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<document ID-DOI="10.11646/zootaxa.5256.1.5" ID-ISSN="1175-5326" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1679054717622" checkinUser="plazi" docAuthor="Billingham, Zacariah D., Theischinger, Gunther &amp; Levesque-Beaudin, Valerie" docDate="2023" docId="03ED0C5F70143019FF1F27A98DF6F879" docLanguage="en" docName="zootaxa.5256.1.5.pdf" docOrigin="Zootaxa 5256 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Molophilus Curtis 1833" docType="treatment" docVersion="1" lastPageNumber="80" masterDocId="FFD474277017301AFF88235B8D01FFEC" masterDocTitle="Two new species of crane fly (Diptera: Limoniidae) from Tasmania, Australia" masterLastPageNumber="83" masterPageNumber="77" pageNumber="80" updateTime="1679312955712" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title>Two new species of crane fly (Diptera: Limoniidae) from Tasmania, Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Billingham, Zacariah D.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6163-9073</mods:nameIdentifier>
<mods:affiliation>Department of Ecology, Environment &amp; Evolution, La Trobe University, Bundoora, Vic 3086, Australia.</mods:affiliation>
<mods:nameIdentifier type="email">zac.billingham@ghd.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Theischinger, Gunther</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5207-2626</mods:nameIdentifier>
<mods:affiliation>Australian Museum, Entomology, 6 College Street, Sydney, NSW 2010, Australia. &amp; theischingergunther @ gmail. com; http: // orcid. org / 0000 - 0002 - 5207 - 2626</mods:affiliation>
<mods:nameIdentifier type="email">theischingergunther@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Levesque-Beaudin, Valerie</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6053-0949</mods:nameIdentifier>
<mods:affiliation>Centre for Biodiversity Genomics, University of Guelph, 50 Stone Road East, Guelph, N 1 G 2 W 1, Canada. &amp; valerie @ uoguelph. ca; http: // orcid. org / 0000 - 0002 - 6053 - 0949</mods:affiliation>
<mods:nameIdentifier type="email">valerie@uoguelph.ca</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5256</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>77</mods:start>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.5</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<subSubSection box="[151,538,1266,1293]" pageId="3" pageNumber="80" type="nomenclature">
<paragraph blockId="3.[151,538,1266,1293]" box="[151,538,1266,1293]" pageId="3" pageNumber="80">
<heading bold="true" box="[151,538,1266,1293]" fontSize="11" level="1" pageId="3" pageNumber="80" reason="1">
<emphasis bold="true" box="[151,538,1266,1293]" pageId="3" pageNumber="80">
Genus
<taxonomicName ID-CoL="87HLS" ID-ENA="700853" authority="Curtis, 1833" authorityName="Curtis" authorityYear="1833" box="[238,538,1266,1293]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[238,377,1266,1293]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
Curtis, 1833
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="80" type="materials_examined">
<paragraph blockId="3.[151,1437,1339,1941]" box="[151,999,1339,1365]" pageId="3" pageNumber="80">
<typeStatus box="[151,207,1339,1365]" pageId="3" pageNumber="80">Type</typeStatus>
species:
<taxonomicName authority="Curtis, 1833" authorityName="Curtis" authorityYear="1833" box="[309,722,1339,1365]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis box="[309,574,1339,1365]" italics="true" pageId="3" pageNumber="80">Molophilus brevipennis</emphasis>
Curtis, 1833
</taxonomicName>
, by original designation.
</paragraph>
<paragraph blockId="3.[151,1437,1339,1941]" pageId="3" pageNumber="80">
Diagnosis. Rostrum short, little developed; meron between mesocoxa and metacoxa extensive, seen from lateral aspect larger than mesocoxa; setae of legs simple;
<collectionCode box="[735,754,1411,1437]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="80">R</collectionCode>
<subScript attach="left" box="[753,758,1429,1443]" fontSize="6" pageId="3" pageNumber="80">l</subScript>
longer than 2/3 wing length; Rs ending in cell r
<subScript attach="left" box="[1317,1325,1429,1443]" fontSize="6" pageId="3" pageNumber="80">3</subScript>
;
<collectionCode box="[1342,1361,1411,1437]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="80">R</collectionCode>
<subScript attach="left" box="[1361,1386,1429,1443]" fontSize="6" pageId="3" pageNumber="80">2+3</subScript>
and
<collectionCode box="[151,170,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="80">R</collectionCode>
<subScript attach="left" box="[170,196,1465,1479]" fontSize="6" pageId="3" pageNumber="80">4+5</subScript>
distinct;
<collectionCode box="[301,320,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="80">R</collectionCode>
<subScript attach="left" box="[319,327,1465,1479]" fontSize="6" pageId="3" pageNumber="80">5</subScript>
not fused with M
<subScript attach="left" box="[528,554,1465,1479]" fontSize="6" pageId="3" pageNumber="80">1+2</subScript>
; r-m present; CuA
<subScript attach="left" box="[766,774,1465,1479]" fontSize="6" pageId="3" pageNumber="80">2</subScript>
and A
<subScript attach="left" box="[848,853,1465,1479]" fontSize="6" pageId="3" pageNumber="80">l</subScript>
not fused distally; a
<subScript attach="left" box="[1081,1089,1465,1479]" fontSize="6" pageId="3" pageNumber="80">2</subScript>
long and wide (modified from
<bibRefCitation author="Theischinger, G." box="[151,364,1483,1509]" pageId="3" pageNumber="80" pagination="5 - 150" refId="ref4026" refString="Theischinger, G. (1992) The Limoniinae (Diptera: Tipulidae) of Australia. II. The genus Molophilus Curtis. Stapfia, 27, 5 - 150." type="journal article" year="1992">Theischinger 1992</bibRefCitation>
).
</paragraph>
<paragraph blockId="3.[151,1437,1339,1941]" pageId="3" pageNumber="80">
The genus
<taxonomicName authorityName="Curtis" authorityYear="1833" box="[323,451,1519,1545]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis box="[323,451,1519,1545]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
</taxonomicName>
includes over 1,000 species, present on all continents except
<collectingCountry box="[1140,1260,1519,1545]" name="Antarctica" pageId="3" pageNumber="80">Antarctica</collectingCountry>
. Their diversity is greatest in the Australasian region, with 464 species occurring across
<collectingCountry box="[970,1076,1555,1581]" name="Australia" pageId="3" pageNumber="80">Australia</collectingCountry>
,
<collectingCountry box="[1088,1265,1555,1581]" name="New Caledonia" pageId="3" pageNumber="80">New Caledonia</collectingCountry>
,
<collectingCountry box="[1278,1432,1555,1581]" name="New Zealand" pageId="3" pageNumber="80">New Zealand</collectingCountry>
,
<collectingCountry box="[151,365,1591,1617]" name="Papua New Guinea" pageId="3" pageNumber="80">Papua New Guinea</collectingCountry>
and West Papua (
<bibRefCitation author="Oosterbroek, P." box="[559,766,1591,1617]" pageId="3" pageNumber="80" refId="ref3847" refString="Oosterbroek, P. (2023) Catalogue of the craneflies of the World (CCW). Available online at ccw. naturalis. nl (accessed January 2023)" type="journal volume" year="2023">Oosterbroek 2023</bibRefCitation>
).
<collectingCountry box="[782,885,1591,1617]" name="Australia" pageId="3" pageNumber="80">Australia</collectingCountry>
boasts more species of
<taxonomicName authorityName="Curtis" authorityYear="1833" box="[1145,1273,1591,1617]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis box="[1145,1273,1591,1617]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
</taxonomicName>
than any other country or continent with 325 species recorded, including the species newly described here. Except for a single species,
<taxonomicName authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" box="[246,721,1663,1689]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="flavonotatus" subGenus="Molophilus">
<emphasis box="[246,277,1663,1688]" italics="true" pageId="3" pageNumber="80">M.</emphasis>
(
<emphasis box="[292,420,1663,1689]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
)
<emphasis box="[436,574,1663,1689]" italics="true" pageId="3" pageNumber="80">flavonotatus</emphasis>
Skuse, 1890
</taxonomicName>
, which is also known from
<collectingCountry box="[1029,1181,1663,1689]" name="New Zealand" pageId="3" pageNumber="80">New Zealand</collectingCountry>
, the Australian species of
<taxonomicName authorityName="Curtis" authorityYear="1833" box="[182,310,1699,1725]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis box="[182,310,1699,1725]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
</taxonomicName>
are all endemic.
</paragraph>
<paragraph blockId="3.[151,1437,1339,1941]" pageId="3" pageNumber="80">
<bibRefCitation author="Theischinger, G." box="[199,429,1735,1761]" pageId="3" pageNumber="80" pagination="5 - 150" refId="ref4026" refString="Theischinger, G. (1992) The Limoniinae (Diptera: Tipulidae) of Australia. II. The genus Molophilus Curtis. Stapfia, 27, 5 - 150." type="journal article" year="1992">Theischinger (1992)</bibRefCitation>
undertook a comprehensive review of the Australian
<taxonomicName authorityName="Curtis" authorityYear="1833" box="[1048,1176,1735,1761]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis box="[1048,1176,1735,1761]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
</taxonomicName>
, describing many new species and providing a key to the subgenera and species groups. Following Theischingers review additional new species have been described over numerous works (
<bibRefCitation author="Billingham, Z. D. &amp; Theischinger, G." box="[789,1178,1807,1833]" pageId="3" pageNumber="80" pagination="321 - 332" refId="ref3274" refString="Billingham, Z. D. &amp; Theischinger, G. (2018 a) Three new species of crane fly from Australia (Diptera: Tipuloidea). Linzer biologische Beitrage, 50 (1), 321 - 332." type="journal article" year="2018">Billingham &amp; Theischinger 2018a</bibRefCitation>
,
<bibRefCitation author="Billingham, Z. D. &amp; Theischinger, G." box="[1191,1263,1807,1833]" pageId="3" pageNumber="80" pagination="75 - 92" refId="ref3316" refString="Billingham, Z. D. &amp; Theischinger, G. (2018 b) New and poorly known species of Molophilus Curtis crane flies from Australia (Diptera: Limoniidae). Australian Entomologist, 45 (1), 75 - 92." type="journal article" year="2018">2018b</bibRefCitation>
,
<bibRefCitation author="Billingham, Z. D. &amp; Theischinger, G." box="[1275,1346,1807,1833]" pageId="3" pageNumber="80" pagination="803 - 809" refId="ref3361" refString="Billingham, Z. D. &amp; Theischinger, G. (2019 a) Two new species of Molophilus Curtis from Australia (Diptera: Limoniidae). Linzer biologische Beitrage, 51 (2), 803 - 809." type="journal article" year="2019">2019a</bibRefCitation>
,
<bibRefCitation author="Billingham, Z. D. &amp; Theischinger, G." box="[1358,1432,1807,1833]" pageId="3" pageNumber="80" pagination="93 - 98" refId="ref3403" refString="Billingham, Z. D. &amp; Theischinger, G. (2019 b) A new species of Molophilus (Superbomolophilus) Theischinger (Diptera: Limoniidae) from the Barrington Tops region of New South Wales, Australia. Australian Entomologist, 46 (2), 93 - 98." type="journal article" year="2019">2019b</bibRefCitation>
;
<bibRefCitation author="Theischinger, G." box="[151,363,1843,1869]" pageId="3" pageNumber="80" pagination="1 - 36" refId="ref4090" refString="Theischinger, G. (1994) New and insufficiently known species of Gynoplistia Macquart and Molophilus Curtis (Diptera: Tipulidae: Limoniidae) from Australia. Stapfia, 36, 1 - 36." type="journal article" year="1994">Theischinger 1994</bibRefCitation>
,
<bibRefCitation author="Theischinger, G." box="[374,446,1843,1869]" pageId="3" pageNumber="80" pagination="1 - 18" refId="ref4126" refString="Theischinger, G. (1996 a) The Limoniinae (Diptera: Tipulidae) of Australia. VI. New and insufficiently known species of Toxorhina Loew, Limonia Meigen, Atarba Osten Sacken, Amphineurus Skuse, Gonomyia Meigen and Molophilus Curtis. Stapfia, 44, 1 - 18." type="journal article" year="1996">1996a</bibRefCitation>
,
<bibRefCitation author="Theischinger, G." box="[457,516,1843,1869]" pageId="3" pageNumber="80" pagination="493 - 502" refId="ref4214" refString="Theischinger, G. (1999) The Limoniinae (Diptera: Tipulidae) of Australia. IX. New taxa of Gynoplistia Macquart, Gonomyia Meigen and Molophilus Curtis. Linzer biologische Beitrage, 31 (1), 493 - 502." type="journal article" year="1999">1999</bibRefCitation>
,
<bibRefCitation author="Theischinger, G." box="[527,586,1843,1869]" pageId="3" pageNumber="80" pagination="1181 - 1190" refId="ref4258" refString="Theischinger, G. (2000) The Limoniinae (Diptera: Tipulidae) of Australia. X. New species of Toxorhina Loew, Limonia Meigen, Austrolimnophila Alexander, Gynoplistia Macquart and Molophilus Curtis. Linzer biologische Beitrage, 32 (2), 1181 - 1190." type="journal article" year="2000">2000</bibRefCitation>
,
<bibRefCitation author="Theischinger, G." box="[597,656,1843,1869]" pageId="3" pageNumber="80" pagination="275 - 278" refId="ref4308" refString="Theischinger, G. (2012) Two remarkable new species of Molophilus Curtis from Australia (Insecta: Diptera: Tipuloidea, Limoniidae). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 148 / 149, 275 - 278." type="journal article" year="2012">2012</bibRefCitation>
, 2017).
</paragraph>
<paragraph blockId="3.[151,1437,1339,1941]" pageId="3" pageNumber="80">
<taxonomicName authorityName="Curtis" authorityYear="1833" box="[199,327,1879,1905]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis box="[199,327,1879,1905]" italics="true" pageId="3" pageNumber="80">Molophilus</emphasis>
</taxonomicName>
is readily distinguished from other Australian Limoniid genera using the key provided in
<bibRefCitation author="Theischinger, G." pageId="3" pageNumber="80" pagination="117 - 144" refId="ref4179" refString="Theischinger, G. (1996 b) The Limoniinae (Diptera: Tipulidae) of Australia. VIII. Preliminary key to the genus-group taxa. Stapfia, 44, 117 - 144." type="journal article" year="1996">Theischinger (1996b)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>