159 lines
19 KiB
XML
159 lines
19 KiB
XML
<document id="8EDE7BF98BBEBD0C80A03F8C95CB3266" ID-DOI="10.5281/zenodo.6120089" ID-ISSN="0035-418X" ID-Zenodo-Dep="6120089" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645039399653" checkinUser="carolina" docAuthor="Makranczy, György" docDate="2014" docId="03E287D4FFA5FF8D24A4597BFBDCFC6B" docLanguage="en" docName="RevSuisseZool.121.3.319-347.pdf.imf" docOrigin="Revue suisse de Zoologie 121 (3)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Thinobius paraminor Makranczy & H- 2014, sp. nov." docType="treatment" docUuid="DF7BBF6A-A8AC-449D-88F9-3780BBECC3E2" docUuidSource="ZooBank" docVersion="4" lastPageNumber="332" masterDocId="FFDBFFACFFAEFF8024335818FFFBFFA0" masterDocTitle="The Far Eastern species of Thinobius Kiesenwetter, 1844 (Coleoptera: Staphylinidae, Oxytelinae) lacking female modified genital appendage" masterLastPageNumber="347" masterPageNumber="319" pageNumber="330" updateTime="1699297514242" updateUser="plazi">
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<mods:title id="C1E128E6774305C161D76A8BBF62EA22">The Far Eastern species of Thinobius Kiesenwetter, 1844 (Coleoptera: Staphylinidae, Oxytelinae) lacking female modified genital appendage</mods:title>
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<mods:name id="7F51F434B2CEC380F3D7CE5F3591291C" type="personal">
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<mods:roleTerm id="5FD905FC14B853603A8AB480E86C6C3F">Author</mods:roleTerm>
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<mods:namePart id="8E1EE1FBF5D6E3283D9C988E40CD2FC3">Makranczy, György</mods:namePart>
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<mods:title id="EE74D2BD082D69287FC586914586B8E9">Revue suisse de Zoologie</mods:title>
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<mods:part id="F5465E009833535713A17EA22EC0390F">
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<mods:date id="C76F4D7564E249D49D08862D6AD388A8">2014</mods:date>
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<mods:number id="E352AB50CF01D40D051AD970EFAFCA11">2014-09-30</mods:number>
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<mods:detail id="361DC313E5D1AD653F9FF837B7673D29" type="volume">
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<mods:number id="ADC28919D0ECBC24EB8849A115B23B81">121</mods:number>
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<mods:number id="A4A1A0BC5A1D4A7806F61C9A8EC3669A">3</mods:number>
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<mods:start id="20B95D7D730F73B485F07FDE3B187B1F">319</mods:start>
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<mods:classification id="CF81348800F2BF8623282948AF45EB93">journal article</mods:classification>
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<mods:identifier id="684FF9370D9D926FCFA1A7B835D97AFD" type="DOI">10.5281/zenodo.6120089</mods:identifier>
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<mods:identifier id="6F0B7ED2121782C2FA0DBA488B40A082" type="ISSN">0035-418X</mods:identifier>
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<mods:identifier id="EE17E55E61819DE6AC391BE95156DA53" type="Zenodo-Dep">6120089</mods:identifier>
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<mods:identifier id="9D11D1516C2C31FFFCD3B2B8EB78D782" type="ZooBank">AC5D93F0-A61E-4550-8973-C4BF90E30A14</mods:identifier>
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<treatment id="03E287D4FFA5FF8D24A4597BFBDCFC6B" LSID="urn:lsid:zoobank.org:act:DF7BBF6A-A8AC-449D-88F9-3780BBECC3E2" httpUri="http://treatment.plazi.org/id/03E287D4FFA5FF8D24A4597BFBDCFC6B" lastPageId="13" lastPageNumber="332" pageId="11" pageNumber="330">
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<subSubSection id="C3516549FFA5FF8B24A4597BFBC9FEDB" box="[151,1074,355,379]" pageId="11" pageNumber="330" type="nomenclature">
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<paragraph id="8BF436C2FFA5FF8B24A4597BFBC9FEDB" blockId="11.[151,1074,355,379]" box="[151,1074,355,379]" pageId="11" pageNumber="330">
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<heading id="D0BC81AEFFA5FF8B24A4597BFBC9FEDB" box="[151,1074,355,379]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="330" reason="2">
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<taxonomicName id="4C4B4D41FFA5FF8B24A4597BFE7FFEDB" ID-CoL="56GVK" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[151,388,355,379]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="330" phylum="Arthropoda" rank="species" species="paraminor" status="sp. nov.">
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<emphasis id="B93FEAD0FFA5FF8B24A4597BFE7FFEDB" bold="true" box="[151,388,355,379]" italics="true" pageId="11" pageNumber="330">Thinobius paraminor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFA5FF8B25BF597CFE24FEDB" box="[396,479,356,379]" pageId="11" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
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Figs 2, 18-19, 25, 32, 53, 55-57, 78
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFA5FF8B24D0599FFD48FE4D" pageId="11" pageNumber="330" type="materials_examined">
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<paragraph id="8BF436C2FFA5FF8B24D0599FFD48FE4D" blockId="11.[151,1073,390,493]" pageId="11" pageNumber="330">
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<materialsCitation id="3B233C9FFFA5FF8B24D0599FFD56FE4D" collectingDate="2010-09-08" collectionCode="HNHM, MHNG" county="Naerincheon river" elevation="200" latitude="38.066387" location="Gangwon-do" longLatPrecision="19" longitude="128.19084" municipality="Injae-gun district" pageId="11" pageNumber="330" specimenCount="2" typeStatus="holotype">
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TYPE MATERIAL:
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<collectionCode id="ED5AAE07FFA5FF8B25B9599FFE28FE3C" box="[394,467,391,412]" pageId="11" pageNumber="330">HNHM</collectionCode>
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,
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<typeStatus id="54F08860FFA5FF8B25D0599EFDCCFE3C" box="[483,567,390,412]" pageId="11" pageNumber="330" type="holotype">holotype</typeStatus>
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(3); “S-KOREA,
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<location id="8E946019FFA5FF8B26C4599EFC8FFE3C" LSID="urn:lsid:plazi:treatment:03E287D4FFA5FF8D24A4597BFBDCFC6B:8E946019FFA5FF8B26C4599EFC8FFE3C" box="[759,884,390,412]" county="Naerincheon river" latitude="38.066387" longLatPrecision="19" longitude="128.19084" municipality="Injae-gun district" name="Gangwon-do" pageId="11" pageNumber="330">Gangwon-do</location>
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,
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<collectingMunicipality id="6B90ACB8FFA5FF8B27B0599EFBD7FE3C" box="[899,1068,390,412]" pageId="11" pageNumber="330">Injae-gun district</collectingMunicipality>
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,
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<locationDeviation id="9A462FE0FFA5FF8B24A45987FEF6FE15" box="[151,269,415,437]" location="Inje" pageId="11" pageNumber="330">
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<quantity id="4CB39B27FFA5FF8B24A45987FF31FE15" box="[151,202,415,437]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="330" unit="km" value="2.0">2 km</quantity>
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E Inje
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</locationDeviation>
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,
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<collectingCounty id="62954E4EFFA5FF8B25285987FE31FE15" box="[283,458,415,437]" pageId="11" pageNumber="330">Naerincheon river</collectingCounty>
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, sand at W end of bridge,
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<quantity id="4CB39B27FFA5FF8B26E75987FCE8FE15" box="[724,787,415,437]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="330" unit="m" value="200.0">
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<elevation id="0066D1F1FFA5FF8B26E75987FCE8FE15" box="[724,787,415,437]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="330" unit="m" value="200.0">200 m</elevation>
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</quantity>
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, gravel/sand border at water, flotation (2B),
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<geoCoordinate id="EE7F5005FFA5FF8B251B59A1FE63FE6E" box="[296,408,441,463]" degrees="38" direction="north" minutes="03" orientation="latitude" pageId="11" pageNumber="330" precision="15" seconds="59" value="38.066387">38°03'59"N</geoCoordinate>
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,
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<geoCoordinate id="EE7F5005FFA5FF8B259659A1FDE4FE6E" box="[421,543,441,463]" degrees="128" direction="east" minutes="11" orientation="longitude" pageId="11" pageNumber="330" precision="15" seconds="27" value="128.19084">128°11'27"E</geoCoordinate>
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;
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<date id="FFF51002FFA5FF8B261F59A1FD62FE6F" box="[556,665,441,463]" pageId="11" pageNumber="330" value="2010-09-08">
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<collectingDate id="EFB1E9EAFFA5FF8B261F59A1FD62FE6F" box="[556,665,441,463]" pageId="11" pageNumber="330" value="2010-09-08">08.IX.2010</collectingDate>
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</date>
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; [leg.] Makranczy & al.”.
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<typeStatus id="54F08860FFA5FF8B24D059C0FEBFFE4C" box="[227,324,472,493]" pageId="11" pageNumber="330" type="paratype">PARATYPE</typeStatus>
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:
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<collectionCode id="ED5AAE07FFA5FF8B256259CFFE61FE4D" box="[337,410,471,493]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="330" type="Museum">MHNG</collectionCode>
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, 13, Same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFA5FF8D24D059E6FEDFFD65" lastPageId="13" lastPageNumber="332" pageId="11" pageNumber="330" type="description">
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<paragraph id="8BF436C2FFA5FF8B24D059E6FCC9FAB5" blockId="11.[151,1074,509,1653]" pageId="11" pageNumber="330">DESCRIPTION: Habitus as in Fig. 20. Measurements (n=2): HW = 0.20 (0.195- 0.20); TW = 0.20 (0.195-0.205); PW = 0.21 (0.21-0.215); SW = 0.22 (0.21-0.22); AW = 0.26 (0.25-0.26); HL = 0.19 (0.18-0.19); EL = 0.07 (0.07-0.07); TL = 0.06 (0.06- 0.06); PL = 0.17 (0.16-0.17); SL = 0.30 (0.29-0.305); SC = 0.27 (0.265-0.27); FB = 0.64 (0.63–0.64); BL = 1.22 (1.21–1.23) mm. BODY very slightly bicoloured. Head and pronotum reddish medium to dark brown, abdomen dark brown, apically more blackish; elytra medium brown, slightly darker at shoulders and scutellum. Legs light brown, mouthparts and antennae medium brown, basal antennomeres slightly lighter. Fine and dense punctation of body parts intermixed with (and thereby obscured by) microsculpture, but still well observable on forebody, body appearing weakly lustrous. Coriaceous or at places rather imbricate microsculpture covering all body parts (strongest on abdomen), but neck conspicuously smooth and shiny, transversally substrigulate. Compared to preceding plates, tergite VII notably shinier towards apex, as also tergite VIII (but barely exposed). Pubescence on forebody fine, very short and depressed, dust-like; on abdomen medium dense short setae and on apical edges of tergites a row of conspicuously long setae. Abdominal sides and apex with a few darker, medium long, stronger bristles. Strong dark bristle on outer side of supraantennal prominence at anterior border of eye and on posterior edge of vertex adjacent to temples, similar ones right behind anterior pronotal corner and on side at 3/5 length. Direction of setation on head and pronotum almost exclusively anterior, except posterior part of vertex where mixed. Elytra and abdomen with posteriorly directed setae. Antennae, legs and mouthparts with very short, not conspicuous setation, except for the stiff, darker bristle slightly distad from middle of each tibia and ones near apices of femora. Sides of elytra with three stiffer setae equally distributed on side (anterior one little behind shoulder, posterior little before outer corner).</paragraph>
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<paragraph id="8BF436C2FFA5FF8B24D05D05FCB6F9B5" blockId="11.[151,1074,509,1653]" pageId="11" pageNumber="330">Head (Fig. 25) 1.25x wider than long, temples (Fig. 18) usually straight, with a gentle curve (if any) anteriorly, rather broadly rounded posteriorly. Vertex rather flat, supraantennal prominences weakly developed. Anterior border of neck not marked with groove, only by change of microsculpture. Frontoclypeal suture appearing as fine, often shinier groove and dark line connecting supraantennal tubercles, similarly darker spots situated at both sides on middle of vertex. ANTENNAE with antennomeres 4 and 6 appearing smaller and much more transverse than adjacent (more or less as long as broad) ones (Fig. 53), articles 9-10 somewhat longer than broad.</paragraph>
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<paragraph id="8BF436C2FFA5FF8B24D05E06FBCAF9D5" blockId="11.[151,1074,509,1653]" pageId="11" pageNumber="330">Pronotum (Fig. 19) slightly transverse, 1.33x wider than long, just a little wider than head, anterior margin gently arched, with even a slight concavity before anterior corners, latter narrowly rounded. Sides and posterior marging form an almost perfect</paragraph>
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<paragraph id="8BF436C2FFA2FF8C26135DE6FD71F9B3" blockId="12.[136,1059,1533,1662]" box="[544,650,1533,1555]" pageId="12" pageNumber="331">FIGS 25-31</paragraph>
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<footnote id="E8502ACCFFA2FF8C24BB5E04FEC2F9DE" pageId="12" pageNumber="331">
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<paragraph id="8BF436C2FFA2FF8C24BB5E04FEC2F9DE" blockId="12.[136,1059,1533,1662]" pageId="12" pageNumber="331">
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(25)
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<taxonomicName id="4C4B4D41FFA2FF8C248B5E04FE7CF992" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[184,391,1564,1586]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="331" phylum="Arthropoda" rank="species" species="paraminor" status="sp. nov.">
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<emphasis id="B93FEAD0FFA2FF8C248B5E04FE7CF992" box="[184,391,1564,1586]" italics="true" pageId="12" pageNumber="331">Thinobius paraminor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFA2FF8C25BC5E05FE3EF992" box="[399,453,1565,1586]" pageId="12" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
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; head and pronotum. (26-29)
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<taxonomicName id="4C4B4D41FFA2FF8C26C35E05FC40F992" authority="Gildenkov" authorityName="Gildenkov" authorityYear="1998" box="[752,955,1564,1586]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="331" phylum="Arthropoda" rank="species" species="zerchei">
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<emphasis id="B93FEAD0FFA2FF8C26C35E05FCB6F992" box="[752,845,1564,1586]" italics="true" pageId="12" pageNumber="331">T. zerchei</emphasis>
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Gildenkov
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</taxonomicName>
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; head and pronotum, 3 (26), elytra (27), proleg (28), baseoftibia (29), antennomeres 4-8 (30). (31)
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<taxonomicName id="4C4B4D41FFA2FF8C24BB5E57FEE4F9C5" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[136,287,1615,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="331" phylum="Arthropoda" rank="species" species="injae" status="sp. nov.">
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<emphasis id="B93FEAD0FFA2FF8C24BB5E57FEE4F9C5" box="[136,287,1615,1637]" italics="true" pageId="12" pageNumber="331">Thinobius injae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFA2FF8C25155E57FEA0F9C4" box="[294,347,1615,1636]" pageId="12" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
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; apex of scutellum. Scale bar = 0.075 mm for 31, 0.1 mm for 25, 29-30, 0.2 mmfor 26-28.
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</paragraph>
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</footnote>
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<paragraph id="8BF436C2FFA3FF8D24A458FBFEE4FE7B" blockId="13.[151,1074,227,571]" pageId="13" pageNumber="332">arch of circle, posterior corners inconspicuous. pronotum with traces of impressions on middle of disc and around middle of sides. Pronotal marginal bead visible on posterior margin as a thin line. Scutellum large, setose and dull. ELYTRA (Fig. 32) long, one elytron more than twice as long as broad, gently curved in cross-section and parallelsided (very little dilation posteriorly). Shoulders moderately developed. Along suture without marginal bead, sutural corners broadly rounded. LEGS of medium length, tarsal lobes thin and appear quite short, tibiae strongly fusiform, appear a lot wider in middle than at ends.</paragraph>
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<paragraph id="8BF436C2FFA3FF8D24D059FBFDE0FD9B" blockId="13.[151,1074,227,571]" pageId="13" pageNumber="332">Abdomen very weakly fusiform, sides more or less straight and parallel; widest in the middle, very slightly less wide than elytra at broadest point. Posterior margin of tergite VII with thin palisade fringe.</paragraph>
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<paragraph id="8BF436C2FFA3FF8D24D05A56FEDFFD65" blockId="13.[151,1074,589,709]" pageId="13" pageNumber="332">PRIMARY AND SECONDARY SEXUAL FEATURES: Posterior corners of tergite VIII (similar in both sexes) gently protruding, posterior edge otherwise straight. Male sternite VIII as in Fig. 55, tergite IX as in Fig. 56, sternite IX as in Fig. 57. Aedeagus as in Fig. 78.</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFA3FF8D24D05AC1FEABFCB0" pageId="13" pageNumber="332" type="etymology">
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<paragraph id="8BF436C2FFA3FF8D24D05AC1FBCAFD50" blockId="13.[151,1073,728,784]" box="[227,1073,728,752]" pageId="13" pageNumber="332">ETYMOLOGY: The specific epithet refers to the confusing external similarity to</paragraph>
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<paragraph id="8BF436C2FFA3FF8D24A45AE0FEABFCB0" blockId="13.[151,1073,728,784]" box="[151,336,760,784]" pageId="13" pageNumber="332">
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<taxonomicName id="4C4B4D41FFA3FF8D24A45AE0FEB2FCB0" authorityName="Mulsant & Rey" authorityYear="1870" box="[151,329,760,784]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="332" phylum="Arthropoda" rank="species" species="minor">
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<emphasis id="B93FEAD0FFA3FF8D24A45AE0FEB2FCB0" box="[151,329,760,784]" italics="true" pageId="13" pageNumber="332">Thinobius minor</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFA3FF8D24D05B39FE6EFCD8" pageId="13" pageNumber="332" type="discussion">
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<paragraph id="8BF436C2FFA3FF8D24D05B39FE6EFCD8" blockId="13.[151,1074,800,889]" pageId="13" pageNumber="332">
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REMARKS: This taxon before the dissection appeared remarkably similar to
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<taxonomicName id="4C4B4D41FFA3FF8D24A45B59FE18FCF8" authority="Mulsant & Rey, 1870" authorityName="Mulsant & Rey" authorityYear="1870" box="[151,483,832,857]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="332" phylum="Arthropoda" rank="species" species="minor">
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<emphasis id="B93FEAD0FFA3FF8D24A45B59FF08FCF8" box="[151,243,833,856]" italics="true" pageId="13" pageNumber="332">T. minor</emphasis>
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Mulsant & Rey, 1870
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</taxonomicName>
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. However, based on the genital structures, it is closest to
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<taxonomicName id="4C4B4D41FFA3FF8D24805B79FE76FCD8" authority="Pyot, 1874" authorityName="Pyot" authorityYear="1874" box="[179,397,864,889]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="332" phylum="Arthropoda" rank="species" species="ligeris">
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<emphasis id="B93FEAD0FFA3FF8D24805B79FEE9FCD8" box="[179,274,864,888]" italics="true" pageId="13" pageNumber="332">T. ligeris</emphasis>
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Pyot, 1874
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFA3FF8D24D05B91FBDCFC6B" pageId="13" pageNumber="332" type="distribution">
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<paragraph id="8BF436C2FFA3FF8D24D05B91FBEBFC00" blockId="13.[227,1040,904,929]" box="[227,1040,904,929]" pageId="13" pageNumber="332">
|
||
DISTRIBUTION: The species is known only from the
|
||
<typeStatus id="54F08860FFA3FF8D27215B91FCBBFC01" box="[786,832,905,929]" pageId="13" pageNumber="332">type</typeStatus>
|
||
locality (Figs 3-4).
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA3FF8D24D05BACFBDCFC6B" blockId="13.[227,1063,947,972]" box="[227,1063,947,972]" pageId="13" pageNumber="332">
|
||
BIONOMICS: Collected with
|
||
<taxonomicName id="4C4B4D41FFA3FF8D26225BACFD9AFC6B" authorityName="Makranczy & H-" authorityYear="2014" box="[529,609,948,971]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="332" phylum="Arthropoda" rank="species" species="injae">
|
||
<emphasis id="B93FEAD0FFA3FF8D26225BACFD9AFC6B" box="[529,609,948,971]" italics="true" pageId="13" pageNumber="332">T. injae</emphasis>
|
||
</taxonomicName>
|
||
, at a lower river, open, sunny gravelbank.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |