347 lines
50 KiB
XML
347 lines
50 KiB
XML
<document id="C01CE6B6C7DE70565EB8798D15CA56FE" ID-DOI="10.5281/zenodo.184289" ID-GBIF-Dataset="7e689522-86cf-40aa-8dfe-00f5d5068f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="184289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459967162582" checkinUser="plazi" docAuthor="Chagas-Junior, Amazonas, Edgecombe, Gregory D. & Minelli, Alessandro" docDate="2008" docId="03E287A82F0E6579FF74F8C4FAC8FB63" docLanguage="en" docName="zt01888p046.pdf" docOrigin="Zootaxa 1888" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Scolopendropsis duplicata Chagas-Junior, Edgecombe & Minelli, 2008, n. sp." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="FFDBFFD02F0A6570FFE3FFF7FFE3FFF9" masterDocTitle="Variability in trunk segmentation in the centipede order Scolopendromorpha: a remarkable new species of Scolopendropsis Brandt (Chilopoda: Scolopendridae) from Brazil" masterLastPageNumber="46" masterPageNumber="36" pageNumber="40" updateTime="1698231150171" updateUser="plazi">
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<mods:mods id="2A5526159D55CD43E703E0CCF7A6BF9E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3547913AC63C23C31E77DB18F2842FBB">
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<mods:title id="FB31DE52862EDEE58070225E5678D12B">Variability in trunk segmentation in the centipede order Scolopendromorpha: a remarkable new species of Scolopendropsis Brandt (Chilopoda: Scolopendridae) from Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="1AA659787645121B4F1183F077FD3633" type="personal">
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<mods:role id="AB57AA924516064137A41E2373F2919A">
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<mods:roleTerm id="5940207D01BB8AB4602C8F21936CBC21">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="730D96617B4FC88F5E8664DCC568458D">Chagas-Junior, Amazonas</mods:namePart>
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</mods:name>
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<mods:name id="B9D9102C418938806EF66A539BA695CA" type="personal">
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<mods:roleTerm id="C2D09B6E33D11B9236B99E6FD8AC072A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1767823CE46C12CA70697D0E367017A3">Edgecombe, Gregory D.</mods:namePart>
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</mods:name>
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||
<mods:name id="FDC52BA94FEC3B8D928755D354759443" type="personal">
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<mods:role id="5679B19EB92C8410AAC63BC815DAA1A6">
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<mods:roleTerm id="421D54781D6213F9FCF308FEA9E5C594">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="7E3A0CECE0218BDF9E47E39C04239412">Minelli, Alessandro</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E159A6A6B2CFFB7BBAE9FAC9C70031AC">text</mods:typeOfResource>
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<mods:relatedItem id="9795F30469678B0BA2946DF3476C8F9C" type="host">
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<mods:titleInfo id="D23B7B7101C706ED9E916D1ED7465CED">
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<mods:title id="966EBB3FDDE6B672E09C0526B3A945C3">Zootaxa</mods:title>
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||
</mods:titleInfo>
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||
<mods:part id="37C6510A149F29618F48B738CB504064">
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||
<mods:date id="D566C7F8C7FFF1CB11FBEEDD10B355A3">2008</mods:date>
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||
<mods:detail id="E10B9165A33991FF43AEA663329488D3" type="volume">
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||
<mods:number id="50F687D6FDAF40DE20EB0D76650C4169">1888</mods:number>
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</mods:detail>
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<mods:extent id="9ED1DC3DA3BA88D1DAC9D2425C67DED9" unit="page">
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<mods:start id="1EDD9EDDF11B301DF5D7C168BA55ED72">36</mods:start>
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<mods:end id="8C0E2D6CBC384B6A413ACD99F531C939">46</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:classification id="DEE6B03C8C9DD59EFF7533B3307E5A03">journal article</mods:classification>
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<mods:identifier id="A633787BBA5DAA798F895C9A94CDFA50" type="DOI">10.5281/zenodo.184289</mods:identifier>
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<mods:identifier id="21FC81E85F404B80427F524A74A588B0" type="GBIF-Dataset">7e689522-86cf-40aa-8dfe-00f5d5068f3b</mods:identifier>
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<mods:identifier id="5249927C75F86938CE7E19B7F32C19A5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="049272F31EA9C4ADFDF7078EF0A9BE5D" type="Zenodo-Dep">184289</mods:identifier>
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</mods:mods>
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<treatment id="03E287A82F0E6579FF74F8C4FAC8FB63" ID-DOI="http://doi.org/10.5281/zenodo.6235447" ID-GBIF-Taxon="119377449" ID-Zenodo-Dep="6235447" LSID="urn:lsid:plazi:treatment:03E287A82F0E6579FF74F8C4FAC8FB63" httpUri="http://treatment.plazi.org/id/03E287A82F0E6579FF74F8C4FAC8FB63" lastPageId="9" lastPageNumber="45" pageId="4" pageNumber="40">
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||
<subSubSection id="C35165352F0E6574FF74F8C4FEF9F889" pageId="4" pageNumber="40" type="nomenclature">
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||
<paragraph id="8BF436BE2F0E6574FF74F8C4FDFEF8B4" blockId="4.[151,541,1843,1904]" box="[151,541,1843,1869]" pageId="4" pageNumber="40">
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||
<heading id="D0BC81D22F0E6574FF74F8C4FDFEF8B4" bold="true" box="[151,541,1843,1869]" fontSize="11" level="1" pageId="4" pageNumber="40" reason="1">
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||
<emphasis id="B93FEAAC2F0E6574FF74F8C4FDFEF8B4" bold="true" box="[151,541,1843,1869]" pageId="4" pageNumber="40">
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||
<taxonomicName id="4C4B4D3D2F0E6574FF74F8C4FE30F8B4" box="[151,467,1843,1869]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="40" phylum="Arthropoda" rank="species" species="duplicata" status="sp. nov.">
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||
<emphasis id="B93FEAAC2F0E6574FF74F8C4FE30F8B4" bold="true" box="[151,467,1843,1869]" italics="true" pageId="4" pageNumber="40">Scolopendropsis duplicata</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A20C57D72F0E6574FE39F8C4FDFEF8B4" box="[474,541,1843,1869]" pageId="4" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BF436BE2F0E6574FF74F8A1FEF9F889" blockId="4.[151,541,1843,1904]" box="[151,282,1878,1904]" pageId="4" pageNumber="40">
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||
<figureCitation id="13702A3B2F0E6574FF74F8A1FEF6F889" box="[151,277,1878,1904]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 5" captionStart-2="FIGURES 6 – 11" captionStart-3="FIGURES 12 – 14" captionStartId-0="2.[151,255,1249,1273]" captionStartId-1="4.[151,269,1624,1648]" captionStartId-2="6.[151,269,1933,1957]" captionStartId-3="7.[151,269,1968,1992]" captionTargetBox-0="[191,1412,300,1241]" captionTargetBox-1="[193,1389,204,1587]" captionTargetBox-2="[322,1264,206,1891]" captionTargetBox-3="[339,1249,205,1929]" captionTargetId-0="figure@2.[191,1422,300,1244]" captionTargetId-1="figure@4.[185,1405,194,1599]" captionTargetId-2="figure@6.[306,1280,194,1902]" captionTargetId-3="figure@7.[339,1249,194,1935]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Scolopendropsis duplicata n. sp. Paratype MNRJ 15306. Lateral view. Scale bar, 1 mm." captionText-1="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." captionText-2="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." captionText-3="FIGURES 12 – 14. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 12, trunk segments 3 – 5, lateral view; 13, trunk segments 7 – 12, dorsal view; 14, trunk segments 7 – 11, ventral view. Scale bars, 1 mm." httpUri-0="https://zenodo.org/record/184290/files/figure.png" httpUri-1="https://zenodo.org/record/184291/files/figure.png" httpUri-2="https://zenodo.org/record/184292/files/figure.png" httpUri-3="https://zenodo.org/record/184293/files/figure.png" pageId="4" pageNumber="40">Figs. 1–14</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35165352F0E6575FF74F852FE7EFEAB" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40" type="diagnosis">
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<paragraph id="8BF436BE2F0E6575FF74F852FE7EFEAB" blockId="4.[151,1436,1957,2024]" lastBlockId="5.[151,1436,152,2018]" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40">
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||
<emphasis id="B93FEAAC2F0E6574FF74F852FEFAF846" bold="true" box="[151,281,1957,1983]" pageId="4" pageNumber="40">
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<emphasis id="B93FEAAC2F0E6574FF74F852FEF3F846" bold="true" box="[151,272,1957,1983]" italics="true" pageId="4" pageNumber="40">Diagnosis</emphasis>
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:
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</emphasis>
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Thirty-nine or 43 trunk segments; paramedian sutures of T1 incomplete anteriorly, extending ¾ of length; posterior border of coxopleuron usually bearing single short spine caudal to posterolateral corner of pore field; ultimate leg prefemur with two ventromedial and one to three dorsomedial spines in longitudinal rows (when multiple), the dorsomedial row aligned with prefemoral process; latter with two or three apical spines; flattened medial face of prefemur variably bearing one to six small spines that, when maximally developed, are aligned in two rows; posterior halves of ultimate leg prefemur and femur with sulcus-like longitudinal depression dorsad.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35165352F0F6575FF25FEA8FE35FCF3" pageId="5" pageNumber="41" type="materials_examined">
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||
<paragraph id="8BF436BE2F0F6575FF25FEA8FE35FCF3" blockId="5.[151,1436,152,2018]" pageId="5" pageNumber="41">
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<emphasis id="B93FEAAC2F0F6575FF25FEA8FE73FE80" bold="true" box="[198,400,351,377]" pageId="5" pageNumber="41">
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||
<emphasis id="B93FEAAC2F0F6575FF25FEA8FE65FE80" bold="true" box="[198,390,351,377]" italics="true" pageId="5" pageNumber="41">
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<typeStatus id="54F0881C2F0F6575FF25FEA8FEE3FE80" box="[198,256,351,377]" pageId="5" pageNumber="41">Type</typeStatus>
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specimens
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</emphasis>
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:
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</emphasis>
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Segment number (39 or 43) indicated in parentheses.
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<typeStatus id="54F0881C2F0F6575FBC8FE97FB7BFE83" box="[1067,1176,352,378]" pageId="5" pageNumber="41" type="holotype">Holotype</typeStatus>
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||
<collectionCode id="ED5AAE7B2F0F6575FB40FE97FB12FE83" box="[1187,1265,352,378]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="41">MNRJ</collectionCode>
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15258, 1 ex., Tocantins State, Porto Nacional,
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<geoCoordinate id="EE7F50792F0F6575FDC7FE7FFD7CFE5B" box="[548,671,392,418]" direction="south" orientation="latitude" pageId="5" pageNumber="41" precision="5" value="-10.0589">10.0589°S</geoCoordinate>
|
||
<geoCoordinate id="EE7F50792F0F6575FD49FE7FFCC2FE5B" box="[682,801,392,418]" direction="west" orientation="longitude" pageId="5" pageNumber="41" precision="55" value="-48.412">48.412°W</geoCoordinate>
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||
, 22L 0 783671 UTM 8886955,
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||
<date id="FFF5107E2F0F6575FB7CFE7FFB5DFE5B" box="[1183,1214,392,418]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29</date>
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.IX–07.X.2001, D. Pavan (43).
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<typeStatus id="54F0881C2F0F6575FEC6FE47FE74FE33" box="[293,407,432,458]" pageId="5" pageNumber="41" type="paratype">Paratypes</typeStatus>
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all from
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<typeStatus id="54F0881C2F0F6575FDE0FE47FDD6FE33" box="[515,565,432,458]" pageId="5" pageNumber="41">type</typeStatus>
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locality, leg. D. Pavan unless indicated otherwise:
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<typeStatus id="54F0881C2F0F6575FB69FE47FB0CFE33" box="[1162,1263,432,458]" pageId="5" pageNumber="41" type="paratype">paratype</typeStatus>
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||
<collectionCode id="ED5AAE7B2F0F6575FB15FE47FAA7FE33" box="[1270,1348,432,458]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="41">MNRJ</collectionCode>
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15259, 1 ex.,
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<date id="FFF5107E2F0F6575FF0FFE2FFE59FE0B" box="[236,442,472,498]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX–07.X.2001</date>
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||
(39);
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<collectionCode id="ED5AAE7B2F0F6575FDECFE2FFDBEFE0B" box="[527,605,472,498]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="41">MNRJ</collectionCode>
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||
15305, 1 ex.,
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<date id="FFF5107E2F0F6575FCC3FE2FFC07FE0B" box="[800,996,472,498]" pageId="5" pageNumber="41" value="2001-10-29" valueMax="2001-10-07" valueMin="2001-10-29">29.X–07.X.2001</date>
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||
(43);
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||
<collectionCode id="ED5AAE7B2F0F6575FBDAFE2FFB64FE0B" box="[1081,1159,472,498]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="41">MNRJ</collectionCode>
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15306, 6 ex.,
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<date id="FFF5107E2F0F6575FAA9FE2FFEF1FDE3" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX– 07.X.2001</date>
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(39);
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<collectionCode id="ED5AAE7B2F0F6575FE82FDF7FE7CFDE3" box="[353,415,512,538]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="5" pageNumber="41">IBSP</collectionCode>
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1519, 1 ex., Ribeirão
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<collectingCountry id="F35C762E2F0F6575FD55FDF7FCABFDE3" box="[694,840,512,538]" name="Saint Lucia" pageId="5" pageNumber="41">Santa Luzia</collectingCountry>
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||
, Fazenda Sandoval, 2000, Equipe Investco/Ulbra (43);
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||
<collectionCode id="ED5AAE7B2F0F6575FF34FDDFFEF6FDBB" box="[215,277,552,578]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="5" pageNumber="41">IBSP</collectionCode>
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||
1498, 1 ex.,
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||
<date id="FFF5107E2F0F6575FE4CFDDFFD95FDBB" box="[431,630,552,578]" pageId="5" pageNumber="41" value="2000-08-07" valueMax="2000-08-11" valueMin="2000-08-07">07–11.VIII.2000</date>
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||
, I. Knysak, R. Martins & G. Puorto (43);
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||
<collectionCode id="ED5AAE7B2F0F6575FBBCFDDFFB7EFDBB" box="[1119,1181,552,578]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="5" pageNumber="41">IBSP</collectionCode>
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||
2406, 1 ex., Ribeirão
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||
<collectingCountry id="F35C762E2F0F6575FF74FDA7FEC6FD93" box="[151,293,592,618]" name="Saint Lucia" pageId="5" pageNumber="41">Santa Luzia</collectingCountry>
|
||
, Fazenda Sandoval,
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||
<date id="FFF5107E2F0F6575FDF1FDA7FD2FFD93" box="[530,716,592,618]" pageId="5" pageNumber="41" value="2000-07-10" valueMax="2000-07-13" valueMin="2000-07-10">10–13.VII.2000</date>
|
||
, Equipe Investco/Ulbra (39);
|
||
<collectionCode id="ED5AAE7B2F0F6575FBC4FDA7FB86FD93" box="[1063,1125,592,618]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="5" pageNumber="41">IBSP</collectionCode>
|
||
2843, 1 ex., Fazenda Sandoval,
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||
<date id="FFF5107E2F0F6575FF0FFD8FFE64FD6B" box="[236,391,632,658]" pageId="5" pageNumber="41" value="2000-08-11">11.VIII.2000</date>
|
||
, I. Knysak, R. Martins & G. Puorto (43);
|
||
<collectionCode id="ED5AAE7B2F0F6575FC42FD8FFC3CFD6B" box="[929,991,632,658]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="5" pageNumber="41">IBSP</collectionCode>
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||
2392, 2 ex., Fazenda Sandoval,
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||
<date id="FFF5107E2F0F6575FA9CFD8FFEF2FD43" pageId="5" pageNumber="41" value="2000-05-05" valueMax="2000-05-10" valueMin="2000-05-05">5– 10.V.2000</date>
|
||
, I. Knysak, R. Martins & G. Puorto (43);
|
||
<collectionCode id="ED5AAE7B2F0F6575FD10FD57FCB3FD43" box="[755,848,672,698]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="41">NMNH</collectionCode>
|
||
, 1 ex.,
|
||
<date id="FFF5107E2F0F6575FC42FD57FB8CFD43" box="[929,1135,672,698]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX–07.X.2001</date>
|
||
(39);
|
||
<collectionCode id="ED5AAE7B2F0F6575FB56FD57FB1BFD43" LSID="urn:lsid:biocol.org:col:33791" box="[1205,1272,672,698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="41">MCZ</collectionCode>
|
||
, 1 ex.,
|
||
<date id="FFF5107E2F0F6575FAA9FD57FEF1FD1B" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX– 07.X.2001</date>
|
||
(39);
|
||
<collectionCode id="ED5AAE7B2F0F6575FEB9FD3FFE53FD1B" box="[346,432,712,738]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="5" pageNumber="41">NCSM</collectionCode>
|
||
, 2 ex.,
|
||
<date id="FFF5107E2F0F6575FDE7FD3FFD32FD1B" box="[516,721,712,738]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX–07.X.2001</date>
|
||
(43, 39);
|
||
<collectionCode id="ED5AAE7B2F0F6575FCA6FD3FFC72FD1B" box="[837,913,712,738]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="41">MZSP</collectionCode>
|
||
, 3 ex.,
|
||
<date id="FFF5107E2F0F6575FC06FD3FFB50FD1B" box="[997,1203,712,738]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX–07.X.2001</date>
|
||
(43, 39);
|
||
<collectionCode id="ED5AAE7B2F0F6575FAC4FD3FFA61FD1B" box="[1319,1410,712,738]" pageId="5" pageNumber="41">BMNH</collectionCode>
|
||
, 2 ex.,
|
||
<date id="FFF5107E2F0F6575FF2BFD07FE75FCF3" box="[200,406,752,778]" pageId="5" pageNumber="41" value="2001-09-29" valueMax="2001-10-07" valueMin="2001-09-29">29.IX–07.X.2001</date>
|
||
(39).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165352F0F6575FF25FCE0FE06FCA3" pageId="5" pageNumber="41" type="etymology">
|
||
<paragraph id="8BF436BE2F0F6575FF25FCE0FE06FCA3" blockId="5.[151,1436,152,2018]" pageId="5" pageNumber="41">
|
||
<emphasis id="B93FEAAC2F0F6575FF25FCE0FEB3FCC8" bold="true" box="[198,336,791,817]" italics="true" pageId="5" pageNumber="41">Etymology:</emphasis>
|
||
The specific name refers to the near doubling of trunk segment numbers compared to the most closely related species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165352F0F6578FF25FC90FCB5F868" lastPageId="8" lastPageNumber="44" pageId="5" pageNumber="41" type="description">
|
||
<paragraph id="8BF436BE2F0F6578FF25FC90FCB5F868" blockId="5.[151,1436,152,2018]" lastBlockId="8.[151,1437,1631,2017]" lastPageId="8" lastPageNumber="44" pageId="5" pageNumber="41">
|
||
<emphasis id="B93FEAAC2F0F6575FF25FC90FEBFFC78" bold="true" box="[198,348,871,897]" pageId="5" pageNumber="41">
|
||
<emphasis id="B93FEAAC2F0F6575FF25FC90FEB1FC78" bold="true" box="[198,338,871,897]" italics="true" pageId="5" pageNumber="41">Description</emphasis>
|
||
:
|
||
</emphasis>
|
||
Length of 39-segmented individuals ranging from
|
||
<quantity id="4CB39B5B2F0F6575FC50FC9FFBD6FC7B" box="[947,1077,872,898]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="7.4" metricValueMin="3.1" pageId="5" pageNumber="41" unit="mm" value="52.5" valueMax="74.0" valueMin="31.0">31–74 mm</quantity>
|
||
; that of 43-segmented individuals from
|
||
<quantity id="4CB39B5B2F0F6575FEEDFC67FE6CFC53" box="[270,399,912,938]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="7.8" metricValueMin="5.3" pageId="5" pageNumber="41" unit="mm" value="65.5" valueMax="78.0" valueMin="53.0">53–78 mm</quantity>
|
||
. Cephalic plate, trunk and legs yellow in specimens preserved in alcohol; forcipulae and ultimate legs orange.
|
||
<emphasis id="B93FEAAC2F0F6575FE71FC4FFDDEFC28" box="[402,573,952,977]" italics="true" pageId="5" pageNumber="41">Cephalic plate</emphasis>
|
||
: smooth, 1.3 times as long as wide, distinctly narrower than T1, with four ocelli on each side. Posterior longitudinal median suture extending for ½–¾ of length (
|
||
<figureCitation id="13702A3B2F0F6575FB5AFC17FAE0FC03" box="[1209,1283,992,1018]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">Fig. 6</figureCitation>
|
||
), difficult to detect in juveniles. Posterior border overlain by T1; basal plates at posterolateral corners delimited by continuous suture across posteromedial margin.
|
||
<emphasis id="B93FEAAC2F0F6575FD6CFBC7FD1DFBB0" box="[655,766,1072,1097]" italics="true" pageId="5" pageNumber="41">Antennae</emphasis>
|
||
: short, usually reaching backwards to midlength of T1; with 17 articles, basal six glabrous/sparsely hirsute.
|
||
<emphasis id="B93FEAAC2F0F6575FD15FBAFFC71FB88" box="[758,914,1112,1137]" italics="true" pageId="5" pageNumber="41">Coxosternum</emphasis>
|
||
: tooth plates about 1.3 times as long as wide (
|
||
<figureCitation id="13702A3B2F0F6575FF7CFB77FF11FB63" box="[159,242,1152,1178]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="5" pageNumber="41">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3B2F0F6575FF1CFB77FEF3FB63" box="[255,272,1152,1178]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">7</figureCitation>
|
||
); inner teeth fused, with two ill-defined cusps, lateral teeth isolated, apices slightly caudad to inner teeth; basal sutures approximately transverse. Longitudinal median suture bifurcating posteriorly into inverted Y-shape, crossed at or near division point by a subtransverse suture curving anteriad laterally and extending to forcipular bases (
|
||
<figureCitation id="13702A3B2F0F6575FE87FB0FFE5AFAEB" box="[356,441,1272,1298]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="5" pageNumber="41">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3B2F0F6575FE24FB0FFE3BFAEB" box="[455,472,1272,1298]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">7</figureCitation>
|
||
). Process of forcipular trochanteroprefemur with two blunt apical denticles.
|
||
<emphasis id="B93FEAAC2F0F6575FA8AFB0FFEC7FAC0" italics="true" pageId="5" pageNumber="41">Second maxilla</emphasis>
|
||
: symmetrical accessory claws on each side of apical claw. Coxosternum with complete median sulcus.
|
||
<emphasis id="B93FEAAC2F0F6575FF13FABFFEAFFA98" box="[240,332,1352,1377]" italics="true" pageId="5" pageNumber="41">Tergites</emphasis>
|
||
: smooth; T1 with paramedian sutures on posterior ¾ (
|
||
<figureCitation id="13702A3B2F0F6575FC3AFABFFBC0FA9B" box="[985,1059,1352,1378]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">Fig. 6</figureCitation>
|
||
); TT2-38 or TT2-42 with complete paramedian sutures; ultimate tergite (T39 or T43) about 1.5 times as long as penultimate, with complete median longitudinal suture (
|
||
<figureCitation id="13702A3B2F0F6575FE03FA6FFDD2FA4B" box="[480,561,1432,1458]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="5" pageNumber="41">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3B2F0F6575FDDDFA6FFDADFA4B" box="[574,590,1432,1458]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">8</figureCitation>
|
||
). Margination on ultimate tergite only. Pretergites usually present on the posterior segments.
|
||
<emphasis id="B93FEAAC2F0F6575FE65FA37FE08FA20" box="[390,491,1472,1497]" italics="true" pageId="5" pageNumber="41">Sternites</emphasis>
|
||
: smooth; complete paramedian sutures present on SS2-38 or SS2-42 (
|
||
<figureCitation id="13702A3B2F0F6575FAD6FA37FA6DFA23" box="[1333,1422,1472,1498]" captionStart="FIGURES 12 – 14" captionStartId="7.[151,269,1968,1992]" captionTargetBox="[339,1249,205,1929]" captionTargetId="figure@7.[339,1249,194,1935]" captionTargetPageId="7" captionText="FIGURES 12 – 14. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 12, trunk segments 3 – 5, lateral view; 13, trunk segments 7 – 12, dorsal view; 14, trunk segments 7 – 11, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184293/files/figure.png" pageId="5" pageNumber="41">Fig. 14</figureCitation>
|
||
). Ultimate sternite considerably longer than wide (
|
||
<figureCitation id="13702A3B2F0F6575FD3CFA1FFCD7F9FB" box="[735,820,1512,1538]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="5" pageNumber="41">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3B2F0F6575FCA0FA1FFCB7F9FB" box="[835,852,1512,1538]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="5" pageNumber="41">9</figureCitation>
|
||
), lateral margins gently converging caudad, with median longitudinal depression; posterior margin gently convex or straight.
|
||
<emphasis id="B93FEAAC2F0F6575FBEEF9E7FB99F9D0" box="[1037,1146,1552,1577]" italics="true" pageId="5" pageNumber="41">Spiracles</emphasis>
|
||
: on segments 3, 5, 8, 10, 12, 14, 16, 18, 20, 22, 24, 26, 28, 30, 32, 34, 36, 38 of all specimens; those with 43 trunk segments additionally bear spiracles on segments 40 and 42. Anteriormost spiracle enlarged relative to others, nearly half length of T3; second spiracle (segment 5) only slightly larger than those immediately more posterior. Longitudinal pleurites parallel to trunk axis (
|
||
<figureCitation id="13702A3B2F0F6575FDE1F947FDBAF933" box="[514,601,1712,1738]" captionStart="FIGURES 12 – 14" captionStartId="7.[151,269,1968,1992]" captionTargetBox="[339,1249,205,1929]" captionTargetId="figure@7.[339,1249,194,1935]" captionTargetPageId="7" captionText="FIGURES 12 – 14. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 12, trunk segments 3 – 5, lateral view; 13, trunk segments 7 – 12, dorsal view; 14, trunk segments 7 – 11, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184293/files/figure.png" pageId="5" pageNumber="41">Fig. 12</figureCitation>
|
||
).
|
||
<emphasis id="B93FEAAC2F0F6575FD8EF947FD14F930" box="[621,759,1712,1737]" italics="true" pageId="5" pageNumber="41">Coxopleura</emphasis>
|
||
: longer than wide, pore field half as wide as coxopleuron, separated from pore-free part of pleuron by well-incised longitudinal sulcus; posterior border of coxopleuron bearing short spine immediately caudal to posterolateral corner of pore field (
|
||
<figureCitation id="13702A3B2F0F6575FBC9F8F7FB91F8E3" box="[1066,1138,1792,1818]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="5" pageNumber="41">Fig. 4</figureCitation>
|
||
), usually present on only one side, sometimes absent in specimens with 39 leg-bearing segments. Posterior border of coxopleuron abruptly truncated, coxopleural process absent.
|
||
<emphasis id="B93FEAAC2F0F6575FD27F8A7FD18F890" box="[708,763,1872,1897]" italics="true" pageId="5" pageNumber="41">Legs</emphasis>
|
||
: tarsi composed of two articles. Tarsus 1 shorter than tarsus 2. Spur present on tarsus 1 of legs 1–37 or 1–41; ultimate and penultimate legs lacking spur. Pretarsus longer or equal in length to tarsus 2; proximal 1/3 pale, distal 2/3 strongly pigmented with concave ventral surface bounded on each side by sharp marginal ridge; pretarsi of legs 1–38 or 1–42 with two accessory spurs about ¼ as long as pretarsi on anterior segments, increasing to about 1/3 length of pretarsi on posterior segments.
|
||
<emphasis id="B93FEAAC2F026578FF0EF97FFE6FF958" box="[237,396,1672,1697]" italics="true" pageId="8" pageNumber="44">Ultimate legs</emphasis>
|
||
: thickened, forcipulate, with relatively short articles; prefemora with two ventromedial and one to three dorsomedial spines in longitudinal rows (when multiple), the dorsomedial row aligned with distomedial process (
|
||
<figureCitation id="13702A3B2F026578FE71F920FE05F908" box="[402,486,1751,1777]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1624,1648]" captionTargetBox="[193,1389,204,1587]" captionTargetId="figure@4.[185,1405,194,1599]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Scolopendropsis duplicata n. sp. Holotype MNRJ 15258. 2, forcipulae; 3, tooth plates (tp) and processes of trochanteroprefemora (pr) of forcipulae; 4, sternite and coxopleurae of segment 43; 5, segment 43 and ultimate legs, dorsal view. Scale bars, 0.5 mm. Additional abbreviations: dp, distomedial prefemoral process; ld, longitudinal depression on femur; ls, longitudinal sulcus delimiting pore field; sp, spine at posterior border of coxopleuron." httpUri="https://zenodo.org/record/184291/files/figure.png" pageId="8" pageNumber="44">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3B2F026578FE17F920FDF7F908" box="[500,532,1751,1777]" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="8" pageNumber="44">10</figureCitation>
|
||
); the latter conical, bearing two or three small apical spines. Medial face of prefemur flattened, variably bearing one to six small spines that, when maximally developed, are aligned in two rows. Posterior halves of prefemur and femur with sulci-like longitudinal depressions on dorsal side (
|
||
<figureCitation id="13702A3B2F026578FA8FF8D0FF5BF890" captionStart="FIGURES 6 – 11" captionStartId="6.[151,269,1933,1957]" captionTargetBox="[322,1264,206,1891]" captionTargetId="figure@6.[306,1280,194,1902]" captionTargetPageId="6" captionText="FIGURES 6 – 11. Scolopendropsis duplicata n. sp. Paratype MNRJ 2392. 6, cephalic plate and T 1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm." httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="8" pageNumber="44">Fig. 10</figureCitation>
|
||
). Pretarsus with one basal accessory spur, shorter than those on ambulatory legs; ventral surface of pretarsus with serrated ridge bearing numerous small, sharp teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3466362F0C6576FF74F87AFCCEF813" httpUri="https://zenodo.org/record/184292/files/figure.png" pageId="6" pageNumber="42" targetBox="[322,1264,206,1891]" targetPageId="6">
|
||
<paragraph id="8BF436BE2F0C6576FF74F87AFCCEF813" blockId="6.[151,1436,1933,2026]" pageId="6" pageNumber="42">
|
||
<emphasis id="B93FEAAC2F0C6576FF74F87AFEB3F85C" bold="true" box="[151,336,1933,1957]" pageId="6" pageNumber="42">FIGURES 6–11.</emphasis>
|
||
<taxonomicName id="4C4B4D3D2F0C6576FEB9F879FD94F85C" box="[346,631,1934,1957]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="duplicata" status="sp. nov.">
|
||
<emphasis id="B93FEAAC2F0C6576FEB9F879FD94F85C" box="[346,631,1934,1957]" italics="true" pageId="6" pageNumber="42">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAAC2F0C6576FD63F87AFD5DF85C" bold="true" box="[640,702,1933,1957]" pageId="6" pageNumber="42">
|
||
<taxonomicNameLabel id="A20C57D72F0C6576FD63F87AFD5DF85C" box="[640,702,1933,1957]" pageId="6" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Paratype MNRJ 2392. 6, cephalic plate and T1; 7, forcipular coxosternum; 8, segment 43 and ultimate legs, dorsal view; 9, segment 43 and ultimate legs, ventral view; 10, ultimate legs, dorsal view; 11, ultimate legs, ventral view. Scale bars, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466362F0D6577FF74F847FB8EF813" httpUri="https://zenodo.org/record/184293/files/figure.png" pageId="7" pageNumber="43" targetBox="[339,1249,205,1929]" targetPageId="7">
|
||
<paragraph id="8BF436BE2F0D6577FF74F847FB8EF813" blockId="7.[151,1436,1968,2026]" pageId="7" pageNumber="43">
|
||
<emphasis id="B93FEAAC2F0D6577FF74F847FE83F831" bold="true" box="[151,352,1968,1992]" pageId="7" pageNumber="43">FIGURES 12–14.</emphasis>
|
||
<taxonomicName id="4C4B4D3D2F0D6577FE89F846FD6BF831" box="[362,648,1969,1992]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="duplicata" status="sp. nov.">
|
||
<emphasis id="B93FEAAC2F0D6577FE89F846FD6BF831" box="[362,648,1969,1992]" italics="true" pageId="7" pageNumber="43">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAAC2F0D6577FD72F847FD33F831" bold="true" box="[657,720,1968,1992]" pageId="7" pageNumber="43">
|
||
<taxonomicNameLabel id="A20C57D72F0D6577FD72F847FD33F831" box="[657,720,1968,1992]" pageId="7" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Paratype MNRJ 2392. 12, trunk segments 3–5, lateral view; 13, trunk segments 7–12, dorsal view; 14, trunk segments 7–11, ventral view. Scale bars, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466362F026578FF74FA05FB20F9D2" httpUri="https://zenodo.org/record/184294/files/figure.png" pageId="8" pageNumber="44" targetBox="[154,1411,217,1457]" targetPageId="8">
|
||
<paragraph id="8BF436BE2F026578FF74FA05FB20F9D2" blockId="8.[151,1436,1522,1580]" pageId="8" pageNumber="44">
|
||
<emphasis id="B93FEAAC2F026578FF74FA05FECAF9F3" bold="true" box="[151,297,1522,1546]" pageId="8" pageNumber="44">FIGURE 15.</emphasis>
|
||
South America, with dot indicating type locality of
|
||
<taxonomicName id="4C4B4D3D2F026578FC8FFA05FB69F9F0" box="[876,1162,1522,1545]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="duplicata" status="sp. nov.">
|
||
<emphasis id="B93FEAAC2F026578FC8FFA05FB69F9F0" box="[876,1162,1522,1545]" italics="true" pageId="8" pageNumber="44">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAAC2F026578FB71FA05FB32F9F3" bold="true" box="[1170,1233,1522,1546]" pageId="8" pageNumber="44">
|
||
<taxonomicNameLabel id="A20C57D72F026578FB71FA05FB32F9F3" box="[1170,1233,1522,1546]" pageId="8" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Brazil: Tocantins State: Porto Nacional). The distribution of
|
||
<taxonomicName id="4C4B4D3D2F026578FDBCF9E2FC61F9D2" authority="Brandt, 1841" authorityName="Brandt" authorityYear="1841" box="[607,898,1556,1580]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="bahiensis">
|
||
<emphasis id="B93FEAAC2F026578FDBCF9E2FD3CF9D5" box="[607,735,1557,1580]" italics="true" pageId="8" pageNumber="44">S. bahiensis</emphasis>
|
||
(Brandt, 1841)
|
||
</taxonomicName>
|
||
is indicated by black squares.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35165352F026578FF25F868FCFFF818" pageId="8" pageNumber="44" type="distribution">
|
||
<paragraph id="8BF436BE2F026578FF25F868FCFFF818" blockId="8.[151,1437,1631,2017]" pageId="8" pageNumber="44">
|
||
<emphasis id="B93FEAAC2F026578FF25F868FE81F840" bold="true" box="[198,354,1951,1977]" italics="true" pageId="8" pageNumber="44">Distribution:</emphasis>
|
||
<collectingCountry id="F35C762E2F026578FE88F868FE55F840" box="[363,438,1951,1977]" name="Brazil" pageId="8" pageNumber="44">Brazil</collectingCountry>
|
||
, Tocantins State.
|
||
<taxonomicName id="4C4B4D3D2F026578FD66F857FC5EF840" box="[645,957,1952,1977]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="44" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F026578FD66F857FC5EF840" box="[645,957,1952,1977]" italics="true" pageId="8" pageNumber="44">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
is restricted to typical “cerrado” vegetation, and is known only from its
|
||
<typeStatus id="54F0881C2F026578FDF6F830FDA4F818" box="[533,583,1991,2017]" pageId="8" pageNumber="44">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13702A3B2F026578FD54F830FCEDF818" box="[695,782,1991,2017]" captionStart="FIGURE 15" captionStartId="8.[151,255,1522,1546]" captionTargetBox="[154,1411,217,1457]" captionTargetId="figure@8.[151,1436,194,1498]" captionTargetPageId="8" captionText="FIGURE 15. South America, with dot indicating type locality of Scolopendropsis duplicata n. sp. (Brazil: Tocantins State: Porto Nacional). The distribution of S. bahiensis (Brandt, 1841) is indicated by black squares." httpUri="https://zenodo.org/record/184294/files/figure.png" pageId="8" pageNumber="44">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165352F036579FF25FF60FAC8FB63" pageId="9" pageNumber="45" type="discussion">
|
||
<paragraph id="8BF436BE2F036579FF25FF60FE61FDE3" blockId="9.[151,1436,151,1178]" pageId="9" pageNumber="45">
|
||
<emphasis id="B93FEAAC2F036579FF25FF60FEB6FF48" bold="true" box="[198,341,151,177]" italics="true" pageId="9" pageNumber="45">Discussion:</emphasis>
|
||
Despite the fact that this new species differs from all other
|
||
<taxonomicName id="4C4B4D3D2F036579FBF4FF6FFAE6FF4B" box="[1047,1285,152,178]" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
|
||
with respect to a character (trunk segment number) that is routinely used to diagnose genera or families within the order, the highly detailed similarity to
|
||
<taxonomicName id="4C4B4D3D2F036579FDFAFF1FFCB1FEF8" box="[537,850,232,257]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="bahiensis">
|
||
<emphasis id="B93FEAAC2F036579FDFAFF1FFCB1FEF8" box="[537,850,232,257]" italics="true" pageId="9" pageNumber="45">Scolopendropsis bahiensis</emphasis>
|
||
</taxonomicName>
|
||
in all morphological characters apart from trunk segmentation leads us to favour its congeneric classification. We note the paradox that variability in scolopendromorph segmentation is a remarkable discovery, and yet
|
||
<taxonomicName id="4C4B4D3D2F036579FC65FECFFBF1FEA8" box="[902,1042,312,337]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F036579FC65FECFFBF1FEA8" box="[902,1042,312,337]" italics="true" pageId="9" pageNumber="45">S. duplicata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D3D2F036579FBAFFECFFB3BFEA8" box="[1100,1240,312,337]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="bahiensis">
|
||
<emphasis id="B93FEAAC2F036579FBAFFECFFB3BFEA8" box="[1100,1240,312,337]" italics="true" pageId="9" pageNumber="45">S. bahiensis</emphasis>
|
||
</taxonomicName>
|
||
are so similar in other respects and their sister group relationship so highly corroborated that generic separation is unwarranted. The discovery of segmental polymorphism in
|
||
<taxonomicName id="4C4B4D3D2F036579FCECFE7FFC7FFE58" box="[783,924,392,417]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F036579FCECFE7FFC7FFE58" box="[783,924,392,417]" italics="true" pageId="9" pageNumber="45">S. duplicata</emphasis>
|
||
</taxonomicName>
|
||
suggests that
|
||
<bibRefCitation id="EFDA4B4F2F036579FBA1FE7FFAEEFE5B" author="Schileyko" box="[1090,1293,392,418]" pageId="9" pageNumber="46" refString="Schileyko, A. A. (2006) Redescription of Scolopendropsis bahiensis (Brandt, 1841), the relations between Scolopendropsis and Rhoda, and notes on some characters used in scolopendromorph taxonomy (Chilopoda: Scolopendromorpha). Arthropoda Selecta, 15, 9 - 17." type="journal article" year="2006">Schileyko (2006)</bibRefCitation>
|
||
was correct in interpreting
|
||
<taxonomicName id="4C4B4D3D2F036579FEA5FE47FE30FE30" box="[326,467,432,457]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="bahiensis">
|
||
<emphasis id="B93FEAAC2F036579FEA5FE47FE30FE30" box="[326,467,432,457]" italics="true" pageId="9" pageNumber="45">S. bahiensis</emphasis>
|
||
</taxonomicName>
|
||
as also polymorphic. More evidence is now available demonstrating that segmental polymorphism is indeed possible in a scolopendromorph species and more specifically is known within this restricted clade.
|
||
</paragraph>
|
||
<paragraph id="8BF436BE2F036579FF25FDDFFDCAFCCB" blockId="9.[151,1436,151,1178]" pageId="9" pageNumber="45">
|
||
Because the genitalia are concealed beneath the genital sternite, we have not amassed an adequate sample size to determine how segmental polymorphism correlates with sex. Dissected individuals with 39 segments include ones with spermatophores, confirming that at least some are males. Spermatophores are lacking in dissected 43-segmented specimens, but we have not ascertained whether they are invariably females. Even if some exceptions are eventually found to the possible rule that males in this species have 39 segments and females have 43,
|
||
<taxonomicName id="4C4B4D3D2F036579FE80FD07FE0CFCF0" box="[355,495,752,777]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F036579FE80FD07FE0CFCF0" box="[355,495,752,777]" italics="true" pageId="9" pageNumber="45">S. duplicata</emphasis>
|
||
</taxonomicName>
|
||
is likely to represent the first example of sexual dimorphism in segment number in non-geophilomorph centipedes.
|
||
</paragraph>
|
||
<paragraph id="8BF436BE2F036579FF25FCB7FAC8FB63" blockId="9.[151,1436,151,1178]" pageId="9" pageNumber="45">
|
||
Most specimens of
|
||
<taxonomicName id="4C4B4D3D2F036579FE51FCB7FDA3FCA0" box="[434,576,832,857]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F036579FE51FCB7FDA3FCA0" box="[434,576,832,857]" italics="true" pageId="9" pageNumber="45">S. duplicata</emphasis>
|
||
</taxonomicName>
|
||
were found in pitfall traps for reptiles and amphibians in the dry, xeric “cerrado”, a vegetation typical of central
|
||
<collectingCountry id="F35C762E2F036579FD6DFC9FFD39FC7B" box="[654,730,872,898]" name="Brazil" pageId="9" pageNumber="45">Brazil</collectingCountry>
|
||
. All specimens were collected before flooding of the Luis Eduardo Magalhães hydroelectric power plant, in the Tocantins River, and the
|
||
<typeStatus id="54F0881C2F036579FBBFFC67FB6CFC53" box="[1116,1167,912,938]" pageId="9" pageNumber="45">type</typeStatus>
|
||
locality is now under water. Vegetation around the lake is the same as that at the now submerged
|
||
<typeStatus id="54F0881C2F036579FBF2FC4FFBA7FC2B" box="[1041,1092,952,978]" pageId="9" pageNumber="45">type</typeStatus>
|
||
locality. An expedition organized by the first author in
|
||
<date id="FFF5107E2F036579FE3FFC17FDB5FC03" box="[476,598,992,1018]" pageId="9" pageNumber="45" value="2007-06">June 2007</date>
|
||
failed to discover any specimens of
|
||
<taxonomicName id="4C4B4D3D2F036579FBEFFC17FB79FC00" box="[1036,1178,992,1017]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B93FEAAC2F036579FBEFFC17FB79FC00" box="[1036,1178,992,1017]" italics="true" pageId="9" pageNumber="45">S. duplicata</emphasis>
|
||
</taxonomicName>
|
||
, even though a forest patch
|
||
<quantity id="4CB39B5B2F036579FF3EFBFFFEC5FBDB" box="[221,294,1032,1058]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="45" unit="m" value="500.0">500 m</quantity>
|
||
away from the
|
||
<typeStatus id="54F0881C2F036579FE3FFBFFFDECFBDB" box="[476,527,1032,1058]" pageId="9" pageNumber="45">type</typeStatus>
|
||
locality was sampled. Thus, the original habitat of this species may have been impacted by the flooding of the hydroelectric power plant, and further expeditions are needed to seek additional individuals of this remarkable Brazilian species. Also noteworthy is the fact that a closely related species,
|
||
<taxonomicName id="4C4B4D3D2F036579FF30FB77FDB8FB63" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[211,603,1152,1178]" class="Chilopoda" family="Scolopendridae" genus="Rhoda" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="45" phylum="Arthropoda" rank="species" species="spinifer">
|
||
<emphasis id="B93FEAAC2F036579FF30FB77FE62FB60" box="[211,385,1152,1177]" italics="true" pageId="9" pageNumber="45">Rhoda spinifer</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B4F2F036579FE73FB77FDB1FB63" author="Kraepelin" box="[400,594,1152,1178]" pageId="9" pageNumber="46" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, (2) 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, occurs sympatrically on the left bank of the Tocantins River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |