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<document id="01CEA93BDE9D87F21489307D9D63EC06" ID-CLB-Dataset="6550" ID-DOI="10.5281/zenodo.4503189" ID-GBIF-Dataset="44ee08a6-22ce-4f43-be98-4ac76aad2c24" ID-ISSN="2345-7600" ID-Zenodo-Dep="4503189" ID-ZooBank="CBB28174-9B04-4A65-8C7C-5EB4B3608D98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612459944989" checkinUser="carolina" docAuthor="Ng, Peter K. L., Devi, Suvarna &amp; Kumar, Appukuttannair Biju" docDate="2018" docId="03E3224DFFB32827FBA6F843FD16AA0E" docLanguage="en" docName="RafflesBZool.66.300-319.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Neparilia Ng &amp; Devi &amp; Kumar 2018, new genus" docType="treatment" docUuid="B8804A57-EA6B-4271-AA80-A4BD65B13AC0" docUuidSource="ZooBank" docVersion="6" lastPageNumber="317" masterDocId="FFDA5A35FFB82836FFADFFE8FFF0A231" masterDocTitle="The genus Parilia Wood-Mason, in Wood-Mason &amp; Alcock, 1891, with description of a new species and establishment of a new genus for P. tuberculata Sakai, 1961 (Crustacea, Brachyura, Leucosiidae)" masterLastPageNumber="319" masterPageNumber="300" pageNumber="311" updateTime="1698926984719" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8760E39D7404B7FB0F0A50A9E05CD7C2">The genus Parilia Wood-Mason, in Wood-Mason &amp; Alcock, 1891, with description of a new species and establishment of a new genus for P. tuberculata Sakai, 1961 (Crustacea, Brachyura, Leucosiidae)</mods:title>
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<mods:namePart id="638E1B0D37BF7D8FA288B5D50E735006">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="0256CC50519E6158ECB9894CDAAD2534">Devi, Suvarna</mods:namePart>
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<mods:namePart id="9FFF27DA8F2F431AC4EF617D1BBA0D96">Kumar, Appukuttannair Biju</mods:namePart>
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<mods:title id="7B37133F59C9FBF83D8952F9DF88413F">Raffles Bulletin of Zoology</mods:title>
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<treatment id="03E3224DFFB32827FBA6F843FD16AA0E" ID-DOI="http://doi.org/10.5281/zenodo.4502853" ID-GBIF-Taxon="182253769" ID-Zenodo-Dep="4502853" LSID="urn:lsid:zoobank.org:act:B8804A57-EA6B-4271-AA80-A4BD65B13AC0" httpUri="http://treatment.plazi.org/id/03E3224DFFB32827FBA6F843FD16AA0E" lastPageId="17" lastPageNumber="317" pageId="11" pageNumber="311">
<subSubSection id="C350C0D0FFB3283DFBA6F843FB0AA5F2" box="[1035,1274,1963,1987]" pageId="11" pageNumber="311" type="nomenclature">
<paragraph id="8BF5935BFFB3283DFBA6F843FB0AA5F2" blockId="11.[1035,1274,1963,1987]" box="[1035,1274,1963,1987]" pageId="11" pageNumber="311">
<heading id="D0BD2437FFB3283DFBA6F843FB0AA5F2" bold="true" box="[1035,1274,1963,1987]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="311" reason="2">
<emphasis id="B93E4F49FFB3283DFBA6F843FB0AA5F2" bold="true" box="[1035,1274,1963,1987]" pageId="11" pageNumber="311">
<taxonomicName id="4C4AE8D8FFB3283DFBA6F843FB83A5F2" ID-CoL="847XH" authority="Ng &amp; Devi &amp; Kumar, 2018" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[1035,1139,1963,1987]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93E4F49FFB3283DFBA6F843FB83A5F2" bold="true" box="[1035,1139,1963,1987]" italics="true" pageId="11" pageNumber="311">Neparilia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20DF232FFB3283DFB2FF843FB0AA5F2" box="[1154,1274,1963,1987]" pageId="11" pageNumber="311" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFB3283AFCEEF803FB38AA10" lastPageId="12" lastPageNumber="312" pageId="11" pageNumber="311" type="diagnosis">
<paragraph id="8BF5935BFFB3283AFCEEF803FB38AA10" blockId="11.[835,1474,2027,2115]" lastBlockId="12.[813,1452,1864,2081]" lastPageId="12" lastPageNumber="312" pageId="11" pageNumber="311">
<emphasis id="B93E4F49FFB3283DFCEEF803FC4BAA32" bold="true" box="[835,955,2027,2051]" pageId="11" pageNumber="311">Diagnosis.</emphasis>
Carapace ovate; dorsal surface covered large rounded tubercles and granules. Cardiac region with low rounded tubercle; intestinal region with 2 short spines. Exopod of third maxilliped broad, distinctly wider than ischium and merus, with outer margin strongly convex; coxa wide, not semicircular in shape, not bracketing base of exopod. Suborbital region prominently compressed by expanded buccal cavity. Adult male chelipeds about 2 times longer than carapace length; chela not elongate, fingers longer than palm, in large male individuals, closing tightly. Male pleon with somites 1 and 2 free, somites 35 completely fused, somite 6 separated from somite 5 by suture but immobile, telson free; distal and proximal surfaces of somite 6 each with a distinct cup-shaped tubercle, that on distal part larger; telson elongate, linguiform, longer than somite 6. G1 slender, distal part tapering without lateral projection or flanges. G2 distal segment as long as basal segment, with curved flagelliform tip.
</paragraph>
</subSubSection>
<caption id="DF35C3D3FFB4283AFFDCF937FE3AA520" ID-DOI="http://doi.org/10.5281/zenodo.4503203" ID-Zenodo-Dep="4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="12" pageNumber="312" startId="12.[113,148,1759,1781]" targetBox="[113,1451,189,1743]" targetPageId="12">
<paragraph id="8BF5935BFFB4283AFFDCF937FE3AA520" blockId="12.[113,1452,1759,1809]" pageId="12" pageNumber="312">
Fig. 10.
<taxonomicName id="4C4AE8D8FFB4283AFF6BF937FDD6A4C4" authority="(Sakai, 1961)" baseAuthorityName="Sakai" baseAuthorityYear="1961" box="[198,550,1759,1781]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFB4283AFF6BF937FE69A4C5" box="[198,409,1759,1780]" italics="true" pageId="12" pageNumber="312">Neparilia tuberculata</emphasis>
(
<bibRefCitation id="EFDBEEAAFFB4283AFE05F937FDD0A4C4" author="Sakai T" box="[424,544,1759,1781]" pageId="12" pageNumber="312" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
)
</taxonomicName>
, male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped.
</paragraph>
</caption>
<caption id="DF35C3D3FFB5283BFF25F876FE0EA5DF" ID-DOI="http://doi.org/10.5281/zenodo.4503205" ID-Zenodo-Dep="4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="13" pageNumber="313" startId="13.[136,171,1950,1972]" targetBox="[136,1475,273,1935]" targetPageId="13">
<paragraph id="8BF5935BFFB5283BFF25F876FE0EA5DF" blockId="13.[136,1474,1950,2030]" pageId="13" pageNumber="313">
Fig. 11.
<taxonomicName id="4C4AE8D8FFB5283BFF76F876FDC9A585" authority="(Sakai, 1961)" baseAuthorityName="Sakai" baseAuthorityYear="1961" box="[219,569,1950,1972]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="313" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFB5283BFF76F876FE5DA582" box="[219,429,1950,1971]" italics="true" pageId="13" pageNumber="313">Neparilia tuberculata</emphasis>
(
<bibRefCitation id="EFDBEEAAFFB5283BFE16F876FDC3A585" author="Sakai T" box="[443,563,1950,1972]" pageId="13" pageNumber="313" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
)
</taxonomicName>
, male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela.
</paragraph>
</caption>
<caption id="DF35C3D3FFB62838FFDCFBCCFAD0A667" ID-DOI="http://doi.org/10.5281/zenodo.4503207" ID-Zenodo-Dep="4503207" httpUri="https://zenodo.org/record/4503207/files/figure.png" pageId="14" pageNumber="314" startId="14.[113,148,1060,1082]" targetBox="[93,1481,174,1055]" targetPageId="14">
<paragraph id="8BF5935BFFB62838FFDCFBCCFAD0A667" blockId="14.[113,1451,1059,1110]" pageId="14" pageNumber="314">
Fig. 12.
<taxonomicName id="4C4AE8D8FFB62838FF68FBCCFDD5A60B" authority="(Sakai, 1961)" baseAuthorityName="Sakai" baseAuthorityYear="1961" box="[197,549,1060,1082]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFB62838FF68FBCCFE69A608" box="[197,409,1060,1081]" italics="true" pageId="14" pageNumber="314">Neparilia tuberculata</emphasis>
(
<bibRefCitation id="EFDBEEAAFFB62838FE0AFBCCFDEFA60B" author="Sakai T" box="[423,543,1060,1082]" pageId="14" pageNumber="314" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
)
</taxonomicName>
, male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales: A, D = 1.0 mm; B, C = 0.5 mm.
</paragraph>
</caption>
<subSection id="E2C58AB0FFB62827FFDCFB65FD16AA0E" lastPageId="17" lastPageNumber="317" pageId="14" pageNumber="314" type="multiple">
<subSubSection id="C350C0D0FFB62838FFDCFB65FE89A6F4" pageId="14" pageNumber="314" type="type_taxon">
<paragraph id="8BF5935BFFB62838FFDCFB65FE89A6F4" blockId="14.[113,752,1165,1221]" pageId="14" pageNumber="314">
<emphasis id="B93E4F49FFB62838FFDCFB65FEF6A694" bold="true" box="[113,262,1165,1189]" pageId="14" pageNumber="314">
<typeStatus id="54F12DF9FFB62838FFDCFB65FF5BA694" box="[113,171,1165,1189]" pageId="14" pageNumber="314">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C4AE8D8FFB62838FEA6FB65FDABA694" ID-CoL="7XYDG" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[267,603,1165,1189]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFB62838FEA6FB65FE27A694" box="[267,471,1165,1189]" italics="true" pageId="14" pageNumber="314">Parilia tuberculata</emphasis>
<bibRefCitation id="EFDBEEAAFFB62838FE71FB65FDABA694" author="Sakai T" box="[476,603,1165,1189]" pageId="14" pageNumber="314" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
, by monotypy and present designation.
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFB62838FFDCFB05FE7AA774" pageId="14" pageNumber="314" type="etymology">
<paragraph id="8BF5935BFFB62838FFDCFB05FE7AA774" blockId="14.[113,752,1261,1349]" pageId="14" pageNumber="314">
<emphasis id="B93E4F49FFB62838FFDCFB05FF04A734" bold="true" box="[113,244,1261,1285]" pageId="14" pageNumber="314">Etymology.</emphasis>
From the Latin “ne-” for not, alluding to the present realisation that the
<typeStatus id="54F12DF9FFB62838FE3AFAE6FE35A717" box="[407,453,1294,1318]" pageId="14" pageNumber="314">type</typeStatus>
species is not a member of
<taxonomicName id="4C4AE8D8FFB62838FFDCFAC5FF4DA774" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[113,189,1325,1349]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FFDCFAC5FF4DA774" box="[113,189,1325,1349]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C350C0D0FFB62827FFDCFA85FD16AA0E" lastPageId="17" lastPageNumber="317" pageId="14" pageNumber="314" type="discussion">
<paragraph id="8BF5935BFFB62838FFDCFA85FBC6A777" blockId="14.[113,754,1389,2117]" lastBlockId="14.[813,1452,1166,1350]" pageId="14" pageNumber="314">
<emphasis id="B93E4F49FFB62838FFDCFA85FF2FA7B4" bold="true" box="[113,223,1389,1413]" pageId="14" pageNumber="314">Remarks.</emphasis>
<bibRefCitation id="EFDBEEAAFFB62838FF4EFA85FE99A7B4" author="Sakai T" box="[227,361,1389,1413]" pageId="14" pageNumber="314" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai (1961)</bibRefCitation>
placed this species in
<taxonomicName id="4C4AE8D8FFB62838FDE0FA85FD67A7B4" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[589,663,1389,1413]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FDE0FA85FD67A7B4" box="[589,663,1389,1413]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
because it possessed a very broad exopod of the third maxilliped. It is, however, not possible to retain
<taxonomicName id="4C4AE8D8FFB62838FE4BFA46FD70A7F4" authorityName="Sakai" authorityYear="1961" box="[486,640,1453,1477]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B93E4F49FFB62838FE4BFA46FD70A7F4" box="[486,640,1453,1477]" italics="true" pageId="14" pageNumber="314">P. tuberculata</emphasis>
</taxonomicName>
in
<taxonomicName id="4C4AE8D8FFB62838FD0BFA45FD02A7F4" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[678,754,1453,1477]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FD0BFA45FD02A7F4" box="[678,754,1453,1477]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
because the coxa of the third maxilliped is not substantially laterally expanded or semicircular (
<figureCitation id="13718FDEFFB62838FE49FA05FDBDA434" box="[484,589,1517,1541]" captionStart="Fig" captionStartId="12.[113,148,1759,1781]" captionTargetBox="[113,1451,189,1743]" captionTargetId="figure-8@12.[113,1451,189,1743]" captionTargetPageId="12" captionText="Fig. 10. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="14" pageNumber="314">Figs. 10C</figureCitation>
,
<figureCitation id="13718FDEFFB62838FDFAFA05FD74A434" box="[599,644,1517,1541]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">11B</figureCitation>
) (vs. coxa prominently expanded laterally, appearing semicircular in
<taxonomicName id="4C4AE8D8FFB62838FFDCF9C5FF4CA474" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[113,188,1581,1605]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FFDCF9C5FF4CA474" box="[113,188,1581,1605]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FF60F9C5FEEAA474" box="[205,282,1581,1605]" captionStart="Fig" captionStartId="5.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1685]" captionTargetId="figure-8@5.[136,1474,189,1685]" captionText="Fig. 4. Frontal view of cephalothorax and third maxillipeds. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778829" httpUri="https://zenodo.org/record/4778829/files/figure.png" pageId="14" pageNumber="314">Figs. 4</figureCitation>
,
<figureCitation id="13718FDEFFB62838FE85F9C5FEC7A474" box="[296,311,1581,1605]" captionStart="Fig" captionStartId="7.[136,171,1910,1932]" captionTargetBox="[163,1448,189,1893]" captionTargetId="figure-8@7.[163,1448,189,1893]" captionText="Fig. 5. Right third maxilliped. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, young male (24.8 × 23.4 mm) (ZRC 2017.216), India; C, P. alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India; H, P. pattersoni, new species, paratype male (72.5 × 69.8 mm) (DABFUK), India; I, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778837" httpUri="https://zenodo.org/record/4778837/files/figure.png" pageId="14" pageNumber="314">5</figureCitation>
); the adult male chelipeds are relatively short (about 2 times carapace length), the chela are not elongate with the fingers longer than the palm (
<figureCitation id="13718FDEFFB62838FDD0F985FD1CA4B4" box="[637,748,1645,1669]" captionStart="Fig" captionStartId="12.[113,148,1759,1781]" captionTargetBox="[113,1451,189,1743]" captionTargetId="figure-8@12.[113,1451,189,1743]" captionTargetPageId="12" captionText="Fig. 10. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="14" pageNumber="314">Figs. 10A</figureCitation>
,
<figureCitation id="13718FDEFFB62838FFDCF965FF51A494" box="[113,161,1677,1701]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">11G</figureCitation>
) (vs. adult male chelipeds more than 3 times longer than carapace length, chela prominently elongate with fingers much shorter than palm in
<taxonomicName id="4C4AE8D8FFB62838FE3AF925FE13A4D4" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[407,483,1741,1765]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FE3AF925FE13A4D4" box="[407,483,1741,1765]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FE5FF925FDCBA4D4" box="[498,571,1741,1765]" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="14" pageNumber="314">Figs. 1</figureCitation>
,
<figureCitation id="13718FDEFFB62838FDE6F925FDABA4D4" box="[587,603,1741,1765]" captionStart="Fig" captionStartId="8.[113,148,1410,1432]" captionTargetBox="[113,1451,189,1393]" captionTargetId="figure-279@8.[113,1451,189,1393]" captionText="Fig. 6. Outer view of right chela. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778841" httpUri="https://zenodo.org/record/4778841/files/figure.png" pageId="14" pageNumber="314">6</figureCitation>
); the surfaces of the ambulatory meri are covered with short, sharp spines (
<figureCitation id="13718FDEFFB62838FFD4F8E5FF15A514" box="[121,229,1805,1829]" captionStart="Fig" captionStartId="12.[113,148,1759,1781]" captionTargetBox="[113,1451,189,1743]" captionTargetId="figure-8@12.[113,1451,189,1743]" captionTargetPageId="12" captionText="Fig. 10. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="14" pageNumber="314">Figs. 10A</figureCitation>
,
<figureCitation id="13718FDEFFB62838FF5DF8E5FECEA514" box="[240,318,1805,1829]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">11C, D</figureCitation>
) (vs. surface of ambulatory meri smooth or covered with granules, never spiniform in
<taxonomicName id="4C4AE8D8FFB62838FDCDF8C5FD5CA574" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[608,684,1837,1861]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FDCDF8C5FD5CA574" box="[608,684,1837,1861]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FD11F8C5FF8FA554" captionStart="Fig" captionStartId="2.[113,148,1269,1291]" captionTargetBox="[113,1451,189,1253]" captionTargetId="figure-9@2.[113,1451,189,1253]" captionTargetPageId="2" captionText="Fig. 1. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; B, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; C, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; D, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503191" httpUri="https://zenodo.org/record/4503191/files/figure.png" pageId="14" pageNumber="314">Figs. 1</figureCitation>
,
<figureCitation id="13718FDEFFB62838FF3DF8A5FF50A554" box="[144,160,1869,1893]" captionStart="Fig" captionStartId="3.[136,171,1290,1312]" captionTargetBox="[136,1474,189,1275]" captionTargetId="figure-8@3.[136,1474,189,1275]" captionTargetPageId="3" captionText="Fig. 2. Overall view. A, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, female (45.2 × 37.0 mm) (DABFUK), India; B, P. major Sakai, 1961, female (48.3 × 49.2 mm) (ZRC 2001.552), Philippines; C, P. pattersoni, new species, paratype female (58.2 × 55.8 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4503193" httpUri="https://zenodo.org/record/4503193/files/figure.png" pageId="14" pageNumber="314">2</figureCitation>
); the male pleonal somites 5 and 6 are functionally fused even though there is a distinct suture separating them and somite 6 has a cup-like tubercle on both the distal and proximal surfaces, with the distal one more prominent (
<figureCitation id="13718FDEFFB62838FD68F845FF28A5D4" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">Fig. 11C, E, F</figureCitation>
) (vs. male pleonal somites 5 and 6 mobile, surface of somite 6 smooth in
<taxonomicName id="4C4AE8D8FFB62838FEC4F805FE45AA34" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[361,437,2029,2053]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FEC4F805FE45AA34" box="[361,437,2029,2053]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FE68F805FD8DAA34" box="[453,637,2029,2053]" captionStart="Fig" captionStartId="9.[135,170,950,972]" captionTargetBox="[136,1474,189,933]" captionTargetId="figure-310@9.[136,1474,189,933]" captionText="Fig. 7. Male thoracic sternum and pleon. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778845" httpUri="https://zenodo.org/record/4778845/files/figure.png" pageId="14" pageNumber="314">Fig. 7A, C, E, G</figureCitation>
); the male telson is linguiform and much longer than somite 6 (
<figureCitation id="13718FDEFFB62838FD68F7E5FF4EAA74" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">Fig. 11C, E</figureCitation>
) (vs. telson triangular, subequal in length to somite
<quantity id="4CB23EBEFFB62838FC80FB66FCAAA697" box="[813,858,1166,1190]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="14" pageNumber="314" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="4C4AE8D8FFB62838FCCBFB66FC45A697" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[870,949,1166,1190]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FCCBFB66FC45A697" box="[870,949,1166,1190]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FC6AFB66FB7FA697" box="[967,1167,1166,1190]" captionStart="Fig" captionStartId="9.[135,170,950,972]" captionTargetBox="[136,1474,189,933]" captionTargetId="figure-310@9.[136,1474,189,933]" captionText="Fig. 7. Male thoracic sternum and pleon. A, B, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; C, D, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; E, F, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; G, H, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India." figureDoi="http://doi.org/10.5281/zenodo.4778845" httpUri="https://zenodo.org/record/4778845/files/figure.png" pageId="14" pageNumber="314">Fig. 7A, C, E, G</figureCitation>
); and the G1 is straight, without a subdistal lateral projection, not flared and lacks serrulate flanges distally (
<figureCitation id="13718FDEFFB62838FB92FB26FB7AA6D7" box="[1087,1162,1230,1254]" captionStart="Fig" captionStartId="14.[113,148,1060,1082]" captionTargetBox="[93,1481,174,1055]" captionTargetId="figure-566@14.[93,1481,174,1055]" captionTargetPageId="14" captionText="Fig. 12. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4503207" httpUri="https://zenodo.org/record/4503207/files/figure.png" pageId="14" pageNumber="314">Fig. 12</figureCitation>
AC) (vs. distal part of G1 bifurcated, with prominent lateral projection, each projection fared with serrulate or denticulate flanges in
<taxonomicName id="4C4AE8D8FFB62838FA8EFAE6FA9FA717" authorityName="Wood-Mason, in Wood-Mason &amp; Alcock" authorityYear="1891" box="[1315,1391,1294,1318]" class="Malacostraca" family="Leucosiidae" genus="Parilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FA8EFAE6FA9FA717" box="[1315,1391,1294,1318]" italics="true" pageId="14" pageNumber="314">Parilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FA2CFAE6FCC9A777" captionStart="Fig" captionStartId="10.[113,148,1974,1996]" captionTargetBox="[115,1453,190,1952]" captionTargetId="figure-8@10.[99,1459,176,1964]" captionText="Fig. 8. Gonopods. AD, Parilia alcocki Wood-Mason, in Wood-Mason &amp; Alcock, 1891, male (66.1 × 57.4 mm) (ZRC 2017.218), India; EI, P. major Sakai, 1961, male (70.0 × 69.0 mm) (ZRC 2001.552), Philippines; JM, P. ovata Chen, 1984, male (31.6 × 43.3 mm) (ZRC 2017.219), Philippines; NS, P. pattersoni, new species, holotype male (53.0 × 52.6 mm) (DABFUK), India. A, E, J, N, left G1 (ventral view); O, left G1 (dorsal view); B, F, K, P, distal part of left G1 (ventral view); C, G, H, L, Q, R, distal part of left G1 (dorsal view); D, I, M, S, left G2. Scales: A, D, E, I, J, M, N, O, S = 2.0 mm; B, C, FH, K, L, PR = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4778849" httpUri="https://zenodo.org/record/4778849/files/figure.png" pageId="14" pageNumber="314">Fig. 8</figureCitation>
AC, EH, JL, NR).
</paragraph>
<paragraph id="8BF5935BFFB62827FC80FA86FEE8A40E" blockId="14.[811,1452,1390,2118]" lastBlockId="17.[136,775,1254,1599]" lastPageId="17" lastPageNumber="317" pageId="14" pageNumber="314">
The carapace and general external features of
<taxonomicName id="4C4AE8D8FFB62838FAEFFA86FA5CA7B7" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[1346,1452,1390,1414]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FAEFFA86FA5CA7B7" box="[1346,1452,1390,1414]" italics="true" pageId="14" pageNumber="314">Neparilia</emphasis>
</taxonomicName>
resemble
<taxonomicName id="4C4AE8D8FFB62838FC3EFA66FAEBA797" authority="Ng &amp; Richer de Forges, 2007" authorityName="Ng &amp; Richer de Forges" authorityYear="2007" box="[915,1307,1422,1446]" class="Malacostraca" family="Leucosiidae" genus="Galilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FC3EFA66FC2FA797" box="[915,991,1422,1446]" italics="true" pageId="14" pageNumber="314">Galilia</emphasis>
<bibRefCitation id="EFDBEEAAFFB62838FC49FA66FAEBA797" author="Ng PKL &amp; Richer de Forges B" box="[996,1307,1422,1446]" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54F12DF9FFB62838FA84FA66FAA8A797" box="[1321,1368,1422,1446]" pageId="14" pageNumber="314">type</typeStatus>
species
<taxonomicName id="4C4AE8D8FFB62838FC80FA46FAEFA7F7" authority="Ng &amp; Richer de Forges, 2007" authorityName="Ng &amp; Richer de Forges" authorityYear="2007" box="[813,1311,1454,1478]" class="Malacostraca" family="Leucosiidae" genus="Galilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="narusei">
<emphasis id="B93E4F49FFB62838FC80FA46FC21A7F4" box="[813,977,1454,1478]" italics="true" pageId="14" pageNumber="314">Galilia narusei</emphasis>
<bibRefCitation id="EFDBEEAAFFB62838FC74FA46FAEFA7F7" author="Ng PKL &amp; Richer de Forges B" box="[985,1311,1454,1478]" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
</taxonomicName>
), but several features easily distinguish them. In
<taxonomicName id="4C4AE8D8FFB62838FB32FA26FB1AA7D7" authorityName="Ng &amp; Richer de Forges" authorityYear="2007" box="[1183,1258,1486,1510]" class="Malacostraca" family="Leucosiidae" genus="Galilia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FB32FA26FB1AA7D7" box="[1183,1258,1486,1510]" italics="true" pageId="14" pageNumber="314">Galilia</emphasis>
</taxonomicName>
, the exopod of the third maxilliped is transversely narrower than the endopod (
<bibRefCitation id="EFDBEEAAFFB62838FC99F9E6FB87A417" author="Ng PKL &amp; Richer de Forges B" box="[820,1143,1550,1574]" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 2E;
<bibRefCitation id="EFDBEEAAFFB62838FB7AF9E6FC96A477" author="Komai T &amp; Tsuchida S" pageId="14" pageNumber="314" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: figs. 19B, 21A) (vs. exopod is distinctly broader than the endopod in
<taxonomicName id="4C4AE8D8FFB62838FC73F9A6FBB6A457" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[990,1094,1614,1638]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FC73F9A6FBB6A457" box="[990,1094,1614,1638]" italics="true" pageId="14" pageNumber="314">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FBF5F9A6FB36A457" box="[1112,1222,1614,1638]" captionStart="Fig" captionStartId="12.[113,148,1759,1781]" captionTargetBox="[113,1451,189,1743]" captionTargetId="figure-8@12.[113,1451,189,1743]" captionTargetPageId="12" captionText="Fig. 10. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="14" pageNumber="314">Figs. 10C</figureCitation>
,
<figureCitation id="13718FDEFFB62838FB78F9A6FAF5A457" box="[1237,1285,1614,1638]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">11B</figureCitation>
); the intestinal region has two large rounded tubercles (
<bibRefCitation id="EFDBEEAAFFB62838FB59F986FC4EA497" author="Ng PKL &amp; Richer de Forges B" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 2F;
<bibRefCitation id="EFDBEEAAFFB62838FB89F966FACEA497" author="Komai T &amp; Tsuchida S" box="[1060,1342,1678,1702]" pageId="14" pageNumber="314" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: fig. 19A, C) (vs. intestinal region with two short spines in
<taxonomicName id="4C4AE8D8FFB62838FA93F946FA54A4F7" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[1342,1444,1710,1734]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FA93F946FA54A4F7" box="[1342,1444,1710,1734]" italics="true" pageId="14" pageNumber="314">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FC80F926FC6BA4D7" box="[813,923,1742,1766]" captionStart="Fig" captionStartId="12.[113,148,1759,1781]" captionTargetBox="[113,1451,189,1743]" captionTargetId="figure-8@12.[113,1451,189,1743]" captionTargetPageId="12" captionText="Fig. 10. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, overall view; B, dorsal view of carapace; C, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4503203" httpUri="https://zenodo.org/record/4503203/files/figure.png" pageId="14" pageNumber="314">Figs. 10B</figureCitation>
,
<figureCitation id="13718FDEFFB62838FC04F926FC2BA4D7" box="[937,987,1742,1766]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">11D</figureCitation>
); the fingers of the chela are shorter than the palm (
<bibRefCitation id="EFDBEEAAFFB62838FC37F906FB10A537" author="Ng PKL &amp; Richer de Forges B" box="[922,1248,1774,1798]" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 3DF;
<bibRefCitation id="EFDBEEAAFFB62838FACEF906FC09A517" author="Komai T &amp; Tsuchida S" pageId="14" pageNumber="314" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: fig. 20A) (vs. fingers distinctly longer than the palm in
<taxonomicName id="4C4AE8D8FFB62838FC42F8C6FBA6A577" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[1007,1110,1838,1862]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FC42F8C6FBA6A577" box="[1007,1110,1838,1862]" italics="true" pageId="14" pageNumber="314">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FBC5F8C6FB3EA577" box="[1128,1230,1838,1862]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">Fig. 11G</figureCitation>
); the male telson is prominently elongate with the male sternopleonal cavity reaching to the suture between thoracic sternites 2 and 3 (
<bibRefCitation id="EFDBEEAAFFB62838FA27F886FBBCA597" author="Ng PKL &amp; Richer de Forges B" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 3A, B;
<bibRefCitation id="EFDBEEAAFFB62838FB78F866FC99A5F7" author="Komai T &amp; Tsuchida S" pageId="14" pageNumber="314" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: figs. 19F, 21B) (vs. male telson proportionately shorter and the male sternopleonal cavity reaches only to the midpoint of sternite
<quantity id="4CB23EBEFFB62838FB90F806FB9AAA37" box="[1085,1130,2030,2054]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="314" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C4AE8D8FFB62838FBD9F806FB2BAA37" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[1140,1243,2030,2054]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFB62838FBD9F806FB2BAA37" box="[1140,1243,2030,2054]" italics="true" pageId="14" pageNumber="314">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFB62838FB40F806FA83AA37" box="[1261,1395,2030,2054]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="14" pageNumber="314">Fig. 11C, E</figureCitation>
); the surface of male pleonal somite 6 has a truncate tubercle on the distal margin (
<bibRefCitation id="EFDBEEAAFFB62838FC55F7C6FAB9AA77" author="Ng PKL &amp; Richer de Forges B" box="[1016,1353,2094,2118]" pageId="14" pageNumber="314" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 3A, B;
<bibRefCitation id="EFDBEEAAFFA92827FF07FB0EFE32A6CE" author="Komai T &amp; Tsuchida S" box="[170,450,1254,1279]" pageId="17" pageNumber="317" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: fig. 19F) (vs. the surface has a cup-like tubercle on both the proximal and distal surfaces in
<taxonomicName id="4C4AE8D8FFA92827FF0AFACEFEFEA70F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[167,270,1318,1342]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FF0AFACEFEFEA70F" box="[167,270,1318,1342]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFA92827FE8DFACEFE56A70E" box="[288,422,1318,1343]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="17" pageNumber="317">Fig. 11C, E</figureCitation>
); male pleonal somites 5 and 6 are completely fused, separated at most by a shallow suture (
<bibRefCitation id="EFDBEEAAFFA92827FF3DFA8FFE21A74E" author="Ng PKL &amp; Richer de Forges B" box="[144,465,1382,1407]" pageId="17" pageNumber="317" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFDBEEAAFFA92827FD99FA8EFF31A7AE" author="Komai T &amp; Tsuchida S" pageId="17" pageNumber="317" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: fig. 19F) (vs. somites 5 and 6 are free and separated by a deep suture in
<taxonomicName id="4C4AE8D8FFA92827FEC9FA4EFE3AA78F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[356,458,1446,1470]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FEC9FA4EFE3AA78F" box="[356,458,1446,1470]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFA92827FE76FA4EFDADA78E" box="[475,605,1446,1471]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="17" pageNumber="317">Fig. 11C, E</figureCitation>
); and the distal third of the G1 is prominently bent (
<bibRefCitation id="EFDBEEAAFFA92827FDAAFA2FFF32A7CE" author="Ng PKL &amp; Richer de Forges B" pageId="17" pageNumber="317" pagination="15 - 24" refId="ref10734" refString="Ng PKL &amp; Richer de Forges B (2007) A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa, 1662: 15 - 24." type="journal article" year="2007">Ng &amp; Richer de Forges, 2007</bibRefCitation>
: fig. 4G, H;
<bibRefCitation id="EFDBEEAAFFA92827FEF9FA0EFD84A7CE" author="Komai T &amp; Tsuchida S" box="[340,628,1510,1535]" pageId="17" pageNumber="317" pagination="1 - 46" refId="ref10444" refString="Komai T &amp; Tsuchida S (2014) Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;. Zootaxa, 3764: 279 - 316. Krishnamoorthy P (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of the Zoological Survey of India, Occasional Paper, 304: 1 - 46." type="journal article" year="2014">Komai &amp; Tsuchida, 2014</bibRefCitation>
: fig. 21C, D; Shih et al., 2013: fig. 3A, B) (vs. G1 straight in
<taxonomicName id="4C4AE8D8FFA92827FD34F9EEFD0FA42F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[665,767,1542,1566]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FD34F9EEFD0FA42F" box="[665,767,1542,1566]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFA92827FF25F9CEFF25A40E" box="[136,213,1574,1599]" captionStart="Fig" captionStartId="14.[113,148,1060,1082]" captionTargetBox="[93,1481,174,1055]" captionTargetId="figure-566@14.[93,1481,174,1055]" captionTargetPageId="14" captionText="Fig. 12. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4503207" httpUri="https://zenodo.org/record/4503207/files/figure.png" pageId="17" pageNumber="317">Fig. 12</figureCitation>
AC).
</paragraph>
<caption id="DF35C3D3FFB72839FF25F824FA33A5CF" ID-DOI="http://doi.org/10.5281/zenodo.4503213" ID-Zenodo-Dep="4503213" httpUri="https://zenodo.org/record/4503213/files/figure.png" pageId="15" pageNumber="315" startId="15.[136,171,1996,2018]" targetBox="[136,1474,253,1980]" targetPageId="15">
<paragraph id="8BF5935BFFB72839FF25F824FA33A5CF" blockId="15.[136,1475,1996,2046]" pageId="15" pageNumber="315">Fig. 13. Freshly collected male (not available for examination) from southern Philippines by LUMIWAN 2008 cruise. A, overall view; B, C, frontal view of cephalothorax D, ventral view showing third maxillipeds and anterior thoracic sternum. Photographs by T.-Y. Chan.</paragraph>
</caption>
<caption id="DF35C3D3FFA82826FFDCF876FBEEA5DF" ID-DOI="http://doi.org/10.5281/zenodo.4503215" ID-Zenodo-Dep="4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="16" pageNumber="316" startId="16.[113,148,1950,1972]" targetBox="[113,1451,274,1935]" targetPageId="16">
<paragraph id="8BF5935BFFA82826FFDCF876FBEEA5DF" blockId="16.[113,1452,1950,2030]" pageId="16" pageNumber="316">
Fig. 14.
<taxonomicName id="4C4AE8D8FFA82826FF68F876FDD7A585" authority="(Sakai, 1961)" baseAuthorityName="Sakai" baseAuthorityYear="1961" box="[197,551,1950,1972]" class="Malacostraca" family="Leucosiidae" genus="Urashima" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="316" phylum="Arthropoda" rank="species" species="pustuloides">
<emphasis id="B93E4F49FFA82826FF68F876FE6AA582" box="[197,410,1950,1971]" italics="true" pageId="16" pageNumber="316">Urashima pustuloides</emphasis>
(
<bibRefCitation id="EFDBEEAAFFA82826FE04F876FDD0A585" author="Sakai T" box="[425,544,1950,1972]" pageId="16" pageNumber="316" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
)
</taxonomicName>
, male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela.
</paragraph>
</caption>
<caption id="DF35C3D3FFA92827FF25FB88FD39A69E" ID-DOI="http://doi.org/10.5281/zenodo.4503219" ID-Zenodo-Dep="4503219" httpUri="https://zenodo.org/record/4503219/files/figure.png" pageId="17" pageNumber="317" startId="17.[136,171,1120,1142]" targetBox="[141,1474,193,1105]" targetPageId="17">
<paragraph id="8BF5935BFFA92827FF25FB88FD39A69E" blockId="17.[136,1474,1120,1199]" pageId="17" pageNumber="317">
Fig. 15.
<taxonomicName id="4C4AE8D8FFA92827FF71FB88FDCEA647" authority="(Sakai, 1961)" baseAuthorityName="Sakai" baseAuthorityYear="1961" box="[220,574,1120,1142]" class="Malacostraca" family="Leucosiidae" genus="Urashima" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="species" species="pustuloides">
<emphasis id="B93E4F49FFA92827FF71FB88FE41A644" box="[220,433,1120,1141]" italics="true" pageId="17" pageNumber="317">Urashima pustuloides</emphasis>
(
<bibRefCitation id="EFDBEEAAFFA92827FE6DFB88FDC7A647" author="Sakai T" box="[448,567,1120,1142]" pageId="17" pageNumber="317" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
)
</taxonomicName>
, male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, left G1 (ventral view); B, distal chitinous part of left G1 (ventral view); C, distal chitinous part of left G1 (subventral view); D, distal chitinous part of left G1 (dorsal view); E, left G2. Scales: A, E = 2.0 mm; BD = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BF5935BFFA92827FF25F98EFD16AA0E" blockId="17.[136,777,1638,2111]" pageId="17" pageNumber="317">
<taxonomicName id="4C4AE8D8FFA92827FF25F98EFF1EA44F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[136,238,1638,1662]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FF25F98EFF1EA44F" box="[136,238,1638,1662]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
also superficially resembles
<taxonomicName id="4C4AE8D8FFA92827FDB2F98EFCF6A44E" authority="Galil, 2003" authorityName="Galil" authorityYear="2003" box="[543,774,1638,1663]" class="Malacostraca" family="Leucosiidae" genus="Urashima" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FDB2F98EFD7AA44F" box="[543,650,1638,1662]" italics="true" pageId="17" pageNumber="317">Urashima</emphasis>
Galil, 2003
</taxonomicName>
(
<typeStatus id="54F12DF9FFA92827FF22F96FFF4EA4AE" box="[143,190,1671,1695]" pageId="17" pageNumber="317">type</typeStatus>
species
<taxonomicName id="4C4AE8D8FFA92827FEBAF96EFD73A4AF" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[279,643,1670,1694]" class="Malacostraca" family="Leucosiidae" genus="Randallia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="species" species="pustuloides">
<emphasis id="B93E4F49FFA92827FEBAF96EFE0FA4AF" box="[279,511,1670,1694]" italics="true" pageId="17" pageNumber="317">Randallia pustuloides</emphasis>
<bibRefCitation id="EFDBEEAAFFA92827FDA8F96EFD73A4AF" author="Sakai T" box="[517,643,1670,1694]" pageId="17" pageNumber="317" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
) (
<figureCitation id="13718FDEFFA92827FD34F96EFD0BA4AE" box="[665,763,1670,1695]" captionStart="Fig" captionStartId="16.[113,148,1950,1972]" captionTargetBox="[113,1451,274,1935]" captionTargetId="figure-9@16.[113,1451,274,1935]" captionTargetPageId="16" captionText="Fig. 14. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="17" pageNumber="317">Fig. 14A</figureCitation>
), but in
<taxonomicName id="4C4AE8D8FFA92827FF7CF94EFECCA48F" authorityName="Galil" authorityYear="2003" box="[209,316,1702,1726]" class="Malacostraca" family="Leucosiidae" genus="Urashima" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FF7CF94EFECCA48F" box="[209,316,1702,1726]" italics="true" pageId="17" pageNumber="317">Urashima</emphasis>
</taxonomicName>
, the third maxilliped has the exopod only slightly broader than the exopod (
<figureCitation id="13718FDEFFA92827FE54F92EFD8FA4EE" box="[505,639,1734,1759]" captionStart="Fig" captionStartId="16.[113,148,1950,1972]" captionTargetBox="[113,1451,274,1935]" captionTargetId="figure-9@16.[113,1451,274,1935]" captionTargetPageId="16" captionText="Fig. 14. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="17" pageNumber="317">Fig. 14C, D</figureCitation>
); the fingers of the chela are shorter than the palm (
<figureCitation id="13718FDEFFA92827FDE6F90EFD44A4CE" box="[587,692,1766,1791]" captionStart="Fig" captionStartId="16.[113,148,1950,1972]" captionTargetBox="[113,1451,274,1935]" captionTargetId="figure-9@16.[113,1451,274,1935]" captionTargetPageId="16" captionText="Fig. 14. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="17" pageNumber="317">Fig. 14G</figureCitation>
;
<bibRefCitation id="EFDBEEAAFFA92827FD6EF90EFF31A52F" author="Sakai T" pageId="17" pageNumber="317" pagination="131 - 150" refId="ref10919" refString="Sakai T (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2): 131 - 150, figs. 1 - 4, pls. 3, 4." type="journal article" year="1961">Sakai, 1961</bibRefCitation>
: pl. 3 fig. 4;
<bibRefCitation id="EFDBEEAAFFA92827FEF4F8EEFE2FA52F" author="Sakai T" box="[345,479,1798,1822]" pageId="17" pageNumber="317" pagination="728 - 738" refId="ref10963" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo. Samouelle G (1819) The Entomologist's Useful Compendium, or an Introduction to the Knowledge of the British Insects. London, Thomas Boys, Shih Y-J, Ho P-H, Chan T-Y &amp; Naruse T (2013) New records of leucosiid crabs (Decapoda, Brachyura) from Taiwan. Crustaceana, 86: 728 - 738." type="book chapter" year="1976">Sakai, 1976</bibRefCitation>
: pl. 30 fig. 1; Galil, 2003: fig. 2E, F;
<bibRefCitation id="EFDBEEAAFFA92827FEAEF8CFFE48A50E" author="Tan SH &amp; Wu S-H &amp; Huang J-F" box="[259,440,1830,1855]" pageId="17" pageNumber="317" pagination="141 - 147" refId="ref11219" refString="Tan SH, Wu S-H &amp; Huang J-F (2000) Seven new records of marine crabs from Taiwan (Crustacea: Decapoda: Brachyura: Calappidae, Lencosiidae and Cancridae). In: Hwang J-S, Wang C-H &amp; Chan T-Y (eds.) Proceedings of the International Symposium on Marine Biology in Taiwan - Crustacean and Zooplankton Taxonomy, Ecology and Living Resources. National Taiwan Museum Special Publication Series, No. 10: 141 - 147." type="book chapter" year="2000">Tan et al., 2000</bibRefCitation>
: fig. 3b) (vs. fingers distinctly longer than the palm in
<taxonomicName id="4C4AE8D8FFA92827FE24F8AEFE1FA56F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[393,495,1862,1886]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FE24F8AEFE1FA56F" box="[393,495,1862,1886]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFA92827FE53F8AEFD90A56E" box="[510,608,1862,1887]" captionStart="Fig" captionStartId="13.[136,171,1950,1972]" captionTargetBox="[136,1475,273,1935]" captionTargetId="figure-7@13.[136,1475,273,1935]" captionTargetPageId="13" captionText="Fig. 11. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, B, frontal view of cephalothorax and third maxillipeds; C, D, thoracic sternum and pleon; E, pleon; F, pleonal somite 6 showing cup-like tubercles on distal and proximal surfaces; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503205" httpUri="https://zenodo.org/record/4503205/files/figure.png" pageId="17" pageNumber="317">Fig. 11G</figureCitation>
); the surface of male pleonal somite 6 has a spine on the distal margin (
<figureCitation id="13718FDEFFA92827FD71F88EFF46A5AE" captionStart="Fig" captionStartId="16.[113,148,1950,1972]" captionTargetBox="[113,1451,274,1935]" captionTargetId="figure-9@16.[113,1451,274,1935]" captionTargetPageId="16" captionText="Fig. 14. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="17" pageNumber="317">Fig. 14E</figureCitation>
; Galil, 2003: 416); and male pleonal somites 5 and 6 are completely fused (
<figureCitation id="13718FDEFFA92827FEDEF84EFE21A58E" box="[371,465,1958,1983]" captionStart="Fig" captionStartId="16.[113,148,1950,1972]" captionTargetBox="[113,1451,274,1935]" captionTargetId="figure-9@16.[113,1451,274,1935]" captionTargetPageId="16" captionText="Fig. 14. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, overall view; B, C, frontal view of cephalothorax and third maxillipeds; D, right third maxilliped; E, anterior thoracic sternum, telson and pleonal somites 5 and 6 showing spine on distal margin of somite 6; F, thoracic sternum and pleon; G, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4503215" httpUri="https://zenodo.org/record/4503215/files/figure.png" pageId="17" pageNumber="317">Fig. 14F</figureCitation>
; Galil, 2003: 416). While the G1s of the two genera are superficially similar in general shape, that of
<taxonomicName id="4C4AE8D8FFA92827FE80F80EFE6AA5CF" authorityName="Galil" authorityYear="2003" box="[301,410,2022,2046]" class="Malacostraca" family="Leucosiidae" genus="Urashima" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FE80F80EFE6AA5CF" box="[301,410,2022,2046]" italics="true" pageId="17" pageNumber="317">Urashima</emphasis>
</taxonomicName>
has the distal quarter distinctly chitinised and spatuliform (
<figureCitation id="13718FDEFFA92827FE1FF7EEFE0DAA2F" box="[434,509,2054,2078]" captionStart="Fig" captionStartId="17.[136,171,1120,1142]" captionTargetBox="[141,1474,193,1105]" captionTargetId="figure-8@17.[122,1489,178,1115]" captionTargetPageId="17" captionText="Fig. 15. Urashima pustuloides (Sakai, 1961), male (44.5 × 37.6 mm) (ZRC 2017.1069), Philippines. A, left G1 (ventral view); B, distal chitinous part of left G1 (ventral view); C, distal chitinous part of left G1 (subventral view); D, distal chitinous part of left G1 (dorsal view); E, left G2. Scales: A, E = 2.0 mm; BD = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4503219" httpUri="https://zenodo.org/record/4503219/files/figure.png" pageId="17" pageNumber="317">Fig. 15</figureCitation>
AD) (vs. the distal part is not chitinised and conical in
<taxonomicName id="4C4AE8D8FFA92827FE4DF7CEFDB6AA0F" authorityName="Ng &amp; Devi &amp; Kumar" authorityYear="2018" box="[480,582,2086,2110]" class="Malacostraca" family="Leucosiidae" genus="Neparilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4F49FFA92827FE4DF7CEFDB6AA0F" box="[480,582,2086,2110]" italics="true" pageId="17" pageNumber="317">Neparilia</emphasis>
</taxonomicName>
;
<figureCitation id="13718FDEFFA92827FDFBF7CEFD53AA0E" box="[598,675,2086,2111]" captionStart="Fig" captionStartId="14.[113,148,1060,1082]" captionTargetBox="[93,1481,174,1055]" captionTargetId="figure-566@14.[93,1481,174,1055]" captionTargetPageId="14" captionText="Fig. 12. Neparilia tuberculata (Sakai, 1961), male (37.0 × 34.4 mm) (ZRC 2017.1052), Philippines. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsal view); D, left G2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4503207" httpUri="https://zenodo.org/record/4503207/files/figure.png" pageId="17" pageNumber="317">Fig. 12</figureCitation>
AC).
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>