196 lines
39 KiB
XML
196 lines
39 KiB
XML
<document id="2552F0198B887DA44175931001D233FD" ID-CLB-Dataset="8324" ID-DOI="10.11646/zootaxa.4920.4.3" ID-GBIF-Dataset="3f6c02fc-66e9-4ad6-a759-ef23da595c48" ID-ISSN="1175-5326" ID-Zenodo-Dep="4491181" ID-ZooBank="22E8AB54-654D-4882-9084-41F0D229D010" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1612249726991" checkinUser="plazi" docAuthor="Ruschel, Tatiana P. & Sanborn, Allen F." docDate="2021" docId="03D787C8EC46FFC3FFE05EC0FDA431E5" docLanguage="en" docName="zootaxa.4920.4.3.pdf" docOrigin="Zootaxa 4920 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhaeboepelis Ruschel & Sanborn 2021, n. gen." docType="treatment" docVersion="8" lastPageNumber="522" masterDocId="FFEEFFB0EC4AFFCEFF775816FFDA377F" masterDocTitle="Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)" masterLastPageNumber="527" masterPageNumber="509" pageNumber="521" updateTime="1698926442746" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F5A46C7162CACDA57EACACBA66094D5D">
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<mods:title id="976B6F25DF452A07DBCCA43EB88EDE96">Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)</mods:title>
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<mods:name id="C8093A8B80370DE67202BE53DE986FAF" type="personal">
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<mods:roleTerm id="A50E0915408163C2D4ED5B3011A33BC3">Author</mods:roleTerm>
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<mods:namePart id="1CD0F191D9C9BDAB2730FE5FB7FA23AA">Ruschel, Tatiana P.</mods:namePart>
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<mods:nameIdentifier id="AABC2207503BCFFC7F480B822E7649C9" type="ORCID">0000-0002-9052-1760</mods:nameIdentifier>
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<mods:affiliation id="1025FA43EE7278EA48A0B08C74763606">Plazi, Rio Grande do Sul, Porto Alegre, RS, Brazil. urn: lsid: zoobank. org: author: 9 D 376223 - 5637 - 4376 - 9 CBC- 9 F 98441455 A 6 tatiana. ruschel @ gmail. com; https: // orcid. org / 0000 - 0002 - 9052 - 1760</mods:affiliation>
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<mods:nameIdentifier id="6921A23D1C8A2B295E4D65C84A32EE7F" type="email">tatiana.ruschel@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6CA0A29D452637905DF7E88A5B83935E">Sanborn, Allen F.</mods:namePart>
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<mods:affiliation id="24B3F56A03F9338DD93DC328CCF979A0">Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161 - 6695, USA. urn: lsid: zoobank. org: author: 9544 FF 3 F- 9 B 1 F- 457 A- 90 A 2 - 8 A 1 C 0 C 1 F 22 A 0</mods:affiliation>
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<mods:title id="5C538B0B4D7A0507CA6EC6245E1B501D">Zootaxa</mods:title>
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<mods:part id="F25521EC7D407250AEEFD4C95F51BBA2">
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<mods:date id="B4C0B27F51BDC7C1FF39BD80F6F10F04">2021</mods:date>
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<mods:number id="C25815CD3E15EDF6FB3E3EBBA694305C">2021-02-02</mods:number>
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<mods:number id="8CDB6840B0289E5971A4786A3BC6CEF8">4920</mods:number>
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<mods:identifier id="35B8F528CD417E779A50A258265C5409" type="DOI">10.11646/zootaxa.4920.4.3</mods:identifier>
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<treatment id="03D787C8EC46FFC3FFE05EC0FDA431E5" ID-DOI="http://doi.org/10.5281/zenodo.4491194" ID-GBIF-Taxon="177317072" ID-Zenodo-Dep="4491194" LSID="urn:lsid:plazi:treatment:03D787C8EC46FFC3FFE05EC0FDA431E5" httpUri="http://treatment.plazi.org/id/03D787C8EC46FFC3FFE05EC0FDA431E5" lastPageId="13" lastPageNumber="522" pageId="12" pageNumber="521">
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<subSubSection id="C3646555EC46FFC2FFE05EC0FE48318E" box="[151,402,1750,1777]" pageId="12" pageNumber="521" type="nomenclature">
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<paragraph id="8BC136DEEC46FFC2FFE05EC0FE48318E" blockId="12.[151,402,1750,1813]" box="[151,402,1750,1777]" pageId="12" pageNumber="521">
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<heading id="D08981B2EC46FFC2FFE05EC0FE48318E" bold="true" box="[151,402,1750,1777]" fontSize="11" level="1" pageId="12" pageNumber="521" reason="1">
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<emphasis id="B90AEACCEC46FFC2FFE05EC0FE48318E" bold="true" box="[151,402,1750,1777]" pageId="12" pageNumber="521">
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<taxonomicName id="4C7E4D5DEC46FFC2FFE05EC0FEE2318E" ID-CoL="8KDYS" authority="Ruschel & Sanborn, 2021" authorityName="Ruschel & Sanborn" authorityYear="2021" box="[151,312,1750,1777]" class="Insecta" family="Cicadidae" genus="Rhaeboepelis" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90AEACCEC46FFC2FFE05EC0FEE2318E" bold="true" box="[151,312,1750,1777]" italics="true" pageId="12" pageNumber="521">Rhaeboepelis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957B7EC46FFC2FE375EC1FE48318E" box="[320,402,1751,1777]" pageId="12" pageNumber="521" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC46FFC2FFE05EECFEC8306A" box="[151,274,1786,1813]" pageId="12" pageNumber="521" type="description">
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<paragraph id="8BC136DEEC46FFC2FFE05EECFEC8306A" blockId="12.[151,402,1750,1813]" box="[151,274,1786,1813]" pageId="12" pageNumber="521">
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(
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<figureCitation id="13452A5BEC46FFC2FFE85EECFED0306A" box="[159,266,1786,1813]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="14.[151,250,1507,1531]" captionStartId-1="15.[151,250,1636,1660]" captionTargetBox-0="[151,1435,181,1483]" captionTargetBox-1="[151,1436,186,1612]" captionTargetId-0="figure-23@14.[151,1435,181,1483]" captionTargetId-1="figure-19@15.[151,1436,181,1612]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." captionText-1="FIGURE 9. Rhaeboepelis takiyaae n. gen., n. sp., male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in latero-ventral view; D, phallus in left view; E, phallus in right view.Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4491204" figureDoi-1="http://doi.org/10.5281/zenodo.4491206" httpUri-0="https://zenodo.org/record/4491204/files/figure.png" httpUri-1="https://zenodo.org/record/4491206/files/figure.png" pageId="12" pageNumber="521">Figs. 8–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC46FFC2FFE05F29FD463029" box="[151,668,1855,1879]" pageId="12" pageNumber="521" type="type_taxon">
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<paragraph id="8BC136DEEC46FFC2FFE05F29FD463029" blockId="12.[151,668,1855,1879]" box="[151,668,1855,1879]" pageId="12" pageNumber="521">
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<emphasis id="B90AEACCEC46FFC2FFE05F29FEF93028" bold="true" box="[151,291,1855,1879]" pageId="12" pageNumber="521">
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<typeStatus id="54C5887CEC46FFC2FFE05F29FF163028" box="[151,204,1855,1879]" pageId="12" pageNumber="521">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C7E4D5DEC46FFC2FE5D5F29FDD53028" ID-CoL="8KLB6" authority="Ruschel & Sanborn, 2021" authorityName="Ruschel & Sanborn" authorityYear="2021" box="[298,527,1855,1879]" class="Insecta" family="Cicadidae" genus="Rhaeboepelis" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="521" phylum="Arthropoda" rank="species" species="takiyaae" status="n. gen., n. sp.">
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<emphasis id="B90AEACCEC46FFC2FE5D5F29FDD53028" box="[298,527,1855,1879]" italics="true" pageId="12" pageNumber="521">Rhaeboepelis takiyaae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957B7EC46FFC2FD625F29FD463029" box="[533,668,1855,1879]" pageId="12" pageNumber="521" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC46FFC2FFE05F93FCD330E0" box="[151,777,1925,1952]" pageId="12" pageNumber="521" type="taxon_list">
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||
<paragraph id="8BC136DEEC46FFC2FFE05F93FCD330E0" blockId="12.[151,1436,1925,2023]" box="[151,777,1925,1952]" pageId="12" pageNumber="521">
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<emphasis id="B90AEACCEC46FFC2FFE05F93FE8630E0" bold="true" box="[151,348,1925,1951]" pageId="12" pageNumber="521">Species included</emphasis>
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.
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<taxonomicName id="4C7E4D5DEC46FFC2FE1D5F93FDBD30E0" authority="Ruschel & Sanborn, 2021" authorityName="Ruschel & Sanborn" authorityYear="2021" box="[362,615,1925,1951]" class="Insecta" family="Cicadidae" genus="Rhaeboepelis" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="521" phylum="Arthropoda" rank="species" species="takiyaae" status="n. gen., n. sp.">
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<emphasis id="B90AEACCEC46FFC2FE1D5F93FDBD30E0" box="[362,615,1925,1951]" italics="true" pageId="12" pageNumber="521">Rhaeboepelis takiyaae</emphasis>
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</taxonomicName>
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<emphasis id="B90AEACCEC46FFC2FD195F93FCD330E0" bold="true" box="[622,777,1925,1951]" pageId="12" pageNumber="521">
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<taxonomicNameLabel id="A23957B7EC46FFC2FD195F93FCD330E0" box="[622,777,1925,1951]" pageId="12" pageNumber="521" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC46FFC2FFB05FBFFE043098" pageId="12" pageNumber="521" type="discussion">
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<paragraph id="8BC136DEEC46FFC2FFB05FBFFE043098" blockId="12.[151,1436,1925,2023]" pageId="12" pageNumber="521">
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<emphasis id="B90AEACCEC46FFC2FFB05FBFFEE130BC" bold="true" box="[199,315,1961,1987]" pageId="12" pageNumber="521">Remarks.</emphasis>
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The genus has a similar appearance to
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<taxonomicName id="4C7E4D5DEC46FFC2FD995FBCFC9630BC" authorityName="Stal" authorityYear="1864" box="[750,844,1962,1987]" class="Insecta" family="Cicadidae" genus="Proarna" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="521" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEACCEC46FFC2FD995FBCFC9630BC" box="[750,844,1962,1987]" italics="true" pageId="12" pageNumber="521">Proarna</emphasis>
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</taxonomicName>
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but differs from all species of that genus in the shape of the opercula and genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC47FFC3FFB05881FB6437AA" pageId="13" pageNumber="522" type="etymology">
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<paragraph id="8BC136DEEC47FFC3FFB05881FB6437AA" blockId="13.[151,1437,151,1690]" pageId="13" pageNumber="522">
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<emphasis id="B90AEACCEC47FFC3FFB05881FE9437CE" bold="true" box="[199,334,151,177]" pageId="13" pageNumber="522">Etymology.</emphasis>
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The name is a combination of
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<taxonomicName id="4C7E4D5DEC47FFC3FDD05881FD2737CE" authorityName="Cope" authorityYear="1862" box="[679,765,151,177]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="13" pageNumber="522" phylum="Chordata" rank="genus">
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<emphasis id="B90AEACCEC47FFC3FDD05881FD2737CE" box="[679,765,151,177]" italics="true" pageId="13" pageNumber="522">Rhaebo</emphasis>
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</taxonomicName>
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- (Gr. rhaibos, bent, crooked) and –
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<emphasis id="B90AEACCEC47FFC3FBFC5881FB1637CE" box="[1163,1228,151,177]" italics="true" pageId="13" pageNumber="522">epelis</emphasis>
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(Gr. epelis, cover, lid) in reference to the sinuous posterior margin of the male operculum. The genus is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646555EC47FFC3FFB058C8FEBB33BA" pageId="13" pageNumber="522" type="description">
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<paragraph id="8BC136DEEC47FFC3FFB058C8FCA63326" blockId="13.[151,1437,151,1690]" pageId="13" pageNumber="522">
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<emphasis id="B90AEACCEC47FFC3FFB058C8FE833787" bold="true" box="[199,345,222,248]" pageId="13" pageNumber="522">Description.</emphasis>
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Head wider than mesonotum. Eyes protruding beyond anterior pronotum, lateral margin of eye wider than anterolateral margin of pronotal collar. Vertex at area of ocelli longer than front, front angled ventrally between vertex and frontoclypeal suture. Ocellus well developed, lateral ocelli closer to each other than to eyes, higher than median ocellus in frontal view, ocular tubercle well developed. Supra–antennal plates slightly prominent relative to anterior margin of head, reaching half distance to eyes (
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<figureCitation id="13452A5BEC47FFC3FCF95979FC3D36F6" box="[910,999,367,393]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8B</figureCitation>
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). Postclypeus slightly more prominent than anterior margin of head, anterior margin arched in dorsal view, central sulcus narrow (
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<figureCitation id="13452A5BEC47FFC3FBF35985FB0036D2" box="[1156,1242,403,429]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8B</figureCitation>
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), sub-rectangular in ventral view (
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<figureCitation id="13452A5BEC47FFC3FE2459A1FE7136AE" box="[339,427,439,465]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8C</figureCitation>
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), ventral margin straight in lateral view. Anteclypeus ventral surface with lateroposterior margins forming an acute angle, posterolateral flaps absent (
|
||
<figureCitation id="13452A5BEC47FFC3FC4959CDFC4D368A" box="[830,919,475,501]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8C</figureCitation>
|
||
). Labium long, reaching abdominal sternite II (
|
||
<figureCitation id="13452A5BEC47FFC3FFD759E9FF203566" box="[160,250,511,537]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8D</figureCitation>
|
||
). Pronotal collar narrow in dorsal view, lateral angle expanding, projected laterally, lateral margin slightly convex, narrow, wider than lateral margin of mesonotum (
|
||
<figureCitation id="13452A5BEC47FFC3FC5A5A35FC5F3542" box="[813,901,547,573]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8B</figureCitation>
|
||
). Mesonotum covering dorsal metanotum (
|
||
<figureCitation id="13452A5BEC47FFC3FA185A35FF67351E" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8A</figureCitation>
|
||
). Cruciform elevation with central area swollen, lateral areas flat, posterior arms almost transversely oriented, posterior margin slightly emarginated (
|
||
<figureCitation id="13452A5BEC47FFC3FD205A7DFD6935FA" box="[599,691,619,645]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8A</figureCitation>
|
||
). Anterior basisternum 3 obtusely angled, not prominent relative to mesocoxae (
|
||
<figureCitation id="13452A5BEC47FFC3FE335A99FE7A35D6" box="[324,416,655,681]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8D</figureCitation>
|
||
). Three segmented tarsi. Wings hyaline, forewings narrow, about 3.25 longer than broad, costal margin curved, radial and radiomedial crossveins not parallel, apical cell 2 about half-length of apical cell 1, pterostigma present (
|
||
<figureCitation id="13452A5BEC47FFC3FED25AC1FDDB358E" box="[421,513,727,753]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8A</figureCitation>
|
||
). Male operculum sub-triangular with posterolateral extension, lateral and posterior margins sinuous, opercula not meeting medially (
|
||
<figureCitation id="13452A5BEC47FFC3FC745AEDFC87346A" box="[771,861,763,789]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8E</figureCitation>
|
||
). Meracanthus an elongated triangle, not reaching half operculum length (
|
||
<figureCitation id="13452A5BEC47FFC3FED55B09FE203446" box="[418,506,799,825]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8E</figureCitation>
|
||
). Male abdomen subcylindrical, segment 2 widest, subsequent segments narrower than preceding segment, tergite 1 narrow, anterodorsal region covered by cruciform elevation (
|
||
<figureCitation id="13452A5BEC47FFC3FB955B55FAE13421" box="[1250,1339,835,862]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8A</figureCitation>
|
||
).Timbal cover flat, incomplete, timbal exposed dorsolaterally (
|
||
<figureCitation id="13452A5BEC47FFC3FD835B71FC9334FE" box="[756,841,871,897]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8F</figureCitation>
|
||
). Male sternite I twice as long as metacoxae, swollen (
|
||
<figureCitation id="13452A5BEC47FFC3FFD75B9DFF2834D9" box="[160,242,907,934]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig.8D</figureCitation>
|
||
), sternite VII sub–rectangular (
|
||
<figureCitation id="13452A5BEC47FFC3FD255B9DFD7734DA" box="[594,685,907,933]" captionStart="FIGURE 8" captionStartId="14.[151,250,1507,1531]" captionTargetBox="[151,1435,181,1483]" captionTargetId="figure-23@14.[151,1435,181,1483]" captionTargetPageId="14" captionText="FIGURE 8. Rhaeboepelis takiyaae n. gen., n. sp. A, holotype male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491204" httpUri="https://zenodo.org/record/4491204/files/figure.png" pageId="13" pageNumber="522">Fig. 8G</figureCitation>
|
||
). Pygofer distal shoulder anterior margin terminating in acute projection; dorsal beak absent; pygofer upper lobe absent; pygofer basal lobe well-developed (
|
||
<figureCitation id="13452A5BEC47FFC3FBEE5BB9FB2B34B6" box="[1177,1265,943,969]" captionStart="FIGURE 9" captionStartId="15.[151,250,1636,1660]" captionTargetBox="[151,1436,186,1612]" captionTargetId="figure-19@15.[151,1436,181,1612]" captionTargetPageId="15" captionText="FIGURE 9. Rhaeboepelis takiyaae n. gen., n. sp., male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in latero-ventral view; D, phallus in left view; E, phallus in right view.Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491206" httpUri="https://zenodo.org/record/4491206/files/figure.png" pageId="13" pageNumber="522">Fig. 9C</figureCitation>
|
||
). Uncus longer than basal lobe in lateral view, uncal dorsal crest flattened, divided medially; lateral branches of uncus with distal margin arched and apex directed anteriorly; ventral apophyses formed from lateral branches of uncus fused (
|
||
<figureCitation id="13452A5BEC47FFC3FA185BE1FF06334A" captionStart="FIGURE 9" captionStartId="15.[151,250,1636,1660]" captionTargetBox="[151,1436,186,1612]" captionTargetId="figure-19@15.[151,1436,181,1612]" captionTargetPageId="15" captionText="FIGURE 9. Rhaeboepelis takiyaae n. gen., n. sp., male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in latero-ventral view; D, phallus in left view; E, phallus in right view.Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491206" httpUri="https://zenodo.org/record/4491206/files/figure.png" pageId="13" pageNumber="522">Fig. 9A, B</figureCitation>
|
||
). Theca tubular opened medially, apex with long, slender vesica, cornuti present, spine of vesica present, thecal and vesical processes absent (
|
||
<figureCitation id="13452A5BEC47FFC3FD7B5C29FD5C3326" box="[524,646,1087,1113]" captionStart="FIGURE 9" captionStartId="15.[151,250,1636,1660]" captionTargetBox="[151,1436,186,1612]" captionTargetId="figure-19@15.[151,1436,181,1612]" captionTargetPageId="15" captionText="FIGURE 9. Rhaeboepelis takiyaae n. gen., n. sp., male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in latero-ventral view; D, phallus in left view; E, phallus in right view.Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491206" httpUri="https://zenodo.org/record/4491206/files/figure.png" pageId="13" pageNumber="522">Fig. 9D, E</figureCitation>
|
||
). Female is unknown.
|
||
</paragraph>
|
||
<paragraph id="8BC136DEEC47FFC3FFB05C75FEBB33BA" blockId="13.[151,1437,151,1690]" pageId="13" pageNumber="522">
|
||
<emphasis id="B90AEACCEC47FFC3FFB05C75FE1D3303" bold="true" box="[199,455,1122,1149]" pageId="13" pageNumber="522">Measurements (mm).</emphasis>
|
||
Length of body: 18.45; length of forewing: 26.35; width of forewing: 8.10; length of head: 1.81; width of head including eyes: 6.50; width of pronotum including suprahumeral plates: 6.50; width of mesonotum: 6.10.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3646555EC47FFC3FFB05CD8FE1C310A" pageId="13" pageNumber="522" type="diagnosis">
|
||
<paragraph id="8BC136DEEC47FFC3FFB05CD8FE1C310A" blockId="13.[151,1437,151,1690]" pageId="13" pageNumber="522">
|
||
<emphasis id="B90AEACCEC47FFC3FFB05CD8FE983397" bold="true" box="[199,322,1230,1256]" pageId="13" pageNumber="522">Diagnosis.</emphasis>
|
||
The genus can be distinguished from all other genera in
|
||
<taxonomicName id="4C7E4D5DEC47FFC3FCC95CD9FBF13396" authorityName="Distant" authorityYear="1905" box="[958,1067,1231,1257]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="tribe" tribe="Fidicinini">Fidicinini</taxonomicName>
|
||
by the following combination of characters: head including eyes broader than mesonotum; eyes protruding beyond anterior pronotum, postclypeus with dorsal surface tumid, lateroposterior margins forming an acute angle, anteclypeus posterolateral flaps absent, longitudinal groove slender; labium long, reaching sternite II; anterior basisternum 3 obtusely angled, not prominent relative to mesocoxae; male operculum sub-triangular with posterolateral extension, lateral and posterior margins sinuous; three segmented tarsi; uncal dorsal crest divided medially and projected posteriorly; lateral branches of uncus with distal margin arched and apex anteriorly directed; theca a long tube opened medially, thecal and vesical processes absent, cornuti present, spine of vesica present. Lateral branches of uncus with distal margin arched similar to
|
||
<taxonomicName id="4C7E4D5DEC47FFC3FE7A5DF9FD1D3176" authority="Torres, 1963" authorityName="Torres" authorityYear="1963" box="[269,711,1519,1545]" class="Insecta" family="Cicadidae" genus="Prasinosoma" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="species" species="fuembuenai">
|
||
<emphasis id="B90AEACCEC47FFC3FE7A5DF9FDF73176" box="[269,557,1519,1545]" italics="true" pageId="13" pageNumber="522">Prasinosoma fuembuenai</emphasis>
|
||
<bibRefCitation id="EFEF4B2FEC47FFC3FD425DF9FD1D3176" author="Torres" box="[565,711,1519,1545]" firstAuthor="Torres" pageId="13" pageNumber="522" pagination="13 - 23" refId="ref10943" refString="Torres, B. A. (1963) Desmembracion en los generos Proarna Stal y Tympanoterpes Stal. Creacion del nuevo genero: Prasinosoma (Homoptera, Cicadidae). Revista Sociedad Uruguaya Entomologica, 5, 13 - 23." type="journal article" year="1963">Torres, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
, however the new genus differs in the expanded and unparallel lateral margin of pronotal collar, eyes protruding beyond the anterior pronotum, and the uncal dorsal crest being divided medially, all of which differ from the
|
||
<taxonomicName id="4C7E4D5DEC47FFC3FDE85E21FC12312E" authority="Torres, 1963" authorityName="Torres" authorityYear="1963" box="[671,968,1591,1617]" class="Insecta" family="Cicadidae" genus="Prasinosoma" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEACCEC47FFC3FDE85E21FCE9312F" box="[671,819,1591,1616]" italics="true" pageId="13" pageNumber="522">Prasinosoma</emphasis>
|
||
<bibRefCitation id="EFEF4B2FEC47FFC3FC4D5E21FC12312E" author="Torres" box="[826,968,1591,1617]" firstAuthor="Torres" pageId="13" pageNumber="522" pagination="13 - 23" refId="ref10943" refString="Torres, B. A. (1963) Desmembracion en los generos Proarna Stal y Tympanoterpes Stal. Creacion del nuevo genero: Prasinosoma (Homoptera, Cicadidae). Revista Sociedad Uruguaya Entomologica, 5, 13 - 23." type="journal article" year="1963">Torres, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
species. The operculum shape appears to be unique in the
|
||
<taxonomicName id="4C7E4D5DEC47FFC3FE255E4DFE19310A" authorityName="Distant" authorityYear="1905" box="[338,451,1627,1653]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="tribe" tribe="Fidicinini">Fidicinini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3646555EC47FFC3FFB05E68FDA431E5" box="[199,638,1662,1690]" pageId="13" pageNumber="522" type="distribution">
|
||
<paragraph id="8BC136DEEC47FFC3FFB05E68FDA431E5" blockId="13.[151,1437,151,1690]" box="[199,638,1662,1690]" pageId="13" pageNumber="522">
|
||
<emphasis id="B90AEACCEC47FFC3FFB05E68FEBB31E7" bold="true" box="[199,353,1662,1688]" pageId="13" pageNumber="522">Distribution.</emphasis>
|
||
<collectingRegion id="49BAF83CEC47FFC3FE105E69FE3831E6" box="[359,482,1663,1689]" country="Costa Rica" name="Puntarenas" pageId="13" pageNumber="522">Puntarenas</collectingRegion>
|
||
(
|
||
<collectingCountry id="F369764EEC47FFC3FE865E69FDAB31E5" box="[497,625,1663,1690]" name="Costa Rica" pageId="13" pageNumber="522">Costa Rica</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |