.png" pageId="26" pageNumber="327">Figs 60, 62</figureCitation>
|
||
). Form and extent of exopodal and peritrematal structures much as in female (
|
||
<figureCitation id="13702A7E1A7CCE42EE51FCF3FB47D368" box="[1084,1171,836,861]" captionStart="FIGURES 60 – 62" captionStartId="35.[151,264,1331,1354]" captionTargetBox="[156,1429,206,1292]" captionTargetId="figure@35.[151,1436,193,1310]" captionTargetPageId="35" captionText="FIGURES 60 – 62. Hispiniphis spp., adult male. 60 – 61. Hispiniphis parva n. sp.: 60. Idiosoma, ventral aspect; 61. Subcapitulum. 62. Hispiniphis sp., detail of sternitigenital region." httpUri="https://zenodo.org/record/244203/files/figure.png" pageId="26" pageNumber="327">Fig. 60</figureCitation>
|
||
). Opisthogastric region much as in female, with same complement of poroids but only four or five pairs of opisthogastric setae (
|
||
<emphasis id="B93FEAE91A7CCE42EF42FCDFFA8FD3B5" box="[1327,1371,872,896]" italics="true" pageId="26" pageNumber="327">JV1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EF04FCDFFA41D3B5" box="[1385,1429,872,896]" italics="true" pageId="26" pageNumber="327">JV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EAFAFC3BFF12D391" box="[151,198,908,932]" italics="true" pageId="26" pageNumber="327">ZV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EABEFC3AFF2BD391" box="[211,255,909,932]" italics="true" pageId="26" pageNumber="327">JV5</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7CCE42EBE7FC3BFE62D391" box="[394,438,908,932]" italics="true" pageId="26" pageNumber="327">JV3</emphasis>
|
||
). All ventral idiosomatic setae smooth, similar in shape and length as female. Anal shield similar in form and structure as in female.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7CCE42EAAAFC64FBB0D5F5" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="327">
|
||
<emphasis id="B93FEAE91A7CCE42EAAAFC64FEB6D3D8" bold="true" box="[199,354,979,1005]" pageId="26" pageNumber="327">
|
||
<taxonomicName id="4C4B4D781A7CCE42EAAAFC64FE8FD3D9" box="[199,347,979,1004]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="26" pageNumber="327" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7CCE42EAAAFC64FE8FD3D9" bold="true" box="[199,347,979,1004]" italics="true" pageId="26" pageNumber="327">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B93FEAE91A7CCE42EB1DFC63FD95D3D8" box="[368,577,980,1005]" italics="true" pageId="26" pageNumber="327">Female and male</emphasis>
|
||
. Gnathotectum with anterior margin convex, denticulate (
|
||
<figureCitation id="13702A7E1A7CCE42EE93FC63FA89D3D8" box="[1278,1373,980,1005]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Figs 52</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7CCE42EF1DFC63FA44D3D8" box="[1392,1424,980,1005]" captionStart="FIGURES 56 – 59" captionStartId="34.[151,264,1557,1580]" captionTargetBox="[151,1424,400,1529]" captionTargetId="figure@34.[151,1436,391,1544]" captionTargetPageId="34" captionText="FIGURES 56 – 59. Hispiniphis parva n. sp., adult male: 56. Idiosoma, dorsal aspect; 57. Gnathotectum; 58. Chelicera, lateral view; 59. Detail of movable cheliceral digit and spermatodactyl." httpUri="https://zenodo.org/record/244202/files/figure.png" pageId="26" pageNumber="327">57</figureCitation>
|
||
). Chelicerae generally as described for the genus-group description. Fixed cheliceral digit sparsely multidentate (offset subapical tooth and six to nine teeth), with pilus dentilis reduced to alveolar remnant or indiscernible; movable digit of female tridentate, with a small pointed process on mid-ventral face (
|
||
<figureCitation id="13702A7E1A7CCE42EE36FB88FB64D46D" box="[1115,1200,1087,1112]" captionStart="FIGURES 41 – 44" captionStartId="31.[151,264,1905,1928]" captionTargetBox="[162,1426,449,1859]" captionTargetId="figure@31.[151,1436,445,1885]" captionTargetPageId="31" captionText="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." httpUri="https://zenodo.org/record/244198/files/figure.png" pageId="26" pageNumber="327">Fig. 44</figureCitation>
|
||
). Movable cheliceral digit of male with one small tooth, and lacking small process on ventral face; spermatodactyl digit-like, variously sinuous, directed anteriorly, not recurved basally (
|
||
<figureCitation id="13702A7E1A7CCE42E88AFB30FCA4D495" box="[743,880,1159,1184]" captionStart="FIGURES 56 – 59" captionStartId="34.[151,264,1557,1580]" captionTargetBox="[151,1424,400,1529]" captionTargetId="figure@34.[151,1436,391,1544]" captionTargetPageId="34" captionText="FIGURES 56 – 59. Hispiniphis parva n. sp., adult male: 56. Idiosoma, dorsal aspect; 57. Gnathotectum; 58. Chelicera, lateral view; 59. Detail of movable cheliceral digit and spermatodactyl." httpUri="https://zenodo.org/record/244202/files/figure.png" pageId="26" pageNumber="327">Figs 58–59</figureCitation>
|
||
); male chela lacking any arthrodial process on paraxial surface at base of movable digit. Corniculi of female and male similar in form, short, stout, inserted at level of insertions of setae
|
||
<emphasis id="B93FEAE91A7CCE42EBB6FB78FDD2D4DD" box="[475,518,1231,1256]" italics="true" pageId="26" pageNumber="327">hp1</emphasis>
|
||
(
|
||
<figureCitation id="13702A7E1A7CCE42E874FB78FD77D4DD" box="[537,675,1231,1256]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Figs 51, 53</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7CCE42E8D9FB78FD07D4DD" box="[692,723,1231,1256]" captionStart="FIGURES 60 – 62" captionStartId="35.[151,264,1331,1354]" captionTargetBox="[156,1429,206,1292]" captionTargetId="figure@35.[151,1436,193,1310]" captionTargetPageId="35" captionText="FIGURES 60 – 62. Hispiniphis spp., adult male. 60 – 61. Hispiniphis parva n. sp.: 60. Idiosoma, ventral aspect; 61. Subcapitulum. 62. Hispiniphis sp., detail of sternitigenital region." httpUri="https://zenodo.org/record/244203/files/figure.png" pageId="26" pageNumber="327">61</figureCitation>
|
||
); paraxial process at base of corniculus slightly shorter than corniculus (
|
||
<figureCitation id="13702A7E1A7CCE42EB76FB43FEBBD538" box="[283,367,1268,1293]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Fig. 51</figureCitation>
|
||
); bifid internal malae elongate, extending nearly to or beyond apex of palpgenu (
|
||
<figureCitation id="13702A7E1A7CCE42EE93FB43FA86D538" box="[1278,1362,1268,1293]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Fig. 53</figureCitation>
|
||
); apex of labrum extending to palptibia. Subcapitulum with a pair of pore-like structures at level of insertion of corniculi (
|
||
<figureCitation id="13702A7E1A7CCE42EAF2FA8BFF27D560" box="[159,243,1340,1365]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Fig. 53</figureCitation>
|
||
). Subcapitular setae smooth, with
|
||
<emphasis id="B93FEAE91A7CCE42E81DFA8BFD4DD560" box="[624,665,1340,1365]" italics="true" pageId="26" pageNumber="327">hp1</emphasis>
|
||
clearly longer and thicker than other pairs. Deutosternum with seven moderately narrow, multidenticulate rows of denticles, connected by lateral margins; basal two rows not narrowed, sometimes more finely multidenticulate. Palpal setation and form of setae as described for genus-group description (
|
||
<figureCitation id="13702A7E1A7CCE42EAF2FA10FF26D5F5" box="[159,242,1447,1472]" captionStart="FIGURES 50 – 55" captionStartId="33.[151,264,1480,1503]" captionTargetBox="[156,1427,349,1450]" captionTargetId="figure@33.[151,1436,337,1460]" captionTargetPageId="33" captionText="FIGURES 50 – 55. Hispiniphis spp., adult female. 50 – 54. Hispiniphis parva n. sp.: 50. Detail of corniculus and salivary stylus, dorsal aspect; 51. Corniculus with basal process, ventral aspect; 52. Gnathotectum, variants; 53. Subcapitulum; 54 a. Palp, dorsal view with detail of several palpal setae (left call-outs) (dorsal setae in solid black color); 54 b. Palptarsus, ventral view; 54 c. Palptarsus dorsal view. 55. Hispiniphis sp., detail of spermatheca." httpUri="https://zenodo.org/record/244200/files/figure.png" pageId="26" pageNumber="327">Fig. 54</figureCitation>
|
||
a–c); palptrochanter with inner seta somewhat longer than external setae (54a).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7CCE43EAAAFA7BFECFD0E0" blockId="26.[151,1437,151,2013]" lastBlockId="27.[151,1437,151,2013]" lastPageId="27" lastPageNumber="328" pageId="26" pageNumber="327">
|
||
<emphasis id="B93FEAE91A7CCE42EAAAFA7BFED0D5D0" bold="true" box="[199,260,1484,1509]" pageId="26" pageNumber="327">
|
||
<emphasis id="B93FEAE91A7CCE42EAAAFA7BFF29D5D6" bold="true" box="[199,253,1484,1507]" italics="true" pageId="26" pageNumber="327">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B93FEAE91A7CCE42EB62FA7BFE0ED5D0" box="[271,474,1484,1509]" italics="true" pageId="26" pageNumber="327">Female and male</emphasis>
|
||
. Legs moderately short, similar in length to dorsal shield but clearly shorter than idiosoma. Legs I and IV similar in length and thickness to legs II, III (
|
||
<figureCitation id="13702A7E1A7CCE42E987FA58FBA6D63D" box="[1002,1138,1519,1544]" captionStart="FIGURES 63 – 74" captionStartId="36.[151,264,1680,1703]" captionTargetBox="[151,1433,305,1644]" captionTargetId="figure@36.[151,1436,301,1659]" captionTargetPageId="36" captionText="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." httpUri="https://zenodo.org/record/244204/files/figure.png" pageId="26" pageNumber="327">Figs 63–70</figureCitation>
|
||
). Tarsus I with sensilla
|
||
<emphasis id="B93FEAE91A7CCE42EFFCFA46FA48D63D" box="[1425,1436,1521,1544]" italics="true" pageId="26" pageNumber="327">s</emphasis>
|
||
inconspicuous, short and acute (
|
||
<figureCitation id="13702A7E1A7CCE42EB92F9A3FD87D619" box="[511,595,1556,1581]" captionStart="FIGURES 63 – 74" captionStartId="36.[151,264,1680,1703]" captionTargetBox="[151,1433,305,1644]" captionTargetId="figure@36.[151,1436,301,1659]" captionTargetPageId="36" captionText="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." httpUri="https://zenodo.org/record/244204/files/figure.png" pageId="26" pageNumber="327">Fig. 72</figureCitation>
|
||
a), and with ventrodistal acuminate process extending under pretarsus (
|
||
<figureCitation id="13702A7E1A7CCE42EF02F9A3FF62D665" captionStart="FIGURES 63 – 74" captionStartId="36.[151,264,1680,1703]" captionTargetBox="[151,1433,305,1644]" captionTargetId="figure@36.[151,1436,301,1659]" captionTargetPageId="36" captionText="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." httpUri="https://zenodo.org/record/244204/files/figure.png" pageId="26" pageNumber="327">Fig. 72</figureCitation>
|
||
b). Pretarsus of legs I to IV with slender claws and large, rounded pulvillus; those of legs II–IV with short, inconspicuous paradactyli formed as narrow sclerotised shafts flanking claws (
|
||
<figureCitation id="13702A7E1A7CCE42EE7AF9EBFBB9D641" box="[1047,1133,1628,1653]" captionStart="FIGURES 63 – 74" captionStartId="36.[151,264,1680,1703]" captionTargetBox="[151,1433,305,1644]" captionTargetId="figure@36.[151,1436,301,1659]" captionTargetPageId="36" captionText="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." httpUri="https://zenodo.org/record/244204/files/figure.png" pageId="26" pageNumber="327">Fig. 74</figureCitation>
|
||
). Legs II to IV with tarsus (excluding pretarsus) about twice as long as tibia. Tarsi II–IV with apical setal processes
|
||
<emphasis id="B93FEAE91A7CCE42EE14F9C8FB79D6AD" box="[1145,1197,1663,1688]" italics="true" pageId="26" pageNumber="327">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EED6F9C8FB3AD6AD" box="[1211,1262,1663,1688]" italics="true" pageId="26" pageNumber="327">pd -1</emphasis>
|
||
inconspicuous, shorter than short pretarsi, and with apical ventral process short, triangular, more retrorse in male than in female (
|
||
<figureCitation id="13702A7E1A7CCE42EAF2F970FEF4D6D5" box="[159,288,1735,1760]" captionStart="FIGURES 63 – 74" captionStartId="36.[151,264,1680,1703]" captionTargetBox="[151,1433,305,1644]" captionTargetId="figure@36.[151,1436,301,1659]" captionTargetPageId="36" captionText="FIGURES 63 – 74. Hispiniphis parva n. sp.: 63 – 66. Adult female, right legs I to IV, excepting coxae and tarsi, dorsal views (dorsal setae in solid black color): 63. Leg I; 64. Leg II; 65. Leg III; 66 Leg IV; 67 - 70. Adult female, coxae I to IV, ventral views: 67. Coxa I; 68. Coxa II; 69. Coxa III; 70. Coxa IV; 71 - 72. Tarsus I, right leg, adult male: 71. Dorsal view, (basal dorsal setae in solid black color); 72 a. Detail of sensorial apical setae; 72 b. Detail of ventro-apical process; 73 - 74. Tarsus II, right leg, adult female: 73. Dorsal view, (dorsal setae in solid black color); 74. Detail of acuminate ventroapical process extending beneath pretarsus." httpUri="https://zenodo.org/record/244204/files/figure.png" pageId="26" pageNumber="327">Figs 73, 74</figureCitation>
|
||
). Setation and its ontogeny on segments of legs I to IV deficient from full complement of
|
||
<taxonomicName id="4C4B4D781A7CCE42EF65F970FA48D6D5" box="[1288,1436,1735,1760]" class="Arachnida" family="Ascidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="family">Melicharidae</taxonomicName>
|
||
(as presented by
|
||
<bibRefCitation id="EFDA4B0A1A7CCE42EB3BF95BFDBFD730" author="Lindquist" box="[342,619,1772,1797]" pageId="26" pageNumber="327" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans 1965</bibRefCitation>
|
||
for Melicharini), coxae, 2-2-2-1; trochanters, 5-5-5-5; femora, 11 (2 2/1 2/2 2) – 9 or 10 (1 2 or 3/1 2/2 1) – 6 (1 2/1 2/0 0) – 6 (1 2/1 2/0 0); genua, 11 (2 3/1 2/1 2) – 8 (1 3/1 2/0 1) – 7 (1 2/1 2/0 1) – 7 (1 2/0 3/0 1); tibiae, 10 (2 2/1 2/1 2) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1); trochanter I seta
|
||
<emphasis id="B93FEAE91A7CCE42EAA1F8E0FF3CD745" box="[204,232,1879,1904]" italics="true" pageId="26" pageNumber="327">ad</emphasis>
|
||
vestigial, only alveolus discernible. In addition to setal absences noted for family: femur I, tibia I, and sometimes femur II lacking seta
|
||
<emphasis id="B93FEAE91A7CCE42E863F8CBFD95D7A1" box="[526,577,1916,1941]" italics="true" pageId="26" pageNumber="327">ad -3</emphasis>
|
||
; genu II lacking seta
|
||
<emphasis id="B93FEAE91A7CCE42E951F8CBFCBDD7A1" box="[828,873,1916,1941]" italics="true" pageId="26" pageNumber="327">al -2</emphasis>
|
||
; however, genua II–III retaining
|
||
<emphasis id="B93FEAE91A7CCE42EE88F8CAFB2AD7A1" box="[1253,1278,1917,1940]" italics="true" pageId="26" pageNumber="327">av</emphasis>
|
||
, and genu IV retaining
|
||
<emphasis id="B93FEAE91A7CCE42EB69F817FEE3D78D" box="[260,311,1952,1977]" italics="true" pageId="26" pageNumber="327">pd -3</emphasis>
|
||
. Several setae moderately thickened, spinelike:
|
||
<emphasis id="B93FEAE91A7CCE42E90CF817FC40D78D" box="[865,916,1952,1977]" italics="true" pageId="26" pageNumber="327">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42E9C8F817FC0CD78D" box="[933,984,1952,1977]" italics="true" pageId="26" pageNumber="327">pd -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42E985F817FBC8D78D" box="[1000,1052,1952,1977]" italics="true" pageId="26" pageNumber="327">pd -2</emphasis>
|
||
on femora I–II and
|
||
<emphasis id="B93FEAE91A7CCE42EF64F817FAE7D78C" box="[1289,1331,1952,1977]" italics="true" pageId="26" pageNumber="327">ad1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7CCE42EF2EF817FAA2D78D" box="[1347,1398,1952,1977]" italics="true" pageId="26" pageNumber="327">pd -1</emphasis>
|
||
on femora III–IV;
|
||
<emphasis id="B93FEAE91A7CCE42EB21F873FE54D7E9" box="[332,384,1988,2013]" italics="true" pageId="26" pageNumber="327">ad -2</emphasis>
|
||
on femur IV. Seta
|
||
<emphasis id="B93FEAE91A7CCE42E802F872FD5DD7E9" box="[623,649,1989,2012]" italics="true" pageId="26" pageNumber="327">av</emphasis>
|
||
on femur III and
|
||
<emphasis id="B93FEAE91A7CCE42E906F872FC49D7E9" box="[875,925,1988,2012]" italics="true" pageId="26" pageNumber="327">pv -2</emphasis>
|
||
on trochanters I–IV (on female and male) attenuated. Other leg setae simple, not strongly differentiated on either sex (male leg setal dimorphism not developed).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFF68FE81D198" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFF68FEB7D0CD" bold="true" box="[199,355,223,248]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFF68FE88D0CD" bold="true" box="[199,348,223,248]" italics="true" pageId="27" pageNumber="328">Deutonymph</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B93FEAE91A7DCE43EB06FF68FD9FD0CD" box="[363,587,223,248]" italics="true" pageId="27" pageNumber="328">Idiosomatic dorsum</emphasis>
|
||
. Discoidal form. Dorsal shield poorly sclerotised, without lateral incisions, lightly lineated, with
|
||
<emphasis id="B93FEAE91A7DCE43EBE8FEB2FE4BD129" box="[389,415,261,284]" italics="true" pageId="27" pageNumber="328">ca</emphasis>
|
||
. 35 pairs of similarly short setae except more posterior ones smaller: 17 pairs (
|
||
<emphasis id="B93FEAE91A7DCE43EF7CFEB3FAF3D129" box="[1297,1319,260,284]" italics="true" pageId="27" pageNumber="328">j1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EF58FEB3FABAD128" box="[1333,1390,260,285]" italics="true" pageId="27" pageNumber="328">j3–j6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EF11FEB3FA41D129" box="[1404,1429,260,284]" italics="true" pageId="27" pageNumber="328">z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EAFAFE90FF02D175" box="[151,214,295,320]" italics="true" pageId="27" pageNumber="328">z4–z6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EA89FE9FFF29D175" box="[228,253,296,320]" italics="true" pageId="27" pageNumber="328">s1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EB66FE9FFEF0D175" box="[267,292,296,320]" italics="true" pageId="27" pageNumber="328">s2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EB5CFE90FEA5D175" box="[305,369,295,320]" italics="true" pageId="27" pageNumber="328">s4–s6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EB12FE9FFE4CD175" box="[383,408,296,320]" italics="true" pageId="27" pageNumber="328">r2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EBC8FE9FFE6BD175" box="[421,447,296,320]" italics="true" pageId="27" pageNumber="328">r3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EBA1FE9FFE31D10A" box="[460,485,296,319]" italics="true" pageId="27" pageNumber="328">r5</emphasis>
|
||
) on podonotal region, and a maximum 18 pairs (
|
||
<emphasis id="B93FEAE91A7DCE43EE6AFE9FFBF5D175" box="[1031,1057,296,320]" italics="true" pageId="27" pageNumber="328">J1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE45FE9FFB96D175" box="[1064,1090,296,320]" italics="true" pageId="27" pageNumber="328">J3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE3DFE9FFBBFD175" box="[1104,1131,296,320]" italics="true" pageId="27" pageNumber="328">J4</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE15FE9FFB47D10A" box="[1144,1171,296,319]" italics="true" pageId="27" pageNumber="328">J5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EECCFE9FFB3ED175" box="[1185,1258,296,320]" italics="true" pageId="27" pageNumber="328">Z2–Z5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE9AFE9FFAE9D175" box="[1271,1341,296,320]" italics="true" pageId="27" pageNumber="328">S1–S5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EF26FE9FFA47D175" box="[1355,1427,295,320]" italics="true" pageId="27" pageNumber="328">R1-R6</emphasis>
|
||
) on opisthonotal region (
|
||
<figureCitation id="13702A7E1A7DCE43EBC7FEFBFDD1D150" box="[426,517,332,357]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Figs 75</figureCitation>
|
||
); many of these setae displaced more marginally on dorsal shield than in adult. Ventrolateral soft cuticle nearly devoid of setae. Dorsal idiosomatic complement of poroids and gland-pores similar to that in female.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFE00FD33D30D" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFE00FE4DD1E5" box="[199,409,439,464]" italics="true" pageId="27" pageNumber="328">Idiosomatic venter</emphasis>
|
||
. Sternal region lightly sclerotised, unornamented, without endopodal extensions between coxae, with three pairs of platelets bearing sternal setae
|
||
<emphasis id="B93FEAE91A7DCE43E974FE6BFC89D1C1" box="[793,861,476,500]" italics="true" pageId="27" pageNumber="328">s1-st3</emphasis>
|
||
and poroids
|
||
<emphasis id="B93FEAE91A7DCE43E99EFE6BFBC1D1C1" box="[1011,1045,476,500]" italics="true" pageId="27" pageNumber="328">
|
||
<date id="FFF5103B1A7DCE43E99EFE6BFBC1D1C1" box="[1011,1045,476,500]" pageId="27" pageNumber="328">iv1</date>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE48FE6BFB93D1C1" box="[1061,1095,476,500]" italics="true" pageId="27" pageNumber="328">
|
||
<date id="FFF5103B1A7DCE43EE48FE6BFB93D1C1" box="[1061,1095,476,500]" pageId="27" pageNumber="328">iv2</date>
|
||
</emphasis>
|
||
; setae
|
||
<emphasis id="B93FEAE91A7DCE43EEF5FE6BFB37D1C1" box="[1176,1251,476,500]" italics="true" pageId="27" pageNumber="328">st4-st5</emphasis>
|
||
inserted on soft cuticle between coxae III and IV (
|
||
<figureCitation id="13702A7E1A7DCE43E82DFE48FD48D22D" box="[576,668,511,536]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Fig. 76</figureCitation>
|
||
); poroids
|
||
<emphasis id="B93FEAE91A7DCE43E977FDB7FCE8D222" box="[794,828,512,535]" italics="true" pageId="27" pageNumber="328">
|
||
<date id="FFF5103B1A7DCE43E977FDB7FCE8D222" box="[794,828,512,535]" pageId="27" pageNumber="328">iv5</date>
|
||
</emphasis>
|
||
absent. Anal shield prominent, lightly sclerotised, unornamented, roughly rounded, with circumanal setae similar in length; adanal gland pores (
|
||
<emphasis id="B93FEAE91A7DCE43EEDFFD93FB0ED209" box="[1202,1242,548,572]" italics="true" pageId="27" pageNumber="328">gv3</emphasis>
|
||
) at level of paraanal setae; cribrum as in adult female. Opisthogaster with four to six pairs of ventral setae (the four larval
|
||
<emphasis id="B93FEAE91A7DCE43EF5DFDFFFA88D255" box="[1328,1372,584,608]" italics="true" pageId="27" pageNumber="328">JV1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EF04FDFFFA41D255" box="[1385,1429,584,608]" italics="true" pageId="27" pageNumber="328">JV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EAFAFDDAFF17D2B1" box="[151,195,621,644]" italics="true" pageId="27" pageNumber="328">JV5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EABCFDDBFED4D2B1" box="[209,256,620,644]" italics="true" pageId="27" pageNumber="328">ZV2</emphasis>
|
||
, sometimes plus two minute
|
||
<emphasis id="B93FEAE91A7DCE43E821FDDBFDACD2B1" box="[588,632,620,644]" italics="true" pageId="27" pageNumber="328">JV3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E8EAFDDBFD67D2B1" box="[647,691,620,644]" italics="true" pageId="27" pageNumber="328">JV4</emphasis>
|
||
), sometimes not flanked by further setae on soft cuticle (
|
||
<figureCitation id="13702A7E1A7DCE43EF57FDDBFA44D2B0" box="[1338,1424,620,645]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Fig. 76</figureCitation>
|
||
). Peritrematal shields and peritremes shortened, paedomorphic, as in protonymphal instar, developed only alongside level between coxae III–IV (
|
||
<figureCitation id="13702A7E1A7DCE43EBB7FD03FDFBD2F8" box="[474,559,692,717]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Fig. 76</figureCitation>
|
||
). Peritrematal gland pore
|
||
<emphasis id="B93FEAE91A7DCE43E93FFD03FCA9D2F9" box="[850,893,692,716]" italics="true" pageId="27" pageNumber="328">gp2</emphasis>
|
||
and poroid
|
||
<emphasis id="B93FEAE91A7DCE43EE69FD03FBFCD2F9" box="[1028,1064,692,716]" italics="true" pageId="27" pageNumber="328">ip2</emphasis>
|
||
on soft cuticle between levels of coxae II and III; poststigmatic poroid
|
||
<emphasis id="B93FEAE91A7DCE43E804FD6EFD5ED2C5" box="[617,650,729,752]" italics="true" pageId="27" pageNumber="328">ips</emphasis>
|
||
adherent to stigma on posterior edge of peritrematal shield, and poststigmatic gland pore
|
||
<emphasis id="B93FEAE91A7DCE43EBDEFD4BFE0AD321" box="[435,478,764,788]" italics="true" pageId="27" pageNumber="328">gp3</emphasis>
|
||
and poroid
|
||
<emphasis id="B93FEAE91A7DCE43E80AFD4BFD5ED321" box="[615,650,764,788]" italics="true" pageId="27" pageNumber="328">ip3</emphasis>
|
||
on soft cuticle. Rim of exopodal plate behind coxa IV inconspicuous or indiscernible, with gland pore
|
||
<emphasis id="B93FEAE91A7DCE43E866FC97FDE7D30D" box="[523,563,800,824]" italics="true" pageId="27" pageNumber="328">gv2</emphasis>
|
||
not discernible.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFCF3FBC7D3B5" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<taxonomicName id="4C4B4D781A7DCE43EAAAFCF3FE82D368" box="[199,342,836,861]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="27" pageNumber="328" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFCF3FE82D368" box="[199,342,836,861]" italics="true" pageId="27" pageNumber="328">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Gnathotectum, chelicerae and other mouthpart structures, corniculi and adjacent structures as in adult female; subcapitular and palpal structures similar to those in adult female.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFC3AFE6AD46D" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFC3AFF28D391" box="[199,252,909,932]" italics="true" pageId="27" pageNumber="328">Legs</emphasis>
|
||
. Pretarsal structures, and chaetotaxy of legs I–IV as in adult, including these additions to protonymphal setal complement:
|
||
<emphasis id="B93FEAE91A7DCE43EB01FC06FE49D3FD" box="[364,413,944,968]" italics="true" pageId="27" pageNumber="328">av -2</emphasis>
|
||
on trochanters I–IV;
|
||
<emphasis id="B93FEAE91A7DCE43E8E2FC18FD68D3FD" box="[655,700,943,968]" italics="true" pageId="27" pageNumber="328">al -2</emphasis>
|
||
and
|
||
<emphasis id="B93FEAE91A7DCE43E899FC18FCF6D3FD" box="[756,802,943,968]" italics="true" pageId="27" pageNumber="328">pl -2</emphasis>
|
||
on genu I,
|
||
<emphasis id="B93FEAE91A7DCE43E9CEFC18FC03D3FD" box="[931,983,943,968]" italics="true" pageId="27" pageNumber="328">ad -3</emphasis>
|
||
on genua I–II,
|
||
<emphasis id="B93FEAE91A7DCE43EEE8FC18FB6DD3FD" box="[1157,1209,943,968]" italics="true" pageId="27" pageNumber="328">pd -3</emphasis>
|
||
on genu IV,
|
||
<emphasis id="B93FEAE91A7DCE43EF26FC18FAACD3FD" box="[1355,1400,943,968]" italics="true" pageId="27" pageNumber="328">pl -1</emphasis>
|
||
on genu IV,
|
||
<emphasis id="B93FEAE91A7DCE43EA91FC62FEF9D3D9" box="[252,301,980,1004]" italics="true" pageId="27" pageNumber="328">av -1</emphasis>
|
||
on genua II–IV,
|
||
<emphasis id="B93FEAE91A7DCE43EB84FC62FDCED3D9" box="[489,538,980,1004]" italics="true" pageId="27" pageNumber="328">pv -1</emphasis>
|
||
on genu II,
|
||
<emphasis id="B93FEAE91A7DCE43E8CCFC63FD1AD3D9" box="[673,718,980,1005]" italics="true" pageId="27" pageNumber="328">al -2</emphasis>
|
||
and
|
||
<emphasis id="B93FEAE91A7DCE43E969FC63FCE5D3D9" box="[772,817,980,1005]" italics="true" pageId="27" pageNumber="328">pl -2</emphasis>
|
||
on tibia I. Form and shape of leg setae similar to those on adult female, except seta
|
||
<emphasis id="B93FEAE91A7DCE43EB8DFC40FE22D425" box="[480,502,1015,1040]" italics="true" pageId="27" pageNumber="328">pl</emphasis>
|
||
on femur, genu, tibia, basitarsus of leg IV strongly developed and recurved/bent. Legs IV directed posteriorly, as in protonymph. Tarsi II–IV with form of setae and acuminate form of apical ventral process as in protonymph.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFBD4FB8FD54D" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFBD4FD9FD448" box="[199,587,1123,1149]" italics="true" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFBD4FE88D449" bold="true" box="[199,348,1123,1148]" italics="true" pageId="27" pageNumber="328">Protonymph.</emphasis>
|
||
Idiosomatic dorsum
|
||
</emphasis>
|
||
. Discoidal to ovoidal form. Dorsal shielding weakly sclerotised, with or without delineation between podonotal and pygidial shields, and with faint lateral, mostly podonotal ornamentation; mesonotal scutellae not evident (
|
||
<figureCitation id="13702A7E1A7DCE43E8D2FB1BFCC3D4F1" box="[703,791,1196,1221]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Figs 77</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7DCE43E948FB1BFC90D4F1" box="[805,836,1196,1220]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="27" pageNumber="328">89</figureCitation>
|
||
); dorsum maximally with 24 to 33 pairs of similarly short setae except more posterior ones smaller,
|
||
<emphasis id="B93FEAE91A7DCE43E8CAFB67FD16D4D2" box="[679,706,1232,1255]" italics="true" pageId="27" pageNumber="328">J5</emphasis>
|
||
minute (protonymphal setae
|
||
<emphasis id="B93FEAE91A7DCE43EE65FB67FBF5D4D2" box="[1032,1057,1232,1255]" italics="true" pageId="27" pageNumber="328">s5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE43FB67FB9CD4DD" box="[1070,1096,1232,1256]" italics="true" pageId="27" pageNumber="328">r2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE38FB67FBBAD4DD" box="[1109,1134,1232,1256]" italics="true" pageId="27" pageNumber="328">r3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE11FB67FB41D4D2" box="[1148,1173,1232,1255]" italics="true" pageId="27" pageNumber="328">r5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EECEFB67FB14D4DD" box="[1187,1216,1232,1256]" italics="true" pageId="27" pageNumber="328">Z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EEA3FB67FB3DD4DD" box="[1230,1257,1232,1256]" italics="true" pageId="27" pageNumber="328">S2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EE9AFB67FAC3D4DD" box="[1271,1303,1232,1256]" italics="true" pageId="27" pageNumber="328">R1</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7DCE43EAFAFB43FF65D539" box="[151,177,1268,1292]" italics="true" pageId="27" pageNumber="328">J1</emphasis>
|
||
added to larval complement,
|
||
<emphasis id="B93FEAE91A7DCE43EB91FB43FDC5D539" box="[508,529,1268,1292]" italics="true" pageId="27" pageNumber="328">j2</emphasis>
|
||
absent); setae
|
||
<emphasis id="B93FEAE91A7DCE43E8DBFB43FD04D539" box="[694,720,1268,1292]" italics="true" pageId="27" pageNumber="328">J3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E8B3FB42FD2CD539" box="[734,760,1269,1292]" italics="true" pageId="27" pageNumber="328">J5</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E96BFB43FCF0D539" box="[774,804,1268,1292]" italics="true" pageId="27" pageNumber="328">Z4</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E95CFB42FC9BD539" box="[817,847,1269,1292]" italics="true" pageId="27" pageNumber="328">Z5</emphasis>
|
||
sometimes on pygidial shield if delineated (
|
||
<figureCitation id="13702A7E1A7DCE43EF51FB43FA5BD539" box="[1340,1423,1268,1293]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="27" pageNumber="328">Fig. 89</figureCitation>
|
||
). Sometimes as many as eight pairs of dorsal deutonymphal setae (
|
||
<emphasis id="B93FEAE91A7DCE43E91AFAAFFC45D505" box="[887,913,1304,1328]" italics="true" pageId="27" pageNumber="328">s1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E9F2FAA0FC6CD505" box="[927,952,1303,1328]" italics="true" pageId="27" pageNumber="328">z6</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E9ABFAAFFC37D505" box="[966,995,1304,1328]" italics="true" pageId="27" pageNumber="328">S1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E99CFAAFFBECD505" box="[1009,1080,1303,1328]" italics="true" pageId="27" pageNumber="328">R2-R6</emphasis>
|
||
) may accelerate their ontogeny to this instar (
|
||
<figureCitation id="13702A7E1A7DCE43EB5CFA8BFE60D561" box="[305,436,1340,1365]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Figs 77, 78</figureCitation>
|
||
). Body dorsum with complement of 11 pairs of discernible pore-like structures, of which six non-secretory (poroids) and five superficially appear to be secretory (gland pores).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAFA33FE2CD71D" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAFA33FE4DD5A9" box="[199,409,1412,1437]" italics="true" pageId="27" pageNumber="328">Idiosomatic venter</emphasis>
|
||
. Sternal region with two or three pairs of weakly sclerotised platelets (without endopodal elements) bearing setae
|
||
<emphasis id="B93FEAE91A7DCE43EBC7FA1FFE1FD5F5" box="[426,459,1448,1472]" italics="true" pageId="27" pageNumber="328">st1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EBB7FA1FFE28D5F5" box="[474,508,1448,1472]" italics="true" pageId="27" pageNumber="328">st2</emphasis>
|
||
and poroids
|
||
<emphasis id="B93FEAE91A7DCE43E8FFFA1FFD60D5F5" box="[658,692,1448,1472]" italics="true" pageId="27" pageNumber="328">
|
||
<date id="FFF5103B1A7DCE43E8FFFA1FFD60D5F5" box="[658,692,1448,1472]" pageId="27" pageNumber="328">iv1</date>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E8A9FA1FFD32D5F5" box="[708,742,1448,1472]" italics="true" pageId="27" pageNumber="328">
|
||
<date id="FFF5103B1A7DCE43E8A9FA1FFD32D5F5" box="[708,742,1448,1472]" pageId="27" pageNumber="328">iv2</date>
|
||
</emphasis>
|
||
(
|
||
<date id="FFF5103B1A7DCE43E895FA1FFCCFD5F5" box="[760,795,1448,1472]" pageId="27" pageNumber="328">
|
||
<emphasis id="B93FEAE91A7DCE43E895FA1FFCCFD5F5" box="[760,795,1448,1472]" italics="true" pageId="27" pageNumber="328">iv2</emphasis>
|
||
</date>
|
||
added to larval complement); setae
|
||
<emphasis id="B93FEAE91A7DCE43EED6FA1FFB08D5F5" box="[1211,1244,1448,1472]" italics="true" pageId="27" pageNumber="328">st3</emphasis>
|
||
on either a third pair of weak platelets or on soft cuticle; setae
|
||
<emphasis id="B93FEAE91A7DCE43E8DCFA7AFD06D5D1" box="[689,722,1485,1508]" italics="true" pageId="27" pageNumber="328">st5</emphasis>
|
||
rudimentary, paragenital poroids indiscernible. Opisthogastric region reduced to small area behind coxae IV. Anal shield faintly sclerotised, unornamented, yet prominent, about as wide as long, with postanal seta longer than para-anal setae; para-anal setae inserted at mid-level of anal opening, roughly in transverse line with adanal gland pores; cribrum similar in development as in deutonymph. Opisthogaster with four or six pairs of ventral setae;
|
||
<emphasis id="B93FEAE91A7DCE43E963F9EBFCEDD641" box="[782,825,1628,1652]" italics="true" pageId="27" pageNumber="328">JV1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E921F9EBFCACD641" box="[844,888,1628,1652]" italics="true" pageId="27" pageNumber="328">JV2</emphasis>
|
||
, sometimes
|
||
<emphasis id="B93FEAE91A7DCE43EE60F9EBFBE9D641" box="[1037,1085,1628,1652]" italics="true" pageId="27" pageNumber="328">ZV2</emphasis>
|
||
anterior to anal shield; larval complement sometimes augmented by normally deutonymphal setae
|
||
<emphasis id="B93FEAE91A7DCE43E9D5F937FC37D6AD" box="[952,995,1664,1688]" italics="true" pageId="27" pageNumber="328">JV3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E998F937FBF5D6AD" box="[1013,1057,1664,1688]" italics="true" pageId="27" pageNumber="328">JV4</emphasis>
|
||
on posterior margin, along with larval
|
||
<emphasis id="B93FEAE91A7DCE43EAB2F912FEDFD689" box="[223,267,1701,1724]" italics="true" pageId="27" pageNumber="328">JV5</emphasis>
|
||
(
|
||
<figureCitation id="13702A7E1A7DCE43EB77F913FEA6D689" box="[282,370,1700,1725]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="27" pageNumber="328">Figs 78</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7DCE43EBEDF913FE74D689" box="[384,416,1700,1724]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="27" pageNumber="328">90</figureCitation>
|
||
). Peritrematal region on each side with poroid and gland pore (
|
||
<emphasis id="B93FEAE91A7DCE43EE1DF913FB47D689" box="[1136,1171,1700,1724]" italics="true" pageId="27" pageNumber="328">ip2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43EECEF913FB18D689" box="[1187,1228,1700,1724]" italics="true" pageId="27" pageNumber="328">gp2</emphasis>
|
||
) on soft cuticle at mid-level of coxae III, anterior to apex of weakly formed peritreme between coxae III and IV, and with poststigmatic poroid and gland pore (
|
||
<emphasis id="B93FEAE91A7DCE43E853F95BFDB5D731" box="[574,609,1772,1796]" italics="true" pageId="27" pageNumber="328">ip3</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7DCE43E81CF95BFD4FD731" box="[625,667,1772,1796]" italics="true" pageId="27" pageNumber="328">gp3</emphasis>
|
||
) on soft cuticle behind peritreme. Exopodal rim and gland pore
|
||
<emphasis id="B93FEAE91A7DCE43EF19F95BFA48D731" box="[1396,1436,1772,1796]" italics="true" pageId="27" pageNumber="328">gv2</emphasis>
|
||
behind coxa IV not discernible.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7DCE43EAAAF883FBC9D7E9" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="328">
|
||
<taxonomicName id="4C4B4D781A7DCE43EAAAF883FE82D778" box="[199,342,1844,1869]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="27" pageNumber="328" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7DCE43EAAAF883FE82D778" box="[199,342,1844,1869]" italics="true" pageId="27" pageNumber="328">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Form of gnathotectum, corniculus and adjacent paraxial process, elongated internal malae (
|
||
<figureCitation id="13702A7E1A7DCE43EF02F883FF62D745" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="27" pageNumber="328">Fig. 85</figureCitation>
|
||
), and other gnathosomatic structures similar to those in deutonymph, except palpi with normal protonymphal complement of setae (see
|
||
<bibRefCitation id="EFDA4B0A1A7DCE43EBA5F8CAFD87D7A0" author="Evans" box="[456,595,1916,1941]" pageId="27" pageNumber="328" refString="Evans, G. O. (1964) Some observations on the chaetotaxy of the pedipalps in the Mesostigmata (Acari). Annals and Magazine of Natural History, Series 13, 6, 513 - 527." type="journal article" year="1964">Evans 1964</bibRefCitation>
|
||
), including only one trochanter seta. Cheliceral movable digit bi- or tridentate, with mid-ventral projection rudimentary or indiscernible; fixed digit weakly multidenticulate, with offset subapical tooth and sparse (five or six) teeth along masticatory surface (
|
||
<figureCitation id="13702A7E1A7DCE43E9D1F873FBDBD7E9" box="[956,1039,1988,2013]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="27" pageNumber="328">Fig. 87</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7ACE44EAAAFF2EFEB6D22D" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFF2EFF28D085" box="[199,252,153,176]" italics="true" pageId="28" pageNumber="329">Legs</emphasis>
|
||
. Legs I to IV with pretarsi, well-developed claws, and normal protonymphal complement of setae as described for
|
||
<taxonomicName id="4C4B4D781A7ACE44EB42FF0BFE5DD0E0" box="[303,393,188,213]" class="Arachnida" family="Ascidae" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="329" phylum="Arthropoda" rank="family">Ascidae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EBDEFF0BFD02D0E0" author="Lindquist" box="[435,726,188,213]" pageId="28" pageNumber="329" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans (1965)</bibRefCitation>
|
||
except for absence of second ventral seta on femur II: coxae, 2- 2-2-1; trochanters 4-4-4-4 (1 0/2 1 on I, II, 1 1/2 0 on III, IV); femora 10 (2 2/1 2/1 2) – 7 (1 2/1 2/0 1) – 5 (1 2/1 1/ 0 0) – 4 or 5 (1 2/0 1/0 0 or 1); genua 8 or 9 (1 2 or 3/1 2/1 1) – 6 or 7 (1 2 or 3/0 2/0 1) – 6(1 2/0 2/0 1) – 5 or 6 (1 2/0 2/0 0 or 1); tibiae 8 (1 2/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1); normally deutonymphal seta
|
||
<emphasis id="B93FEAE91A7ACE44EAA7FEFBFF23D151" box="[202,247,332,357]" italics="true" pageId="28" pageNumber="329">pl -1</emphasis>
|
||
sometimes accelerated in appearance on femur and genu IV, and
|
||
<emphasis id="B93FEAE91A7ACE44E9B0FEFBFBC4D151" box="[989,1040,332,357]" italics="true" pageId="28" pageNumber="329">ad -3</emphasis>
|
||
similarly on genua I and II. Coxae I–II distal rims with serrated posteroventral strips. Leg setae generally simple. Legs IV directed posteriorly, with seta
|
||
<emphasis id="B93FEAE91A7ACE44EAA4FE23FF0BD198" box="[201,223,404,429]" italics="true" pageId="28" pageNumber="329">pl</emphasis>
|
||
on femur, genu, tibia, and to less degree on tarsus IV slightly to clearly thicker and longer than other setae of segments (
|
||
<figureCitation id="13702A7E1A7ACE44EB62FE00FEB0D1E5" box="[271,356,439,464]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Fig. 79</figureCitation>
|
||
). Tarsi II–IV with setae
|
||
<emphasis id="B93FEAE91A7ACE44E81AFE00FD7ED1E5" box="[631,682,439,464]" italics="true" pageId="28" pageNumber="329">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44E8D4FE00FD39D1E5" box="[697,749,439,464]" italics="true" pageId="28" pageNumber="329">pd -1</emphasis>
|
||
small but as long as short pretarsus;
|
||
<emphasis id="B93FEAE91A7ACE44EEFCFE00FB67D1E5" box="[1169,1203,439,464]" italics="true" pageId="28" pageNumber="329">md</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EEAFFE00FB3BD1E5" box="[1218,1263,439,464]" italics="true" pageId="28" pageNumber="329">al -2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EE93FE0EFAE4D1E5" box="[1278,1328,440,464]" italics="true" pageId="28" pageNumber="329">av -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EF53FE00FABFD1E5" box="[1342,1387,439,464]" italics="true" pageId="28" pageNumber="329">pl -1</emphasis>
|
||
and
|
||
<emphasis id="B93FEAE91A7ACE44EAFAFE6BFF10D1C1" box="[151,196,476,501]" italics="true" pageId="28" pageNumber="329">pl -2</emphasis>
|
||
longer, thinner than other tarsal setae, except
|
||
<emphasis id="B93FEAE91A7ACE44E8A9FE6AFD22D1C1" box="[708,758,476,500]" italics="true" pageId="28" pageNumber="329">av -2</emphasis>
|
||
similarly attenuate on tarsus IV; apical ventral tarsal process acutely triangular.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7ACE44EAAAFD93FE1FD320" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFD93FDDDD208" box="[199,521,548,573]" italics="true" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFD93FEC6D20E" bold="true" box="[199,274,548,571]" italics="true" pageId="28" pageNumber="329">Larva.</emphasis>
|
||
Idiosomatic dorsum
|
||
</emphasis>
|
||
. Body with dorsal shielding weakly sclerotised, not clearly delimited, less discernibly ornamented than on nymphal instars; mesonotal scutellae indiscernible (
|
||
<figureCitation id="13702A7E1A7ACE44EE0CFDF0FB68D255" box="[1121,1212,583,608]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Figs 80</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7ACE44EEA7FDFFFB3DD255" box="[1226,1257,584,608]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="28" pageNumber="329">91</figureCitation>
|
||
). Body dorsum with 17 pairs of setae: nine podonotal pairs, and eight opisthonotal pairs (
|
||
<emphasis id="B93FEAE91A7ACE44EE6FFDDBFBC8D2B1" box="[1026,1052,620,644]" italics="true" pageId="28" pageNumber="329">J2</emphasis>
|
||
); discernible pore-like structures include eight non-secretory poroids (three podonotal, five opisthonotal) and three apparently secretory gland pores (two podonotal, one opisthonotal). Peritrematal region on each side with gland pore
|
||
<emphasis id="B93FEAE91A7ACE44EE22FD03FBADD2F9" box="[1103,1145,692,716]" italics="true" pageId="28" pageNumber="329">gp1</emphasis>
|
||
at level of setae
|
||
<emphasis id="B93FEAE91A7ACE44EF54FD03FA84D2F9" box="[1337,1360,692,716]" italics="true" pageId="28" pageNumber="329">j3</emphasis>
|
||
, gland pore
|
||
<emphasis id="B93FEAE91A7ACE44EABCFD6FFF2FD2C5" box="[209,251,728,752]" italics="true" pageId="28" pageNumber="329">gp2</emphasis>
|
||
and poroid
|
||
<emphasis id="B93FEAE91A7ACE44EBE5FD6FFE78D2C5" box="[392,428,728,752]" italics="true" pageId="28" pageNumber="329">ip2</emphasis>
|
||
at level above coxae II–III, and poststigmatic gland pore
|
||
<emphasis id="B93FEAE91A7ACE44EE28FD6FFBBBD2C5" box="[1093,1135,728,752]" italics="true" pageId="28" pageNumber="329">gp3</emphasis>
|
||
and poroid
|
||
<emphasis id="B93FEAE91A7ACE44EE96FD6FFACBD2C5" box="[1275,1311,728,752]" italics="true" pageId="28" pageNumber="329">ip3</emphasis>
|
||
at level of anal shield anterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7ACE44EAAAFCA8FD27D46D" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFCA8FE43D30D" box="[199,407,799,824]" italics="true" pageId="28" pageNumber="329">Idiosomatic venter</emphasis>
|
||
. Tritosternum normally developed, similar in form as in subsequent instars. Sternal region with two or three pairs of weakly (hardly discernible) sclerotised platelets bearing setae
|
||
<emphasis id="B93FEAE91A7ACE44EEE7FCF3FB7ED369" box="[1162,1194,836,860]" italics="true" pageId="28" pageNumber="329">st1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EED7FCF3FB0ED369" box="[1210,1242,836,860]" italics="true" pageId="28" pageNumber="329">st2</emphasis>
|
||
and poroids
|
||
<emphasis id="B93FEAE91A7ACE44EF1FFCF3FA40D369" box="[1394,1428,836,860]" italics="true" pageId="28" pageNumber="329">
|
||
<date id="FFF5103B1A7ACE44EF1FFCF3FA40D369" box="[1394,1428,836,860]" pageId="28" pageNumber="329">iv1</date>
|
||
</emphasis>
|
||
; setae
|
||
<emphasis id="B93FEAE91A7ACE44EAB7FCDFFF2FD3B5" box="[218,251,872,896]" italics="true" pageId="28" pageNumber="329">st3</emphasis>
|
||
on soft cuticle or on a third pair of platelets; sometimes protonymphal setae
|
||
<emphasis id="B93FEAE91A7ACE44EEE2FCDFFB7BD34A" box="[1167,1199,872,895]" italics="true" pageId="28" pageNumber="329">st5</emphasis>
|
||
accelerately present between coxae III (
|
||
<figureCitation id="13702A7E1A7ACE44EB00FC3BFE17D391" box="[365,451,908,933]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Figs 81</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7ACE44EBA3FC3BFE3AD391" box="[462,494,908,932]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="28" pageNumber="329">92</figureCitation>
|
||
); pair of subcutaneous structures sometimes discernible at level of posterior margins of coxae III. Anal shield faintly sclerotised, elongate or as wide as long; para-anal setae inserted at level slightly posterior to anal opening, and slightly longer than postanal seta; adanal gland pores (
|
||
<emphasis id="B93FEAE91A7ACE44EE3FFC63FBAED3D9" box="[1106,1146,980,1004]" italics="true" pageId="28" pageNumber="329">gv3</emphasis>
|
||
) at level of postanal seta; cribrum rudimentary or absent. Venter with six pairs of opisthogastric setae on soft cuticle around anal shield, including
|
||
<emphasis id="B93FEAE91A7ACE44EB7BFBABFE95D401" box="[278,321,1052,1076]" italics="true" pageId="28" pageNumber="329">JV1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EB30FBABFE5DD401" box="[349,393,1052,1076]" italics="true" pageId="28" pageNumber="329">JV2</emphasis>
|
||
, sometimes rudimentary
|
||
<emphasis id="B93FEAE91A7ACE44E8BDFBABFD2BD401" box="[720,767,1052,1076]" italics="true" pageId="28" pageNumber="329">ZV2</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44E976FBAAFC93D401" box="[795,839,1053,1076]" italics="true" pageId="28" pageNumber="329">JV5</emphasis>
|
||
anterolaterally and ventrally displaced
|
||
<emphasis id="B93FEAE91A7ACE44EF25FBABFAB0D401" box="[1352,1380,1052,1076]" italics="true" pageId="28" pageNumber="329">S4</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EFEDFBAAFA48D401" box="[1408,1436,1053,1076]" italics="true" pageId="28" pageNumber="329">S5</emphasis>
|
||
posterolaterally (sometimes
|
||
<emphasis id="B93FEAE91A7ACE44EBB9FBF7FDD4D462" box="[468,512,1088,1111]" italics="true" pageId="28" pageNumber="329">JV5</emphasis>
|
||
and
|
||
<emphasis id="B93FEAE91A7ACE44E85BFBF7FD86D46D" box="[566,594,1088,1112]" italics="true" pageId="28" pageNumber="329">S4</emphasis>
|
||
or
|
||
<emphasis id="B93FEAE91A7ACE44E81AFBF7FD40D462" box="[631,660,1088,1111]" italics="true" pageId="28" pageNumber="329">S5</emphasis>
|
||
absent).
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7ACE44EAAAFBD3FD80D505" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<taxonomicName id="4C4B4D781A7ACE44EAAAFBD3FE82D448" box="[199,342,1124,1149]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="28" pageNumber="329" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFBD3FE82D448" box="[199,342,1124,1149]" italics="true" pageId="28" pageNumber="329">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Gnathotectum with anterior margin less denticulate than in subsequent instars (
|
||
<figureCitation id="13702A7E1A7ACE44EF5EFBD3FA5BD449" box="[1331,1423,1124,1149]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Fig. 83</figureCitation>
|
||
). Cheliceral movable digit with two teeth, and a rudimentary mid-ventral projection (
|
||
<figureCitation id="13702A7E1A7ACE44EE54FB30FB59D495" box="[1081,1165,1159,1184]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Fig. 84</figureCitation>
|
||
); dentition of fixed digit weakly developed, with offset subapical tooth and masticatory ridge with
|
||
<emphasis id="B93FEAE91A7ACE44E99BFB1AFBC4D4F1" box="[1014,1040,1197,1220]" italics="true" pageId="28" pageNumber="329">ca</emphasis>
|
||
. two to four weak teeth. Form of elongated internal malae, corniculus and adjacent acuminate process much as in subsequent instars (
|
||
<figureCitation id="13702A7E1A7ACE44EF5BFB78FA5BD4DD" box="[1334,1423,1231,1256]" captionStart="FIGURES 79 – 87" captionStartId="38.[151,264,1700,1723]" captionTargetBox="[151,1436,197,1664]" captionTargetId="figure@38.[151,1436,193,1679]" captionTargetPageId="38" captionText="FIGURES 79 – 87. Hispiniphis parva n. sp.: 79. Protonymph, leg IV, right leg, dorsal view (dorsal setae in solid black color); 80 - 84, 86. Larva: 80. Idiosoma, dorsal aspect; 81. Idiosoma, ventral aspect; 82. Detail of subcapitulum apex (distal margin of deutosternum not shown); 83. Gnathotectum; 84. Chelicera, lateral view; 86. Tarsus I, apical region with ventro-apical process; 85, 87. Protonymph: 85. Detail of subcapitulum apex; 87. Chelicera, lateral view." httpUri="https://zenodo.org/record/244206/files/figure.png" pageId="28" pageNumber="329">Fig. 82</figureCitation>
|
||
); deutosternum with rows of denticles similar to those in nymphs. Palpus with normal larval complement of setae (see
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EAA6FAAFFE8DD505" author="Evans" box="[203,345,1303,1328]" pageId="28" pageNumber="329" refString="Evans, G. O. (1964) Some observations on the chaetotaxy of the pedipalps in the Mesostigmata (Acari). Annals and Magazine of Natural History, Series 13, 6, 513 - 527." type="journal article" year="1964">Evans, 1964</bibRefCitation>
|
||
); palptrochanter nude.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7ACE44EAAAFA8AFC08D63D" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAFA8AFF28D561" box="[199,252,1341,1364]" italics="true" pageId="28" pageNumber="329">Legs</emphasis>
|
||
. Legs I to III with pretarsi, well-developed claws, and with normal larval complement of setae as described for
|
||
<taxonomicName id="4C4B4D781A7ACE44EB5EFAE8FE59D54D" box="[307,397,1375,1400]" class="Arachnida" family="Ascidae" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="329" phylum="Arthropoda" rank="family">Ascidae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EBD7FAE8FD36D54D" author="Lindquist" box="[442,738,1375,1400]" pageId="28" pageNumber="329" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans (1965)</bibRefCitation>
|
||
except for retardation of appearance of seta
|
||
<emphasis id="B93FEAE91A7ACE44EEB3FAE8FAC6D54D" box="[1246,1298,1375,1400]" italics="true" pageId="28" pageNumber="329">ad -2</emphasis>
|
||
on femur II until protonymph: coxae, 2-2-2; trochanters 4-4-4; femora 10 (2 2/1 2/1 2) – 7 (1 2/1 2/0 1) – 5 (1 2/1 1/0 0); genua 8 (1 2/1 2/1 1) – 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1); tibiae 8 (1 2/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1). Coxae I- III distal rims smooth. Leg setae generally simple, not markedly differentiated. Tarsi II and III with setae
|
||
<emphasis id="B93FEAE91A7ACE44EF58FA7BFABCD5D1" box="[1333,1384,1484,1509]" italics="true" pageId="28" pageNumber="329">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B93FEAE91A7ACE44EF1BFA7BFF71D63D" italics="true" pageId="28" pageNumber="329">pd - 1</emphasis>
|
||
as long as short pretarsus; apical ventral tarsal process acutely triangular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7ACE44EAAAF9A3FCC2D640" pageId="28" pageNumber="329" type="etymology">
|
||
<paragraph id="8BF436FB1A7ACE44EAAAF9A3FCC2D640" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAF9A3FE9AD618" bold="true" box="[199,334,1556,1581]" pageId="28" pageNumber="329">Etymology.</emphasis>
|
||
The name of the genus, feminine in gender, is a Latinised combination of the term
|
||
<emphasis id="B93FEAE91A7ACE44EF55F9A3FA41D618" box="[1336,1429,1556,1581]" italics="true" pageId="28" pageNumber="329">hispidus</emphasis>
|
||
, meaning bristly, upon which the family-group name of the beetles is based, and
|
||
<emphasis id="B93FEAE91A7ACE44EE2CF980FBA3D665" box="[1089,1143,1591,1616]" italics="true" pageId="28" pageNumber="329">iphis</emphasis>
|
||
, meaning stalwart, and a suffix commonly used among names for gamasine mites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7ACE44EAAAF9C8FD55D745" pageId="28" pageNumber="329" type="distribution">
|
||
<paragraph id="8BF436FB1A7ACE44EAAAF9C8FD55D745" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAF9C8FDD5D6AD" bold="true" box="[199,513,1663,1688]" pageId="28" pageNumber="329">Distribution and habitats.</emphasis>
|
||
The genus is currently based on one newly described species and on additional material representing five undescribed species. All instars of mites of the genus
|
||
<taxonomicName id="4C4B4D781A7ACE44EE56F913FB68D688" box="[1083,1212,1700,1725]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7ACE44EE56F913FB68D688" box="[1083,1212,1700,1725]" italics="true" pageId="28" pageNumber="329">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
are known only as subelytral symbionts of adult hispine beetles of the genus
|
||
<taxonomicName id="4C4B4D781A7ACE44E946F970FC60D6D5" box="[811,948,1735,1760]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7ACE44E946F970FC60D6D5" box="[811,948,1735,1760]" italics="true" pageId="28" pageNumber="329">Cephaloleia</emphasis>
|
||
</taxonomicName>
|
||
, tribe Immatidiini (or
|
||
<taxonomicName id="4C4B4D781A7ACE44EED8F970FA85D6D5" box="[1205,1361,1735,1760]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="tribe" tribe="Cephaloleiini">Cephaloleiini</taxonomicName>
|
||
). Over 200 species of the neotropical genus
|
||
<taxonomicName id="4C4B4D781A7ACE44E821F95BFD01D730" box="[588,725,1772,1797]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEAE91A7ACE44E821F95BFD01D730" box="[588,725,1772,1797]" italics="true" pageId="28" pageNumber="329">Cephaloleia</emphasis>
|
||
</taxonomicName>
|
||
are recognised, including at least 40 sympatric species from lowland tropical rain forest in the La Selva Biological Station,
|
||
<collectingCountry id="F35C766B1A7ACE44E931F8A7FC0DD71D" box="[860,985,1807,1832]" name="Costa Rica" pageId="28" pageNumber="329">Costa Rica</collectingCountry>
|
||
, alone; larvae of the majority of species feed on unfurled leaves of various families of the Order
|
||
<taxonomicName id="4C4B4D781A7ACE44E967F883FC42D778" box="[778,918,1844,1869]" class="Liliopsida" kingdom="Animalia" order="Zingiberales" pageId="28" pageNumber="329" phylum="Tracheophyta" rank="order">Zingiberales</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44E9C9F883FBECD778" author="Staines" box="[932,1080,1844,1869]" pageId="28" pageNumber="329" refString="Staines, C. L. (1996) The genus Cephaloleia (Coleotera: Chrysomelidae) in Central America and the West Indies. Revista Biologia Tropical, Special Number, 3, 3 - 87." type="journal article" year="1996">Staines 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EE2EF883FBAAD779" author="Staines" box="[1091,1150,1844,1868]" pageId="28" pageNumber="329" refString="Garcia-Robledo, C. & Staines, C. L. (2008) Herbivory in gingers from latest Cretaceous to present: is the ichnogenus Cephaloleichnites (Hispinae, Coleoptera) a rolled-leaf beetle? Journal of Paleontology, 82, 1035 - 1037. http: // dx. doi. org / 10.1666 / 07 - 089.1" type="journal article" year="2008">2008</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EEE4F883FB10D779" author="Staines" box="[1161,1220,1844,1868]" pageId="28" pageNumber="329" refString="Staines, C. L. (2009) A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa, 2033, 58 - 68." type="journal article" year="2009">2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EEA3F883FF23D745" author="Garcia-Robledo" pageId="28" pageNumber="329" refString="Garcia-Robledo, C., Horvitz, C. C. & Staines, C. L. (2010) Larval morphology and development, host plants, adult longevity, sexual dimorphism and notes on natural history in Cephaloleia rolled-leaf beetles (Coleoptera: Chrysomelidae: Hispinae). Zootaxa, 2610, 50 - 68." type="journal article" year="2010">
|
||
Garcia-Robledo
|
||
<emphasis id="B93FEAE91A7ACE44EFE5F882FF79D745" italics="true" pageId="28" pageNumber="329">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="EFDA4B0A1A7ACE44EB67F8E0FDA1D745" author="Staines" box="[266,629,1879,1904]" pageId="28" pageNumber="329" refString="Staines, C. L. & Garcia-Robledo, C. (2014) The genus Cephaloleia Chevrolat, 1836 (Coleoptera: Chrysomelidae: Cassidinae). Zookeys, 436, 1 - 355. http: // dx. doi. org / 10.3897 / zookeys. 436.5766" type="journal article" year="2014">Staines & Garcia-Robledo 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165701A7ACE45EAAAF8CBFDA4D320" lastPageId="29" lastPageNumber="330" pageId="28" pageNumber="329" type="discussion">
|
||
<paragraph id="8BF436FB1A7ACE45EAAAF8CBFD7FD0C2" blockId="28.[151,1436,151,2013]" lastBlockId="29.[151,1437,151,789]" lastPageId="29" lastPageNumber="330" pageId="28" pageNumber="329">
|
||
<emphasis id="B93FEAE91A7ACE44EAAAF8CBFEEFD7A0" bold="true" box="[199,315,1916,1941]" pageId="28" pageNumber="329">Remarks.</emphasis>
|
||
<emphasis id="B93FEAE91A7ACE44EB2AF8CBFD04D7A0" box="[327,720,1916,1941]" italics="true" pageId="28" pageNumber="329">Reduction in size of opisthosoma</emphasis>
|
||
. Among species of
|
||
<taxonomicName id="4C4B4D781A7ACE44E9AAF8CBFB9CD7A0" box="[967,1096,1916,1941]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7ACE44E9AAF8CBFB9CD7A0" box="[967,1096,1916,1941]" italics="true" pageId="28" pageNumber="329">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
for which specimens of the nymphs are available, a smallness of the opisthosomatic region is notable (
|
||
<figureCitation id="13702A7E1A7ACE44E98DF817FBE3D78D" box="[992,1079,1952,1977]" captionStart="FIGURES 75 – 78" captionStartId="37.[151,264,1869,1892]" captionTargetBox="[151,1436,375,1834]" captionTargetId="figure@37.[151,1436,373,1849]" captionTargetPageId="37" captionText="FIGURES 75 – 78. Hispiniphis parva n. sp.: 75. Deutonymph, idiosoma, dorsal aspect; 76. Deutonymph, idiosoma, ventral aspect; 77. Protonymph, idiosoma, dorsal aspect; 78. Protonymph, idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244205/files/figure.png" pageId="28" pageNumber="329">Figs 78</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7ACE44EE29F817FBB0D78D" box="[1092,1124,1952,1976]" captionStart="FIGURES 89 – 92" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[152,1436,301,1850]" captionTargetId="figure@40.[151,1436,301,1864]" captionTargetPageId="40" captionText="FIGURES 89 – 92. Hispiniphis sp. 3: Protonymph: 89. Idiosoma, dorsal aspect; 90. Idiosoma, ventral aspect. Larva: 91. Idiosoma, dorsal aspect; 92. Idiosoma, ventral aspect." httpUri="https://zenodo.org/record/244208/files/figure.png" pageId="28" pageNumber="329">90</figureCitation>
|
||
). There is a compactness of body behind the legs IV in both nymphal instars, leaving the anal shield and adjacent structures squeezed into a relatively small space. Perhaps this may be correlated with evolution towards a rapid life cycle, and lessened need for enlargement of the body region not involved in mobility. This reduction is not evident in the nymphs of the one known species of the sister-genus,
|
||
<taxonomicName id="4C4B4D781A7BCE45E871FF68FD70D0CD" box="[540,676,223,248]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="311" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7BCE45E871FF68FD70D0CD" box="[540,676,223,248]" italics="true" pageId="29" pageNumber="330">Makarovaia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7BCE45EAAAFEB3FE1BD1E5" blockId="29.[151,1437,151,789]" pageId="29" pageNumber="330">
|
||
<emphasis id="B93FEAE91A7BCE45EAAAFEB3FCF1D128" box="[199,805,260,285]" italics="true" pageId="29" pageNumber="330">Interspecific variation in adult dorsal shield attributes</emphasis>
|
||
. The shape and ornamentation of the adult dorsal shield, as well as the complement and size of its setae, vary markedly among the five species of
|
||
<taxonomicName id="4C4B4D781A7BCE45EE12FE90FAD5D175" box="[1151,1281,295,320]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7BCE45EE12FE90FAD5D175" box="[1151,1281,295,320]" italics="true" pageId="29" pageNumber="330">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
for which we have material at hand (
|
||
<figureCitation id="13702A7E1A7BCE45EBF1FEFBFE25D151" box="[412,497,332,357]" captionStart="FIGURE 88" captionStartId="39.[151,250,1491,1513]" captionTargetBox="[156,1429,204,1457]" captionTargetId="figure@39.[151,1436,193,1469]" captionTargetPageId="39" captionText="FIGURE 88. Hispiniphis spp., chaetotaxy of dorsal shield, female: 88 a. Hispiniphis parva; 88 b. Hispiniphis sp. 4; 88 c. Hispiniphis sp. 2; 88 d. Hispiniphis sp. 5; 88 e. Hispiniphis sp. 3." httpUri="https://zenodo.org/record/244207/files/figure.png" pageId="29" pageNumber="330">Fig. 88</figureCitation>
|
||
). In general, the description of the genus accounts for these differences, until such time as other of these species are described subsequently. Such readily discernible morphological differences may be quite useful in determining whatever level of host specificity there may be between species of this genus of mites and their beetle hosts.
|
||
</paragraph>
|
||
<paragraph id="8BF436FB1A7BCE45EAAAFE6AFDA4D320" blockId="29.[151,1437,151,789]" pageId="29" pageNumber="330">
|
||
<emphasis id="B93FEAE91A7BCE45EAAAFE6AFD61D1C0" box="[199,693,476,501]" italics="true" pageId="29" pageNumber="330">Symmetry of dorsal idiosomatic hypertrichy</emphasis>
|
||
. Adults (to some extent preceded by nymphs) of some species of
|
||
<taxonomicName id="4C4B4D781A7BCE45EAFAFE48FECCD22D" box="[151,280,511,536]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7BCE45EAFAFE48FECCD22D" box="[151,280,511,536]" italics="true" pageId="29" pageNumber="330">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
display a pattern of hypertrichy that appears to be a duplicative neotrichy derived secondarily within the genus. This is indicated to a slight extent by outline of setal duplicates in
|
||
<figureCitation id="13702A7E1A7BCE45EE7FFD93FBBED209" box="[1042,1130,548,573]" captionStart="FIGURES 41 – 44" captionStartId="31.[151,264,1905,1928]" captionTargetBox="[162,1426,449,1859]" captionTargetId="figure@31.[151,1436,445,1885]" captionTargetPageId="31" captionText="FIGURES 41 – 44. Hispiniphis parva n. sp., adult female: 41. Detail of separate views of dorsal setae denoted; 42. Idiosoma, dorsal aspect; 43. Detail of peritrematal shield, peritreme, and pore – like structures; 44. Chelicera, paraxial lateral view and detail of dorsal cheliceral seta and dentition." httpUri="https://zenodo.org/record/244198/files/figure.png" pageId="29" pageNumber="330">Figs 42</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7BCE45EE1AFD93FB42D208" box="[1143,1174,548,573]" captionStart="FIGURES 45 – 49" captionStartId="32.[151,264,1577,1600]" captionTargetBox="[163,1431,199,1538]" captionTargetId="figure@32.[151,1436,193,1556]" captionTargetPageId="32" captionText="FIGURES 45 – 49. Hispiniphis parva n. sp., adult female: 45. Idiosoma, ventral aspect; 46. Detail of separate view of ventral setae denoted; 47. Tritosternum; 48. Epigynal shield; 49. Anal shield." httpUri="https://zenodo.org/record/244199/files/figure.png" pageId="29" pageNumber="330">45</figureCitation>
|
||
,
|
||
<figureCitation id="13702A7E1A7BCE45EECFFD93FB6AD208" box="[1186,1214,548,573]" captionStart="FIGURES 56 – 59" captionStartId="34.[151,264,1557,1580]" captionTargetBox="[151,1424,400,1529]" captionTargetId="figure@34.[151,1436,391,1544]" captionTargetPageId="34" captionText="FIGURES 56 – 59. Hispiniphis parva n. sp., adult male: 56. Idiosoma, dorsal aspect; 57. Gnathotectum; 58. Chelicera, lateral view; 59. Detail of movable cheliceral digit and spermatodactyl." httpUri="https://zenodo.org/record/244202/files/figure.png" pageId="29" pageNumber="330">56</figureCitation>
|
||
of
|
||
<taxonomicName id="4C4B4D781A7BCE45EE8AFD92FA99D209" box="[1255,1357,549,572]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="parva">
|
||
<emphasis id="B93FEAE91A7BCE45EE8AFD92FB2FD209" box="[1255,1275,549,572]" italics="true" pageId="29" pageNumber="330">H</emphasis>
|
||
.
|
||
<emphasis id="B93FEAE91A7BCE45EF61FD92FA99D209" box="[1292,1357,549,572]" italics="true" pageId="29" pageNumber="330">parva</emphasis>
|
||
</taxonomicName>
|
||
, but is much more evident among adults of a couple of other species, to be described subsequently. This form of neotrichy falls within the conceptual framework of cosmiotrichy, as distinguished by Grandjean (1965), in which the setae occupying a neotrichous territory are arranged in recogniseable order, usually in linear files, such that they are idiotrichous, may be assigned sigla, indicating homologies. Rarely noted as such among gamsine mites (but see
|
||
<bibRefCitation id="EFDA4B0A1A7BCE45EAFAFD60FE18D2C5" author="Lindquist" box="[151,460,727,752]" pageId="29" pageNumber="330" refString="Lindquist, E. E. & Moraza, M. L. (1999) Observations on homologies of idiosomal setae in Zerconidae (Acari: Mesostigmata), with modified notation for some posterior body setae. Acarologia, 39, 203 - 226." type="journal article" year="1999">Lindquist & Moraza 1999</bibRefCitation>
|
||
), this phenomenon will be discussed further in a subsequent paper, in which other species of
|
||
<taxonomicName id="4C4B4D781A7BCE45EB60FD4BFE5AD320" box="[269,398,764,789]" class="Arachnida" family="Ascidae" genus="Hispiniphis" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93FEAE91A7BCE45EB60FD4BFE5AD320" box="[269,398,764,789]" italics="true" pageId="29" pageNumber="330">Hispiniphis</emphasis>
|
||
</taxonomicName>
|
||
are to be described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |