treatments-xml/data/03/E3/22/03E3224DFFBA2832FC72F988FAF1AA30.xml

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<mods:title id="40C7C01F80621E29C40E71E6EA08B82D">The genus Parilia Wood-Mason, in Wood-Mason &amp; Alcock, 1891, with description of a new species and establishment of a new genus for P. tuberculata Sakai, 1961 (Crustacea, Brachyura, Leucosiidae)</mods:title>
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<mods:namePart id="B2B7F71A35E922D826D62A4FEED46A55">Ng, Peter K. L.</mods:namePart>
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<treatment id="03E3224DFFBA2832FC72F988FAF1AA30" ID-DOI="http://doi.org/10.5281/zenodo.4778996" ID-GBIF-Taxon="182253772" ID-Zenodo-Dep="4778996" LSID="urn:lsid:plazi:treatment:03E3224DFFBA2832FC72F988FAF1AA30" httpUri="http://treatment.plazi.org/id/03E3224DFFBA2832FC72F988FAF1AA30" lastPageId="4" lastPageNumber="304" pageId="2" pageNumber="302">
<subSubSection id="C350C0D0FFBA2834FC72F988FB08A449" box="[991,1272,1632,1656]" pageId="2" pageNumber="302" type="nomenclature">
<paragraph id="8BF5935BFFBA2834FC72F988FB08A449" blockId="2.[896,1368,1632,1688]" box="[991,1272,1632,1656]" pageId="2" pageNumber="302">
<heading id="D0BD2437FFBA2834FC72F988FB08A449" bold="true" box="[991,1272,1632,1656]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="302" reason="2">
<taxonomicName id="4C4AE8D8FFBA2834FC72F988FB08A449" ID-CoL="75RLN" authority="Chen, 1984" authorityName="Chen" authorityYear="1984" box="[991,1272,1632,1656]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBA2834FC72F988FB08A449" bold="true" box="[991,1272,1632,1656]" pageId="2" pageNumber="302">
<emphasis id="B93E4F49FFBA2834FC72F988FB9EA449" bold="true" box="[991,1134,1632,1656]" italics="true" pageId="2" pageNumber="302">Parilia ovata</emphasis>
Chen, 1984
</emphasis>
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</subSubSection>
<subSubSection id="C350C0D0FFBA2834FC2DF968FAA8A4A9" box="[896,1368,1664,1688]" pageId="2" pageNumber="302" type="description">
<paragraph id="8BF5935BFFBA2834FC2DF968FAA8A4A9" blockId="2.[896,1368,1632,1688]" box="[896,1368,1664,1688]" pageId="2" pageNumber="302">
(
<figureCitation id="13718FDEFFBA2834FC2AF968FC16A4A9" box="[903,998,1664,1688]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="2" pageNumber="302">Figs. 1C</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FC59F968FBE5A4A9" box="[1012,1045,1664,1688]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="2" pageNumber="302">3C</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FB8EF968FB91A4A9" box="[1059,1121,1664,1688]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="2" pageNumber="302">4E, F</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FBC3F968FB7CA4A9" box="[1134,1164,1664,1688]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="2" pageNumber="302">5F</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FB34F968FB4AA4A9" box="[1177,1210,1664,1688]" captionStart="Fig" captionStartId="8.[113,148,1410,1432]" captionTargetBox="[113,1451,189,1393]" captionTargetId="figure-279@8.[113,1451,189,1393]" captionText="Fig. 6. Outer view of right chela. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778841" httpUri="https://zenodo.org/record/4778841/files/figure.png" pageId="2" pageNumber="302">6C</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FB65F969FAF6A4A9" box="[1224,1286,1665,1688]" captionStart="Fig" captionStartId="9.[135,170,950,972]" captionTargetBox="[136,1474,189,933]" captionTargetId="figure-310@9.[136,1474,189,933]" captionText="Fig. 7. Male thoracic sternum and pleon. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778845" httpUri="https://zenodo.org/record/4778845/files/figure.png" pageId="2" pageNumber="302">7E, F</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FABEF968FAEEA4A9" box="[1299,1310,1664,1688]" captionStart="Fig" captionStartId="10.[113,148,1974,1996]" captionTargetBox="[115,1453,190,1952]" captionTargetId="figure-8@10.[99,1459,176,1964]" captionText="Fig. 8. Gonopods. A–D, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; E–I, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; J–M, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; N–S, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India. A, E, J, N, left G1 (ventral view); O, left G1 (dorsal view); B, F, K, P, distal part of left G1 (ventral view); C, G, H, L, Q, R, distal part of left G1 (dorsal view); D, I, M, S, left G2. Scales: A, D, E, I, J, M, N, O, S = 2.0 mm; B, C, F–H, K, L, P–R = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4778849" httpUri="https://zenodo.org/record/4778849/files/figure.png" pageId="2" pageNumber="302">8</figureCitation>
J–M)
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFBA2834FC80F957FB3BA51A" pageId="2" pageNumber="302" type="reference_group">
<paragraph id="8BF5935BFFBA2834FC80F957FBB6A53F" blockId="2.[813,1452,1727,1835]" pageId="2" pageNumber="302">
<taxonomicName id="4C4AE8D8FFBA2834FC80F957FB90A4E5" ID-CoL="75RLN" authority="Chen, 1984: 482" authorityName="Chen" authorityPageNumber="482" authorityYear="1984" box="[813,1120,1727,1749]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBA2834FC80F957FC5FA4E4" box="[813,943,1727,1749]" italics="true" pageId="2" pageNumber="302">Parilia ovata</emphasis>
Chen, 1984: 482
</taxonomicName>
, figs. 1–6; Chen, 1989: 235, text fig. 30d; 
<bibRefCitation id="EFDBEEAAFFBA2834FC03F934FBA5A4C0" author="Tan CGS" box="[942,1109,1756,1778]" pageId="2" pageNumber="302" pagination="1021 - 1058" refId="ref11174" refString="Tan CGS (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30 (7): 1021 - 1058, figs. 1 - 8." type="journal article" year="1996">Tan, 1996: 1048</bibRefCitation>
; Chen &amp; Sun, 2002: 366, text fig. 163; 
<bibRefCitation id="EFDBEEAAFFBA2834FC28F911FBB2A53F" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[901,1090,1785,1807]" pageId="2" pageNumber="302" pagination="1 - 286" refId="ref10633" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 92</bibRefCitation>
.
</paragraph>
<paragraph id="8BF5935BFFBA2834FC80F8FEFB3BA51A" blockId="2.[813,1452,1727,1835]" box="[813,1227,1813,1835]" pageId="2" pageNumber="302">
<taxonomicName id="4C4AE8D8FFBA2834FC80F8FEFB76A51A" ID-CoL="456NS" authority="Zarenkov, 1990: 62" authorityName="Zarenkov" authorityPageNumber="62" authorityYear="1990" box="[813,1158,1813,1835]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="302" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBA2834FC80F8FEFC4CA51B" box="[813,956,1813,1835]" italics="true" pageId="2" pageNumber="302">Myra anomala</emphasis>
Zarenkov, 1990: 62
</taxonomicName>
, fig. 5.
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFBA2834FC80F8BBFC17A59A" pageId="2" pageNumber="302" type="materials_examined">
<paragraph id="8BF5935BFFBA2834FC80F8BBFC17A59A" blockId="2.[813,1452,1875,1963]" pageId="2" pageNumber="302">
<emphasis id="B93E4F49FFBA2834FC80F8BBFBD1A55A" bold="true" box="[813,1057,1875,1899]" pageId="2" pageNumber="302">Material examined.</emphasis>
<materialsCitation id="3B229906FFBA2834FB80F8BCFC6DA59A" ID-GBIF-Occurrence="3115212317" collectionCode="ZRC" collectorName="J. Arbasto" country="Philippines" location="northwestern Panglao" pageId="2" pageNumber="302" specimenCode="ZRC 2017.219" specimenCount="1" specimenCount-male="1" stateProvince="Bohol">
<specimenCount id="9D4C58D2FFBA2834FB80F8BCFB8DA55A" box="[1069,1149,1875,1900]" pageId="2" pageNumber="302" type="male">1 male</specimenCount>
(31.6 × 
<quantity id="4CB23EBEFFBA2834FB40F8BCFAA8A55D" box="[1261,1368,1876,1900]" metricMagnitude="-2" metricUnit="m" metricValue="4.33" pageId="2" pageNumber="302" unit="mm" value="43.3">43.3 mm</quantity>
) (
<specimenCode id="DBEC3B20FFBA2834FAD8F8BCFC61A5BD" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="302">ZRC 2017.219</specimenCode>
), 
<location id="8E95C580FFBA2834FC04F89BFB67A5BA" LSID="urn:lsid:plazi:treatment:03E3224DFFBA2832FC72F988FAF1AA30:8E95C580FFBA2834FC04F89BFB67A5BA" box="[937,1175,1907,1931]" country="Philippines" name="northwestern Panglao" pageId="2" pageNumber="302" stateProvince="Bohol">northwestern Panglao</location>
, 
<collectingRegion id="498E5DB9FFBA2834FB09F89BFB18A5BA" box="[1188,1256,1907,1931]" country="Philippines" name="Bohol" pageId="2" pageNumber="302">Bohol</collectingRegion>
, 
<collectingCountry id="F35DD3CBFFBA2834FB58F89BFA80A5BA" box="[1269,1392,1907,1931]" name="Philippines" pageId="2" pageNumber="302">Philippines</collectingCountry>
, coll. 
<collectorName id="26BFF68DFFBA2834FC80F87CFC6DA59A" box="[813,925,1939,1963]" pageId="2" pageNumber="302">J. Arbasto</collectorName>
</materialsCitation>
, 2006.
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFBA2835FC80F83BFE06A538" lastPageId="3" lastPageNumber="303" pageId="2" pageNumber="302" type="diagnosis">
<paragraph id="8BF5935BFFBA2835FC80F83BFE06A538" blockId="2.[813,1451,2003,2091]" lastBlockId="3.[136,775,1425,1801]" lastPageId="3" lastPageNumber="303" pageId="2" pageNumber="302">
<emphasis id="B93E4F49FFBA2834FC80F83BFC53A5DA" bold="true" box="[813,931,2003,2027]" pageId="2" pageNumber="302">Diagnosis.</emphasis>
Carapace longitudinally ovate (
<figureCitation id="13718FDEFFBA2834FAA7F83BFA9BA5DD" box="[1290,1387,2003,2028]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="2" pageNumber="302">Figs. 1C</figureCitation>
, 
<figureCitation id="13718FDEFFBA2834FAD6F83CFA6EA5DD" box="[1403,1438,2004,2028]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="2" pageNumber="302">3C</figureCitation>
); dorsal surface smooth (
<figureCitation id="13718FDEFFBA2834FBEDF81BFB6CAA3D" box="[1088,1180,2035,2060]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="2" pageNumber="302">Fig. 3C</figureCitation>
); branchial regions not swollen laterally and dorsally, lateral margin of carapace entire, smooth (
<figureCitation id="13718FDEFFBB2835FE99FA79FE7AA798" box="[308,394,1425,1449]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="3" pageNumber="303">Fig. 3C</figureCitation>
); dome-shape in frontal view (
<figureCitation id="13718FDEFFBB2835FD71FA79FF5AA7F8" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="3" pageNumber="303">Fig. 4E</figureCitation>
); frontal region protruding prominently anteriorly with buccal cavity and third maxillipeds not visible in dorsal view, frontal lobes rounded (
<figureCitation id="13718FDEFFBB2835FE60FA19FDD5A438" box="[461,549,1521,1545]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="3" pageNumber="303">Fig. 3C</figureCitation>
); cardiac spine very long (
<figureCitation id="13718FDEFFBB2835FF67F9F9FED2A418" box="[202,290,1553,1577]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="3" pageNumber="303">Fig. 3C</figureCitation>
); outer surface of third maxillipeds smooth (
<figureCitation id="13718FDEFFBB2835FF3CF9D9FEE9A478" box="[145,281,1585,1609]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="3" pageNumber="303">Figs. 4E, F</figureCitation>
, 
<figureCitation id="13718FDEFFBB2835FE84F9D9FEBBA478" box="[297,331,1585,1609]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="3" pageNumber="303">5F</figureCitation>
); exopod of third maxilliped in adult broad but not foliaceous (
<figureCitation id="13718FDEFFBB2835FE09F9B9FE08A458" box="[420,504,1617,1641]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="3" pageNumber="303">Fig. 5F</figureCitation>
); surfaces of ambulatory legs and chelipeds completely smooth (
<figureCitation id="13718FDEFFBB2835FD90F999FD6DA4B8" box="[573,669,1649,1673]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="3" pageNumber="303">Figs. 1C</figureCitation>
, 
<figureCitation id="13718FDEFFBB2835FD00F999FD3FA4B8" box="[685,719,1649,1673]" captionStart="Fig" captionStartId="8.[113,148,1410,1432]" captionTargetBox="[113,1451,189,1393]" captionTargetId="figure-279@8.[113,1451,189,1393]" captionText="Fig. 6. Outer view of right chela. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778841" httpUri="https://zenodo.org/record/4778841/files/figure.png" pageId="3" pageNumber="303">6C</figureCitation>
); G1 proportionately shorter, distal part flared, subdistal lateral projection large, very wide, fan-like, laterally flattened (
<figureCitation id="13718FDEFFBB2835FD71F959FF64A4D8" captionStart="Fig" captionStartId="10.[113,148,1974,1996]" captionTargetBox="[115,1453,190,1952]" captionTargetId="figure-8@10.[99,1459,176,1964]" captionText="Fig. 8. Gonopods. A–D, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; E–I, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; J–M, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; N–S, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India. A, E, J, N, left G1 (ventral view); O, left G1 (dorsal view); B, F, K, P, distal part of left G1 (ventral view); C, G, H, L, Q, R, distal part of left G1 (dorsal view); D, I, M, S, left G2. Scales: A, D, E, I, J, M, N, O, S = 2.0 mm; B, C, F–H, K, L, P–R = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4778849" httpUri="https://zenodo.org/record/4778849/files/figure.png" pageId="3" pageNumber="303">Fig. 8</figureCitation>
J–L); female sternopleonal cavity condition not known. South 
<collectingCountry id="F35DD3CBFFBB2835FF62F919FEFFA538" box="[207,271,1777,1801]" name="China" pageId="3" pageNumber="303">China</collectingCountry>
Sea and 
<collectingCountry id="F35DD3CBFFBB2835FEDBF919FE02A538" box="[374,498,1777,1801]" name="Philippines" pageId="3" pageNumber="303">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF35C3D3FFBB2835FF25FAE2FE57A76B" ID-DOI="http://doi.org/10.5281/zenodo.4503193" ID-Zenodo-Dep="4503193" httpUri="https://zenodo.org/record/4503193/files/figure.png" pageId="3" pageNumber="303" startId="3.[136,171,1290,1312]" targetBox="[136,1474,189,1275]" targetPageId="3">
<paragraph id="8BF5935BFFBB2835FF25FAE2FE57A76B" blockId="3.[136,1475,1290,1370]" pageId="3" pageNumber="303">
Fig. 2. Overall view. A, 
<taxonomicName id="4C4AE8D8FFBB2835FE2DFAE2FC08A711" authority="Wood-Mason, in Wood-Mason &amp; Alcock, 1891" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[384,1016,1290,1312]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B93E4F49FFBB2835FE2DFAE2FDE2A72E" box="[384,530,1290,1311]" italics="true" pageId="3" pageNumber="303">Parilia alcocki</emphasis>
Wood-Mason, in 
<bibRefCitation id="EFDBEEAAFFBB2835FD64FAE2FC08A711" author="Wood-Mason J &amp; Alcock A" box="[713,1016,1290,1312]" pageId="3" pageNumber="303" pagination="258 - 272" refId="ref11528" refString="Wood-Mason J &amp; Alcock A (1891) Natural History Notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;, Commander R. F. Hoskyn, R. N., commanding. No. 21. On the Results of the last Season's Deepsea Dredging. Annals and Magazine of Natural History, March 1891, (6) 7 (39): 258 - 272. [For an elaboration on the authorship of the taxa described in this series of papers, see HUYS et al., 2014.]" type="journal article" year="1891">Wood-Mason &amp; Alcock, 1891</bibRefCitation>
</taxonomicName>
, female (45.2 × 37.0 mm) (DABFUK), India; B, 
<taxonomicName id="4C4AE8D8FFBB2835FF08FAC0FE88A70C" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[165,376,1319,1341]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B93E4F49FFBB2835FF08FAC0FF0BA70C" box="[165,251,1320,1341]" italics="true" pageId="3" pageNumber="303">P. major</emphasis>
<bibRefCitation id="EFDBEEAAFFBB2835FEAEFACFFE88A70C" author="Sakai T" box="[259,376,1319,1341]" pageId="3" pageNumber="303" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; C, 
<taxonomicName id="4C4AE8D8FFBB2835FC64FAC0FBB7A70C" authority="Ng &amp; Devi &amp; Kumar, 2018" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[969,1095,1320,1341]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="pattersoni" status="sp. nov.">
<emphasis id="B93E4F49FFBB2835FC64FAC0FBB7A70C" box="[969,1095,1320,1341]" italics="true" pageId="3" pageNumber="303">P. pattersoni</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20DF232FFBB2835FBF8FAC0FB3CA70C" box="[1109,1228,1319,1341]" pageId="3" pageNumber="303" rank="species">new species</taxonomicNameLabel>
, paratype female (58.2 × 55.8 mm) (DABFUK), India.
</paragraph>
</caption>
<subSubSection id="C350C0D0FFBB2832FF25F8D9FD45AA10" lastPageId="4" lastPageNumber="304" pageId="3" pageNumber="303" type="discussion">
<paragraph id="8BF5935BFFBB2835FF25F8D9FEBAAA38" blockId="3.[136,775,1841,2057]" pageId="3" pageNumber="303">
<emphasis id="B93E4F49FFBB2835FF25F8D9FF07A578" bold="true" box="[136,247,1841,1865]" pageId="3" pageNumber="303">Remarks.</emphasis>
This is not a common species, with only a few records. The original description was by Chen (1984) from one male and one female specimens from 
<quantity id="4CB23EBEFFBB2835FDDFF899FD4AA5B8" box="[626,698,1905,1929]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="303" unit="m" value="173.0">173 m</quantity>
in the South 
<collectingCountry id="F35DD3CBFFBB2835FF7FF879FEE3A598" box="[210,275,1937,1961]" name="China" pageId="3" pageNumber="303">China</collectingCountry>
Sea, with a subsequent record of a juvenile from the 
<collectingCountry id="F35DD3CBFFBB2835FF5CF859FE99A5F8" box="[241,361,1969,1993]" name="Philippines" pageId="3" pageNumber="303">Philippines</collectingCountry>
by 
<bibRefCitation id="EFDBEEAAFFBB2835FE3AF85AFDE5A5F8" author="Tan CGS" box="[407,533,1969,1993]" pageId="3" pageNumber="303" pagination="1021 - 1058" refId="ref11174" refString="Tan CGS (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30 (7): 1021 - 1058, figs. 1 - 8." type="journal article" year="1996">Tan (1996)</bibRefCitation>
. We here confirm the record from the 
<collectingCountry id="F35DD3CBFFBB2835FE91F839FE44A5D8" box="[316,436,2001,2025]" name="Philippines" pageId="3" pageNumber="303">Philippines</collectingCountry>
with the discovery of an adult male from 
<collectingRegion id="498E5DB9FFBB2835FEACF819FEB6AA38" box="[257,326,2033,2057]" country="Philippines" name="Bohol" pageId="3" pageNumber="303">Bohol</collectingRegion>
.
</paragraph>
<paragraph id="8BF5935BFFBB2832FCEEFA79FD45AA10" blockId="3.[835,1477,1425,2089]" lastBlockId="4.[113,753,1865,2081]" lastPageId="4" lastPageNumber="304" pageId="3" pageNumber="303">
Zarenkov (1990) described 
<taxonomicName id="4C4AE8D8FFBB2835FBDFFA7AFAE2A798" authorityName="Zarenkov" authorityYear="1990" box="[1138,1298,1425,1449]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FBDFFA7AFAE2A798" box="[1138,1298,1425,1449]" italics="true" pageId="3" pageNumber="303">Myra anomala</emphasis>
</taxonomicName>
from one small juvenile male 
<quantity id="4CB23EBEFFBB2835FC77FA59FBC9A7F8" box="[986,1081,1457,1481]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="303" unit="mm" value="12.3">12.3 mm</quantity>
in carapace length from 
<collectingCountry id="F35DD3CBFFBB2835FAEEFA59FA60A7F8" box="[1347,1424,1457,1481]" name="Vietnam" pageId="3" pageNumber="303">Tonkin</collectingCountry>
Bay in northern 
<collectingCountry id="F35DD3CBFFBB2835FC6FFA39FBEEA7D8" box="[962,1054,1489,1513]" name="Vietnam" pageId="3" pageNumber="303">Vietnam</collectingCountry>
at a depth of 
<specimenCount id="9D4C58D2FFBB2835FB19FA39FAF0A7D8" box="[1204,1280,1489,1513]" pageId="3" pageNumber="303" type="generic">
<quantity id="4CB23EBEFFBB2835FB19FA39FB0CA7D8" box="[1204,1276,1489,1513]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="303" unit="m" value="180.0">180 m</quantity>
.
</specimenCount>
<bibRefCitation id="EFDBEEAAFFBB2835FAAAFA39FA32A7D8" author="PariliaInvestigator Galil BS" box="[1287,1474,1489,1513]" pageId="3" pageNumber="303" pagination="395 - 422" refId="ref9922" refString="PariliaInvestigator Galil BS (2001) A revision of Myra Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75 (24): 409 - 446, figs. 1 - 19. Galil BS (2003) Four new genera of leucosiid crabs (Crustacea: Brachyura: Leucosiidae) for three new species and nine species previously in the genus Randallia Stimpson, 1857, with a redescription of the type species, R. ornata (Randall, 1939). Proceedings of the Biological Society of Washington, 116 (2): 395 - 422." type="journal article" year="2001">Galil (2001: 414)</bibRefCitation>
noted that this species “does not belong within 
<taxonomicName id="4C4AE8D8FFBB2835FA90FA1AFA87A438" authorityName="Leach" authorityYear="1817" box="[1341,1399,1522,1545]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFBB2835FA90FA1AFA87A438" box="[1341,1399,1522,1545]" italics="true" pageId="3" pageNumber="303">Myra</emphasis>
</taxonomicName>
, as the male abdomen is depicted with an articulate sixth segment.” All 
<taxonomicName id="4C4AE8D8FFBB2835FCDDF9DAFC5AA478" authorityName="Leach" authorityYear="1817" box="[880,938,1586,1609]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFBB2835FCDDF9DAFC5AA478" box="[880,938,1586,1609]" italics="true" pageId="3" pageNumber="303">Myra</emphasis>
</taxonomicName>
species have male pleonal somite 6 completely fused with somite 5 and their G1 structures are very different (
<bibRefCitation id="EFDBEEAAFFBB2835FCE6F999FC39A4B8" author="PariliaInvestigator Galil BS" box="[843,969,1649,1673]" pageId="3" pageNumber="303" pagination="395 - 422" refId="ref9922" refString="PariliaInvestigator Galil BS (2001) A revision of Myra Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75 (24): 409 - 446, figs. 1 - 19. Galil BS (2003) Four new genera of leucosiid crabs (Crustacea: Brachyura: Leucosiidae) for three new species and nine species previously in the genus Randallia Stimpson, 1857, with a redescription of the type species, R. ornata (Randall, 1939). Proceedings of the Biological Society of Washington, 116 (2): 395 - 422." type="journal article" year="2001">Galil, 2001</bibRefCitation>
). 
<bibRefCitation id="EFDBEEAAFFBB2835FC71F999FB92A4B8" author="PariliaInvestigator Galil BS" box="[988,1122,1649,1673]" pageId="3" pageNumber="303" pagination="395 - 422" refId="ref9922" refString="PariliaInvestigator Galil BS (2001) A revision of Myra Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75 (24): 409 - 446, figs. 1 - 19. Galil BS (2003) Four new genera of leucosiid crabs (Crustacea: Brachyura: Leucosiidae) for three new species and nine species previously in the genus Randallia Stimpson, 1857, with a redescription of the type species, R. ornata (Randall, 1939). Proceedings of the Biological Society of Washington, 116 (2): 395 - 422." type="journal article" year="2001">Galil (2001)</bibRefCitation>
, however, did not reassign 
<taxonomicName id="4C4AE8D8FFBB2835FA25F99AFC51A498" authorityName="Zarenkov" authorityYear="1990" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FA25F99AFC51A498" italics="true" pageId="3" pageNumber="303">Myra anomala</emphasis>
</taxonomicName>
to another genus. 
<bibRefCitation id="EFDBEEAAFFBB2835FBD8F97AFADAA498" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1141,1322,1681,1706]" pageId="3" pageNumber="303" pagination="1 - 286" refId="ref10633" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al. (2008)</bibRefCitation>
synonymised 
<taxonomicName id="4C4AE8D8FFBB2835FCEEF95AFB69A4F8" authority="Zarenkov, 1990" authorityName="Zarenkov" authorityYear="1990" box="[835,1177,1713,1737]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FCEEF95AFC13A4F8" box="[835,995,1713,1737]" italics="true" pageId="3" pageNumber="303">Myra anomala</emphasis>
Zarenkov, 1990
</taxonomicName>
, under 
<taxonomicName id="4C4AE8D8FFBB2835FB46F959FC8DA4D8" authority="Chen, 1984" authorityName="Chen" authorityYear="1984" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBB2835FB46F959FA8AA4F8" box="[1259,1402,1713,1737]" italics="true" pageId="3" pageNumber="303">Parilia ovata</emphasis>
Chen, 1984
</taxonomicName>
, without comment. From Zarenkov’s (1990: 62, fig. 5-5–10) description and figures, 
<taxonomicName id="4C4AE8D8FFBB2835FB1CF91AFAA4A538" authorityName="Zarenkov" authorityYear="1990" box="[1201,1364,1777,1801]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FB1CF91AFAA4A538" box="[1201,1364,1777,1801]" italics="true" pageId="3" pageNumber="303">Myra anomala</emphasis>
</taxonomicName>
is almost certainly a junior synonym of 
<taxonomicName id="4C4AE8D8FFBB2835FB33F8FAFB0CA518" authorityName="Chen" authorityYear="1984" box="[1182,1276,1810,1833]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBB2835FB33F8FAFB0CA518" box="[1182,1276,1810,1833]" italics="true" pageId="3" pageNumber="303">P. ovata</emphasis>
</taxonomicName>
. The carapace of 
<taxonomicName id="4C4AE8D8FFBB2835FCEEF8DAFC38A578" authorityName="Zarenkov" authorityYear="1990" box="[835,968,1841,1865]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FCEEF8DAFC38A578" box="[835,968,1841,1865]" italics="true" pageId="3" pageNumber="303">M. anomala</emphasis>
</taxonomicName>
(cf. Zarenkov, 1990: fig. 5-5) resembles that of 
<taxonomicName id="4C4AE8D8FFBB2835FCC9F8BAFC31A558" authorityName="Chen" authorityYear="1984" box="[868,961,1874,1897]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBB2835FCC9F8BAFC31A558" box="[868,961,1874,1897]" italics="true" pageId="3" pageNumber="303">P. ovata</emphasis>
</taxonomicName>
(
<figureCitation id="13718FDEFFBB2835FC7EF8B9FBD9A558" box="[979,1065,1873,1897]" captionStart="Fig" captionStartId="4.[113,148,1730,1752]" captionTargetBox="[113,1451,189,1715]" captionTargetId="figure-8@4.[113,1451,189,1715]" captionTargetPageId="4" captionText="Fig. 3. Dorsal view of carapace. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="3" pageNumber="303">Fig. 3C</figureCitation>
) except that the lateral tubercles are relatively more prominent which is expected of a juvenile specimen. The exopod of the third maxilliped of 
<taxonomicName id="4C4AE8D8FFBB2835FAEEF87AFA32A598" authorityName="Zarenkov" authorityYear="1990" box="[1347,1474,1937,1961]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FAEEF87AFA32A598" box="[1347,1474,1937,1961]" italics="true" pageId="3" pageNumber="303">M. anomala</emphasis>
</taxonomicName>
(Zarenkov, 1990: fig. 5-9) is not as wide as that of the adult (
<figureCitation id="13718FDEFFBB2835FC24F839FC2AA5D8" box="[905,986,2001,2025]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="3" pageNumber="303">Fig. 5F</figureCitation>
) but this has also been observed for juvenile 
<taxonomicName id="4C4AE8D8FFBB2835FCEEF81AFC43AA38" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[835,947,2033,2057]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B93E4F49FFBB2835FCEEF81AFC43AA38" box="[835,947,2033,2057]" italics="true" pageId="3" pageNumber="303">P. alcocki</emphasis>
</taxonomicName>
in which the structure is proportionately more slender (see earlier). The male pleon of 
<taxonomicName id="4C4AE8D8FFBB2835FAA0F7FAFA64AA18" authorityName="Zarenkov" authorityYear="1990" box="[1293,1428,2065,2089]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBB2835FAA0F7FAFA64AA18" box="[1293,1428,2065,2089]" italics="true" pageId="3" pageNumber="303">M. anomala</emphasis>
</taxonomicName>
has the shape of typical 
<taxonomicName id="4C4AE8D8FFBC2832FEFEF8A1FE6FA550" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[339,415,1865,1889]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFBC2832FEFEF8A1FE6FA550" box="[339,415,1865,1889]" italics="true" pageId="4" pageNumber="304">Parilia</emphasis>
</taxonomicName>
, with somite 6 and telson free and the surfaces unarmed (Zarenkov, 1990: fig. 5-10); and the fingers of the chela are shorter than the palm (Zarenkov, 1990: fig. 5-7). The G1 of 
<taxonomicName id="4C4AE8D8FFBC2832FE3CF842FDE2A5F0" authorityName="Zarenkov" authorityYear="1990" box="[401,530,1961,1985]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBC2832FE3CF842FDE2A5F0" box="[401,530,1961,1985]" italics="true" pageId="4" pageNumber="304">M. anomala</emphasis>
</taxonomicName>
is that of a juvenile, being not distinctly elongate, and the subdistal part appears to be damaged, although it appears that there was a lateral projection at that junction (Zarenkov, 1990: fig. 5-6).
</paragraph>
</subSubSection>
<caption id="DF35C3D3FFBC2832FFDCF92AFB39A523" ID-DOI="http://doi.org/10.5281/zenodo.4503197" ID-Zenodo-Dep="4503197" httpUri="https://zenodo.org/record/4503197/files/figure.png" pageId="4" pageNumber="304" startId="4.[113,148,1730,1752]" targetBox="[113,1451,189,1715]" targetPageId="4">
<paragraph id="8BF5935BFFBC2832FFDCF92AFB39A523" blockId="4.[113,1452,1730,1810]" pageId="4" pageNumber="304">
Fig. 3. Dorsal view of carapace. A, 
<taxonomicName id="4C4AE8D8FFBC2832FE6CF92AFBEFA4E9" authority="Wood-Mason, in Wood-Mason &amp; Alcock, 1891" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[449,1055,1730,1752]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B93E4F49FFBC2832FE6CF92AFDA0A4E6" box="[449,592,1730,1751]" italics="true" pageId="4" pageNumber="304">Parilia alcocki</emphasis>
Wood-Mason, in 
<bibRefCitation id="EFDBEEAAFFBC2832FD51F92AFBEFA4E9" author="Wood-Mason J &amp; Alcock A" box="[764,1055,1730,1752]" pageId="4" pageNumber="304" pagination="258 - 272" refId="ref11528" refString="Wood-Mason J &amp; Alcock A (1891) Natural History Notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;, Commander R. F. Hoskyn, R. N., commanding. No. 21. On the Results of the last Season's Deepsea Dredging. Annals and Magazine of Natural History, March 1891, (6) 7 (39): 258 - 272. [For an elaboration on the authorship of the taxa described in this series of papers, see HUYS et al., 2014.]" type="journal article" year="1891">Wood-Mason &amp; Alcock, 1891</bibRefCitation>
</taxonomicName>
, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, 
<taxonomicName id="4C4AE8D8FFBC2832FF7DF908FE55A4C4" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[208,421,1759,1781]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B93E4F49FFBC2832FF7DF908FED6A4C4" box="[208,294,1760,1781]" italics="true" pageId="4" pageNumber="304">P. major</emphasis>
<bibRefCitation id="EFDBEEAAFFBC2832FE82F937FE55A4C4" author="Sakai T" box="[303,421,1759,1781]" pageId="4" pageNumber="304" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, 
<taxonomicName id="4C4AE8D8FFBC2832FC47F908FB48A4C5" authority="Chen, 1984" authorityName="Chen" authorityYear="1984" box="[1002,1208,1759,1781]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBC2832FC47F908FBCCA4C4" box="[1002,1084,1760,1781]" italics="true" pageId="4" pageNumber="304">P. ovata</emphasis>
Chen, 1984
</taxonomicName>
, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, 
<taxonomicName id="4C4AE8D8FFBC2832FE14F915FDC7A523" authority="Ng &amp; Devi &amp; Kumar, 2018" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[441,567,1789,1810]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="pattersoni" status="sp. nov.">
<emphasis id="B93E4F49FFBC2832FE14F915FDC7A523" box="[441,567,1789,1810]" italics="true" pageId="4" pageNumber="304">P. pattersoni</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20DF232FFBC2832FDE8F914FD4DA523" box="[581,701,1788,1810]" pageId="4" pageNumber="304" rank="species">new species</taxonomicNameLabel>
, holotype male (53.0 × 52.6 mm) (DABFUK), India.
</paragraph>
</caption>
<subSubSection id="C350C0D0FFBC2832FC80F8A1FAF1AA30" pageId="4" pageNumber="304" type="distribution">
<paragraph id="8BF5935BFFBC2832FC80F8A1FAF1AA30" blockId="4.[813,1452,1865,2049]" pageId="4" pageNumber="304">
<emphasis id="B93E4F49FFBC2832FC80F8A1FC4FA550" bold="true" box="[813,959,1865,1889]" pageId="4" pageNumber="304">Distribution.</emphasis>
<taxonomicName id="4C4AE8D8FFBC2832FC65F8A1FBA8A550" authorityName="Chen" authorityYear="1984" box="[968,1112,1865,1889]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B93E4F49FFBC2832FC65F8A1FBA8A550" box="[968,1112,1865,1889]" italics="true" pageId="4" pageNumber="304">Parilia ovata</emphasis>
</taxonomicName>
was described from the South 
<collectingCountry id="F35DD3CBFFBC2832FC80F881FC9DA5B0" box="[813,877,1897,1921]" name="China" pageId="4" pageNumber="304">China</collectingCountry>
Sea (Chen, 1984; Chen &amp; Sun, 2002) but has also been reported from the 
<collectingCountry id="F35DD3CBFFBC2832FB89F861FB6CA590" box="[1060,1180,1929,1953]" name="Philippines" pageId="4" pageNumber="304">Philippines</collectingCountry>
(
<bibRefCitation id="EFDBEEAAFFBC2832FB04F861FAE8A590" author="Tan CGS" box="[1193,1304,1929,1953]" pageId="4" pageNumber="304" pagination="1021 - 1058" refId="ref11174" refString="Tan CGS (1996) Leucosiidae of the Albatross expedition to the Philippines, 1907 - 1910 (Crustacea: Brachyura: Decapoda). Journal of Natural History, 30 (7): 1021 - 1058, figs. 1 - 8." type="journal article" year="1996">Tan, 1996</bibRefCitation>
) and northern 
<collectingCountry id="F35DD3CBFFBC2832FC80F841FC7CA5F0" box="[813,908,1961,1985]" name="Vietnam" pageId="4" pageNumber="304">Vietnam</collectingCountry>
(Zarenkov, 1990, as 
<taxonomicName id="4C4AE8D8FFBC2832FB27F842FAC1A5F0" authorityName="Zarenkov" authorityYear="1990" box="[1162,1329,1961,1985]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="304" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93E4F49FFBC2832FB27F842FAC1A5F0" box="[1162,1329,1961,1985]" italics="true" pageId="4" pageNumber="304">Myra anomala</emphasis>
</taxonomicName>
). The few specimens of this species have been found at depths between 
<quantity id="4CB23EBEFFBC2832FC80F801FC58AA30" box="[813,936,2025,2049]" metricMagnitude="2" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.3" metricValueMin="1.73" pageId="4" pageNumber="304" unit="m" value="201.5" valueMax="230.0" valueMin="173.0">173–230 m</quantity>
(Chen, 1984; Zarenkov, 1990).
</paragraph>
</subSubSection>
</treatment>
</document>