296 lines
40 KiB
XML
296 lines
40 KiB
XML
<document id="BFCCB221A1F44FC4C77812BA570F32C3" ID-CLB-Dataset="2910" ID-DOI="10.11646/zootaxa.5082.3.2" ID-GBIF-Dataset="3997d831-ae29-42d0-ba99-cd20cc8b7fc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5788224" ID-ZooBank="EFB0E4A5-6B60-4B39-A651-1C6470FD7222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639730480321" checkinUser="plazi" docAuthor="Li, Yan-Da, Tihelka, Erik, Dahan, Loïc, Huang, Di-Ying & Cai, Chen-Yang" docDate="2021" docId="03D787A2FFE2EE74FF52FC1D8EE56DA7" docLanguage="en" docName="zootaxa.5082.3.2.pdf" docOrigin="Zootaxa 5082 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaenosodendron Li & Tihelka & Dahan & Huang & Cai 2021, gen. nov." docType="treatment" docVersion="5" lastPageNumber="226" masterDocId="FFEEFFDAFFE0EE77FFC5FFAA8B796C12" masterDocTitle="On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)" masterLastPageNumber="244" masterPageNumber="223" pageNumber="225" updateTime="1699472730423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="28CD833AF75FB153CF622144DB8170C7">On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)</mods:title>
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<mods:namePart id="D307E71680014DE8516DFCE133BC2C12">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="740EBC8029FCE08023868CAFEB8F4C26">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="BFA3439F9ED26FFC4B73346C6F9AA261" type="email">ydli@pku.edu.cn</mods:nameIdentifier>
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<mods:namePart id="C4EA810EB2A9E5538DC908D157FC3461">Tihelka, Erik</mods:namePart>
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<mods:affiliation id="B69D0782E26D91E1A1BFAA981A0F26F3">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355</mods:affiliation>
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<mods:nameIdentifier id="6FB4260C49FD138A273DBD9E1C9F2755" type="email">wn20250@bristol.ac.uk</mods:nameIdentifier>
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<mods:namePart id="7D2E9AEB0309A7CCA41C02E805285D88">Dahan, Loïc</mods:namePart>
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<mods:affiliation id="CC0F705ADC0A89687BC35F07922254DA">O. D. Taxonomy and Phylogeny - Entomology, Royal Belgian Institute of Natural Sciences, Vautierstreet 29, B- 1000 Brussels, Belgium ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X & loicdahan @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5680 - 0304</mods:affiliation>
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<mods:nameIdentifier id="031099D9F592C92022426A46353210A2" type="email">loicdahan@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="32026810804E1D69C11126941A86D8E1">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="345A40B4DD0FEACC8B417430C31B85B7">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<mods:nameIdentifier id="996A659824E8C3D775F5052936E82C6D" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:namePart id="63418ACB0C1343DB5D4306E0954CA419">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="4054EA68663CDF39539C354DCE712A5C">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323</mods:affiliation>
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<mods:nameIdentifier id="5C2EBCC5EB7A641C736C17CC1CB70E12" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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<mods:title id="F7F7DB66D6943E05D75CDE7B7722F5A4">Zootaxa</mods:title>
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<mods:part id="58383A54A1C1AC577836AD6FEDC90455">
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<mods:date id="C945C409BFD47D5800D7BAFD92FBE9D9">2021</mods:date>
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<mods:number id="D48059A2E9FB843D2666AD3085D8F078">2021-12-17</mods:number>
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<mods:number id="E55548088BCB46CC3D5E08447745EE45">5082</mods:number>
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<mods:number id="367B5A35359A3EBBCC6F3E2887D6CA6E">3</mods:number>
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<mods:classification id="E8763087CCAABEDA1217C02A38B5BFC1">journal article</mods:classification>
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<mods:identifier id="D09A26DB8CC2FDB1110A7A98675ECBBE" type="DOI">10.11646/zootaxa.5082.3.2</mods:identifier>
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<treatment id="03D787A2FFE2EE74FF52FC1D8EE56DA7" ID-DOI="http://doi.org/10.5281/zenodo.5789402" ID-GBIF-Taxon="191654781" ID-Zenodo-Dep="5789402" LSID="urn:lsid:plazi:treatment:03D787A2FFE2EE74FF52FC1D8EE56DA7" httpUri="http://treatment.plazi.org/id/03D787A2FFE2EE74FF52FC1D8EE56DA7" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225">
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<subSubSection id="C364653FFFE2EE75FF52FC1D89A96FC3" box="[151,720,951,978]" pageId="2" pageNumber="225" type="nomenclature">
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<paragraph id="8BC136B4FFE2EE75FF52FC1D89A96FC3" blockId="2.[151,720,951,978]" box="[151,720,951,978]" pageId="2" pageNumber="225">
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<heading id="D08981D8FFE2EE75FF52FC1D89A96FC3" bold="true" box="[151,720,951,978]" fontSize="11" level="1" pageId="2" pageNumber="225" reason="1">
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<emphasis id="B90AEAA6FFE2EE75FF52FC1D89A96FC3" bold="true" box="[151,720,951,978]" pageId="2" pageNumber="225">
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Genus
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<taxonomicName id="4C7E4D37FFE2EE75FF2BFC1D89226FC3" authority="Li & Cai" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" box="[238,603,951,978]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90AEAA6FFE2EE75FF2BFC1D8A9D6FC0" bold="true" box="[238,484,951,978]" italics="true" pageId="2" pageNumber="225">Archaenosodendron</emphasis>
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Li & Cai
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</taxonomicName>
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<taxonomicNameLabel id="A23957DDFFE2EE75FDA7FC1D89A96FC3" box="[610,720,951,977]" pageId="2" pageNumber="225" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C364653FFFE2EE75FF52FC5688B66806" box="[151,975,1019,1046]" pageId="2" pageNumber="225" type="type_taxon">
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<paragraph id="8BC136B4FFE2EE75FF52FC5688B66806" blockId="2.[151,975,1019,1046]" box="[151,975,1019,1046]" pageId="2" pageNumber="225">
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<emphasis id="B90AEAA6FFE2EE75FF52FC568A5A6801" bold="true" box="[151,291,1019,1044]" pageId="2" pageNumber="225">
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<typeStatus id="54C58816FFE2EE75FF52FC568BB56806" box="[151,204,1020,1044]" pageId="2" pageNumber="225">Type</typeStatus>
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species.
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||
</emphasis>
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||
<taxonomicName id="4C7E4D37FFE2EE75FEEFFC5688B66806" authority="Deng, Zhou, Slipinski, Ren & Pang, 2018" authorityName="Deng, Zhou, Slipinski, Ren & Pang" authorityYear="2018" box="[298,975,1020,1046]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="cretaceum">
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<emphasis id="B90AEAA6FFE2EE75FEEFFC5689586801" box="[298,545,1020,1044]" italics="true" pageId="2" pageNumber="225">Nosodendron cretaceum</emphasis>
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Deng, Zhou, Ślipiński, Ren & Pang, 2018
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C364653FFFE2EE75FF52FBE88BB86892" pageId="2" pageNumber="225" type="etymology">
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||
<paragraph id="8BC136B4FFE2EE75FF52FBE88BB86892" blockId="2.[151,1437,1090,1836]" pageId="2" pageNumber="225">
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<emphasis id="B90AEAA6FFE2EE75FF52FBE88A67684E" bold="true" box="[151,286,1090,1116]" pageId="2" pageNumber="225">Etymology.</emphasis>
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The generic name is composed of “
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<emphasis id="B90AEAA6FFE2EE75FD77FBE88872684E" box="[690,779,1090,1116]" italics="true" pageId="2" pageNumber="225">Archae-</emphasis>
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”, ancient, and “
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<taxonomicName id="4C7E4D37FFE2EE75FC05FBE88F2F684E" authorityName="Latreille" authorityYear="1804" box="[960,1110,1090,1116]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA6FFE2EE75FC05FBE88F2F684E" box="[960,1110,1090,1116]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
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</taxonomicName>
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”. The name is neuter in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364653FFFE2EE75FF02FB238A0E6902" pageId="2" pageNumber="225" type="diagnosis">
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<paragraph id="8BC136B4FFE2EE75FF02FB238A0E6902" blockId="2.[151,1437,1090,1836]" pageId="2" pageNumber="225">
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<emphasis id="B90AEAA6FFE2EE75FF02FB238A3B68B1" bold="true" box="[199,322,1161,1187]" pageId="2" pageNumber="225">Diagnosis.</emphasis>
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Body elongated oval, generally less than
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<quantity id="4C869B51FFE2EE75FCEDFB20881268B6" box="[808,875,1162,1189]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="225" unit="mm" value="5.0">5 mm</quantity>
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long. Eyes not strongly protuberant. Prosternum in front coxae not extremely shortened, longer than half length of prosternal process, with incomplete antennal grooves. Protibia expanded apically; outer edge of protibia with one row of distinct denticles. Tarsomeres slender, not distinctly lobed.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364653FFFE2EE75FF02FAB08A316B3E" pageId="2" pageNumber="225" type="discussion">
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<paragraph id="8BC136B4FFE2EE75FF02FAB089B76A1E" blockId="2.[151,1437,1090,1836]" pageId="2" pageNumber="225">
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<emphasis id="B90AEAA6FFE2EE75FF02FAB08A426926" bold="true" box="[199,315,1306,1332]" pageId="2" pageNumber="225">Remarks.</emphasis>
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<bibRefCitation id="EFEF4B45FFE2EE75FE84FAB089726926" author="Deng, C. & Zhou, Y. - L. & Slipinski, A. & Ren, D. & Pang, H." box="[321,523,1306,1332]" pageId="2" pageNumber="225" pagination="211 - 215" refId="ref7379" refString="Deng, C., Zhou, Y. - L., Slipinski, A., Ren, D. & Pang, H. (2019) The first wounded-tree beetle (Coleoptera: Nosodendridae) from Cretaceous Burmese amber. Cretaceous Research, 93, 211 - 215. https: // doi. org / 10.1016 / j. cretres. 2018.09.019" type="journal article" year="2019">
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||
Deng
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<emphasis id="B90AEAA6FFE2EE75FE41FAB18AC26926" box="[388,443,1306,1332]" italics="true" pageId="2" pageNumber="225">et al.</emphasis>
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(2019)
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</bibRefCitation>
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||
originally placed
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<taxonomicName id="4C7E4D37FFE2EE75FD16FAB1881F6926" authorityName="Deng, Zhou, Slipinski, Ren & Pang" authorityYear="2018" box="[723,870,1307,1332]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="cretaceum">
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<emphasis id="B90AEAA6FFE2EE75FD16FAB1881F6926" box="[723,870,1307,1332]" italics="true" pageId="2" pageNumber="225">N. cretaceum</emphasis>
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</taxonomicName>
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in the extant genus
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<taxonomicName id="4C7E4D37FFE2EE75FB84FAB08FAE6926" authorityName="Latreille" authorityYear="1804" box="[1089,1239,1306,1332]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA6FFE2EE75FB84FAB08FAE6926" box="[1089,1239,1306,1332]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
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</taxonomicName>
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, since the species differs from the only other extant genus,
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<taxonomicName id="4C7E4D37FFE2EE75FDA8FA94887E694A" authorityName="Hava" authorityYear="2003" box="[621,775,1342,1368]" class="Insecta" family="Nosodendridae" genus="Nosoglobulus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA6FFE2EE75FDA8FA94887E694A" box="[621,775,1342,1368]" italics="true" pageId="2" pageNumber="225">Nosoglobulus</emphasis>
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</taxonomicName>
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, in having a larger body and relatively loosely articulated antennal club. They additionally noted that
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<taxonomicName id="4C7E4D37FFE2EE75FDBBFAC9886B696E" authorityName="Deng, Zhou, Slipinski, Ren & Pang" authorityYear="2018" box="[638,786,1379,1404]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="cretaceum">
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<emphasis id="B90AEAA6FFE2EE75FDBBFAC9886B696E" box="[638,786,1379,1404]" italics="true" pageId="2" pageNumber="225">N. cretaceum</emphasis>
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</taxonomicName>
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differs from both of the extant subgenera in
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<taxonomicName id="4C7E4D37FFE2EE75FAC3FAC88EE5696E" authorityName="Latreille" authorityYear="1804" box="[1286,1436,1378,1404]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA6FFE2EE75FAC3FAC88EE5696E" box="[1286,1436,1378,1404]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
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</taxonomicName>
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in the combination of several characters. Recently, we discovered several additional species related to
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<taxonomicName id="4C7E4D37FFE2EE75FACCFA2D8EE469B2" authorityName="Deng, Zhou, Slipinski, Ren & Pang" authorityYear="2018" box="[1289,1437,1415,1440]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="cretaceum">
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<emphasis id="B90AEAA6FFE2EE75FACCFA2D8EE469B2" box="[1289,1437,1415,1440]" italics="true" pageId="2" pageNumber="225">N. cretaceum</emphasis>
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</taxonomicName>
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from Burmese amber. Based on the examination of the new materials, as well as the
|
||
<typeStatus id="54C58816FFE2EE75FB95FA008FCB69D6" box="[1104,1202,1450,1476]" pageId="2" pageNumber="225" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7E4D37FFE2EE75FB1DFA018E1769D6" authorityName="Deng, Zhou, Slipinski, Ren & Pang" authorityYear="2018" box="[1240,1390,1451,1476]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="cretaceum">
|
||
<emphasis id="B90AEAA6FFE2EE75FB1DFA018E1769D6" box="[1240,1390,1451,1476]" italics="true" pageId="2" pageNumber="225">N. cretaceum</emphasis>
|
||
</taxonomicName>
|
||
, we suggest that the difference between this fossil group and extant
|
||
<taxonomicName id="4C7E4D37FFE2EE75FC92FA64889469FA" authorityName="Latreille" authorityYear="1804" box="[855,1005,1486,1512]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FC92FA64889469FA" box="[855,1005,1486,1512]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
|
||
</taxonomicName>
|
||
is large enough to justify a new genus, which is herein designated as
|
||
<taxonomicName id="4C7E4D37FFE2EE75FE23FA5889BE6A1E" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" box="[486,711,1522,1548]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FE23FA5889BE6A1E" box="[486,711,1522,1548]" italics="true" pageId="2" pageNumber="225">Archaenosodendron</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136B4FFE2EE75FF02F9BC8A316B3E" blockId="2.[151,1437,1090,1836]" pageId="2" pageNumber="225">
|
||
In extant
|
||
<taxonomicName id="4C7E4D37FFE2EE75FEF6F9BC8AB06A22" authorityName="Latreille" authorityYear="1804" box="[307,457,1558,1584]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FEF6F9BC8AB06A22" box="[307,457,1558,1584]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C7E4D37FFE2EE75FDCFF9BC89DD6A22" authorityName="Hava" authorityYear="2003" box="[522,676,1558,1584]" class="Insecta" family="Nosodendridae" genus="Nosoglobulus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FDCFF9BC89DD6A22" box="[522,676,1558,1584]" italics="true" pageId="2" pageNumber="225">Nosoglobulus</emphasis>
|
||
</taxonomicName>
|
||
), the prosternum in front of coxae is extremely shortened, while in
|
||
<taxonomicName id="4C7E4D37FFE2EE75FF52F9908A016A46" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" box="[151,376,1594,1620]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FF52F9908A016A46" box="[151,376,1594,1620]" italics="true" pageId="2" pageNumber="225">Archaenosodendron</emphasis>
|
||
</taxonomicName>
|
||
the prosternum is relatively long. More importantly, in extant
|
||
<taxonomicName id="4C7E4D37FFE2EE75FBEFF9908FB96A46" authorityName="Latreille" authorityYear="1804" box="[1066,1216,1594,1620]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FBEFF9908FB96A46" box="[1066,1216,1594,1620]" italics="true" pageId="2" pageNumber="225">Nosodendron</emphasis>
|
||
</taxonomicName>
|
||
, the lateral portions of prosternum are clearly delimited and are distinctly lower compared to the medial portion (raised mesal shelf; fig. 4B in
|
||
<bibRefCitation id="EFEF4B45FFE2EE75FF19F92889656A8E" author="Nomura, S. & Kamezawa, H." box="[220,540,1666,1692]" pageId="2" pageNumber="225" pagination="21 - 25" refId="ref8052" refString="Nomura, S. & Kamezawa, H. (2014) Records of Nosodendron coenosum (Wollaston, 1873) (Nosodendridae) from Shinagawaku, Tokyo, Japan and SEM observation of its morphological structures. Sayabane, (13), 21 - 25. [in Japanese]" type="journal article" year="2014">Nomura & Kamezawa 2014</bibRefCitation>
|
||
), so that when in repose the fore legs can fold compactly and the protrochanters and profemora could lay exactly on the lowered lateral portions of prosternum. In
|
||
<taxonomicName id="4C7E4D37FFE2EE75FBE1F90C8E7C6AD2" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" box="[1060,1285,1702,1728]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FBE1F90C8E7C6AD2" box="[1060,1285,1702,1728]" italics="true" pageId="2" pageNumber="225">Archaenosodendron</emphasis>
|
||
</taxonomicName>
|
||
, although the lateral portions of prosternum might be somewhat lowered, there is no clear and complete boundary between the lateral and medial portions, and the protrochanters and profemora are unlikely to be able to lay compactly against the prosternum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection id="E2F12F5FFFE2EE74FF52F8D48EE56DA7" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225" type="key">
|
||
<subSubSection id="C364653FFFE2EE75FF52F8D4891D6B8A" box="[151,612,1917,1944]" pageId="2" pageNumber="225" type="description">
|
||
<paragraph id="8BC136B4FFE2EE75FF52F8D4891D6B8A" blockId="2.[151,612,1917,1944]" box="[151,612,1917,1944]" pageId="2" pageNumber="225">
|
||
<heading id="D08981D8FFE2EE75FF52F8D4891D6B8A" bold="true" box="[151,612,1917,1944]" fontSize="11" level="1" pageId="2" pageNumber="225" reason="1">
|
||
<emphasis id="B90AEAA6FFE2EE75FF52F8D4891D6B8A" bold="true" box="[151,612,1917,1944]" pageId="2" pageNumber="225">
|
||
Key to species of
|
||
<taxonomicName id="4C7E4D37FFE2EE75FEABF8D7891D6B8A" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" box="[366,612,1917,1944]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA6FFE2EE75FEABF8D7891D6B8A" bold="true" box="[366,612,1917,1944]" italics="true" pageId="2" pageNumber="225">Archaenosodendron</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364653FFFE2EE75FF52F8698EE56BE7" pageId="2" pageNumber="225" type="multiple">
|
||
<footnote id="E8652ABAFFE2EE75FF52F8698EE56BE7" pageId="2" pageNumber="225">
|
||
<paragraph id="8BC136B4FFE2EE75FF52F8698EE56BE7" blockId="2.[151,1436,1987,2038]" pageId="2" pageNumber="225">
|
||
<emphasis id="B90AEAA6FFE2EE75FF52F8698BD36BCA" bold="true" box="[151,170,1987,2008]" pageId="2" pageNumber="225">1.</emphasis>
|
||
Mentum with somewhat longitudinally arranged larger punctures along the lateral margins and smaller punctures in the medial (anteromedial) region (Fig. 8B)................................................
|
||
<taxonomicName id="4C7E4D37FFE2EE75FB92F84A8E336BE4" authority="Li & Cai" authorityName="Li & Cai" authorityYear="2021" box="[1111,1354,2016,2038]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="explanatum" status="sp. nov.">
|
||
<emphasis id="B90AEAA6FFE2EE75FB92F84A8F916BE4" bold="true" box="[1111,1256,2016,2038]" italics="true" pageId="2" pageNumber="225">A. explanatum</emphasis>
|
||
Li & Cai
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957DDFFE2EE75FA95F84A8EE56BE7" box="[1360,1436,2016,2037]" pageId="2" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</footnote>
|
||
</subSubSection>
|
||
<subSubSection id="C364653FFFE3EE74FF52FF328EE56DA7" pageId="3" pageNumber="226" type="key">
|
||
<paragraph id="8BC136B4FFE3EE74FF52FF328EE56CBE" blockId="3.[151,1437,151,438]" box="[151,1436,151,173]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FF52FF328BE66CBF" bold="true" box="[151,159,152,173]" pageId="3" pageNumber="226">-</emphasis>
|
||
Punctures on the mentum not forming the pattern above.......................................................
|
||
<emphasis id="B90AEAA6FFE3EE74FA55FF3D8EE56CBE" bold="true" box="[1424,1436,151,172]" pageId="3" pageNumber="226">2</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC136B4FFE3EE74FF52FF1F8EE56D17" blockId="3.[151,1437,151,438]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FF52FF1F8BD06CD8" bold="true" box="[151,169,181,202]" pageId="3" pageNumber="226">2.</emphasis>
|
||
Outer edge of protibia with about 14–16 relatively sparsely arranged denticles/stout setae, with the apical two distinctly stouter and more closely situated (
|
||
<figureCitation id="13452A31FFE3EE74FE21FF7889456CFA" box="[484,572,210,232]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li & Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="3" pageNumber="226">Fig. 11G</figureCitation>
|
||
). Medial and lateral portions of prosternum not clearly separated (
|
||
<figureCitation id="13452A31FFE3EE74FB72FF788E746CFA" box="[1207,1293,210,232]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li & Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="3" pageNumber="226">Fig. 11D</figureCitation>
|
||
).....................................................................................
|
||
<emphasis id="B90AEAA6FFE3EE74FB9AFF458EE56D17" bold="true" box="[1119,1436,239,261]" pageId="3" pageNumber="226">
|
||
<taxonomicName id="4C7E4D37FFE3EE74FB9AFF458E336D17" authority="Li & Cai" authorityName="Li & Cai" authorityYear="2021" box="[1119,1354,239,261]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="remotidens" status="sp. nov.">
|
||
<emphasis id="B90AEAA6FFE3EE74FB9AFF458F906D17" bold="true" box="[1119,1257,239,261]" italics="true" pageId="3" pageNumber="226">A. remotidens</emphasis>
|
||
Li & Cai
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957DDFFE3EE74FA95FF5A8EE56D17" box="[1360,1436,240,261]" pageId="3" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC136B4FFE3EE74FF52FEA78EE56D2D" blockId="3.[151,1437,151,438]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FF52FEA78BE66D30" bold="true" box="[151,159,269,290]" pageId="3" pageNumber="226">-</emphasis>
|
||
Outer edge of protibia with more denticles/stout setae; the apical two not forming the pattern above. Medial and lateral portions of prosternum separated by an incomplete ridge (
|
||
<figureCitation id="13452A31FFE3EE74FD7CFE8088716D2D" box="[697,776,298,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1748]" captionTargetId="figure-18@5.[151,1436,181,1748]" captionTargetPageId="5" captionText="FIGURE 4. Details of Archaenosodendron cretaceum (Deng et al., 2018) comb. nov., holotype, CNU-COL-MA-0321, under confocal microscopy. A, Mouthparts, ventral view. B, Antenna, ventral view. C, Prothorax, ventral view, showing the incomplete boundary between the lateral medial portions of prosternum (arrowhead). D, Abdominal base, ventral view. E, Head, anterodorsal view. F, Scutellar shield, dorsal view. G–J, Fore legs. K, Mesotarsus. L, Metatibia. Abbreviations: an8–11, antennomeres 8–11; el, elytron; hd, head; lbp, labial palp; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum; v1–3, ventrites 1–3. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788234" httpUri="https://zenodo.org/record/5788234/files/figure.png" pageId="3" pageNumber="226">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="13452A31FFE3EE74FCD7FE8088456D52" box="[786,828,298,320]" captionStart="FIGURE 14" captionStartId="15.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-18@15.[151,1436,181,1480]" captionTargetPageId="15" captionText="FIGURE 14. Details of Archaenosodendron angulare Li & Cai sp. nov., holotype, NIGP177615, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view. E, Anterolateral depression of metaventrite, ventral view. F, G, Protibia and -tarsus. H, Fore and mid legs. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; lbp, labial palp; msc, mesocoxa; mstb, mesotibia; msv, mesoventrite; mt, mentum; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788263" httpUri="https://zenodo.org/record/5788263/files/figure.png" pageId="3" pageNumber="226">14D</figureCitation>
|
||
).................................................
|
||
<emphasis id="B90AEAA6FFE3EE74FA55FE808EE56D2D" bold="true" box="[1424,1436,298,319]" pageId="3" pageNumber="226">3</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC136B4FFE3EE74FF52FEED8EE56D68" blockId="3.[151,1437,151,438]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FF52FEED8BD06D4E" bold="true" box="[151,169,327,348]" pageId="3" pageNumber="226">3.</emphasis>
|
||
Outer edge of protibia with about 30 densely arranged denticles (
|
||
<figureCitation id="13452A31FFE3EE74FC92FEED88C26D4C" box="[855,955,327,350]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1748]" captionTargetId="figure-18@5.[151,1436,181,1748]" captionTargetPageId="5" captionText="FIGURE 4. Details of Archaenosodendron cretaceum (Deng et al., 2018) comb. nov., holotype, CNU-COL-MA-0321, under confocal microscopy. A, Mouthparts, ventral view. B, Antenna, ventral view. C, Prothorax, ventral view, showing the incomplete boundary between the lateral medial portions of prosternum (arrowhead). D, Abdominal base, ventral view. E, Head, anterodorsal view. F, Scutellar shield, dorsal view. G–J, Fore legs. K, Mesotarsus. L, Metatibia. Abbreviations: an8–11, antennomeres 8–11; el, elytron; hd, head; lbp, labial palp; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum; v1–3, ventrites 1–3. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788234" httpUri="https://zenodo.org/record/5788234/files/figure.png" pageId="3" pageNumber="226">Fig. 4G–J</figureCitation>
|
||
).
|
||
<collectingRegion id="49BAF856FFE3EE74FC09FEE28F646D4F" box="[972,1053,328,349]" country="Myanmar" name="Chin" pageId="3" pageNumber="226">Mentum</collectingRegion>
|
||
with anterior angles not obviously produced (
|
||
<figureCitation id="13452A31FFE3EE74FEEDFECF8A0A6D69" box="[296,371,357,379]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1748]" captionTargetId="figure-18@5.[151,1436,181,1748]" captionTargetPageId="5" captionText="FIGURE 4. Details of Archaenosodendron cretaceum (Deng et al., 2018) comb. nov., holotype, CNU-COL-MA-0321, under confocal microscopy. A, Mouthparts, ventral view. B, Antenna, ventral view. C, Prothorax, ventral view, showing the incomplete boundary between the lateral medial portions of prosternum (arrowhead). D, Abdominal base, ventral view. E, Head, anterodorsal view. F, Scutellar shield, dorsal view. G–J, Fore legs. K, Mesotarsus. L, Metatibia. Abbreviations: an8–11, antennomeres 8–11; el, elytron; hd, head; lbp, labial palp; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum; v1–3, ventrites 1–3. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788234" httpUri="https://zenodo.org/record/5788234/files/figure.png" pageId="3" pageNumber="226">Fig. 4A</figureCitation>
|
||
)..................................................................
|
||
<taxonomicName id="4C7E4D37FFE3EE74FB5DFECF8EE56D68" authority="(Deng et al.)" authorityName="Li & Tihelka & Dahan & Huang & Cai" authorityYear="2021" baseAuthorityName="Li & Tihelka & Dahan & Huang & Cai" baseAuthorityYear="2021" box="[1176,1436,356,378]" class="Insecta" family="Nosodendridae" genus="Nosodendron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="cretaceum">
|
||
<emphasis id="B90AEAA6FFE3EE74FB5DFECF8EE56D68" bold="true" box="[1176,1436,356,378]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FB5DFECF8E626D68" bold="true" box="[1176,1307,357,378]" italics="true" pageId="3" pageNumber="226">A. cretaceum</emphasis>
|
||
(Deng
|
||
<emphasis id="B90AEAA6FFE3EE74FAA1FECF8EED6D68" bold="true" box="[1380,1428,356,378]" italics="true" pageId="3" pageNumber="226">et al.</emphasis>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BC136B4FFE3EE74FF52FE288EE56DA7" blockId="3.[151,1437,151,438]" pageId="3" pageNumber="226">
|
||
<emphasis id="B90AEAA6FFE3EE74FF52FE288BE66D85" bold="true" box="[151,159,386,407]" pageId="3" pageNumber="226">-</emphasis>
|
||
Outer edge of protibia with about 20 denticles/stout setae (
|
||
<figureCitation id="13452A31FFE3EE74FCD8FE2888EB6D8A" box="[797,914,386,408]" captionStart="FIGURE 14" captionStartId="15.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-18@15.[151,1436,181,1480]" captionTargetPageId="15" captionText="FIGURE 14. Details of Archaenosodendron angulare Li & Cai sp. nov., holotype, NIGP177615, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view. E, Anterolateral depression of metaventrite, ventral view. F, G, Protibia and -tarsus. H, Fore and mid legs. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; lbp, labial palp; msc, mesocoxa; mstb, mesotibia; msv, mesoventrite; mt, mentum; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788263" httpUri="https://zenodo.org/record/5788263/files/figure.png" pageId="3" pageNumber="226">Fig. 14F–H</figureCitation>
|
||
).
|
||
<collectingRegion id="49BAF856FFE3EE74FC66FE29888D6D8A" box="[931,1012,387,408]" country="Myanmar" name="Chin" pageId="3" pageNumber="226">Mentum</collectingRegion>
|
||
with anterior angles pointed anterolaterally (
|
||
<figureCitation id="13452A31FFE3EE74FF2CFE358A476DA4" box="[233,318,415,438]" captionStart="FIGURE 14" captionStartId="15.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-18@15.[151,1436,181,1480]" captionTargetPageId="15" captionText="FIGURE 14. Details of Archaenosodendron angulare Li & Cai sp. nov., holotype, NIGP177615, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view. E, Anterolateral depression of metaventrite, ventral view. F, G, Protibia and -tarsus. H, Fore and mid legs. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; lbp, labial palp; msc, mesocoxa; mstb, mesotibia; msv, mesoventrite; mt, mentum; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788263" httpUri="https://zenodo.org/record/5788263/files/figure.png" pageId="3" pageNumber="226">Fig. 14B</figureCitation>
|
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)...................................................................
|
||
<emphasis id="B90AEAA6FFE3EE74FBB6FE358EE56DA7" bold="true" box="[1139,1436,415,437]" pageId="3" pageNumber="226">
|
||
<taxonomicName id="4C7E4D37FFE3EE74FBB6FE358E336DA7" authority="Li & Cai" authorityName="Li & Cai" authorityYear="2021" box="[1139,1354,415,437]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="angulare" status="sp. nov.">
|
||
<emphasis id="B90AEAA6FFE3EE74FBB6FE358F916DA7" bold="true" box="[1139,1256,415,437]" italics="true" pageId="3" pageNumber="226">A. angulare</emphasis>
|
||
Li & Cai
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957DDFFE3EE74FA95FE0A8EE56DA7" box="[1360,1436,416,437]" pageId="3" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |