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<document id="9F4D6CC78342AD6E0C0BFA0FFB2C090A" ID-DOI="10.11646/zootaxa.4750.1.5" ID-GBIF-Dataset="8c28e10d-e12c-4399-9bd1-bac99bac1fb4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3702869" ID-ZooBank="1A3B7CFF-292B-46AB-82E8-C78BA34618EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583849282720" checkinUser="plazi" docAuthor="Duarte, Paulo R. M. &amp; Grossi, Paschoal C." docDate="2020" docId="03E31B7A590EFFB5FF6CFC64BB16FDC2" docLanguage="en" docName="zootaxa.4750.1.5.pdf" docOrigin="Zootaxa 4750 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bothynus condacki Duarte &amp; Grossi 2020, new species" docType="treatment" docVersion="6" lastPageNumber="118" masterDocId="FFDA63025902FFA4FFFBFFD5BE17FB1B" masterDocTitle="Bothynus entellus (LePeletier &amp; Serville) (Coleoptera: Scarabaeidae: Dynastinae) species group: taxonomic revision and description of two new species" masterLastPageNumber="121" masterPageNumber="101" pageNumber="113" updateTime="1698726678812" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="3D2ECB4C669AA415262CE572A6838756" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C82D7C30D6F51E7802C7A45F7B40FC6">
<mods:title id="A5FAFBFC1886D213BE702986ABF2E288">Bothynus entellus (LePeletier &amp; Serville) (Coleoptera: Scarabaeidae: Dynastinae) species group: taxonomic revision and description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="94DB34816EF54EDB7E91CC1CFA5FE543" type="personal">
<mods:role id="6D77A9F178AF9CCD89EF59E153625B6D">
<mods:roleTerm id="EA6CA24C5F346C8351CB9A4AD10A1D85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63D3DF15448077A3808C0FB0D25A0CEF">Duarte, Paulo R. M.</mods:namePart>
</mods:name>
<mods:name id="AED477736DCB32015B5EBC78DC85AA8D" type="personal">
<mods:role id="97353F9DD4C9F41E46554797668EA661">
<mods:roleTerm id="DFE0D9526F694878FCBD7447A4D3F6D5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA7D9B1EC341015AF63FD6DAC364866E">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="419796B10738011A01052EF73F41CF09">text</mods:typeOfResource>
<mods:relatedItem id="83E728842B102B015011D5C137D5C9E1" type="host">
<mods:titleInfo id="9ABDA59299D8C7411C5E3CADA34E740B">
<mods:title id="38A8EDEC717CC0C07E8A900849A1C720">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="58CBE55A36EE878B561E2E664B666E36">
<mods:date id="5A851563C39E0F48EAB6CC8BC2D1AF77">2020</mods:date>
<mods:detail id="605C0664464754F87296CB85BE75FAE6" type="pubDate">
<mods:number id="0C522F8DB63313D0D1767D006E994F23">2020-03-10</mods:number>
</mods:detail>
<mods:detail id="0F09B7F70592D510B194D00BDF4C2D46" type="volume">
<mods:number id="826EA525B47B5316904326E40280251D">4750</mods:number>
</mods:detail>
<mods:detail id="B3BF7D9C23AC3C0976A0E4721ED71B60" type="issue">
<mods:number id="D431D86E2B31BCEEDE1B9688F0E94BB0">1</mods:number>
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<mods:extent id="0284CCDF067CE1F3AFE88027EE2EB070" unit="page">
<mods:start id="434BB7809C95C54B65ABAD3E21E3EACC">101</mods:start>
<mods:end id="48B6546C886DB9E710DBA7B20D72FD7F">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5E1EC07B0A861BD675A95BAECF592D26">journal article</mods:classification>
<mods:identifier id="66B58D983A6E343B7583B7789484213E" type="DOI">10.11646/zootaxa.4750.1.5</mods:identifier>
<mods:identifier id="A200B89D63BAAAFDBC6119510191F110" type="GBIF-Dataset">8c28e10d-e12c-4399-9bd1-bac99bac1fb4</mods:identifier>
<mods:identifier id="D40BF947779554B9038BAC25D28C2028" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="827F973FE0FFC79243786A0768FBAE22" type="Zenodo-Dep">3702869</mods:identifier>
<mods:identifier id="3302A93AD74C05DEDDFB99BC615945E5" type="ZooBank">1A3B7CFF-292B-46AB-82E8-C78BA34618EC</mods:identifier>
</mods:mods>
<treatment id="03E31B7A590EFFB5FF6CFC64BB16FDC2" ID-DOI="http://doi.org/10.5281/zenodo.3706215" ID-GBIF-Taxon="162481354" ID-Zenodo-Dep="3706215" LSID="urn:lsid:plazi:treatment:03E31B7A590EFFB5FF6CFC64BB16FDC2" httpUri="http://treatment.plazi.org/id/03E31B7A590EFFB5FF6CFC64BB16FDC2" lastPageId="17" lastPageNumber="118" pageId="12" pageNumber="113">
<subSubSection id="C350F9E7590EFFA8FF6CFC64BCE2F8D0" box="[151,757,945,972]" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="8BF5AA6C590EFFA8FF6CFC64BCE2F8D0" blockId="12.[151,757,945,1007]" box="[151,757,945,972]" pageId="12" pageNumber="113">
<heading id="D0BD1D00590EFFA8FF6CFC64BCE2F8D0" bold="true" box="[151,757,945,972]" fontSize="11" level="1" pageId="12" pageNumber="113" reason="1">
<emphasis id="B93E767E590EFFA8FF6CFC64BCE2F8D0" bold="true" box="[151,757,945,972]" pageId="12" pageNumber="113">
<taxonomicName id="4C4AD1EF590EFFA8FF6CFC64BC4EF8D0" authority="Duarte &amp; Grossi" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[151,601,945,972]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="condacki" status="sp. nov.">
<emphasis id="B93E767E590EFFA8FF6CFC64BF68F8D7" bold="true" box="[151,383,945,972]" italics="true" pageId="12" pageNumber="113">Bothynus condacki</emphasis>
Duarte &amp; Grossi
</taxonomicName>
,
<taxonomicNameLabel id="A20DCB05590EFFA8FD9EFC64BCE2F8D0" box="[613,757,945,971]" pageId="12" pageNumber="113" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C350F9E7590EFFA8FF6CFC00BFA9F8F4" box="[151,446,981,1007]" pageId="12" pageNumber="113" type="description">
<paragraph id="8BF5AA6C590EFFA8FF6CFC00BFA9F8F4" blockId="12.[151,757,945,1007]" box="[151,446,981,1007]" pageId="12" pageNumber="113">
(
<figureCitation id="1371B6E9590EFFA8FF64FC00BEFAF8F4" box="[159,237,981,1007]" captionStart="FIGURE 6" captionStartId="12.[151,250,1982,2007]" captionTargetBox="[192,1399,1267,1950]" captionTargetId="figure@12.[170,1417,1245,1957]" captionTargetPageId="12" captionText="FIGURE 6. Bothynus condacki. A, Male, dorsal view; B, metatrochanter, ventral view (acute apex, arrow). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702881" httpUri="https://zenodo.org/record/3702881/files/figure.png" pageId="12" pageNumber="113">Figs. 6</figureCitation>
A–B; 8F; 9F; 12F)
</paragraph>
</subSubSection>
<subSubSection id="C350F9E7590EFFA8FF6CFBC9BA4EFF64" pageId="12" pageNumber="113" type="diagnosis">
<paragraph id="8BF5AA6C590EFFA8FF6CFBC9BA4EFF64" blockId="12.[151,1436,1052,1223]" pageId="12" pageNumber="113">
<emphasis id="B93E767E590EFFA8FF6CFBC9BF05FF2C" bold="true" box="[151,274,1052,1079]" pageId="12" pageNumber="113">Diagnosis.</emphasis>
<taxonomicName id="4C4AD1EF590EFFA8FEECFBC8BFFCFF2C" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[279,491,1053,1079]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E590EFFA8FEECFBC8BFFCFF2C" box="[279,491,1053,1079]" italics="true" pageId="12" pageNumber="113">Bothynus condacki</emphasis>
</taxonomicName>
is similar to
<emphasis id="B93E767E590EFFA8FD81FBCBBCEDFF2C" box="[634,762,1053,1079]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="4C4AD1EF590EFFA8FD81FBCBBCE1FF2C" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[634,758,1053,1079]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="stenelus">B. stenelus</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="4C4AD1EF590EFFA8FC89FBCBBDE2FF2C" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[882,1013,1053,1079]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E590EFFA8FC89FBCBBDE2FF2C" box="[882,1013,1053,1079]" italics="true" pageId="12" pageNumber="113">B. condacki</emphasis>
</taxonomicName>
can be distinguished by the following characters: metatrochanter with acute apex (
<figureCitation id="1371B6E9590EFFA8FD78FB94BCCFFF47" box="[643,728,1089,1116]" captionStart="FIGURE 6" captionStartId="12.[151,250,1982,2007]" captionTargetBox="[192,1399,1267,1950]" captionTargetId="figure@12.[170,1417,1245,1957]" captionTargetPageId="12" captionText="FIGURE 6. Bothynus condacki. A, Male, dorsal view; B, metatrochanter, ventral view (acute apex, arrow). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702881" httpUri="https://zenodo.org/record/3702881/files/figure.png" pageId="12" pageNumber="113">Fig. 6B</figureCitation>
) (apex rounded in
<taxonomicName id="4C4AD1EF590EFFA8FC51FB97BA3CFF40" box="[938,1067,1089,1115]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="stenellus">
<emphasis id="B93E767E590EFFA8FC51FB97BA3CFF40" box="[938,1067,1089,1115]" italics="true" pageId="12" pageNumber="113">B. stenellus</emphasis>
</taxonomicName>
); parameres with rounded lateral flaps, and with a strong apical contraction at inner margin of right paramere (
<figureCitation id="1371B6E9590EFFA8FC0DFBB0BA5BFF64" box="[1014,1100,1125,1151]" captionStart="FIGURE 8" captionStartId="14.[151,250,1945,1970]" captionTargetBox="[218,1363,209,1898]" captionTargetId="figure@14.[199,1389,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Parameres in caudal view. A, Bothynus entellus; B, B. stenelus; C, B. tricornis; D, B. lancifer; E, B. araya; F, B. condacki (inner apical contraction, arrow). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702885" httpUri="https://zenodo.org/record/3702885/files/figure.png" pageId="12" pageNumber="113">Fig. 8F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C350F9E7590EFFA8FF3CFB5CBF39FFDC" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8BF5AA6C590EFFA8FF3CFB5CBF39FFDC" blockId="12.[151,1436,1052,1223]" pageId="12" pageNumber="113">
<emphasis id="B93E767E590EFFA8FF3CFB5CBF5FFFB8" bold="true" box="[199,328,1161,1187]" pageId="12" pageNumber="113">Etymology</emphasis>
. The specific epithet is a tribute to the collector of the
<typeStatus id="54F114CE590EFFA8FC53FB5CBA1AFFB8" box="[936,1037,1161,1187]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
, João Condack, who is an expert on Pteridophyta.
</paragraph>
</subSubSection>
<caption id="DF35FAE4590EFFA8FF6CF86BBB9BFCCD" ID-DOI="http://doi.org/10.5281/zenodo.3702881" ID-Zenodo-Dep="3702881" box="[151,1420,1982,2007]" httpUri="https://zenodo.org/record/3702881/files/figure.png" pageId="12" pageNumber="113" startId="12.[151,250,1982,2007]" targetBox="[192,1399,1267,1950]" targetPageId="12">
<paragraph id="8BF5AA6C590EFFA8FF6CF86BBB9BFCCD" blockId="12.[151,1420,1982,2007]" box="[151,1420,1982,2007]" pageId="12" pageNumber="113">
<emphasis id="B93E767E590EFFA8FF6CF86BBF04FCCC" bold="true" box="[151,275,1982,2007]" pageId="12" pageNumber="113">FIGURE 6.</emphasis>
<taxonomicName id="4C4AD1EF590EFFA8FEE2F86ABFCEFCCC" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[281,473,1983,2007]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E590EFFA8FEE2F86ABFCEFCCC" box="[281,473,1983,2007]" italics="true" pageId="12" pageNumber="113">Bothynus condacki</emphasis>
</taxonomicName>
.
<emphasis id="B93E767E590EFFA8FE1DF86BBFEFFCCC" bold="true" box="[486,504,1982,2007]" pageId="12" pageNumber="113">A</emphasis>
, Male, dorsal view;
<emphasis id="B93E767E590EFFA8FD32F86ABCCDFCCC" bold="true" box="[713,730,1983,2007]" pageId="12" pageNumber="113">B</emphasis>
, metatrochanter, ventral view (acute apex, arrow). Scale bar: 2 mm.
</paragraph>
</caption>
<caption id="DF35FAE4590FFFA9FF6CF94EBF07FDCC" ID-DOI="http://doi.org/10.5281/zenodo.3702883" ID-Zenodo-Dep="3702883" httpUri="https://zenodo.org/record/3702883/files/figure.png" pageId="13" pageNumber="114" startId="13.[151,250,1691,1716]" targetBox="[199,1389,196,1649]" targetPageId="13">
<paragraph id="8BF5AA6C590FFFA9FF6CF94EBF07FDCC" blockId="13.[151,1436,1691,1752]" pageId="13" pageNumber="114">
<emphasis id="B93E767E590FFFA9FF6CF94EBF04FDAF" bold="true" box="[151,275,1691,1716]" pageId="13" pageNumber="114">FIGURE 7.</emphasis>
Tergite VII, detail of the stridulatory area (carinae, arrow).
<emphasis id="B93E767E590FFFA9FC95F94EBD97FDAF" bold="true" box="[878,896,1691,1716]" pageId="13" pageNumber="114">A</emphasis>
,
<taxonomicName id="4C4AD1EF590FFFA9FC76F949BA57FDAF" box="[909,1088,1692,1716]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E590FFFA9FC76F949BA57FDAF" box="[909,1088,1692,1716]" italics="true" pageId="13" pageNumber="114">Bothynus entellus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590FFFA9FBB6F949BA49FDAF" bold="true" box="[1101,1118,1692,1716]" pageId="13" pageNumber="114">B</emphasis>
,
<taxonomicName id="4C4AD1EF590FFFA9FB90F948BAD5FDAF" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[1131,1218,1692,1716]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="B93E767E590FFFA9FB90F948BA6DFDAF" box="[1131,1146,1693,1716]" italics="true" pageId="13" pageNumber="114">B</emphasis>
.
<emphasis id="B93E767E590FFFA9FB7CF948BAD5FDAF" box="[1159,1218,1693,1716]" italics="true" pageId="13" pageNumber="114">araya</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590FFFA9FB34F94EBAF6FDAF" bold="true" box="[1231,1249,1691,1716]" pageId="13" pageNumber="114">C</emphasis>
,
<taxonomicName id="4C4AD1EF590FFFA9FB15F948BB4EFDAF" authorityName="Dechambre" authorityYear="1981" box="[1262,1369,1692,1716]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="lancifer">
<emphasis id="B93E767E590FFFA9FB15F948BAEAFDAF" box="[1262,1277,1693,1716]" italics="true" pageId="13" pageNumber="114">B</emphasis>
.
<emphasis id="B93E767E590FFFA9FAF1F949BB4EFDAF" box="[1290,1369,1692,1716]" italics="true" pageId="13" pageNumber="114">lancifer</emphasis>
</taxonomicName>
. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="C350F9E7590FFFA9FF3CF8D3BF9FFC5E" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="8BF5AA6C590FFFA9FF3CF8D3BF9FFC5E" blockId="13.[151,1437,1797,2041]" pageId="13" pageNumber="114">
<emphasis id="B93E767E590FFFA9FF3CF8D3BF66FC3B" bold="true" box="[199,369,1797,1824]" pageId="13" pageNumber="114">Type material</emphasis>
.
<materialsCitation id="3B22A031590FFFA9FE78F8D3BF9FFC5E" ID-GBIF-Occurrence="2575116337" collectingDate="1998-12-28" collectionCode="CERPE" country="Brazil" location="Conceicao do Ibitipoca" pageId="13" pageNumber="114" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="54F114CE590FFFA9FE78F8D3BFFCFC3B" box="[387,491,1798,1824]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
male dissected.
<emphasis id="B93E767E590FFFA9FD54F8D0BCE0FC3B" bold="true" box="[687,759,1797,1824]" pageId="13" pageNumber="114">
<collectingCountry id="F35DEAFC590FFFA9FD54F8D0BCE0FC3B" box="[687,759,1797,1824]" name="Brazil" pageId="13" pageNumber="114">Brasil</collectingCountry>
</emphasis>
:
<emphasis id="B93E767E590FFFA9FCF2F8D0BDBAFC3B" bold="true" box="[777,941,1797,1824]" pageId="13" pageNumber="114">
<collectingRegion id="498E648E590FFFA9FCF2F8D0BDBAFC3B" box="[777,941,1797,1824]" country="Brazil" name="Minas Gerais" pageId="13" pageNumber="114">Minas Gerais</collectingRegion>
</emphasis>
:
<location id="8E95FCB7590FFFA9FC3BF8D3BADAFC3B" LSID="urn:lsid:plazi:treatment:03E31B7A590EFFB5FF6CFC64BB16FDC2:8E95FCB7590FFFA9FC3BF8D3BADAFC3B" box="[960,1229,1798,1824]" country="Brazil" name="Conceicao do Ibitipoca" pageId="13" pageNumber="114" stateProvince="Minas Gerais">Conceição do Ibitipoca</location>
,
<date id="FFF48CAC590FFFA9FB27F8D3BB70FC3B" box="[1244,1383,1798,1824]" pageId="13" pageNumber="114" value="1998-12-28">
<collectingDate id="EFB07544590FFFA9FB27F8D3BB70FC3B" box="[1244,1383,1798,1824]" pageId="13" pageNumber="114" value="1998-12-28">28.XII.1998</collectingDate>
</date>
, J.P. Condack—(
<collectionCode id="ED5B32A9590FFFA9FED9F8FFBF6BFC5E" box="[290,380,1834,1861]" pageId="13" pageNumber="114">CERPE</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C350F9E7590FFFB5FF3CF898BB16FDC2" lastPageId="17" lastPageNumber="118" pageId="13" pageNumber="114" type="description">
<paragraph id="8BF5AA6C590FFFABFF3CF898BC5DF99C" blockId="13.[151,1437,1797,2041]" lastBlockId="15.[151,1437,153,683]" lastPageId="15" lastPageNumber="116" pageId="13" pageNumber="114">
<emphasis id="B93E767E590FFFA9FF3CF898BF46FC73" bold="true" box="[199,337,1869,1896]" pageId="13" pageNumber="114">Description</emphasis>
.
<emphasis id="B93E767E590FFFA9FEA4F89BBC1BFC73" bold="true" box="[351,524,1870,1896]" pageId="13" pageNumber="114">
<typeStatus id="54F114CE590FFFA9FEA4F89BBFDBFC73" box="[351,460,1870,1896]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="1371B6E9590FFFA9FDE1F89BBC63FC72" box="[538,628,1870,1897]" captionStart="FIGURE 6" captionStartId="12.[151,250,1982,2007]" captionTargetBox="[192,1399,1267,1950]" captionTargetId="figure@12.[170,1417,1245,1957]" captionTargetPageId="12" captionText="FIGURE 6. Bothynus condacki. A, Male, dorsal view; B, metatrochanter, ventral view (acute apex, arrow). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702881" httpUri="https://zenodo.org/record/3702881/files/figure.png" pageId="13" pageNumber="114">Fig. 6A</figureCitation>
).
<emphasis id="B93E767E590FFFA9FD7CF89BBD00FC73" bold="true" box="[647,791,1870,1896]" pageId="13" pageNumber="114">Body length</emphasis>
: 24.0 mm.
<emphasis id="B93E767E590FFFA9FC6EF89BBA0AFC73" bold="true" box="[917,1053,1869,1896]" pageId="13" pageNumber="114">Body width</emphasis>
: 13.mm.
<emphasis id="B93E767E590FFFA9FB73F898BADBFC73" bold="true" box="[1160,1228,1869,1896]" pageId="13" pageNumber="114">Color</emphasis>
: Completely dark.
<emphasis id="B93E767E590FFFA9FF6CF8A7BEC0FC97" bold="true" box="[151,215,1906,1932]" pageId="13" pageNumber="114">Head</emphasis>
: Clypeus subpentagonal, densely punctate, glabrous, apex with 2 short teeth. Frontoclypeal suture with 2 small, transverse tubercles. Interocular width equals 3.3 transverse eye diameter; frons with surface coarsely ru- gopunctate, with scarce setae scattered laterally near eyes.
<emphasis id="B93E767E590FFFA9FCC2F86FBDDFFCCF" bold="true" box="[825,968,1978,2004]" pageId="13" pageNumber="114">Mouthparts</emphasis>
: Mandibles tridentate; apical and medial teeth triangular, basal tooth lobed and smaller than previous tooth. Maxilla with quadridentate galea; 2 strong apical teeth, 2 weak basal teeth. Mentum subtriangular, surrounded with setose punctures, discal area slightly convex, glabrous.
<emphasis id="B93E767E590DFFABFF1FFF68BF70FBCC" bold="true" box="[228,359,189,215]" pageId="15" pageNumber="116">Pronotum:</emphasis>
Strongly convex posteriorly in lateral view; anterior area with a small apical tubercle followed by a, rounded shaped concavity, moderately deep. Discal and lateral areas finely punctate; concavity punctate in 2 areas at sides, anteriorly with dense, C-shaped and coalescent punctures, becoming moderately punctate toward posterior area.
<emphasis id="B93E767E590DFFABFEB9FEFDBFECFA58" bold="true" box="[322,507,296,323]" pageId="15" pageNumber="116">Scutellar shield</emphasis>
: Subparabolic shaped, smooth, 1.7 times wider than long.
<emphasis id="B93E767E590DFFABFB60FEFCBAF0FA58" bold="true" box="[1179,1255,297,323]" pageId="15" pageNumber="116">Elytra</emphasis>
: Surface nearly completely smooth, punctures inconspicuous (only observed under 90X magnification), longitudinal striae barely marked.
<emphasis id="B93E767E590DFFABFF06FEA4BF22FA90" bold="true" box="[253,309,369,395]" pageId="15" pageNumber="116">Legs</emphasis>
: Protarsomere V with 1 ventro-basal tooth; protarsomere IV extending ventrally, nearly reaching the apex of the ventro-basal tooth of tarsomere V. Metatrochanter with acute apex (
<figureCitation id="1371B6E9590DFFABFBBAFE40BA8CFAAB" box="[1089,1179,405,432]" captionStart="FIGURE 6" captionStartId="12.[151,250,1982,2007]" captionTargetBox="[192,1399,1267,1950]" captionTargetId="figure@12.[170,1417,1245,1957]" captionTargetPageId="12" captionText="FIGURE 6. Bothynus condacki. A, Male, dorsal view; B, metatrochanter, ventral view (acute apex, arrow). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702881" httpUri="https://zenodo.org/record/3702881/files/figure.png" pageId="15" pageNumber="116">Fig. 6B</figureCitation>
).
<emphasis id="B93E767E590DFFABFB54FE41BB33FAB4" bold="true" box="[1199,1316,404,431]" pageId="15" pageNumber="116">Abdomen</emphasis>
: Ventrite I completely, densely setose, II–VI moderately setose on sides, becoming gradually weak toward disc; discal area glabrous. Tergite VII with stridulatory apparatus formed by a band of innumerous, transverse, finely marked carinae. Tergite VIII nearly smooth, only with transverse, oval shaped punctures, confined to disc.
<emphasis id="B93E767E590DFFABFB5FFDD5BB00F900" bold="true" box="[1188,1303,512,539]" pageId="15" pageNumber="116">Aedeagus</emphasis>
: Parameres in caudal view (
<figureCitation id="1371B6E9590DFFABFEB4FDF0BFB1F924" box="[335,422,549,575]" captionStart="FIGURE 8" captionStartId="14.[151,250,1945,1970]" captionTargetBox="[218,1363,209,1898]" captionTargetId="figure@14.[199,1389,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Parameres in caudal view. A, Bothynus entellus; B, B. stenelus; C, B. tricornis; D, B. lancifer; E, B. araya; F, B. condacki (inner apical contraction, arrow). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702885" httpUri="https://zenodo.org/record/3702885/files/figure.png" pageId="15" pageNumber="116">Fig. 8F</figureCitation>
), nearly symmetric, middle area with narrow outer margins, apical half with lateral flaps distinctly wider than basal half, apex of the right paramere with a strong contraction at inner margin. In lateral view, apex shortened, downcurved (
<figureCitation id="1371B6E9590DFFABFE13FDB8BC2AF99C" box="[488,573,621,647]" captionStart="FIGURE 9" captionStartId="15.[151,250,1966,1991]" captionTargetBox="[176,1404,727,1898]" captionTargetId="figure@15.[170,1417,705,1941]" captionTargetPageId="15" captionText="FIGURE 9. Aedeagus in lateral view. A, Bothynus entellus; B, B. stenelus; C, B. tricornis; D, B. lancifer (lateroventral contraction, arrow); E, B. araya; F, B. condacki. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3702887" httpUri="https://zenodo.org/record/3702887/files/figure.png" pageId="15" pageNumber="116">Fig. 9F</figureCitation>
).
</paragraph>
<caption id="DF35FAE4590CFFAAFF6CF84CBD13FCCE" ID-DOI="http://doi.org/10.5281/zenodo.3702885" ID-Zenodo-Dep="3702885" httpUri="https://zenodo.org/record/3702885/files/figure.png" pageId="14" pageNumber="115" startId="14.[151,250,1945,1970]" targetBox="[218,1363,209,1898]" targetPageId="14">
<paragraph id="8BF5AA6C590CFFAAFF6CF84CBD13FCCE" blockId="14.[151,1436,1945,2006]" pageId="14" pageNumber="115">
<emphasis id="B93E767E590CFFAAFF6CF84CBF03FCA9" bold="true" box="[151,276,1945,1970]" pageId="14" pageNumber="115">FIGURE 8.</emphasis>
Parameres in caudal view.
<emphasis id="B93E767E590CFFAAFDD6F84CBC28FCA9" bold="true" box="[557,575,1945,1970]" pageId="14" pageNumber="115">A</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFDB7F84FBD16FCA9" box="[588,769,1946,1970]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E590CFFAAFDB7F84FBD16FCA9" box="[588,769,1946,1970]" italics="true" pageId="14" pageNumber="115">Bothynus entellus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590CFFAAFCF4F84FBD37FCA9" bold="true" box="[783,800,1946,1970]" pageId="14" pageNumber="115">B</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFCD6F84EBD8BFCA9" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[813,924,1946,1970]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="stenelus">
<emphasis id="B93E767E590CFFAAFCD6F84EBD2BFCA9" box="[813,828,1947,1970]" italics="true" pageId="14" pageNumber="115">B</emphasis>
.
<emphasis id="B93E767E590CFFAAFCB1F84FBD8BFCA9" box="[842,924,1946,1970]" italics="true" pageId="14" pageNumber="115">stenelus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590CFFAAFC52F84CBDACFCA9" bold="true" box="[937,955,1945,1970]" pageId="14" pageNumber="115">C</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFC32F84EBA2AFCA9" authorityName="Arrow" authorityYear="1937" box="[969,1085,1946,1970]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="tricornis">
<emphasis id="B93E767E590CFFAAFC32F84EBDCFFCA9" box="[969,984,1947,1970]" italics="true" pageId="14" pageNumber="115">B</emphasis>
.
<emphasis id="B93E767E590CFFAAFC1DF84EBA2AFCA9" box="[998,1085,1947,1970]" italics="true" pageId="14" pageNumber="115">tricornis</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590CFFAAFBB7F84FBA49FCA9" bold="true" box="[1100,1118,1946,1970]" pageId="14" pageNumber="115">D</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFB90F84EBAC0FCA9" authorityName="Dechambre" authorityYear="1981" box="[1131,1239,1946,1970]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="lancifer">
<emphasis id="B93E767E590CFFAAFB90F84EBA6DFCA9" box="[1131,1146,1947,1970]" italics="true" pageId="14" pageNumber="115">B</emphasis>
.
<emphasis id="B93E767E590CFFAAFB73F84FBAC0FCA9" box="[1160,1239,1946,1970]" italics="true" pageId="14" pageNumber="115">lancifer</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590CFFAAFB1EF84FBAE1FCA9" bold="true" box="[1253,1270,1946,1970]" pageId="14" pageNumber="115">E</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFAF8F84EBB4CFCA9" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[1283,1371,1946,1970]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="B93E767E590CFFAAFAF8F84EBB05FCA9" box="[1283,1298,1947,1970]" italics="true" pageId="14" pageNumber="115">B</emphasis>
.
<emphasis id="B93E767E590CFFAAFADBF84EBB4CFCA9" box="[1312,1371,1947,1970]" italics="true" pageId="14" pageNumber="115">araya</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590CFFAAFA91F84FBB6EFCA9" bold="true" box="[1386,1401,1946,1970]" pageId="14" pageNumber="115">F</emphasis>
,
<taxonomicName id="4C4AD1EF590CFFAAFA7DF84EBEE5FCCD" authorityName="Duarte &amp; Grossi" authorityYear="2020" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E590CFFAAFA7DF84EBB82FCA9" box="[1414,1429,1947,1970]" italics="true" pageId="14" pageNumber="115">B</emphasis>
.
<emphasis id="B93E767E590CFFAAFF6CF86BBEE5FCCD" box="[151,242,1982,2006]" italics="true" pageId="14" pageNumber="115">condacki</emphasis>
</taxonomicName>
(inner apical contraction, arrow). Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BF5AA6C590DFFABFF3CFD44BFB5F9B0" blockId="15.[151,1437,153,683]" box="[199,418,657,683]" pageId="15" pageNumber="116">
<emphasis id="B93E767E590DFFABFF3CFD44BF09F9B0" bold="true" box="[199,286,657,683]" pageId="15" pageNumber="116">Female</emphasis>
. Unknown.
</paragraph>
<caption id="DF35FAE4590DFFABFF6CF87BBCE2FCF1" ID-DOI="http://doi.org/10.5281/zenodo.3702887" ID-Zenodo-Dep="3702887" httpUri="https://zenodo.org/record/3702887/files/figure.png" pageId="15" pageNumber="116" startId="15.[151,250,1966,1991]" targetBox="[176,1404,727,1898]" targetPageId="15">
<paragraph id="8BF5AA6C590DFFABFF6CF87BBCE2FCF1" blockId="15.[151,1436,1966,2027]" pageId="15" pageNumber="116">
<emphasis id="B93E767E590DFFABFF6CF87BBF05FCDC" bold="true" box="[151,274,1966,1991]" pageId="15" pageNumber="116">FIGURE 9.</emphasis>
Aedeagus in lateral view.
<emphasis id="B93E767E590DFFABFDE0F87BBC3AFCDC" bold="true" box="[539,557,1966,1991]" pageId="15" pageNumber="116">A</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFDC1F87ABCFBFCDC" box="[570,748,1967,1991]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E590DFFABFDC1F87ABCFBFCDC" box="[570,748,1967,1991]" italics="true" pageId="15" pageNumber="116">Bothynus entellus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590DFFABFD02F87ABD1DFCDC" bold="true" box="[761,778,1967,1991]" pageId="15" pageNumber="116">B</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFCEDF865BD94FCDC" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[790,899,1967,1991]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="stenelus">
<emphasis id="B93E767E590DFFABFCEDF865BD32FCDC" box="[790,805,1968,1991]" italics="true" pageId="15" pageNumber="116">B</emphasis>
.
<emphasis id="B93E767E590DFFABFCCAF87ABD94FCDC" box="[817,899,1967,1991]" italics="true" pageId="15" pageNumber="116">stenelus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590DFFABFC74F87BBDB6FCDC" bold="true" box="[911,929,1966,1991]" pageId="15" pageNumber="116">C</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFC55F865BA37FCDC" authorityName="Arrow" authorityYear="1937" box="[942,1056,1967,1991]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tricornis">
<emphasis id="B93E767E590DFFABFC55F865BDAAFCDC" box="[942,957,1968,1991]" italics="true" pageId="15" pageNumber="116">B</emphasis>
.
<emphasis id="B93E767E590DFFABFC32F865BA37FCDC" box="[969,1056,1968,1991]" italics="true" pageId="15" pageNumber="116">tricornis</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590DFFABFBD6F87ABA28FCDC" bold="true" box="[1069,1087,1967,1991]" pageId="15" pageNumber="116">D</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFBB0F865BAA1FCDC" authorityName="Dechambre" authorityYear="1981" box="[1099,1206,1967,1991]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="lancifer">
<emphasis id="B93E767E590DFFABFBB0F865BA4DFCDC" box="[1099,1114,1968,1991]" italics="true" pageId="15" pageNumber="116">B</emphasis>
.
<emphasis id="B93E767E590DFFABFB9CF87ABAA1FCDC" box="[1127,1206,1967,1991]" italics="true" pageId="15" pageNumber="116">lancifer</emphasis>
</taxonomicName>
(lateroventral contraction, arrow);
<emphasis id="B93E767E590DFFABFEE7F806BF3AFCF0" bold="true" box="[284,301,2003,2027]" pageId="15" pageNumber="116">E</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFEC1F801BF86FCF0" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[314,401,2003,2027]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="B93E767E590DFFABFEC1F801BF5EFCF0" box="[314,329,2004,2027]" italics="true" pageId="15" pageNumber="116">B</emphasis>
.
<emphasis id="B93E767E590DFFABFEADF801BF86FCF0" box="[342,401,2004,2027]" italics="true" pageId="15" pageNumber="116">araya</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E590DFFABFE65F806BFBAFCF0" bold="true" box="[414,429,2003,2027]" pageId="15" pageNumber="116">F</emphasis>
,
<taxonomicName id="4C4AD1EF590DFFABFE41F801BC25FCF0" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[442,562,2003,2027]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E590DFFABFE41F801BFDEFCF0" box="[442,457,2004,2027]" italics="true" pageId="15" pageNumber="116">B</emphasis>
.
<emphasis id="B93E767E590DFFABFE2CF806BC25FCF0" box="[471,562,2003,2027]" italics="true" pageId="15" pageNumber="116">condacki</emphasis>
</taxonomicName>
. Scale bars: 2 mm.
</paragraph>
</caption>
<caption id="DF35FAE45912FFB4FF6CF85ABF07FCF4" ID-DOI="http://doi.org/10.5281/zenodo.3702889" ID-Zenodo-Dep="3702889" httpUri="https://zenodo.org/record/3702889/files/figure.png" pageId="16" pageNumber="117" startId="16.[151,250,1935,1960]" targetBox="[196,1395,214,1870]" targetPageId="16">
<paragraph id="8BF5AA6C5912FFB4FF6CF85ABF07FCF4" blockId="16.[151,1436,1935,2032]" pageId="16" pageNumber="117">
<emphasis id="B93E767E5912FFB4FF6CF85ABF35FCB3" bold="true" box="[151,290,1935,1960]" pageId="16" pageNumber="117">FIGURE 10.</emphasis>
Pronotal surface of females in dorsal view.
<emphasis id="B93E767E5912FFB4FD0BF85ABD15FCB3" bold="true" box="[752,770,1935,1960]" pageId="16" pageNumber="117">A</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FCEAF85ABDD0FCBC" box="[785,967,1935,1959]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E5912FFB4FCEAF85ABDD0FCBC" box="[785,967,1935,1959]" italics="true" pageId="16" pageNumber="117">Bothynus entellus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E5912FFB4FC2DF845BDF0FCB3" bold="true" box="[982,999,1936,1960]" pageId="16" pageNumber="117">B</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FC0DF845BA71FCBC" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1014,1126,1935,1959]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="stenelus">
<emphasis id="B93E767E5912FFB4FC0DF845BA12FCBC" box="[1014,1029,1936,1959]" italics="true" pageId="16" pageNumber="117">B</emphasis>
.
<emphasis id="B93E767E5912FFB4FBEFF85ABA71FCBC" box="[1044,1126,1935,1959]" italics="true" pageId="16" pageNumber="117">stenelus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E5912FFB4FB8EF85ABA90FCB3" bold="true" box="[1141,1159,1935,1960]" pageId="16" pageNumber="117">C</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FB6DF845BB1BFCBC" authorityName="Arrow" authorityYear="1937" box="[1174,1292,1936,1959]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="tricornis">
<emphasis id="B93E767E5912FFB4FB6DF845BAB2FCBC" box="[1174,1189,1936,1959]" italics="true" pageId="16" pageNumber="117">B</emphasis>
.
<emphasis id="B93E767E5912FFB4FB4EF845BB1BFCBC" box="[1205,1292,1936,1959]" italics="true" pageId="16" pageNumber="117">tricornis</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E5912FFB4FAE0F845BB3AFCB3" bold="true" box="[1307,1325,1936,1960]" pageId="16" pageNumber="117">D</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FAC6F845BB81FCBC" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[1341,1430,1936,1959]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="B93E767E5912FFB4FAC6F845BB5BFCBC" box="[1341,1356,1936,1959]" italics="true" pageId="16" pageNumber="117">B</emphasis>
.
<emphasis id="B93E767E5912FFB4FAA0F845BB81FCBC" box="[1371,1430,1936,1959]" italics="true" pageId="16" pageNumber="117">araya</emphasis>
</taxonomicName>
. Proventrite of females in ventral view.
<emphasis id="B93E767E5912FFB4FDD5F861BC28FCD7" bold="true" box="[558,575,1972,1996]" pageId="16" pageNumber="117">E</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FDB6F866BD14FCD0" box="[589,771,1971,1995]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E5912FFB4FDB6F866BD14FCD0" box="[589,771,1971,1995]" italics="true" pageId="16" pageNumber="117">Bothynus entellus</emphasis>
</taxonomicName>
(small furrows, arrow);
<emphasis id="B93E767E5912FFB4FBF8F861BA05FCD7" bold="true" box="[1027,1042,1972,1996]" pageId="16" pageNumber="117">F</emphasis>
,
<taxonomicName id="4C4AD1EF5912FFB4FBD9F861BA6CFCD0" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[1058,1147,1972,1995]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="B93E767E5912FFB4FBD9F861BA26FCD0" box="[1058,1073,1972,1995]" italics="true" pageId="16" pageNumber="117">B</emphasis>
.
<emphasis id="B93E767E5912FFB4FBBBF861BA6CFCD0" box="[1088,1147,1972,1995]" italics="true" pageId="16" pageNumber="117">araya</emphasis>
</taxonomicName>
(long furrow, arrow). Scale bars: 2 mm.
</paragraph>
</caption>
<caption id="DF35FAE45913FFB5FF6CF9D9BF07FD53" ID-DOI="http://doi.org/10.5281/zenodo.3702891" ID-Zenodo-Dep="3702891" httpUri="https://zenodo.org/record/3702891/files/figure.png" pageId="17" pageNumber="118" startId="17.[151,250,1548,1573]" targetBox="[210,1387,215,1512]" targetPageId="17">
<paragraph id="8BF5AA6C5913FFB5FF6CF9D9BF07FD53" blockId="17.[151,1436,1548,1609]" pageId="17" pageNumber="118">
<emphasis id="B93E767E5913FFB5FF6CF9D9BF08FD3E" bold="true" box="[151,287,1548,1573]" pageId="17" pageNumber="118">FIGURE 11.</emphasis>
Tergite VIII, detail of the apical invagination (arrow).
<emphasis id="B93E767E5913FFB5FCB0F9D9BD4AFD3E" bold="true" box="[843,861,1548,1573]" pageId="17" pageNumber="118">A</emphasis>
,
<taxonomicName id="4C4AD1EF5913FFB5FC91F9D8BA09FD3E" box="[874,1054,1549,1573]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E5913FFB5FC91F9D8BA09FD3E" box="[874,1054,1549,1573]" italics="true" pageId="17" pageNumber="118">Bothynus entellus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E5913FFB5FBD7F9D8BA2AFD3E" bold="true" box="[1068,1085,1549,1573]" pageId="17" pageNumber="118">B</emphasis>
,
<taxonomicName id="4C4AD1EF5913FFB5FBB1F9D8BAAFFD3E" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1098,1208,1549,1573]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="stenelus">
<emphasis id="B93E767E5913FFB5FBB1F9D8BA4EFD3F" box="[1098,1113,1549,1572]" italics="true" pageId="17" pageNumber="118">B</emphasis>
.
<emphasis id="B93E767E5913FFB5FB9DF9D8BAAFFD3E" box="[1126,1208,1549,1573]" italics="true" pageId="17" pageNumber="118">stenelus</emphasis>
</taxonomicName>
;
<emphasis id="B93E767E5913FFB5FB3EF9D9BAC0FD3E" bold="true" box="[1221,1239,1548,1573]" pageId="17" pageNumber="118">C</emphasis>
,
<taxonomicName id="4C4AD1EF5913FFB5FB1EF9D8BB4FFD3F" authorityName="Arrow" authorityYear="1937" box="[1253,1368,1549,1572]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="tricornis">
<emphasis id="B93E767E5913FFB5FB1EF9D8BAE3FD3F" box="[1253,1268,1549,1572]" italics="true" pageId="17" pageNumber="118">B</emphasis>
.
<emphasis id="B93E767E5913FFB5FAFAF9D8BB4FFD3F" box="[1281,1368,1549,1572]" italics="true" pageId="17" pageNumber="118">tricornis</emphasis>
</taxonomicName>
. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BF5AA6C5913FFB5FF3CF9A3BB16FDC2" blockId="17.[151,1437,1654,1754]" pageId="17" pageNumber="118">
<emphasis id="B93E767E5913FFB5FF3CF9A3BFFEFD8A" bold="true" box="[199,489,1654,1681]" pageId="17" pageNumber="118">Geographic distribution</emphasis>
:
<collectingCountry id="F35DEAFC5913FFB5FE0CF9A2BC57FD8A" box="[503,576,1655,1681]" name="Brazil" pageId="17" pageNumber="118">Brazil</collectingCountry>
:
<collectingRegion id="498E648E5913FFB5FDB0F9A2BCF5FD8A" box="[587,738,1655,1681]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="118">Minas Gerais</collectingRegion>
(
<figureCitation id="1371B6E95913FFB5FD0AF9A2BD41FD8A" box="[753,854,1655,1681]" captionStart="FIGURE 12" captionStartId="18.[151,250,1969,1994]" captionTargetBox="[227,1361,230,1892]" captionTargetId="figure@18.[195,1393,181,1944]" captionTargetPageId="18" captionText="FIGURE 12. Distribution map of the Bothynus entellus species group. A, Bothynus entellus; B, B. stenelus; C, B. tricornis; D, B. lancifer; E, B. araya; F, B. condacki." figureDoi="http://doi.org/10.5281/zenodo.3702893" httpUri="https://zenodo.org/record/3702893/files/figure.png" pageId="17" pageNumber="118">Fig. 12F</figureCitation>
).
<taxonomicName id="4C4AD1EF5913FFB5FC91F9A2BA29FD8A" authorityName="Duarte &amp; Grossi" authorityYear="2020" box="[874,1086,1655,1681]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="condacki">
<emphasis id="B93E767E5913FFB5FC91F9A2BA29FD8A" box="[874,1086,1655,1681]" italics="true" pageId="17" pageNumber="118">Bothynus condacki</emphasis>
</taxonomicName>
is known only from southeastern
<collectingCountry id="F35DEAFC5913FFB5FF39F94EBF1EFDAE" box="[194,265,1691,1717]" name="Brazil" pageId="17" pageNumber="118">Brazil</collectingCountry>
, in a locality dominated by Cerrado and open Rupestrian fields. This is the only species of the
<taxonomicName id="4C4AD1EF5913FFB5FADDF949BB8BFDAE" box="[1318,1436,1691,1717]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="entellus">
<emphasis id="B93E767E5913FFB5FADDF949BB8BFDAE" box="[1318,1436,1691,1717]" italics="true" pageId="17" pageNumber="118">B. entellus</emphasis>
</taxonomicName>
species group that occurs in a xeric habitat, the other species are found in very wet, forested habitats.
</paragraph>
</subSubSection>
</treatment>
</document>
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