411 lines
51 KiB
XML
411 lines
51 KiB
XML
<document id="1D1E4A8DEE7C0FCBBDCEC8B3FF151EE6" ID-DOI="10.11646/zootaxa.4996.3.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="5074540" ID-ZooBank="0F2F6614-08E3-46A9-AF21-14F7BB79A4CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625555920056" checkinUser="plazi" docAuthor="Inoue, Shota & Nomura, Shûhei" docDate="2021" docId="03CE87A5FFDD9466FF03A8C163A8278E" docLanguage="en" docName="zootaxa.4996.3.11.pdf" docOrigin="Zootaxa 4996 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudophanias tanintharyiensis Inoue & Nomura 2021, sp. nov." docType="treatment" docUuid="39786CC6-528A-4D66-A3E5-F3C02576DF60" docUuidSource="ZooBank" docVersion="3" lastPageNumber="597" masterDocId="FFF7FFDDFFD99461FF94AC2662362777" masterDocTitle="Two new species of Pseudophanias Raffray from Myanmar (Coleoptera Staphylinidae: Pselaphinae)" masterLastPageNumber="599" masterPageNumber="591" pageNumber="595" updateTime="1698992086504" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CDE5015FE6C199F70246966B8EE36740">
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<mods:title id="BC85F7810D7E2902192145B0BD06D900">Two new species of Pseudophanias Raffray from Myanmar (Coleoptera Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="5482AEAE05A9D19C15EE5E6CF362B6FA" type="personal">
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<mods:roleTerm id="BB1111FAE5ECB7271DB5F885F2539E98">Author</mods:roleTerm>
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<mods:namePart id="5CE93125B85061BFB2EC872F25A3A915">Inoue, Shota</mods:namePart>
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<mods:affiliation id="E27E6ADB1100EDF1E77F6CAD410EB930">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan. & The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:name id="5D96FEF622093C397C5C6BE0DFEDA3C7" type="personal">
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<mods:roleTerm id="9837A31DA8DD0F49FD91D1DE464F7EE7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D6EFF05BB9CDBA5BB8D4846CAF577964">Nomura, Shûhei</mods:namePart>
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<mods:nameIdentifier id="D167AD2A2D6A19AA5172D874A21FA774" type="ORCID">0000-0003-1086-5078</mods:nameIdentifier>
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<mods:affiliation id="C2F180F9995F5D1B7AAF5992524EB96F">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan. nomura @ kahaku. go. jp; https: // orcid. org / 0000 - 0003 - 1086 - 5078</mods:affiliation>
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<mods:nameIdentifier id="0CCD38AC936B7F8A65322F47C6670A2E" type="email">nomura@kahaku.go.jp</mods:nameIdentifier>
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<mods:title id="975A6A69749AB98702732D9BC9245AF5">Zootaxa</mods:title>
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<mods:part id="7DE1D3B91AEE06E6339837C220CDD87D">
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<mods:date id="C267401362C97876A062FDB6551B1A1F">2021</mods:date>
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<mods:number id="512BFFBEC1BFEFB7F36AF750899BE715">2021-07-06</mods:number>
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<mods:number id="03027C4116C3ED8E11E1ED7D6A4BCC68">4996</mods:number>
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<mods:classification id="D78E6519E5D7188F247A00EE721C8D5E">journal article</mods:classification>
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<mods:identifier id="491132143DA543400B4CCD9B2014367A" type="DOI">10.11646/zootaxa.4996.3.11</mods:identifier>
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<paragraph id="8BD836B3FFDD9465FF03A8C160492275" blockId="4.[151,639,1255,1319]" box="[151,639,1255,1282]" pageId="4" pageNumber="595">
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<heading id="D09081DFFFDD9465FF03A8C160492275" bold="true" box="[151,639,1255,1282]" fontSize="11" level="1" pageId="4" pageNumber="595" reason="1">
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<emphasis id="B913EAA1FFDD9465FF03A8C160492275" bold="true" box="[151,639,1255,1282]" pageId="4" pageNumber="595">
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<taxonomicName id="4C674D30FFDD9465FF03A8C1602D2275" ID-CoL="8Y8T4" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[151,539,1255,1282]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis" status="sp. nov.">
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<emphasis id="B913EAA1FFDD9465FF03A8C1602D2275" bold="true" box="[151,539,1255,1282]" italics="true" pageId="4" pageNumber="595">Pseudophanias tanintharyiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22057DAFFDD9465FDB6A8CE60492275" box="[546,639,1256,1282]" pageId="4" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6538FFDD9465FF03A92A633D2250" box="[151,267,1292,1319]" pageId="4" pageNumber="595" type="description">
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<paragraph id="8BD836B3FFDD9465FF03A92A633D2250" blockId="4.[151,639,1255,1319]" box="[151,267,1292,1319]" pageId="4" pageNumber="595">
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(
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<figureCitation id="135C2A36FFDD9465FF0BA92A63352250" box="[159,259,1292,1319]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1281,1305]" captionStartId-1="6.[151,250,1596,1620]" captionStartId-2="7.[151,250,1388,1412]" captionTargetBox-0="[158,1427,354,1253]" captionTargetBox-1="[152,1436,192,1572]" captionTargetBox-2="[163,1427,282,1353]" captionTargetId-0="figure-93@5.[151,1436,345,1258]" captionTargetId-1="figure-27@6.[151,1436,181,1573]" captionTargetId-2="figure-57@7.[151,1436,273,1365]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. Dorsal habitus of Pseudophanias tanintharyiensis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." captionText-1="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." captionText-2="FIGURE 6. Male genitalia of Pseudophanias tanintharyiensis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Tergite VIII. F. Sternite VIII. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5074552" figureDoi-1="http://doi.org/10.5281/zenodo.5074554" figureDoi-2="http://doi.org/10.5281/zenodo.5074558" httpUri-0="https://zenodo.org/record/5074552/files/figure.png" httpUri-1="https://zenodo.org/record/5074554/files/figure.png" httpUri-2="https://zenodo.org/record/5074558/files/figure.png" pageId="4" pageNumber="595">Figs 4–6</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37D6538FFDD9465FF03A97261AD21F9" pageId="4" pageNumber="595" type="materials_examined">
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<paragraph id="8BD836B3FFDD9465FF03A97263DD22AD" blockId="4.[151,1437,1364,2039]" pageId="4" pageNumber="595">
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<emphasis id="B913EAA1FFDD9465FF03A97263722219" bold="true" box="[151,324,1364,1390]" pageId="4" pageNumber="595">Type material.</emphasis>
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<materialsCitation id="3B0F3CEEFFDD9465FEDDA97263DD22AD" collectingDate="2017-01-22" collectorName="S. Nomura" country="Myanmar" determinerName="S. Nomura" elevation="150" latitude="14.738056" location="Winkler" longLatPrecision="21" longitude="98.19444" pageId="4" pageNumber="595" specimenCount="2" specimenCount-male="2" stateProvince="Taninthayi" typeStatus="holotype">
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<emphasis id="B913EAA1FFDD9465FEDDA97263C72219" bold="true" box="[329,497,1364,1390]" pageId="4" pageNumber="595">
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<typeStatus id="54DC8811FFDD9465FEDDA97263DA2219" box="[329,492,1364,1390]" pageId="4" pageNumber="595" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D61FD3AFFDD9465FE63A97360382218" box="[503,526,1365,1391]" pageId="4" pageNumber="595" type="male">♂</specimenCount>
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||
, “Pt. A along the gas pipeline / in
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<collectingRegion id="49A3F851FFDD9465FCEAA97261C92219" box="[894,1023,1364,1390]" country="Myanmar" name="Taninthayi" pageId="4" pageNumber="595">Tanintharyi</collectingRegion>
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<taxonomicNameLabel id="A22057DAFFDD9465FB91A973660C2218" box="[1029,1082,1365,1391]" pageId="4" pageNumber="595" rank="subSpecies">N.R.</taxonomicNameLabel>
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(
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<elevation id="004AD180FFDD9465FBD3A97266BC2218" box="[1095,1162,1364,1391]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="595" unit="m" value="150.0">150m</elevation>
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</quantity>
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) / (by
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<location id="8EB86068FFDD9465FB5BA972671C2219" LSID="urn:lsid:plazi:treatment:03CE87A5FFDD9466FF03A8C163A8278E:8EB86068FFDD9465FB5BA972671C2219" box="[1231,1322,1364,1390]" country="Myanmar" latitude="14.738056" longLatPrecision="21" longitude="98.19444" name="Winkler" pageId="4" pageNumber="595" stateProvince="Taninthayi">Winkler</location>
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extractor) / [
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||
<collectingCountry id="F3707623FFDD9465FF38A951637322E5" box="[172,325,1399,1426]" name="Myanmar" pageId="4" pageNumber="595">MYANMAR</collectingCountry>
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] /
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<geoCoordinate id="EE535074FFDD9465FEF4A95F63D322E3" box="[352,485,1401,1428]" degrees="14" direction="north" minutes="44" orientation="latitude" pageId="4" pageNumber="595" precision="15" seconds="17" value="14.738056">N14°44′17″</geoCoordinate>
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<geoCoordinate id="EE535074FFDD9465FE78A95F605B22E3" box="[492,621,1401,1428]" degrees="98" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="595" precision="15" seconds="40" value="98.19444">E98°11′40″</geoCoordinate>
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/
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<date id="FFD91073FFDD9465FD17A95E60C622E5" box="[643,752,1400,1426]" pageId="4" pageNumber="595" value="2017-01-22">
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<collectingDate id="EF9DE99BFFDD9465FD17A95E60C622E5" box="[643,752,1400,1426]" pageId="4" pageNumber="595" value="2017-01-22">22.i.2017</collectingDate>
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</date>
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,
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<collectorName id="26925365FFDD9465FD6FA95E614222E4" box="[763,884,1400,1427]" pageId="4" pageNumber="595">S. Nomura</collectorName>
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leg. // FAUNA MYANMARICA /
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<taxonomicName id="4C674D30FFDD9465FB6FA95E67AE22E5" box="[1275,1432,1400,1426]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
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,
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<taxonomicName id="4C674D30FFDD9465FF03A9BA632B22C1" authorityName="Latreille" authorityYear="1802" box="[151,285,1436,1462]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
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/
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<taxonomicName id="4C674D30FFDD9465FEBBA9BA602422C1" box="[303,530,1436,1463]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="B913EAA1FFDD9465FEBBA9BA63EC22C1" box="[303,474,1436,1462]" italics="true" pageId="4" pageNumber="595">Pseudophanias</emphasis>
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sp. 2
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</taxonomicName>
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/
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<determinerName id="7E04DA77FFDD9465FDB0A9BA60AA22C0" box="[548,668,1436,1463]" pageId="4" pageNumber="595">S. Nomura</determinerName>
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det., 2018 //
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<typeStatus id="54DC8811FFDD9465FCBCA9BA618A22C1" box="[808,956,1436,1462]" pageId="4" pageNumber="595" type="holotype">HOLOTYPE</typeStatus>
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(red) /
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<specimenCount id="9D61FD3AFFDD9465FB9DA9BB661622C0" box="[1033,1056,1437,1463]" pageId="4" pageNumber="595" type="male">♂</specimenCount>
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,
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<taxonomicName id="4C674D30FFDD9465FBBFA9BA66E022C1" authorityName="Raffray" authorityYear="1890" box="[1067,1238,1436,1462]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
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<emphasis id="B913EAA1FFDD9465FBBFA9BA66E022C1" box="[1067,1238,1436,1462]" italics="true" pageId="4" pageNumber="595">Pseudophanias</emphasis>
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</taxonomicName>
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/
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||
<taxonomicName id="4C674D30FFDD9465FB7CA9BA67AA22C1" authorityName="Inoue & Nomura" authorityYear="2021" box="[1256,1436,1436,1462]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
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<emphasis id="B913EAA1FFDD9465FB7CA9BA67AA22C1" box="[1256,1436,1436,1462]" italics="true" pageId="4" pageNumber="595">tanintharyiensis</emphasis>
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</taxonomicName>
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/ des. Inoue &
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<bibRefCitation id="EFF64B42FFDD9465FEA9A9E763D722AD" author="Nomura, S. & Aung, M. M." box="[317,481,1472,1499]" pageId="4" pageNumber="595" pagination="19 - 29" refId="ref4067" refString="Nomura, S. & Aung, M. M. (2021) Inventory studies on the subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar Part 3: A list of collected species in Tanintharyi region in November 2018. Bulletin of the National Museum of Nature and Science, Series A, 47 (1), 19 - 29." type="journal article" year="2021">Nomura, 2021</bibRefCitation>
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”
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD836B3FFDD9465FF53A9C261AD21F9" blockId="4.[151,1437,1364,2039]" pageId="4" pageNumber="595">
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<emphasis id="B913EAA1FFDD9465FF53A9C2635D2289" bold="true" box="[199,363,1508,1534]" pageId="4" pageNumber="595">
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<typeStatus id="54DC8811FFDD9465FF53A9C2635D2289" box="[199,363,1508,1534]" pageId="4" pageNumber="595" type="paratype">PARATYPES</typeStatus>
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</emphasis>
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(
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<specimenCount id="9D61FD3AFFDD9465FEE9A9C263F02289" box="[381,454,1508,1534]" pageId="4" pageNumber="595" type="male">14♂♂</specimenCount>
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,
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<specimenCount id="9D61FD3AFFDD9465FE40A9C2602A2289" box="[468,540,1508,1534]" pageId="4" pageNumber="595" type="female">19♀♀</specimenCount>
|
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):
|
||
<materialsCitation id="3B0F3CEEFFDD9465FDA0A9C261F02289" box="[564,966,1508,1535]" collectingDate="2017-01-22" collectorName="S. Nomura" country="Myanmar" determinerName="S. Nomura" elevation="150" latitude="14.738056" location="Winkler" longLatPrecision="21" longitude="98.19444" pageId="4" pageNumber="595" specimenCount="16" specimenCount-female="8" specimenCount-male="8" stateProvince="Taninthayi" typeStatus="paratype">
|
||
<specimenCount id="9D61FD3AFFDD9465FDA0A9C260592289" box="[564,623,1508,1534]" pageId="4" pageNumber="595" type="male">8♂♂</specimenCount>
|
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,
|
||
<specimenCount id="9D61FD3AFFDD9465FDE8A9C260832289" box="[636,693,1508,1534]" pageId="4" pageNumber="595" type="female">8♀♀</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3CEEFFDD9465FC47A9C2675E2154" collectingDate="2017-01-22" collectorName="Pt. B & S. Nomura & Winkler" country="Myanmar" elevation="150" latitude="14.738056" location="Nature Reserve" longLatPrecision="21" longitude="98.19444" pageId="4" pageNumber="595" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Taninthayi" typeStatus="paratype">
|
||
<specimenCount id="9D61FD3AFFDD9465FC47A9C2663B2289" box="[979,1037,1508,1534]" pageId="4" pageNumber="595" type="male">3♂♂</specimenCount>
|
||
,
|
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<specimenCount id="9D61FD3AFFDD9465FB8FA9C266622289" box="[1051,1108,1508,1534]" pageId="4" pageNumber="595" type="female">6♀♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49A3F851FFDD9465FBF7A9C266D22289" box="[1123,1252,1508,1534]" country="Myanmar" name="Taninthayi" pageId="4" pageNumber="595">Tanintharyi</collectingRegion>
|
||
<location id="8EB86068FFDD9465FB79A9C367AA2288" LSID="urn:lsid:plazi:treatment:03CE87A5FFDD9466FF03A8C163A8278E:8EB86068FFDD9465FB79A9C367AA2288" box="[1261,1436,1509,1535]" country="Myanmar" latitude="14.738056" longLatPrecision="21" longitude="98.19444" name="Nature Reserve" pageId="4" pageNumber="595" stateProvince="Taninthayi">Nature Reserve</location>
|
||
(
|
||
<quantity id="4C9F9B56FFDD9465FF0BAA2E62DC2154" box="[159,234,1544,1571]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="595" unit="m" value="150.0">
|
||
<elevation id="004AD180FFDD9465FF0BAA2E62DC2154" box="[159,234,1544,1571]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="595" unit="m" value="150.0">150 m</elevation>
|
||
</quantity>
|
||
),
|
||
<collectorName id="26925365FFDD9465FF6BAA2F63012154" box="[255,311,1545,1571]" pageId="4" pageNumber="595">Pt. B</collectorName>
|
||
along the gas pipeline,
|
||
<geoCoordinate id="EE535074FFDD9465FDD4AA2F60FE2153" box="[576,712,1545,1572]" degrees="14" direction="north" minutes="44" orientation="latitude" pageId="4" pageNumber="595" precision="15" seconds="17" value="14.738056">14°44′17″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535074FFDD9465FD47AA2F61612153" box="[723,855,1545,1572]" degrees="98" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="595" precision="15" seconds="40" value="98.19444">98°11′40″E</geoCoordinate>
|
||
,
|
||
<date id="FFD91073FFDD9465FCF6AA2E61E62155" box="[866,976,1544,1571]" pageId="4" pageNumber="595" value="2017-01-22">
|
||
<collectingDate id="EF9DE99BFFDD9465FCF6AA2E61E62155" box="[866,976,1544,1571]" pageId="4" pageNumber="595" value="2017-01-22">22 I 2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925365FFDD9465FC4FAA2E66632154" box="[987,1109,1544,1571]" pageId="4" pageNumber="595">S. Nomura</collectorName>
|
||
leg. (
|
||
<collectorName id="26925365FFDD9465FB07AA2E66C62155" box="[1171,1264,1544,1570]" pageId="4" pageNumber="595">Winkler</collectorName>
|
||
extractor)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3CEEFFDD9465FAE6AA2E67532130" collectingDate="2018-11-16" collectorName="S. Nomura" country="Myanmar" elevation="100" latitude="14.739445" location="Nature Reserve" longLatPrecision="21" longitude="98.195" pageId="4" pageNumber="595" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Taninthayi" typeStatus="paratype">
|
||
<specimenCount id="9D61FD3AFFDD9465FAE6AA2E67AE2155" box="[1394,1432,1544,1570]" pageId="4" pageNumber="595" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D61FD3AFFDD9465FF03AA0A628C2131" box="[151,186,1580,1606]" pageId="4" pageNumber="595" type="female">1♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49A3F851FFDD9465FF5EAA0A637D2131" box="[202,331,1580,1606]" country="Myanmar" name="Taninthayi" pageId="4" pageNumber="595">Tanintharyi</collectingRegion>
|
||
<location id="8EB86068FFDD9465FEC7AA0B60342130" LSID="urn:lsid:plazi:treatment:03CE87A5FFDD9466FF03A8C163A8278E:8EB86068FFDD9465FEC7AA0B60342130" box="[339,514,1581,1607]" country="Myanmar" latitude="14.739445" longLatPrecision="21" longitude="98.195" name="Nature Reserve" pageId="4" pageNumber="595" stateProvince="Taninthayi">Nature Reserve</location>
|
||
(
|
||
<quantity id="4C9F9B56FFDD9465FD80AA0A60572130" box="[532,609,1580,1607]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="595" unit="m" value="100.0">
|
||
<elevation id="004AD180FFDD9465FD80AA0A60572130" box="[532,609,1580,1607]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="595" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
),
|
||
<geoCoordinate id="EE535074FFDD9465FDE3AA0B60C9213F" box="[631,767,1581,1608]" degrees="14" direction="north" minutes="44" orientation="latitude" pageId="4" pageNumber="595" precision="15" seconds="22" value="14.739445">14°44′22″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535074FFDD9465FC98AA0B61A6213F" box="[780,912,1581,1608]" degrees="98" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="595" precision="15" seconds="42" value="98.195">98°11′42″E</geoCoordinate>
|
||
,
|
||
<date id="FFD91073FFDD9465FC09AA0A66122131" box="[925,1060,1580,1607]" pageId="4" pageNumber="595" value="2018-11-16">
|
||
<collectingDate id="EF9DE99BFFDD9465FC09AA0A66122131" box="[925,1060,1580,1607]" pageId="4" pageNumber="595" value="2018-11-16">16 XI 2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925365FFDD9465FBA5AA0A669A2130" box="[1073,1196,1580,1607]" pageId="4" pageNumber="595">S. Nomura</collectorName>
|
||
leg. (leaf litter)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3CEEFFDD9465FAE6AA0A61A121F9" collectingDate="2018-11-17" collectorName="Each" country="Myanmar" elevation="100" latitude="14.739445" location="Nature Reserve" longLatPrecision="21" longitude="98.195" pageId="4" pageNumber="595" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Taninthayi" typeStatus="paratype">
|
||
<specimenCount id="9D61FD3AFFDD9465FAE6AA0A67A12131" box="[1394,1431,1580,1606]" pageId="4" pageNumber="595" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D61FD3AFFDD9465FF03AA7662E6211D" box="[151,208,1616,1642]" pageId="4" pageNumber="595" type="female">4♀♀</specimenCount>
|
||
, same data except
|
||
<date id="FFD91073FFDD9465FE31AA76601F211D" box="[421,553,1616,1643]" pageId="4" pageNumber="595" value="2018-11-17">
|
||
<collectingDate id="EF9DE99BFFDD9465FE31AA76601F211D" box="[421,553,1616,1643]" pageId="4" pageNumber="595" value="2018-11-17">17 XI 2018</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="26925365FFDD9465FDA0AA76605A211D" box="[564,620,1616,1642]" pageId="4" pageNumber="595">Each</collectorName>
|
||
<typeStatus id="54DC8811FFDD9465FDE7AA7760E5211C" box="[627,723,1617,1643]" pageId="4" pageNumber="595" type="paratype">paratype</typeStatus>
|
||
pinned with the following label: “
|
||
<typeStatus id="54DC8811FFDD9465FBC0AA7666D5211D" box="[1108,1251,1616,1642]" pageId="4" pageNumber="595" type="paratype">PARATYPE</typeStatus>
|
||
(yellow) /
|
||
<specimenCount id="9D61FD3AFFDD9465FACBAA776742211C" box="[1375,1396,1617,1643]" pageId="4" pageNumber="595" type="male">♂</specimenCount>
|
||
(or
|
||
<specimenCount id="9D61FD3AFFDD9465FF03AA53629C21F8" box="[151,170,1653,1679]" pageId="4" pageNumber="595" type="female">♀</specimenCount>
|
||
),
|
||
<taxonomicName id="4C674D30FFDD9465FF50AA52635921F9" authorityName="Raffray" authorityYear="1890" box="[196,367,1652,1678]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EAA1FFDD9465FF50AA52635921F9" box="[196,367,1652,1678]" italics="true" pageId="4" pageNumber="595">Pseudophanias</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C674D30FFDD9465FE10AA52600E21F9" authorityName="Inoue & Nomura" authorityYear="2021" box="[388,568,1652,1678]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
|
||
<emphasis id="B913EAA1FFDD9465FE10AA52600E21F9" box="[388,568,1652,1678]" italics="true" pageId="4" pageNumber="595">tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
/ des. Inoue &
|
||
<bibRefCitation id="EFF64B42FFDD9465FD71AA5361BA21F9" author="Nomura, S. & Aung, M. M." box="[741,908,1652,1679]" pageId="4" pageNumber="595" pagination="19 - 29" refId="ref4067" refString="Nomura, S. & Aung, M. M. (2021) Inventory studies on the subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar Part 3: A list of collected species in Tanintharyi region in November 2018. Bulletin of the National Museum of Nature and Science, Series A, 47 (1), 19 - 29." type="journal article" year="2021">Nomura, 2021</bibRefCitation>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6538FFDD9465FF53AABE66572069" pageId="4" pageNumber="595" type="diagnosis">
|
||
<paragraph id="8BD836B3FFDD9465FF53AABE66572069" blockId="4.[151,1437,1364,2039]" pageId="4" pageNumber="595">
|
||
<emphasis id="B913EAA1FFDD9465FF53AABE630C21C5" bold="true" box="[199,314,1688,1714]" pageId="4" pageNumber="595">Diagnosis</emphasis>
|
||
. The new species is readily distinguished from the other members of
|
||
<taxonomicName id="4C674D30FFDD9465FBBAAABE66EF21C5" authorityName="Raffray" authorityYear="1890" box="[1070,1241,1688,1714]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EAA1FFDD9465FBBAAABE66EF21C5" box="[1070,1241,1688,1714]" italics="true" pageId="4" pageNumber="595">Pseudophanias</emphasis>
|
||
</taxonomicName>
|
||
by a combination of the following characteristics: Antennomeres 6 enlarged in male; metatibiae with slight laminate plate at mesal margin in male; tergite IV lacking discal carinae; aedeagus asymmetrical; median lobe broadened apically and curved laterally at apical part; apex longitudinally sub-truncated in dorso-ventral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6538FFDD9466FF53AB0E63A8278E" lastPageId="7" lastPageNumber="598" pageId="4" pageNumber="595" type="description">
|
||
<paragraph id="8BD836B3FFDD9465FF53AB0E62E12010" blockId="4.[151,1437,1364,2039]" pageId="4" pageNumber="595">
|
||
<emphasis id="B913EAA1FFDD9465FF53AB0E63672035" bold="true" box="[199,337,1832,1858]" pageId="4" pageNumber="595">Description</emphasis>
|
||
. Male (
|
||
<figureCitation id="135C2A36FFDD9465FE3BAB0E603A2035" box="[431,524,1832,1858]" captionStart="FIGURE 4" captionStartId="5.[151,250,1281,1305]" captionTargetBox="[158,1427,354,1253]" captionTargetId="figure-93@5.[151,1436,345,1258]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal habitus of Pseudophanias tanintharyiensis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5074552" httpUri="https://zenodo.org/record/5074552/files/figure.png" pageId="4" pageNumber="595">Figs 4A</figureCitation>
|
||
and
|
||
<figureCitation id="135C2A36FFDD9465FDDCAB0E60A72034" box="[584,657,1832,1859]" captionStart="FIGURE 5" captionStartId="6.[151,250,1596,1620]" captionTargetBox="[152,1436,192,1572]" captionTargetId="figure-27@6.[151,1436,181,1573]" captionTargetPageId="6" captionText="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." figureDoi="http://doi.org/10.5281/zenodo.5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="4" pageNumber="595">5A, B</figureCitation>
|
||
). Body length
|
||
<quantity id="4C9F9B56FFDD9465FCD4AB0E61DC2034" box="[832,1002,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" metricValueMax="2.02" metricValueMin="1.9" pageId="4" pageNumber="595" unit="mm" value="1.96" valueMax="2.02" valueMin="1.9">1.90–2.02 mm</quantity>
|
||
. Dorsal surface densely covered with setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDD9464FF53AB56665F2636" blockId="4.[151,1437,1364,2039]" lastBlockId="5.[151,1436,150,321]" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595">
|
||
Head (HL/HW 0.89–1.00) slightly wider than long, densely and coarsely punctate; frons moderately short, with frontal rostrum about 3/4 as wide as HW, with longitudinal, shallow sulcus at middle including a glabrous fovea; vertex with pair of fovea; eyes prominent, small; occipital margin in lateral view nearly straight, with dense tufts of setae; U-shaped setose sulci separate from eyes. Antennae (
|
||
<figureCitation id="135C2A36FFDD9465FCAEABFA61A72080" box="[826,913,2012,2039]" captionStart="FIGURE 5" captionStartId="6.[151,250,1596,1620]" captionTargetBox="[152,1436,192,1572]" captionTargetId="figure-27@6.[151,1436,181,1573]" captionTargetPageId="6" captionText="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." figureDoi="http://doi.org/10.5281/zenodo.5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="4" pageNumber="595">Fig. 5C</figureCitation>
|
||
) length
|
||
<quantity id="4C9F9B56FFDD9465FC64ABFA66A12080" box="[1008,1175,2012,2039]" metricMagnitude="-4" metricUnit="m" metricValue="9.85" metricValueMax="10.5" metricValueMin="9.2" pageId="4" pageNumber="595" unit="mm" value="0.985" valueMax="1.05" valueMin="0.92">0.92–1.05 mm</quantity>
|
||
; antennomeres 1 about as long as 2–3 combined; antennomeres 2–5 moniliform, successively widened apically; antennal clubs formed by antennomeres 6–11, in lateral view slightly concave dorsally; antennomeres 6 enlarged, about as long as 1, slightly convex ventrally at basal part, each with glabrous area at ventral surface; antennomeres 7–11 successively broader; antennomeres 11 largest, sub-oval. Maxillary palpi symmetrical; palpomeres 1 minute; palpomeres 2 elongate, narrowed in basal halves; palpomeres 3 small, widest at apices; palpomeres 4 fusiform.
|
||
</paragraph>
|
||
<caption id="DF18663BFFDC9464FF03A9276732226E" ID-DOI="http://doi.org/10.5281/zenodo.5074552" ID-Zenodo-Dep="5074552" box="[151,1284,1281,1306]" httpUri="https://zenodo.org/record/5074552/files/figure.png" pageId="5" pageNumber="596" startId="5.[151,250,1281,1305]" targetBox="[158,1427,354,1253]" targetPageId="5">
|
||
<paragraph id="8BD836B3FFDC9464FF03A9276732226E" blockId="5.[151,1284,1281,1306]" box="[151,1284,1281,1306]" pageId="5" pageNumber="596">
|
||
<emphasis id="B913EAA1FFDC9464FF03A9276325226E" bold="true" box="[151,275,1281,1305]" pageId="5" pageNumber="596">FIGURE 4.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName id="4C674D30FFDC9464FE5AA9276124226E" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[462,786,1281,1305]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="tanintharyiensis" status="sp. nov.">
|
||
<emphasis id="B913EAA1FFDC9464FE5AA9276124226E" box="[462,786,1281,1305]" italics="true" pageId="5" pageNumber="596">Pseudophanias tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EAA1FFDC9464FC8DA92461BF226E" bold="true" box="[793,905,1281,1306]" pageId="5" pageNumber="596">
|
||
<taxonomicNameLabel id="A22057DAFFDC9464FC8DA924615F226D" box="[793,873,1282,1306]" pageId="5" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A.
|
||
</emphasis>
|
||
Male.
|
||
<emphasis id="B913EAA1FFDC9464FC44A92461D1226D" bold="true" box="[976,999,1282,1306]" pageId="5" pageNumber="596">B.</emphasis>
|
||
Female. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836B3FFDC9464FF53A965667422F6" blockId="5.[151,1437,1346,2020]" pageId="5" pageNumber="596">Pronotum (PL/PW 0.86–1.09) about as long as wide, widest at middle, finely punctate except for antebasal area, with a median and pair of lateral antebasal foveae; antebasal area coarsely punctate.</paragraph>
|
||
<paragraph id="8BD836B3FFDC9464FF53A9AC672421D7" blockId="5.[151,1437,1346,2020]" pageId="5" pageNumber="596">
|
||
Elytra (EL/EW 0.59–0.69) much wider than long, trapezoidal; each elytron with two basal foveae; discal stria distinct, extending to posterior third. Mesoventrite (
|
||
<figureCitation id="135C2A36FFDC9464FD7FA988617022BE" box="[747,838,1454,1481]" captionStart="FIGURE 5" captionStartId="6.[151,250,1596,1620]" captionTargetBox="[152,1436,192,1572]" captionTargetId="figure-27@6.[151,1436,181,1573]" captionTargetPageId="6" captionText="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." figureDoi="http://doi.org/10.5281/zenodo.5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="5" pageNumber="596">Fig. 5D</figureCitation>
|
||
) with median setose area narrow; setose area about 1/5 as wide as mesoventral width; lateral mesoventral foveae glabrous; metaventrite coarsely punctate at middle, convex, but area just above metaventral apex roundly impressed; that impression half as long as metaventral length, about 0.3 times wider than metaventral width; anterior margin of that impression demarcated from other part of metaventrite. Legs with each femora broadest near middle; protibiae straight; mesotibiae arcuate at apical fourth; metatibiae (
|
||
<figureCitation id="135C2A36FFDC9464FE8EAA44635B210A" box="[282,365,1634,1661]" captionStart="FIGURE 5" captionStartId="6.[151,250,1596,1620]" captionTargetBox="[152,1436,192,1572]" captionTargetId="figure-27@6.[151,1436,181,1573]" captionTargetPageId="6" captionText="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." figureDoi="http://doi.org/10.5281/zenodo.5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="5" pageNumber="596">Fig. 5E</figureCitation>
|
||
) nearly straight, each mesal margin with slight laminate plate; each tarsi (
|
||
<figureCitation id="135C2A36FFDC9464FB03AA4466DE210A" box="[1175,1256,1634,1661]" captionStart="FIGURE 5" captionStartId="6.[151,250,1596,1620]" captionTargetBox="[152,1436,192,1572]" captionTargetId="figure-27@6.[151,1436,181,1573]" captionTargetPageId="6" captionText="FIGURE 5. Morphological details of Pseudophanias tanintharyiensis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Metatibia. F. SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F." figureDoi="http://doi.org/10.5281/zenodo.5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="5" pageNumber="596">Fig. 5F</figureCitation>
|
||
) with tarsomere 2 two-thirds as long as tarsomere 3; tarsal claws each asymmetrical in size; posterior claws thin, short.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDC9464FF53AA8C60DD219E" blockId="5.[151,1437,1346,2020]" pageId="5" pageNumber="596">
|
||
Abdomen (AL/AW 0.71–0.83) wider than long; tergite IV longest, twice as long as tergite V, lacking discal carinae; tergite VIII and sternite VIII as in
|
||
<figureCitation id="135C2A36FFDC9464FDE2AAE860DE219E" box="[630,744,1742,1769]" captionStart="FIGURE 6" captionStartId="7.[151,250,1388,1412]" captionTargetBox="[163,1427,282,1353]" captionTargetId="figure-57@7.[151,1436,273,1365]" captionTargetPageId="7" captionText="FIGURE 6. Male genitalia of Pseudophanias tanintharyiensis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Tergite VIII. F. Sternite VIII. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074558" httpUri="https://zenodo.org/record/5074558/files/figure.png" pageId="5" pageNumber="596">Fig. 6E, F</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDC9464FF53AAD46721200F" blockId="5.[151,1437,1346,2020]" pageId="5" pageNumber="596">
|
||
Aedeagus (
|
||
<figureCitation id="135C2A36FFDC9464FEDDAAD463F1207A" box="[329,455,1778,1805]" captionStart="FIGURE 6" captionStartId="7.[151,250,1388,1412]" captionTargetBox="[163,1427,282,1353]" captionTargetId="figure-57@7.[151,1436,273,1365]" captionTargetPageId="7" captionText="FIGURE 6. Male genitalia of Pseudophanias tanintharyiensis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Tergite VIII. F. Sternite VIII. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074558" httpUri="https://zenodo.org/record/5074558/files/figure.png" pageId="5" pageNumber="596">Fig. 6A–D</figureCitation>
|
||
)
|
||
<quantity id="4C9F9B56FFDC9464FE4FAAD460B4207A" box="[475,642,1778,1805]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="5" pageNumber="596" unit="mm" value="0.45999999999999996" valueMax="0.48" valueMin="0.44">0.44–0.48 mm</quantity>
|
||
in length, sclerotized, asymmetrical; median lobe elongate, roundly concave at middle thirds in dorsal view, strongly narrowed at posterior of apical part and then broadened apically and curved laterally at apical part, in lateral view curved ventrally and strongly tapered towards apex; apex longitudinally sub-truncated in dorso-ventral view; parameres elongate, with 4 setae at apices; endophallus indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDC9464FF53ABA461D72093" blockId="5.[151,1437,1346,2020]" pageId="5" pageNumber="596">
|
||
Female (
|
||
<figureCitation id="135C2A36FFDC9464FEB8ABA463B120EB" box="[300,391,1922,1948]" captionStart="FIGURE 4" captionStartId="5.[151,250,1281,1305]" captionTargetBox="[158,1427,354,1253]" captionTargetId="figure-93@5.[151,1436,345,1258]" captionTargetPageId="5" captionText="FIGURE 4. Dorsal habitus of Pseudophanias tanintharyiensis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5074552" httpUri="https://zenodo.org/record/5074552/files/figure.png" pageId="5" pageNumber="596">Fig. 4B</figureCitation>
|
||
). General appearance similar to male; antennae unmodified, successively widened towards apices, length
|
||
<quantity id="4C9F9B56FFDC9464FEA2AB8063ED20B6" box="[310,475,1958,1985]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.9" metricValueMin="7.6" pageId="5" pageNumber="596" unit="mm" value="0.825" valueMax="0.89" valueMin="0.76">0.76–0.89 mm</quantity>
|
||
; antennomeres 6 quadrate, slightly longer than 5 and 7; eyes smaller than in male; each metatibiae lacking laminate plate; metaventrite simple, lacking impression.
|
||
</paragraph>
|
||
<caption id="DF18663BFFDF9467FF03AA1A619E21EB" ID-DOI="http://doi.org/10.5281/zenodo.5074554" ID-Zenodo-Dep="5074554" httpUri="https://zenodo.org/record/5074554/files/figure.png" pageId="6" pageNumber="597" startId="6.[151,250,1596,1620]" targetBox="[152,1436,192,1572]" targetPageId="6">
|
||
<paragraph id="8BD836B3FFDF9467FF03AA1A619E21EB" blockId="6.[151,1437,1596,1693]" pageId="6" pageNumber="597">
|
||
<emphasis id="B913EAA1FFDF9467FF03AA1A633A2122" bold="true" box="[151,268,1596,1621]" pageId="6" pageNumber="597">FIGURE 5</emphasis>
|
||
. Morphological details of
|
||
<taxonomicName id="4C674D30FFDF9467FD82AA1B616F2122" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[534,857,1597,1621]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="tanintharyiensis" status="sp. nov.">
|
||
<emphasis id="B913EAA1FFDF9467FD82AA1B616F2122" box="[534,857,1597,1621]" italics="true" pageId="6" pageNumber="597">Pseudophanias tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EAA1FFDF9467FCCAAA1B61982122" bold="true" box="[862,942,1597,1621]" pageId="6" pageNumber="597">
|
||
<taxonomicNameLabel id="A22057DAFFDF9467FCCAAA1B61982122" box="[862,942,1597,1621]" pageId="6" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="B913EAA1FFDF9467FC63AA1A66262123" bold="true" box="[1015,1040,1596,1620]" pageId="6" pageNumber="597">A.</emphasis>
|
||
Ventral habitus.
|
||
<emphasis id="B913EAA1FFDF9467FB2CAA1B66F92122" bold="true" box="[1208,1231,1597,1621]" pageId="6" pageNumber="597">B.</emphasis>
|
||
Head and pronotum in lateral view.
|
||
<emphasis id="B913EAA1FFDF9467FEA7AA46637D210F" bold="true" box="[307,331,1632,1656]" pageId="6" pageNumber="597">C.</emphasis>
|
||
SEM image of antenna in ventral view.
|
||
<emphasis id="B913EAA1FFDF9467FD77AA4760CA210E" bold="true" box="[739,764,1633,1657]" pageId="6" pageNumber="597">D.</emphasis>
|
||
SEM image of metaventrite in ventral view.
|
||
<emphasis id="B913EAA1FFDF9467FB57AA4766ED210E" bold="true" box="[1219,1243,1633,1657]" pageId="6" pageNumber="597">E.</emphasis>
|
||
Metatibia.
|
||
<emphasis id="B913EAA1FFDF9467FADAAA476757210E" bold="true" box="[1358,1377,1633,1657]" pageId="6" pageNumber="597">F.</emphasis>
|
||
SEM image of protarsus. Scale bars: 1.0 mm in A, B; 0.5 mm in C–E; 0.1 mm in F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836B3FFDF9467FF53AAED67AA2192" blockId="6.[151,1437,1739,2017]" box="[199,1436,1739,1765]" pageId="6" pageNumber="597">
|
||
Measurements. Male (N = 7). BL
|
||
<quantity id="4C9F9B56FFDF9467FDDFAAED60C52192" box="[587,755,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" metricValueMax="2.02" metricValueMin="1.9" pageId="6" pageNumber="597" unit="mm" value="1.96" valueMax="2.02" valueMin="1.9">1.90–2.02 mm</quantity>
|
||
; HL
|
||
<quantity id="4C9F9B56FFDF9467FCB8AAED61E42192" box="[812,978,1739,1765]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="6" pageNumber="597" unit="mm" value="0.385" valueMax="0.4" valueMin="0.37">0.37–0.40 mm</quantity>
|
||
, HW
|
||
<quantity id="4C9F9B56FFDF9467FB81AAED668B2192" box="[1045,1213,1739,1765]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="6" pageNumber="597" unit="mm" value="0.415" valueMax="0.44" valueMin="0.39">0.39–0.44 mm</quantity>
|
||
; PL
|
||
<quantity id="4C9F9B56FFDF9467FB65AAED67AE2192" box="[1265,1432,1739,1765]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="6" pageNumber="597" unit="mm" value="0.41" valueMax="0.44" valueMin="0.38">0.38–0.44 mm</quantity>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDF9467FF03AAC967AB207E" blockId="6.[151,1437,1739,2017]" box="[151,1437,1775,1802]" pageId="6" pageNumber="597">
|
||
PW
|
||
<quantity id="4C9F9B56FFDF9467FF5FAAC96342207E" box="[203,372,1775,1801]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="6" pageNumber="597" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43–0.47 mm</quantity>
|
||
; EL
|
||
<quantity id="4C9F9B56FFDF9467FE3AAAC96060207E" box="[430,598,1775,1802]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="6" pageNumber="597" unit="mm" value="0.53" valueMax="0.56" valueMin="0.5">0.50–0.56 mm</quantity>
|
||
, EW
|
||
<quantity id="4C9F9B56FFDF9467FD0FAAC96172207E" box="[667,836,1775,1801]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.4" metricValueMin="7.7" pageId="6" pageNumber="597" unit="mm" value="0.8049999999999999" valueMax="0.84" valueMin="0.77">0.77–0.84 mm</quantity>
|
||
; AL
|
||
<quantity id="4C9F9B56FFDF9467FCEBAAC9661E207E" box="[895,1064,1775,1802]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="6.5" metricValueMin="5.5" pageId="6" pageNumber="597" unit="mm" value="0.6" valueMax="0.65" valueMin="0.55">0.55–0.65 mm</quantity>
|
||
, AW
|
||
<quantity id="4C9F9B56FFDF9467FBFFAAC96722207E" box="[1131,1300,1775,1801]" metricMagnitude="-4" metricUnit="m" metricValue="7.95" metricValueMax="8.1" metricValueMin="7.8" pageId="6" pageNumber="597" unit="mm" value="0.795" valueMax="0.81" valueMin="0.78">0.78–0.81 mm</quantity>
|
||
. Female (N
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDF9467FF03AB3567AB205A" blockId="6.[151,1437,1739,2017]" box="[151,1437,1811,1838]" pageId="6" pageNumber="597">
|
||
= 7). BL
|
||
<quantity id="4C9F9B56FFDF9467FE92AB356399205A" box="[262,431,1811,1837]" metricMagnitude="-3" metricUnit="m" metricValue="1.905" metricValueMax="1.97" metricValueMin="1.84" pageId="6" pageNumber="597" unit="mm" value="1.905" valueMax="1.97" valueMin="1.84">1.84–1.97 mm</quantity>
|
||
; HL
|
||
<quantity id="4C9F9B56FFDF9467FE7FAB3560A2205A" box="[491,660,1811,1837]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="6" pageNumber="597" unit="mm" value="0.39" valueMax="0.4" valueMin="0.38">0.38–0.40 mm</quantity>
|
||
; HW
|
||
<quantity id="4C9F9B56FFDF9467FD4FAB3561B9205A" box="[731,911,1811,1838]" metricMagnitude="-4" metricUnit="m" metricValue="3.875" metricValueMax="4.05" metricValueMin="3.7" pageId="6" pageNumber="597" unit="mm" value="0.3875" valueMax="0.405" valueMin="0.37">0.37–0.405 mm</quantity>
|
||
(HL/HW 0.96–1.08); PL
|
||
<quantity id="4C9F9B56FFDF9467FB28AB356752205A" box="[1212,1380,1811,1837]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="6" pageNumber="597" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40–0.44 mm</quantity>
|
||
; PW
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDF9467FF03AB1167AA2026" blockId="6.[151,1437,1739,2017]" box="[151,1436,1847,1874]" pageId="6" pageNumber="597">
|
||
<quantity id="4C9F9B56FFDF9467FF03AB11630C2026" box="[151,314,1847,1874]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="6" pageNumber="597" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
|
||
(PL/PW 0.95–1.07); EL
|
||
<quantity id="4C9F9B56FFDF9467FDC5AB1160C12026" box="[593,759,1847,1873]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.9" metricValueMin="4.6" pageId="6" pageNumber="597" unit="mm" value="0.475" valueMax="0.49" valueMin="0.46">0.46–0.49 mm</quantity>
|
||
; EW
|
||
<quantity id="4C9F9B56FFDF9467FCA0AB1161E02026" box="[820,982,1847,1873]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.2" metricValueMin="7.6" pageId="6" pageNumber="597" unit="mm" value="0.79" valueMax="0.82" valueMin="0.76">0.76–0.82 mm</quantity>
|
||
(EL/EW 0.58–0.60); AL
|
||
<quantity id="4C9F9B56FFDF9467FB66AB1167AE2026" box="[1266,1432,1847,1874]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="6" pageNumber="597" unit="mm" value="0.615" valueMax="0.65" valueMin="0.58">0.58–0.65 mm</quantity>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDF9466FF03AB7D63A8278E" blockId="6.[151,1437,1739,2017]" lastBlockId="7.[151,1436,151,249]" lastPageId="7" lastPageNumber="598" pageId="6" pageNumber="597">
|
||
AW
|
||
<quantity id="4C9F9B56FFDF9467FF5EAB7D635B2002" box="[202,365,1883,1910]" metricMagnitude="-4" metricUnit="m" metricValue="7.85" metricValueMax="8.2" metricValueMin="7.5" pageId="6" pageNumber="597" unit="mm" value="0.7849999999999999" valueMax="0.82" valueMin="0.75">0.75–0.82 mm</quantity>
|
||
(AL/AW 0.77–0.82).
|
||
<emphasis id="B913EAA1FFDF9467FF53AB59635720EE" bold="true" box="[199,353,1919,1945]" pageId="6" pageNumber="597">Distribution.</emphasis>
|
||
<collectingCountry id="F3707623FFDF9467FEF3AB5963E220EE" box="[359,468,1919,1945]" name="Myanmar" pageId="6" pageNumber="597">Myanmar</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A3F851FFDF9467FE77AB5960F420ED" box="[483,706,1919,1946]" country="Myanmar" name="Taninthayi" pageId="6" pageNumber="597">Tanintharyi Region</collectingRegion>
|
||
).
|
||
<emphasis id="B913EAA1FFDF9467FF53AB85637820CA" bold="true" box="[199,334,1955,1981]" pageId="6" pageNumber="597">Etymology.</emphasis>
|
||
The specific epithet ‘
|
||
<emphasis id="B913EAA1FFDF9467FDD6AB85613620CA" box="[578,768,1955,1981]" italics="true" pageId="6" pageNumber="597">tanintharyiensis’</emphasis>
|
||
refers to the
|
||
<typeStatus id="54DC8811FFDF9467FC07AB8561F520CA" box="[915,963,1955,1981]" pageId="6" pageNumber="597">type</typeStatus>
|
||
locality,
|
||
<collectingRegion id="49A3F851FFDF9467FBBEAB85669820CA" box="[1066,1198,1955,1981]" country="Myanmar" name="Taninthayi" pageId="6" pageNumber="597">Tanintharyi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3707623FFDF9467FB2DAB85671E20CA" box="[1209,1320,1955,1981]" name="Myanmar" pageId="6" pageNumber="597">Myanmar</collectingCountry>
|
||
.
|
||
<emphasis id="B913EAA1FFDF9467FF53ABE1630D2096" bold="true" box="[199,315,1991,2017]" pageId="6" pageNumber="597">Remarks.</emphasis>
|
||
The female of the new species is similar to the female of
|
||
<taxonomicName id="4C674D30FFDF9467FC4FABE166CB2096" authorityName="Inoue, Nomura & Yin" authorityYear="2020" box="[987,1277,1991,2017]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="excavatus">
|
||
<emphasis id="B913EAA1FFDF9467FC4FABE166CB2096" box="[987,1277,1991,2017]" italics="true" pageId="6" pageNumber="597">Pseudophanias excavatus</emphasis>
|
||
</taxonomicName>
|
||
, but it can be distinguished by the shape of antennomeres (antennomeres 2–10 each strongly transverse in
|
||
<taxonomicName id="4C674D30FFDE9466FB45ACB1676927C7" authorityName="Inoue, Nomura & Yin" authorityYear="2020" box="[1233,1375,151,176]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="excavatus">
|
||
<emphasis id="B913EAA1FFDE9466FB45ACB1676927C7" box="[1233,1375,151,176]" italics="true" pageId="7" pageNumber="598">P. excavatus</emphasis>
|
||
</taxonomicName>
|
||
), the narrower mesoventral median setose area than in
|
||
<emphasis id="B913EAA1FFDE9466FD51AC9D616027A3" box="[709,854,187,212]" italics="true" pageId="7" pageNumber="598">
|
||
<taxonomicName id="4C674D30FFDE9466FD51AC9D616427A3" authorityName="Inoue, Nomura & Yin" authorityYear="2020" box="[709,850,187,212]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="excavatus">P. excavatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and the glabrous lateral mesoventral foveae (setose in that of
|
||
<taxonomicName id="4C674D30FFDE9466FE97ACF963BB278F" authorityName="Inoue, Nomura & Yin" authorityYear="2020" box="[259,397,223,248]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="excavatus">
|
||
<emphasis id="B913EAA1FFDE9466FE97ACF963BB278F" box="[259,397,223,248]" italics="true" pageId="7" pageNumber="598">P. excavatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |