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<document id="54F3ECAE8AE1AFA8CFF3A3FF8EC193B3" ID-CLB-Dataset="9346" ID-DOI="10.11646/zootaxa.4895.1.7" ID-GBIF-Dataset="0f10cb02-b704-404d-a6fe-7c82cce3a800" ID-ISSN="1175-5326" ID-Zenodo-Dep="4321565" ID-ZooBank="BF5B9C48-4F1D-4470-9D2A-E3A10193DE49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607972870813" checkinUser="plazi" docAuthor="Liu, Zhiping &amp; Cuccodoro, Giulio" docDate="2020" docId="03E246146475FF853B8FFF259F8C3D5D" docLanguage="en" docName="zootaxa.4895.1.7.pdf" docOrigin="Zootaxa 4895 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megarthrus budai Liu and Cuccodoro 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="134" masterDocId="FFDB3E6C6473FF8F3B18FFB29E383C38" masterDocTitle="Megarthrus of China. Part 2. Megarthrus basicornis Fauvel, 1904 with description of a new species from Mount Emei (Coleoptera: Staphylinidae: Proteininae)" masterLastPageNumber="134" masterPageNumber="124" pageNumber="130" updateTime="1698847542504" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FACB4FA38BC88B85AE3F724F3AF508BF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2C6DEF09D86C48F5CFA949854E014BC2">
<mods:title id="542E3795FFE41BAF23A27092CDA1F7AD">Megarthrus of China. Part 2. Megarthrus basicornis Fauvel, 1904 with description of a new species from Mount Emei (Coleoptera: Staphylinidae: Proteininae)</mods:title>
</mods:titleInfo>
<mods:name id="D0F51FB4F8D00A9263CD998C2FDBEF5D" type="personal">
<mods:role id="0BF68176086A1C7AEA279C1BA67891D7">
<mods:roleTerm id="533D36321D176DEF0929850E90D9C0BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="883457302B9B79A19B9310583AFA5255">Liu, Zhiping</mods:namePart>
<mods:affiliation id="C8486CD7273F4E18CCC02D396D89BC5E">Institute of Entomology, College of Plant Protection, Southwest University, Chongqing 400715, China.</mods:affiliation>
</mods:name>
<mods:name id="F56044181E88A3218DBFBE55185241A7" type="personal">
<mods:role id="6AC59827DF62C1CDC68061F78D86B023">
<mods:roleTerm id="3AAFBCB646C4FA1215E0B0D0597FC9C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="079EFB44093BF162DE600BD7C012C69F">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="EC1E642E52EE6DCA70C5C4364D9EFAC8">Muséum d’histoire naturelle, Case postale 6434 CH- 1211 Genève 6, Switzerland.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EABA6A9316F2616560FE2258B74A8F48">text</mods:typeOfResource>
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<mods:titleInfo id="C0F38D7376476AFC28A40F9A4EE3CCB6">
<mods:title id="C73B4D3B99C838B6742E9AFB6DEC0D54">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="24415029B50CDB1A5FCCA2774B8691E9">
<mods:date id="3EDC21D73F5B13A4EFCED1E6FA77D20C">2020</mods:date>
<mods:detail id="C716D6E3F9F3F1672211EFF8DFBF7790" type="pubDate">
<mods:number id="627E95A8BF80AEEA07913C06E7C1B2D0">2020-12-14</mods:number>
</mods:detail>
<mods:detail id="73F582E78EB237F28098C5338374F255" type="volume">
<mods:number id="420E9981B3136542C536341028509B29">4895</mods:number>
</mods:detail>
<mods:detail id="5634DAE17877ECACAEE78A8EEB01132A" type="issue">
<mods:number id="2E35B3C04697BBFEAB8D6FFE120B6B25">1</mods:number>
</mods:detail>
<mods:extent id="40F667585BB9F5A15D3AB9612ADD3AA0" unit="page">
<mods:start id="E64CD1379E850DFD2D5B00474113CB89">124</mods:start>
<mods:end id="E95ED447284E98F28544450154D5883D">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6C025BB53A7C2EF3F139CAF512A0BA8F">journal article</mods:classification>
<mods:identifier id="48ECE8D64AE6719F2E65982F1BDB0ED6" type="CLB-Dataset">9346</mods:identifier>
<mods:identifier id="51BA4731B43A9FE7076543ABE504FA1D" type="DOI">10.11646/zootaxa.4895.1.7</mods:identifier>
<mods:identifier id="9D19263E60CE5A09EE5607E24B6D16F3" type="GBIF-Dataset">0f10cb02-b704-404d-a6fe-7c82cce3a800</mods:identifier>
<mods:identifier id="989E06A0776D2E027688618489B4C88E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B9191ECBAB08453C2B4ADC861F8AF82E" type="Zenodo-Dep">4321565</mods:identifier>
<mods:identifier id="735E996C4F561495C46F13DB96688CF2" type="ZooBank">BF5B9C48-4F1D-4470-9D2A-E3A10193DE49</mods:identifier>
</mods:mods>
<treatment id="03E246146475FF853B8FFF259F8C3D5D" ID-DOI="http://doi.org/10.5281/zenodo.4324265" ID-GBIF-Taxon="174795509" ID-Zenodo-Dep="4324265" LSID="urn:lsid:plazi:treatment:03E246146475FF853B8FFF259F8C3D5D" httpUri="http://treatment.plazi.org/id/03E246146475FF853B8FFF259F8C3D5D" lastPageId="10" lastPageNumber="134" pageId="6" pageNumber="130">
<subSubSection id="C351A4896475FF893B8FFF259CE03C89" box="[151,728,151,178]" pageId="6" pageNumber="130" type="nomenclature">
<paragraph id="8BF4F7026475FF893B8FFF259CE03C89" blockId="6.[151,931,151,249]" box="[151,728,151,178]" pageId="6" pageNumber="130">
<heading id="D0BC406E6475FF893B8FFF259CE03C89" bold="true" box="[151,728,151,178]" fontSize="11" level="1" pageId="6" pageNumber="130" reason="1">
<emphasis id="B93F2B106475FF893B8FFF259CE03C89" bold="true" box="[151,728,151,178]" pageId="6" pageNumber="130">
<taxonomicName id="4C4B8C816475FF893B8FFF259C563C89" ID-CoL="8Y69R" authority="Liu and Cuccodoro" authorityName="Liu and Cuccodoro" authorityYear="2020" box="[151,622,151,178]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B106475FF893B8FFF259F4A3C8A" bold="true" box="[151,370,151,178]" italics="true" pageId="6" pageNumber="130">Megarthrus budai</emphasis>
Liu and Cuccodoro
</taxonomicName>
,
<taxonomicNameLabel id="A20C966B6475FF893963FF259CE03C89" box="[635,728,151,177]" pageId="6" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C351A4896475FF893B8FFF099D9B3CC1" pageId="6" pageNumber="130" type="description">
<paragraph id="8BF4F7026475FF893B8FFF099F6D3CED" blockId="6.[151,931,151,249]" box="[151,341,187,213]" pageId="6" pageNumber="130">
(
<figureCitation id="1370EB876475FF893B87FF099EC73CED" box="[159,255,187,213]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="6" pageNumber="130">Figs 8-9</figureCitation>
,
<figureCitation id="1370EB876475FF893A13FF099F753CED" box="[267,333,187,213]" captionStart-0="FIGURES 10-13" captionStart-1="FIGURES 14-16" captionStart-2="FIGURES 17-26" captionStart-3="FIGURES 27-32" captionStart-4="FIGURES 33-38" captionStart-5="FIGURE 39" captionStartId-0="4.[151,264,1812,1836]" captionStartId-1="5.[151,264,1768,1792]" captionStartId-2="6.[151,264,1903,1927]" captionStartId-3="7.[151,264,1943,1967]" captionStartId-4="8.[151,264,1783,1807]" captionStartId-5="9.[151,250,1121,1145]" captionTargetBox-0="[151,1436,848,1788]" captionTargetBox-1="[160,1436,194,1726]" captionTargetBox-2="[158,1431,495,1868]" captionTargetBox-3="[207,1379,190,1906]" captionTargetBox-4="[166,1430,192,1760]" captionTargetBox-5="[151,1436,345,1061]" captionTargetId-0="figure-317@4.[151,1436,848,1788]" captionTargetId-1="figure-34@5.[151,1436,181,1744]" captionTargetId-2="figure-85@6.[151,1436,489,1879]" captionTargetId-3="figure-34@7.[197,1390,181,1919]" captionTargetId-4="figure-17@8.[151,1436,181,1760]" captionTargetId-5="figure-102@9.[151,1436,345,1061]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." captionText-1="FIGURES 14-16. Megarthrus budai sp. n., habitus, male: dorsal (14), ventral (15), and lateral (16) views. Scale bar = 0.5 mm." captionText-2="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." captionText-3="FIGURES 27-32. Megarthrus budai sp. n., male: aedeagus in lateral (27, 28) and ventral (30, 31) views; hemitergite IX (29); antenna (32). Scale bar = 0.1 mm." captionText-4="FIGURES 33-38. Megarthrus budai sp. n., female: genital segments, ventral part in dorsal (33) and lateral (34) views, and dorsal part (35) in ventral view; abdominal sternite VIII in dorsal view (36); abdominal tergite VIII in dorsal (37) and lateral (38) views. Scale bar = 0.1 mm." captionText-5="FIGURE 39. Distribution map of Megarthrus basicornis Fauvel (pink squares) and Megarthrus budai sp. n. (red circle). Scale bar =200 km." figureDoi-0="http://doi.org/10.5281/zenodo.4321576" figureDoi-1="http://doi.org/10.5281/zenodo.4321578" figureDoi-2="http://doi.org/10.5281/zenodo.4321580" figureDoi-3="http://doi.org/10.5281/zenodo.4321582" figureDoi-4="http://doi.org/10.5281/zenodo.4321584" figureDoi-5="http://doi.org/10.5281/zenodo.4321586" httpUri-0="https://zenodo.org/record/4321576/files/figure.png" httpUri-1="https://zenodo.org/record/4321578/files/figure.png" httpUri-2="https://zenodo.org/record/4321580/files/figure.png" httpUri-3="https://zenodo.org/record/4321582/files/figure.png" httpUri-4="https://zenodo.org/record/4321584/files/figure.png" httpUri-5="https://zenodo.org/record/4321586/files/figure.png" pageId="6" pageNumber="130">12-39</figureCitation>
)
</paragraph>
<paragraph id="8BF4F7026475FF893B8FFF539D9B3CC1" blockId="6.[151,931,151,249]" box="[151,931,225,249]" pageId="6" pageNumber="130">
<uri id="FFDAFB006475FF893B8FFF539D9B3CC1" box="[151,931,225,249]" pageId="6" pageNumber="130">
urn:lsid:zoobank.org:act:
<uuid id="FFEDCDD76475FF893A85FF539D9B3CC1" box="[413,931,225,249]" pageId="6" pageNumber="130">836693E2-8AB5-4D0C-99A7-F61FAE36CD25</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C351A4896475FF893B8FFE959F723DE9" pageId="6" pageNumber="130" type="materials_examined">
<paragraph id="8BF4F7026475FF893B8FFE959FF33D5D" blockId="6.[151,1436,295,465]" pageId="6" pageNumber="130">
<emphasis id="B93F2B106475FF893B8FFE959F7C3D79" bold="true" box="[151,324,295,321]" pageId="6" pageNumber="130">Type material.</emphasis>
<materialsCitation id="3B23FD5F6475FF893A53FE959FF33D5D" ID-GBIF-Occurrence="2994520305" collectingDate="2011-05-28" collectionCode="MHNG" collectorName="V. Grebennikov" country="China" latitude="29.55" location="Emei Shan" longLatPrecision="20" longitude="103.36056" pageId="6" pageNumber="130" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54F049A06475FF893A53FE959F8E3D79" box="[331,438,295,321]" pageId="6" pageNumber="130" type="holotype">Holotype</typeStatus>
, male “
<collectingCountry id="F35CB7926475FF89391FFE959C683D79" box="[519,592,295,321]" name="China" pageId="6" pageNumber="130">China</collectingCountry>
:
<collectingRegion id="498F39E06475FF893942FE959C8F3D79" box="[602,695,295,321]" country="China" name="Sichuan" pageId="6" pageNumber="130">Sichuan</collectingRegion>
,
<location id="8E94A1D96475FF8939DAFE959D063D79" LSID="urn:lsid:plazi:treatment:03E246146475FF853B8FFF259F8C3D5D:8E94A1D96475FF8939DAFE959D063D79" box="[706,830,295,321]" country="China" latitude="29.55" longLatPrecision="20" longitude="103.36056" name="Emei Shan" pageId="6" pageNumber="130" stateProvince="Sichuan">Emei Shan</location>
,
<geoCoordinate id="EE7F91C56475FF893851FE959DEC3D79" box="[841,980,295,321]" degrees="29" direction="north" minutes="33" orientation="latitude" pageId="6" pageNumber="130" precision="15" seconds="00" value="29.55">N29°33’00’’</geoCoordinate>
<geoCoordinate id="EE7F91C56475FF8938C2FE959A4D3D79" box="[986,1141,295,321]" degrees="103" direction="east" minutes="21" orientation="longitude" pageId="6" pageNumber="130" precision="15" seconds="38" value="103.36056">E103°21’38’’</geoCoordinate>
,
<date id="FFF5D1C26475FF893F98FE959AD73D79" box="[1152,1263,295,321]" pageId="6" pageNumber="130" value="2011-05-28">
<collectingDate id="EFB1282A6475FF893F98FE959AD73D79" box="[1152,1263,295,321]" pageId="6" pageNumber="130" value="2011-05-28">28.v.2011</collectingDate>
</date>
, 1639m, sift08,
<collectorName id="26BE92D46475FF893B8FFEFE9F703D5D" box="[151,328,331,358]" pageId="6" pageNumber="130">V. Grebennikov</collectorName>
”(
<collectionCode id="ED5A6FC76475FF893A79FEF99F873D5D" box="[353,447,331,357]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="130" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF4F7026475FF893BDFFEC29F723DE9" blockId="6.[151,1436,295,465]" pageId="6" pageNumber="130">
<materialsCitation id="3B23FD5F6475FF893BDFFEC29D2D3DB1" ID-GBIF-Occurrence="2994520301" box="[199,789,367,394]" collectingDate="2011-05-28" collectionCode="SWUC" collectorName="V. Grebennikov" country="China" latitude="29.55" location="Emei Shan" longLatPrecision="20" longitude="103.36056" pageId="6" pageNumber="130" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54F049A06475FF893BDFFEC29F003DB2" box="[199,312,368,394]" pageId="6" pageNumber="130" type="paratype">Paratypes</typeStatus>
: same data as holotype,
<specimenCount id="9D4D3C8B6475FF893948FEDD9CA43DB1" box="[592,668,367,393]" pageId="6" pageNumber="130" type="male">1 male</specimenCount>
in
<collectionCode id="ED5A6FC76475FF8939DCFEDD9D2D3DB1" box="[708,789,367,393]" pageId="6" pageNumber="130">SWUC</collectionCode>
</materialsCitation>
<materialsCitation id="3B23FD5F6475FF893806FEDD9A7C3DB1" ID-GBIF-Occurrence="2994520303" box="[798,1092,367,393]" collectionCode="MHNG" country="China" pageId="6" pageNumber="130" specimenCount="3" specimenCount-female="3" stateProvince="Sichuan" typeStatus="paratype">
and
<specimenCount id="9D4D3C8B6475FF893857FEDD9D873DB1" box="[847,959,367,393]" pageId="6" pageNumber="130" type="female">3 females</specimenCount>
, in
<collectionCode id="ED5A6FC76475FF8938F3FEDD9A7C3DB1" box="[1003,1092,367,393]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="130" type="Museum">MHNG</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B23FD5F6475FF893F49FEDD9B083D95" ID-GBIF-Occurrence="2994520306" collectingDate="2011-05-25" collectionCode="MHNG" collectorName="V. Grebennikov" country="China" latitude="29.551111" location="Emei Shan" longLatPrecision="20" longitude="103.35528" pageId="6" pageNumber="130" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
“
<collectingCountry id="F35CB7926475FF893F43FEDD9A9B3DB1" box="[1115,1187,367,393]" name="China" pageId="6" pageNumber="130">China</collectingCountry>
,
<collectingRegion id="498F39E06475FF893FA8FEDD9B353DB1" box="[1200,1293,367,393]" country="China" name="Sichuan" pageId="6" pageNumber="130">Sichuan</collectingRegion>
,
<location id="8E94A1D96475FF893E01FEDD9BA03DB1" LSID="urn:lsid:plazi:treatment:03E246146475FF853B8FFF259F8C3D5D:8E94A1D96475FF893E01FEDD9BA03DB1" box="[1305,1432,367,393]" country="China" latitude="29.551111" longLatPrecision="20" longitude="103.35528" name="Emei Shan" pageId="6" pageNumber="130" stateProvince="Sichuan">Emei Shan</location>
,
<geoCoordinate id="EE7F91C56475FF893B8FFE219F1A3D95" box="[151,290,403,429]" degrees="29" direction="north" minutes="33" orientation="latitude" pageId="6" pageNumber="130" precision="15" seconds="04" value="29.551111">N29°33’04’’</geoCoordinate>
<geoCoordinate id="EE7F91C56475FF893A32FE219FFC3D95" box="[298,452,403,429]" degrees="103" direction="east" minutes="21" orientation="longitude" pageId="6" pageNumber="130" precision="15" seconds="19" value="103.35528">E103°21’19’’</geoCoordinate>
,
<date id="FFF5D1C26475FF893AC9FE219C783D96" box="[465,576,403,430]" pageId="6" pageNumber="130" value="2011-05-25">
<collectingDate id="EFB1282A6475FF893AC9FE219C783D96" box="[465,576,403,430]" pageId="6" pageNumber="130" value="2011-05-25">25.v.2011</collectingDate>
</date>
, 1729m, sift05,
<collectorName id="26BE92D46475FF8939EFFE269D973D95" box="[759,943,403,430]" pageId="6" pageNumber="130">V. Grebennikov</collectorName>
”,
<specimenCount id="9D4D3C8B6475FF8938DCFE219A283D95" box="[964,1040,403,429]" pageId="6" pageNumber="130" type="male">1 male</specimenCount>
and
<specimenCount id="9D4D3C8B6475FF893F50FE219A8D3D95" box="[1096,1205,403,430]" pageId="6" pageNumber="130" type="female">5 females</specimenCount>
in
<collectionCode id="ED5A6FC76475FF893FC2FE219B083D95" box="[1242,1328,403,429]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="130" type="Museum">MHNG</collectionCode>
</materialsCitation>
<materialsCitation id="3B23FD5F6475FF893E2FFE219F7E3DE9" ID-GBIF-Occurrence="2994520302" collectionCode="SWUC" country="China" pageId="6" pageNumber="130" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="paratype">
and
<specimenCount id="9D4D3C8B6475FF893E7FFE219EF53DE9" pageId="6" pageNumber="130" type="female">1 female</specimenCount>
in
<collectionCode id="ED5A6FC76475FF893BE9FE059F7E3DE9" box="[241,326,439,465]" pageId="6" pageNumber="130">SWUC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF34A78A6475FF893B8FF8DD9CBD3BCA" ID-DOI="http://doi.org/10.5281/zenodo.4321580" ID-Zenodo-Dep="4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="6" pageNumber="130" startId="6.[151,264,1903,1927]" targetBox="[158,1431,495,1868]" targetPageId="6">
<paragraph id="8BF4F7026475FF893B8FF8DD9CBD3BCA" blockId="6.[151,1437,1903,2035]" pageId="6" pageNumber="130">
<emphasis id="B93F2B106475FF893B8FF8DD9F6A3BBF" bold="true" box="[151,338,1903,1927]" pageId="6" pageNumber="130">FIGURES 17-26.</emphasis>
<taxonomicName id="4C4B8C816475FF893A43F8DD9C2D3BBF" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[347,533,1903,1927]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B106475FF893A43F8DD9C2D3BBF" box="[347,533,1903,1927]" italics="true" pageId="6" pageNumber="130">Megarthrus budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B6475FF893905F8DD9C6F3BBE" box="[541,599,1903,1926]" pageId="6" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
, male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm.
</paragraph>
</caption>
<caption id="DF34A78A6474FF883B8FF8259FC83BEB" ID-DOI="http://doi.org/10.5281/zenodo.4321582" ID-Zenodo-Dep="4321582" httpUri="https://zenodo.org/record/4321582/files/figure.png" pageId="7" pageNumber="131" startId="7.[151,264,1943,1967]" targetBox="[207,1379,190,1906]" targetPageId="7">
<paragraph id="8BF4F7026474FF883B8FF8259FC83BEB" blockId="7.[151,1436,1943,2004]" pageId="7" pageNumber="131">
<emphasis id="B93F2B106474FF883B8FF8259F683B97" bold="true" box="[151,336,1943,1967]" pageId="7" pageNumber="131">FIGURES 27-32.</emphasis>
<taxonomicName id="4C4B8C816474FF883A4FF82A9C283B88" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[343,528,1944,1968]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B106474FF883A4FF82A9C283B88" box="[343,528,1944,1968]" italics="true" pageId="7" pageNumber="131">Megarthrus budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B6474FF88390EF82A9C773B97" box="[534,591,1944,1967]" pageId="7" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
, male: aedeagus in lateral (27, 28) and ventral (30, 31) views; hemitergite IX (29); antenna (32). Scale bar = 0.1 mm.
</paragraph>
</caption>
<caption id="DF34A78A647BFF873B8FF9459FE53B6F" ID-DOI="http://doi.org/10.5281/zenodo.4321584" ID-Zenodo-Dep="4321584" httpUri="https://zenodo.org/record/4321584/files/figure.png" pageId="8" pageNumber="132" startId="8.[151,264,1783,1807]" targetBox="[166,1430,192,1760]" targetPageId="8">
<paragraph id="8BF4F702647BFF873B8FF9459FE53B6F" blockId="8.[151,1437,1783,1880]" pageId="8" pageNumber="132">
<emphasis id="B93F2B10647BFF873B8FF9459F6A3B37" bold="true" box="[151,338,1783,1807]" pageId="8" pageNumber="132">FIGURES 33-38.</emphasis>
<taxonomicName id="4C4B8C81647BFF873A42F94A9C2C3B28" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[346,532,1784,1808]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="132" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B10647BFF873A42F94A9C2C3B28" box="[346,532,1784,1808]" italics="true" pageId="8" pageNumber="132">Megarthrus budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B647BFF873904F94A9C6E3B37" box="[540,598,1784,1807]" pageId="8" pageNumber="132" rank="species">sp. n.</taxonomicNameLabel>
, female: genital segments, ventral part in dorsal (33) and lateral (34) views, and dorsal part (35) in ventral view; abdominal sternite VIII in dorsal view (36); abdominal tergite VIII in dorsal (37) and lateral (38) views. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C351A489647BFF853BDFF8379F8C3D5D" lastPageId="10" lastPageNumber="134" pageId="8" pageNumber="132" type="description">
<paragraph id="8BF4F702647BFF863BDFF8379F5B3D79" blockId="8.[151,1436,1925,2024]" lastBlockId="9.[151,1437,151,321]" lastPageId="9" lastPageNumber="133" pageId="8" pageNumber="132">
<emphasis id="B93F2B10647BFF873BDFF8379F613BA7" bold="true" box="[199,345,1925,1951]" pageId="8" pageNumber="132">Description.</emphasis>
Habitus as in
<figureCitation id="1370EB87647BFF873AE5F8349C423B98" box="[509,634,1926,1952]" captionStart="FIGURES 14-16" captionStartId="5.[151,264,1768,1792]" captionTargetBox="[160,1436,194,1726]" captionTargetId="figure-34@5.[151,1436,181,1744]" captionTargetPageId="5" captionText="FIGURES 14-16. Megarthrus budai sp. n., habitus, male: dorsal (14), ventral (15), and lateral (16) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4321578" httpUri="https://zenodo.org/record/4321578/files/figure.png" pageId="8" pageNumber="132">Figs 14-16</figureCitation>
. Combined length of head, pronotum, and elytra
<quantity id="4CB35AE7647BFF873FBFF8349B163B98" box="[1191,1326,1926,1952]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="8" pageNumber="132" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7-1.9 mm</quantity>
; maximal pronotal width
<quantity id="4CB35AE7647BFF873A5CF8189FF23BFC" box="[324,458,1962,1988]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="132" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9-1.1 mm</quantity>
. Body predominantly chestnut brown, with head darker and appendages slightly paler, and lateral portions of pronotum markedly paler. Dorsal pubescence fairly uniform, but denser on basal portion of elytra; pubescence on frons converging, with medial setae directed backward; metaventral setae shorter than proventral setae, becoming longer anteriorly; pubescence on abdominal tergites IV-VII parallel; sternites IV-VII with few longer setae posteromedially, but apparently lacking macrosetae. Frons and vertex finely granulate; pronotum and elytra granulo-fossulate; prohypomeron almost smooth (
<figureCitation id="1370EB87647AFF863820FEB19D433D25" box="[824,891,259,285]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="9" pageNumber="133">Fig. 9</figureCitation>
); metaventrite coarsely punctate laterally, almost smooth in middle.
</paragraph>
<caption id="DF34A78A647AFF863B8FFBD39F1838A5" ID-DOI="http://doi.org/10.5281/zenodo.4321586" ID-Zenodo-Dep="4321586" httpUri="https://zenodo.org/record/4321586/files/figure.png" pageId="9" pageNumber="133" startId="9.[151,250,1121,1145]" targetBox="[151,1436,345,1061]" targetPageId="9">
<paragraph id="8BF4F702647AFF863B8FFBD39F1838A5" blockId="9.[151,1436,1121,1181]" pageId="9" pageNumber="133">
<emphasis id="B93F2B10647AFF863B8FFBD39F213841" bold="true" box="[151,281,1121,1145]" pageId="9" pageNumber="133">FIGURE 39</emphasis>
. Distribution map of
<taxonomicName id="4C4B8C81647AFF863AECFBD39D1F3841" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" box="[500,807,1121,1145]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="133" phylum="Arthropoda" rank="species" species="basicornis">
<emphasis id="B93F2B10647AFF863AECFBD39CE43841" box="[500,732,1121,1145]" italics="true" pageId="9" pageNumber="133">Megarthrus basicornis</emphasis>
Fauvel
</taxonomicName>
(pink squares) and
<taxonomicName id="4C4B8C81647AFF8638F7FBD39A9E3841" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[1007,1190,1121,1145]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="133" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B10647AFF8638F7FBD39A9E3841" box="[1007,1190,1121,1145]" italics="true" pageId="9" pageNumber="133">Megarthrus budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B647AFF863FB4FBD39ADB3840" box="[1196,1251,1121,1144]" pageId="9" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
(red circle). Scale bar =200 km.
</paragraph>
</caption>
<paragraph id="8BF4F702647AFF863BDFFB749FFC39AC" blockId="9.[151,1437,1222,2040]" pageId="9" pageNumber="133">
Frons above clypeus forming sharp ridge, the latter very finely carinate; mesal portion of disc weakly convex in lateral view, evenly; U-shaped frontal impression shallow. Temples slightly expanded just behind eyes, then abruptly narrowed, almost smooth. Occipital ridge indistinct. Antenna (
<figureCitation id="1370EB87647AFF863871FABC9D823910" box="[873,954,1294,1320]" captionStart="FIGURES 27-32" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[207,1379,190,1906]" captionTargetId="figure-34@7.[197,1390,181,1919]" captionTargetPageId="7" captionText="FIGURES 27-32. Megarthrus budai sp. n., male: aedeagus in lateral (27, 28) and ventral (30, 31) views; hemitergite IX (29); antenna (32). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321582" httpUri="https://zenodo.org/record/4321582/files/figure.png" pageId="9" pageNumber="133">Fig. 32</figureCitation>
) with scape fairly conical, not compressed; short and dense pubescence present only on antennomeres 5-11; antennomere 11 rather ovoid, slightly compressed. Pronotum as in
<figureCitation id="1370EB87647AFF863A5FFAE49FB33948" box="[327,395,1366,1392]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="9" pageNumber="133">Fig. 8</figureCitation>
; hypomera lacking transverse ridge (
<figureCitation id="1370EB87647AFF863830FAE49D553948" box="[808,877,1366,1392]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="9" pageNumber="133">Fig. 9</figureCitation>
). Scutellum with anterior margin rounded. Elytral disc without notable relief.
</paragraph>
<paragraph id="8BF4F702647AFF863BDFFA2C9ED83AA8" blockId="9.[151,1437,1222,2040]" pageId="9" pageNumber="133">
<emphasis id="B93F2B10647AFF863BDFFA2C9F393980" box="[199,257,1438,1464]" italics="true" pageId="9" pageNumber="133">Male</emphasis>
. Frontoclypeal area, protarsomere 5, metaventrite, and abdominal sternites IV-VI unmodified. Protarsomere 1 possessing ventral patch of modified adhesive setae (
<figureCitation id="1370EB87647AFF863829FA709D9539E4" box="[817,941,1474,1500]" captionStart="FIGURES 10-13" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[151,1436,848,1788]" captionTargetId="figure-317@4.[151,1436,848,1788]" captionTargetPageId="4" captionText="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321576" httpUri="https://zenodo.org/record/4321576/files/figure.png" pageId="9" pageNumber="133">Figs 12-13</figureCitation>
). Metatarsomere 1 about 1.3 times as long as combined length of metatarsomeres 2-4. Peg-like setae absent from protrochanter, metatrochanter, femur, and protibia; arranged in a row on mesotrochanter (
<figureCitation id="1370EB87647AFF86399EF9B89CE23A1C" box="[646,730,1546,1572]" captionStart="FIGURES 17-26" captionStartId="6.[151,264,1903,1927]" captionTargetBox="[158,1431,495,1868]" captionTargetId="figure-85@6.[151,1436,489,1879]" captionTargetPageId="6" captionText="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="9" pageNumber="133">Fig. 19</figureCitation>
); grouped in a field on mesotibiae (
<figureCitation id="1370EB87647AFF863F74F9B89AD03A1C" box="[1132,1256,1546,1572]" captionStart="FIGURES 17-26" captionStartId="6.[151,264,1903,1927]" captionTargetBox="[158,1431,495,1868]" captionTargetId="figure-85@6.[151,1436,489,1879]" captionTargetPageId="6" captionText="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="9" pageNumber="133">Figs 23-24</figureCitation>
) and metatibiae (
<figureCitation id="1370EB87647AFF863B87F99C9F253A71" box="[159,285,1582,1609]" captionStart="FIGURES 17-26" captionStartId="6.[151,264,1903,1927]" captionTargetBox="[158,1431,495,1868]" captionTargetId="figure-85@6.[151,1436,489,1879]" captionTargetPageId="6" captionText="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="9" pageNumber="133">Figs 25-26</figureCitation>
). Metaventrite markedly swollen in middle. Abdominal tergite VIII as in
<figureCitation id="1370EB87647AFF863F56F99C9AF53A70" box="[1102,1229,1582,1608]" captionStart="FIGURES 17-26" captionStartId="6.[151,264,1903,1927]" captionTargetBox="[158,1431,495,1868]" captionTargetId="figure-85@6.[151,1436,489,1879]" captionTargetPageId="6" captionText="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="9" pageNumber="133">Figs 20, 22</figureCitation>
; sternite VIII as in
<figureCitation id="1370EB87647AFF863B8FF9E09ED13A54" box="[151,233,1618,1644]" captionStart="FIGURES 17-26" captionStartId="6.[151,264,1903,1927]" captionTargetBox="[158,1431,495,1868]" captionTargetId="figure-85@6.[151,1436,489,1879]" captionTargetPageId="6" captionText="FIGURES 17-26. Megarthrus budai sp. n., male: abdominal sternite VIII in ventral view (17); protrochanter and profemur in lateral view (18); mesotrochanter and mesofemur in lateral view (19); metatrochanter and metafemur in lateral view (21); abdominal tergite VIII in lateral (20) and dorsal (22) views. mesotibia in lateral (23) and ventral (24) views; metatibia in lateral (25) and ventral (26) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321580" httpUri="https://zenodo.org/record/4321580/files/figure.png" pageId="9" pageNumber="133">Fig. 17</figureCitation>
; hemitergites IX as in
<figureCitation id="1370EB87647AFF863AFAF9E09C0B3A54" box="[482,563,1618,1644]" captionStart="FIGURES 27-32" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[207,1379,190,1906]" captionTargetId="figure-34@7.[197,1390,181,1919]" captionTargetPageId="7" captionText="FIGURES 27-32. Megarthrus budai sp. n., male: aedeagus in lateral (27, 28) and ventral (30, 31) views; hemitergite IX (29); antenna (32). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321582" httpUri="https://zenodo.org/record/4321582/files/figure.png" pageId="9" pageNumber="133">Fig. 29</figureCitation>
; sternite IX without subbasal medial protuberance. Aedeagus as in
<figureCitation id="1370EB87647AFF863E05F9E09EE43AA8" captionStart="FIGURES 27-32" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[207,1379,190,1906]" captionTargetId="figure-34@7.[197,1390,181,1919]" captionTargetPageId="7" captionText="FIGURES 27-32. Megarthrus budai sp. n., male: aedeagus in lateral (27, 28) and ventral (30, 31) views; hemitergite IX (29); antenna (32). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321582" httpUri="https://zenodo.org/record/4321582/files/figure.png" pageId="9" pageNumber="133">Figs 27-28, 30-31</figureCitation>
.
</paragraph>
<paragraph id="8BF4F702647AFF863BDFF9289FF83AE1" blockId="9.[151,1437,1222,2040]" pageId="9" pageNumber="133">
<emphasis id="B93F2B10647AFF863BDFF9289F233A8C" box="[199,283,1690,1716]" italics="true" pageId="9" pageNumber="133">Female</emphasis>
. Abdominal tergite VIII (
<figureCitation id="1370EB87647AFF86395FF9289CFD3A8C" box="[583,709,1690,1716]" captionStart="FIGURES 33-38" captionStartId="8.[151,264,1783,1807]" captionTargetBox="[166,1430,192,1760]" captionTargetId="figure-17@8.[151,1436,181,1760]" captionTargetPageId="8" captionText="FIGURES 33-38. Megarthrus budai sp. n., female: genital segments, ventral part in dorsal (33) and lateral (34) views, and dorsal part (35) in ventral view; abdominal sternite VIII in dorsal view (36); abdominal tergite VIII in dorsal (37) and lateral (38) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321584" httpUri="https://zenodo.org/record/4321584/files/figure.png" pageId="9" pageNumber="133">Figs 37-38</figureCitation>
) forming a medioapical projection; sternite VIII as in
<figureCitation id="1370EB87647AFF863E5BF9289BA03A8C" box="[1347,1432,1690,1716]" captionStart="FIGURES 33-38" captionStartId="8.[151,264,1783,1807]" captionTargetBox="[166,1430,192,1760]" captionTargetId="figure-17@8.[151,1436,181,1760]" captionTargetPageId="8" captionText="FIGURES 33-38. Megarthrus budai sp. n., female: genital segments, ventral part in dorsal (33) and lateral (34) views, and dorsal part (35) in ventral view; abdominal sternite VIII in dorsal view (36); abdominal tergite VIII in dorsal (37) and lateral (38) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321584" httpUri="https://zenodo.org/record/4321584/files/figure.png" pageId="9" pageNumber="133">Fig. 36</figureCitation>
. Genitalia as in
<figureCitation id="1370EB87647AFF863A58F90C9F843AE1" box="[320,444,1726,1753]" captionStart="FIGURES 33-38" captionStartId="8.[151,264,1783,1807]" captionTargetBox="[166,1430,192,1760]" captionTargetId="figure-17@8.[151,1436,181,1760]" captionTargetPageId="8" captionText="FIGURES 33-38. Megarthrus budai sp. n., female: genital segments, ventral part in dorsal (33) and lateral (34) views, and dorsal part (35) in ventral view; abdominal sternite VIII in dorsal view (36); abdominal tergite VIII in dorsal (37) and lateral (38) views. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321584" httpUri="https://zenodo.org/record/4321584/files/figure.png" pageId="9" pageNumber="133">Figs 33-35</figureCitation>
.
</paragraph>
<paragraph id="8BF4F702647AFF863BDFF9539F7D3BC0" blockId="9.[151,1437,1222,2040]" pageId="9" pageNumber="133">
<emphasis id="B93F2B10647AFF863BDFF9539C5E3AC4" bold="true" box="[199,614,1761,1788]" pageId="9" pageNumber="133">Comparisons and diagnostic notes.</emphasis>
In the Palaearctic and Oriental regions
<taxonomicName id="4C4B8C81647AFF863F39F9509AD43AC4" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[1057,1260,1762,1788]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="133" phylum="Arthropoda" rank="species" species="budai">
<emphasis id="B93F2B10647AFF863F39F9509AD43AC4" box="[1057,1260,1762,1788]" italics="true" pageId="9" pageNumber="133">Megarthrus budai</emphasis>
</taxonomicName>
is easily distinguished from its congeners by its deplanate pronotum in combination with the absence of a hypomeral ridge, the presence of short and dense pubescence only on the antennomeres 5-11, and the scape fairly conical and not compressed. Its sexual characters are also diagnostic. The new species possesses a ventral patch of modified adhesive setae on the male protarsomere (
<figureCitation id="1370EB87647AFF863910F8C09CBE3BB4" box="[520,646,1906,1932]" captionStart="FIGURES 10-13" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[151,1436,848,1788]" captionTargetId="figure-317@4.[151,1436,848,1788]" captionTargetPageId="4" captionText="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321576" httpUri="https://zenodo.org/record/4321576/files/figure.png" pageId="9" pageNumber="133">Figs 12-13</figureCitation>
), a feature uniquely shared in the genus by all the members of the
<taxonomicName id="4C4B8C81647AFF863E66F8C19F3C3B88" baseAuthorityName="Paykull" baseAuthorityYear="1789" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="133" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B93F2B10647AFF863E66F8C19F3C3B88" italics="true" pageId="9" pageNumber="133">M. depressus</emphasis>
</taxonomicName>
supergroup (
<bibRefCitation id="EFDA8AF3647AFF863A80F8249C603B88" author="Cuccodoro, G." box="[408,600,1942,1968]" pageId="9" pageNumber="133" pagination="25 - 92" refId="ref2792" refString="Cuccodoro, G. (2011) Megarthrus of Taiwan, with notes on phylogenetic relationships within the genus (Coleoptera: Staphylinidae: Proteininae). Studies and reports, Taxonomical Series, 7 (1 - 2), 25 - 92." type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
). However, neither the shape of its aedeagus nor the male modifications of the legs resemble any of the species currently included in this main lineage of
<taxonomicName id="4C4B8C81647AFF863F02F8089AA63BEC" baseAuthorityName="Cuccodoro" baseAuthorityYear="2011" box="[1050,1182,1978,2004]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B93F2B10647AFF863F02F8089AA63BEC" box="[1050,1182,1978,2004]" italics="true" pageId="9" pageNumber="133">Megarthrus</emphasis>
</taxonomicName>
, and thus its affinities remain unclear.
</paragraph>
<paragraph id="8BF4F7026479FF853BDFFF259EF43CC1" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="134">
Distribution and natural history.
<taxonomicName id="4C4B8C816479FF85392EFF259D3A3C89" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[566,770,151,177]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="134" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B106479FF85392EFF259D3A3C89" box="[566,770,151,177]" italics="true" pageId="10" pageNumber="134">Megarthrus budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B6479FF853812FF259D593C89" box="[778,865,151,177]" pageId="10" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to Mount Emei of
<collectingRegion id="498F39E06479FF853FA3FF259B473C89" box="[1211,1407,151,177]" country="China" name="Sichuan" pageId="10" pageNumber="134">Sichuan province</collectingRegion>
in
<collectingCountry id="F35CB7926479FF853B8FFF099EE23CED" box="[151,218,187,213]" name="China" pageId="10" pageNumber="134">China</collectingCountry>
(
<figureCitation id="1370EB876479FF853BF1FF099F053CED" box="[233,317,187,213]" captionStart="FIGURE 39" captionStartId="9.[151,250,1121,1145]" captionTargetBox="[151,1436,345,1061]" captionTargetId="figure-102@9.[151,1436,345,1061]" captionTargetPageId="9" captionText="FIGURE 39. Distribution map of Megarthrus basicornis Fauvel (pink squares) and Megarthrus budai sp. n. (red circle). Scale bar =200 km." figureDoi="http://doi.org/10.5281/zenodo.4321586" httpUri="https://zenodo.org/record/4321586/files/figure.png" pageId="10" pageNumber="134">Fig. 39</figureCitation>
), where it was collected from sifted litter, at elevations ranging from approximately
<quantity id="4CB35AE76479FF853FF4FF099BA43CED" box="[1260,1436,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.75" metricValueMin="1.6" pageId="10" pageNumber="134" unit="m" value="1675.0" valueMax="1750.0" valueMin="1600.0">1600 to 1750 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BF4F7026479FF853BDFFEB19F8C3D5D" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="134">Etymology. The new species is named after the tenth-century Buddhist monk Budai, famous for his representations with a laughing face and round stomach, with reference to the swollen metaventrite of the males of this species. Name in apposition.</paragraph>
</subSubSection>
</treatment>
</document>
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