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<document id="09A6C6F006B5CDB27FCCE4079353CFBD" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE78B616DD9E4603FED90164" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius schuelkei S. Tarasov 2017, new species" docType="treatment" docUuid="CA113116-C954-450D-8576-5D92D379C637" docUuidSource="ZooBank" docVersion="12" lastPageNumber="139" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="138" updateTime="1698469310636" updateUser="ExternalLinkService">
<mods:mods id="852EA02CE2CB7A72CC566F7D34BABE4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DA31FE115EECF0651FAA82D846E492F9">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="FFA26492DE1EAFBD02C19711C14D5AAD">Tarasov, Sergei</mods:namePart>
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<mods:date id="9F4267E1F9AEE91D804BEABE81143691">2017</mods:date>
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<mods:number id="0F23FA379D01F38A7DCB17C96B707F8A">2017-10-04</mods:number>
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<mods:identifier id="E98EC4D2C7B7AA7B46618A20419E0CB7" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<treatment id="03C187E1AE78B616DD9E4603FED90164" ID-DOI="http://doi.org/10.5281/zenodo.6010842" ID-GBIF-Taxon="135106125" ID-Zenodo-Dep="6010842" LSID="urn:lsid:zoobank.org:act:CA113116-C954-450D-8576-5D92D379C637" httpUri="http://treatment.plazi.org/id/03C187E1AE78B616DD9E4603FED90164" lastPageId="38" lastPageNumber="139" pageId="37" pageNumber="138">
<subSubSection id="C372657CAE78B615DD9E4603FE9B03F7" pageId="37" pageNumber="138" type="nomenclature">
<paragraph id="8BD736F7AE78B615DD9E4603FD730316" blockId="37.[151,954,332,427]" box="[151,680,332,358]" pageId="37" pageNumber="138">
<heading id="D09F819BAE78B615DD9E4603FD730316" bold="true" box="[151,680,332,358]" fontSize="11" level="1" pageId="37" pageNumber="138" reason="1">
<emphasis id="B91CEAE5AE78B615DD9E4603FD730316" bold="true" box="[151,680,332,358]" pageId="37" pageNumber="138">
<taxonomicName id="4C684D74AE78B615DD9E4603FDCA0316" ID-CoL="6TNLX" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[151,529,332,358]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="B91CEAE5AE78B615DD9E4603FE7D0316" bold="true" box="[151,422,332,358]" italics="true" pageId="37" pageNumber="138">Parachorius schuelkei</emphasis>
Tarasov
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE78B615DF114603FD730316" box="[536,680,332,358]" pageId="37" pageNumber="138" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE78B615DD9E4621FE9B03F7" blockId="37.[151,954,332,427]" box="[151,320,366,391]" pageId="37" pageNumber="138">
(
<figureCitation id="13532A72AE78B615DD964621FF2703F7" box="[159,252,366,391]" captionStart="FIGURE 21" captionStartId="37.[151,250,1909,1931]" captionTargetBox="[151,1436,973,1878]" captionTargetId="figure@37.[151,1436,973,1888]" captionTargetPageId="37" captionText="FIGURE 21. Morphology of Parachorius schuelkei. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype)." httpUri="https://zenodo.org/record/1001959/files/figure.png" pageId="37" pageNumber="138">Figs. 21</figureCitation>
,
<figureCitation id="13532A72AE78B615DC014620FEE303F7" box="[264,312,367,391]" captionStart="FIGURE 27" captionStartId="44.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1748]" captionTargetId="figure@44.[151,1436,193,1748]" captionTargetPageId="44" captionText="FIGURE 27. Distribution maps of Parachorius species. (A) Parachorius peninsularis. (B) Parachorius pseudojavanus. (C) Parachorius schuelkei. (D) Parachorius semsanganus. (E) Parachorius solodovnikovi. (F) Parachorius thomsoni. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study. Literature data — localities retrieved from literature." httpUri="https://zenodo.org/record/1001971/files/figure.png" pageId="37" pageNumber="138">27C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B615DD9E46DDFC6103DB" box="[151,954,402,427]" pageId="37" pageNumber="138" type="description">
<paragraph id="8BD736F7AE78B615DD9E46DDFC6103DB" blockId="37.[151,954,332,427]" box="[151,954,402,427]" pageId="37" pageNumber="138">
urn:lsid:zoobank.org:act:CA
<date id="FFD61037AE78B615DCD946DDFDFC03DB" box="[464,551,402,427]" pageId="37" pageNumber="138" value="16-11-31">113116</date>
-C954-450D-8576-5D92D379C637
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B615DD9E4695FC830383" box="[151,856,474,499]" pageId="37" pageNumber="138" type="materials_examined">
<paragraph id="8BD736F7AE78B615DD9E4695FC830383" blockId="37.[151,1436,474,931]" box="[151,856,474,499]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DD9E4695FEF60383" bold="true" box="[151,301,474,499]" pageId="37" pageNumber="138">Type locality</emphasis>
. Baoshan Pref., Gaoligong Shan, Yunnan, China.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B615DDCE46B1FB7F0067" box="[199,1188,510,535]" pageId="37" pageNumber="138" type="etymology">
<paragraph id="8BD736F7AE78B615DDCE46B1FB7F0067" blockId="37.[151,1436,474,931]" box="[199,1188,510,535]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE46B1FE950067" bold="true" box="[199,334,510,535]" pageId="37" pageNumber="138">Etymology.</emphasis>
This species is names after Michael Schülke the collector of the
<typeStatus id="54D38855AE78B615D92C46B0FB8E0067" box="[1061,1109,511,535]" pageId="37" pageNumber="138">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B615DDCE456DFDCE004A" box="[199,533,546,571]" pageId="37" pageNumber="138" type="distribution">
<paragraph id="8BD736F7AE78B615DDCE456DFDCE004A" blockId="37.[151,1436,474,931]" box="[199,533,546,571]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE456DFEBB004B" bold="true" box="[199,352,546,571]" pageId="37" pageNumber="138">Distribution.</emphasis>
<collectingCountry id="F37F7667AE78B615DC6E456DFE76004B" box="[359,429,546,571]" name="China" pageId="37" pageNumber="138">China</collectingCountry>
,
<collectingRegion id="49ACF815AE78B615DCB1456CFDCA004A" box="[440,529,547,570]" country="China" name="Yunnan" pageId="37" pageNumber="138">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B615DDCE4509FE3F00BB" pageId="37" pageNumber="138" type="diagnosis">
<paragraph id="8BD736F7AE78B615DDCE4509FE3F00BB" blockId="37.[151,1436,474,931]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE4509FE9A002F" bold="true" box="[199,321,582,607]" pageId="37" pageNumber="138">Diagnosis.</emphasis>
The external morphology of this species is most similar to
<taxonomicName id="4C684D74AE78B615DEEA4508FB8D002E" authorityName="Tarasov" authorityYear="2017" box="[995,1110,583,606]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE78B615DEEA4508FB8D002E" box="[995,1110,583,606]" italics="true" pageId="37" pageNumber="138">P. javanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D74AE78B615D9864508FA89002F" authorityName="S. Tarasov" authorityYear="2017" box="[1167,1362,582,607]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE78B615D9864508FA89002F" box="[1167,1362,582,607]" italics="true" pageId="37" pageNumber="138">P. pseudojavanus</emphasis>
</taxonomicName>
, but it can be easily distinguished from both of these species by the black coloration and simple, apically rounded shape of aedeagal parameres. In addition,
<taxonomicName id="4C684D74AE78B615DF3245C0FD6900D6" authorityName="Tarasov" authorityYear="2017" box="[571,690,655,678]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE78B615DF3245C0FD6900D6" box="[571,690,655,678]" italics="true" pageId="37" pageNumber="138">P. javanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D74AE78B615DFF945C0FC6E00D7" authorityName="S. Tarasov" authorityYear="2017" box="[752,949,654,679]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE78B615DFF945C0FC6E00D7" box="[752,949,654,679]" italics="true" pageId="37" pageNumber="138">P. pseudojavanus</emphasis>
</taxonomicName>
are known from
<collectingRegion id="49ACF815AE78B615D98F45C0FB6300D6" box="[1158,1208,655,678]" country="Indonesia" name="Jawa Timur" pageId="37" pageNumber="138">Java</collectingRegion>
and
<taxonomicName id="4C684D74AE78B615D9FF45C0FAA500D7" authorityName="S. Tarasov" authorityYear="2017" box="[1270,1406,654,679]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B91CEAE5AE78B615D9FF45C0FAA500D7" box="[1270,1406,654,679]" italics="true" pageId="37" pageNumber="138">P. schuelkei</emphasis>
</taxonomicName>
is distributed in southern
<collectingCountry id="F37F7667AE78B615DC9345FDFE3A00BB" box="[410,481,690,715]" name="China" pageId="37" pageNumber="138">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE78B616DDCE4599FB7C03F8" lastPageId="38" lastPageNumber="139" pageId="37" pageNumber="138" type="description">
<paragraph id="8BD736F7AE78B615DDCE4599FBEF009F" blockId="37.[151,1436,474,931]" box="[199,1076,726,751]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE4599FE8A009F" bold="true" box="[199,337,726,751]" pageId="37" pageNumber="138">Description</emphasis>
.
<emphasis id="B91CEAE5AE78B615DC564599FE44009F" box="[351,415,726,751]" italics="true" pageId="37" pageNumber="138">Body.</emphasis>
Body black, length
<quantity id="4C909B12AE78B615DF884599FCD0009E" box="[641,779,726,751]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="37" pageNumber="138" unit="mm" value="6.65" valueMax="6.8" valueMin="6.5">6.5–6.8 mm</quantity>
. Elytra uniformly colored.
</paragraph>
<paragraph id="8BD736F7AE78B615DDCE45B5FB6A0163" blockId="37.[151,1436,474,931]" box="[199,1201,762,787]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE45B5FED00163" box="[199,267,762,787]" italics="true" pageId="37" pageNumber="138">Head.</emphasis>
Clypeus with two distinctly produced teeth, their outer margin not notched basally.
</paragraph>
<paragraph id="8BD736F7AE78B615DDCE4450FB76012B" blockId="37.[151,1436,474,931]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE4450FEE60146" box="[199,317,799,822]" italics="true" pageId="37" pageNumber="138">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 0.5– 2.0 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE78B615DDCE4429FD1B010E" blockId="37.[151,1436,474,931]" box="[199,704,870,895]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE4429FECE010F" box="[199,277,870,895]" italics="true" pageId="37" pageNumber="138">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE78B615DDCE44C3FD9401D3" blockId="37.[151,1436,474,931]" box="[199,591,906,931]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DDCE44C3FEC901D3" box="[199,274,908,931]" italics="true" pageId="37" pageNumber="138">Wings.</emphasis>
Wings normally developed.
</paragraph>
<caption id="DF17667FAE78B615DD9E403AFDAC05B9" httpUri="https://zenodo.org/record/1001959/files/figure.png" pageId="37" pageNumber="138" targetBox="[151,1436,973,1878]" targetPageId="37">
<paragraph id="8BD736F7AE78B615DD9E403AFDAC05B9" blockId="37.[151,1437,1909,1993]" pageId="37" pageNumber="138">
<emphasis id="B91CEAE5AE78B615DD9E403AFEFD05FB" bold="true" box="[151,294,1909,1931]" pageId="37" pageNumber="138">FIGURE 21.</emphasis>
Morphology of
<taxonomicName id="4C684D74AE78B615DCD6403AFD1205FB" authorityName="S. Tarasov" authorityYear="2017" box="[479,713,1909,1931]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="138" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B91CEAE5AE78B615DCD6403AFD1205FB" box="[479,713,1909,1931]" italics="true" pageId="37" pageNumber="138">Parachorius schuelkei</emphasis>
</taxonomicName>
. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype).
</paragraph>
</caption>
<paragraph id="8BD736F7AE7BB616DDCE47D6FBBD0330" blockId="38.[151,1437,151,788]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE47D6FED802C0" box="[199,259,153,176]" italics="true" pageId="38" pageNumber="139">Legs.</emphasis>
Profemur with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth almost not modified in males, approximately as wide as second tooth; first and second teeth distinctly separated from each other; first tooth normal, distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, not modified. Metatibia slightly sinuate, inner margin not denticulate, straight, not produced apically.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4602FB1F0315" blockId="38.[151,1437,151,788]" box="[199,1220,332,357]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4602FEB90314" box="[199,354,333,356]" italics="true" pageId="38" pageNumber="139">Metasternum.</emphasis>
Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4620FB7C03F8" blockId="38.[151,1437,151,788]" box="[199,1191,367,392]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4620FEE003F8" box="[199,315,367,392]" italics="true" pageId="38" pageNumber="139">Aedeagus.</emphasis>
Parameres symmetrical, simple, apex widely rounded and inferiorly angulate.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DDCE46DBFED90164" pageId="38" pageNumber="139" type="materials_examined">
<paragraph id="8BD736F7AE7BB616DDCE46DBFC6003DD" blockId="38.[151,1437,151,788]" box="[199,955,404,429]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE46DBFEE003DD" bold="true" box="[199,315,404,429]" pageId="38" pageNumber="139">
<typeStatus id="54D38855AE7BB616DDCE46DBFEED03DD" box="[199,310,404,429]" pageId="38" pageNumber="139" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="13532A72AE7BB616DC8246DBFE2803DC" box="[395,499,404,429]" captionStart="FIGURE 21" captionStartId="37.[151,250,1909,1931]" captionTargetBox="[151,1436,973,1878]" captionTargetId="figure@37.[151,1436,973,1888]" captionTargetPageId="37" captionText="FIGURE 21. Morphology of Parachorius schuelkei. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype)." httpUri="https://zenodo.org/record/1001959/files/figure.png" pageId="38" pageNumber="139">Fig. 21D</figureCitation>
). Profemur with sinuate anterior margin.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE46F8FED90164" blockId="38.[151,1437,151,788]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE46F8FDF703A0" bold="true" box="[199,556,439,464]" pageId="38" pageNumber="139">Material examined. Holotype</emphasis>
:
<materialsCitation id="3B003CAAAE7BB616DF4946F8FAC20068" ID-GBIF-Occurrence="1658411523" collectingDate="2007-05-31" collectionCode="ZMHB" collectorName="M. Schulke" country="China" county="Baoshan Pref." elevation="2200" latitude="24.825556" location="Tengchong" longLatPrecision="21" longitude="98.76833" municipality="Gaoligong Shan" pageId="38" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D6EFD7EAE7BB616DF4946F8FD4803A0" box="[576,659,439,464]" pageId="38" pageNumber="139" type="male">
<emphasis id="B91CEAE5AE7BB616DF4946F8FD4803A0" bold="true" box="[576,659,439,464]" pageId="38" pageNumber="139">1 male</emphasis>
</specimenCount>
, &quot;
<collectingCountry id="F37F7667AE7BB616DFA746F7FCD503A0" box="[686,782,440,464]" name="China" pageId="38" pageNumber="139">CHINA</collectingCountry>
:
<collectingRegion id="49ACF815AE7BB616DE1446F7FCA803BF" box="[797,883,440,463]" country="China" name="Yunnan" pageId="38" pageNumber="139">Yunnan</collectingRegion>
[CH07-13],
<collectingCounty id="62B64E7BAE7BB616D90246F8FB6B03A0" box="[1035,1200,439,464]" pageId="38" pageNumber="139">Baoshan Pref.</collectingCounty>
,
<collectingMunicipality id="6BB3AC8DAE7BB616D9B646F8FAA003A0" box="[1215,1403,439,464]" pageId="38" pageNumber="139">Gaoligong Shan</collectingMunicipality>
, E pass,
<quantity id="4C909B12AE7BB616DDDC4693FECF0385" box="[213,276,476,501]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="38" pageNumber="139" unit="km" value="36.0">36km</quantity>
SE
<location id="8EB7602CAE7BB616DC4B4693FE1A0385" LSID="urn:lsid:plazi:treatment:03C187E1AE78B616DD9E4603FED90164:8EB7602CAE7BB616DC4B4693FE1A0385" box="[322,449,476,501]" country="China" county="Baoshan Pref." latitude="24.825556" longLatPrecision="21" longitude="98.76833" municipality="Gaoligong Shan" name="Tengchong" pageId="38" pageNumber="139" stateProvince="Yunnan">Tengchong</location>
,
<elevation id="0045D1C4AE7BB616DCC54693FDC60384" box="[460,541,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="38" pageNumber="139" unit="m" value="2200.0">
<quantity id="4C909B12AE7BB616DCC54693FDC60384" box="[460,541,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="38" pageNumber="139" unit="m" value="2200.0">2200m</quantity>
</elevation>
,
<geoCoordinate id="EE5C5030AE7BB616DF2E4693FD770384" box="[551,684,476,500]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="15" value="24.825556">24°49'32''N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DFBE4693FC9B0385" box="[695,832,476,501]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="15" value="98.76833">98° 46'06''E</geoCoordinate>
, decid. forest, litter, wood, fungi sifted,
<collectingDate id="EF92E9DFAE7BB616D9F54693FAAA0384" box="[1276,1393,476,500]" pageId="38" pageNumber="139" value="2007-05-31">
<date id="FFD61037AE7BB616D9F54693FAAA0384" box="[1276,1393,476,500]" pageId="38" pageNumber="139" value="2007-05-31">31.V.2007</date>
</collectingDate>
,
<collectorName id="269D5321AE7BB616D8754692FF2A0068" pageId="38" pageNumber="139">M. Schülke</collectorName>
/
<typeStatus id="54D38855AE7BB616DC00454FFE460068" box="[265,413,512,536]" pageId="38" pageNumber="139" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C684D74AE7BB616DCAD46B0FCC20067" authority="S. Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[420,793,511,536]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="species" species="schuelkei">Parachorius schuelkei S. Tarasov</taxonomicName>
det. 2010&quot;,
<geoCoordinate id="EE5C5030AE7BB616DEAD454FFBF10068" box="[932,1066,512,536]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="15" value="24.825556">24°49'32&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616D93E46B0FB710068" box="[1079,1194,511,536]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="15" value="98.76833">98°46'6&quot;E</geoCoordinate>
(
<collectionCode id="ED79AE32AE7BB616D9B3454FFACA0068" box="[1210,1297,512,536]" pageId="38" pageNumber="139">ZMHB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE7BB616D82D46B0FE07004D" ID-GBIF-Occurrence="1658411535" country="China" location="China" pageId="38" pageNumber="139" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE7BB616D82D46B0FE5B004D" bold="true" pageId="38" pageNumber="139">
<typeStatus id="54D38855AE7BB616D82D46B0FA470068" box="[1316,1436,511,536]" pageId="38" pageNumber="139" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D6EFD7EAE7BB616DDA9456BFF2F004D" box="[160,244,548,573]" pageId="38" pageNumber="139" type="male">1 male</specimenCount>
,
<specimenCount id="9D6EFD7EAE7BB616DC0D456BFEA2004D" box="[260,377,548,573]" pageId="38" pageNumber="139" type="female">2 females</specimenCount>
)
</emphasis>
:
<location id="8EB7602CAE7BB616DC9A456BFE07004D" LSID="urn:lsid:plazi:treatment:03C187E1AE78B616DD9E4603FED90164:8EB7602CAE7BB616DC9A456BFE07004D" box="[403,476,548,573]" country="China" name="China" pageId="38" pageNumber="139">
<collectingCountry id="F37F7667AE7BB616DC9A456BFE07004D" box="[403,476,548,573]" name="China" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DC9A456BFE07004D" bold="true" box="[403,476,548,573]" pageId="38" pageNumber="139">China</emphasis>
</collectingCountry>
</location>
</materialsCitation>
:
<materialsCitation id="3B003CAAAE7BB616DCE6456BFEB300F4" ID-GBIF-Occurrence="1658411520" collectingDate="2007-05-30" collectionCode="ZMHB" collectorName="M. Schulke" country="China" elevation="2110" latitude="24.837778" location="Tengchong" longLatPrecision="21" longitude="98.76195" municipality="Xiaheishan" pageId="38" pageNumber="139" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B91CEAE5AE7BB616DCE6456BFD16004D" bold="true" box="[495,717,548,573]" pageId="38" pageNumber="139">
<specimenCount id="9D6EFD7EAE7BB616DCE6456BFD8D004D" box="[495,598,548,573]" pageId="38" pageNumber="139" type="female">1 female</specimenCount>
<typeStatus id="54D38855AE7BB616DF6B456BFD16004D" box="[610,717,548,573]" pageId="38" pageNumber="139" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingCountry id="F37F7667AE7BB616DFD6456BFCE7004C" box="[735,828,548,572]" name="China" pageId="38" pageNumber="139">CHINA</collectingCountry>
:
<collectingRegion id="49ACF815AE7BB616DE43456AFC44004C" box="[842,927,549,572]" country="China" name="Yunnan" pageId="38" pageNumber="139">Yunnan</collectingRegion>
[CH07-11], Baoshan Prefecture, Gaoligong Shan, m.
<collectingMunicipality id="6BB3AC8DAE7BB616DC084508FE590010" box="[257,386,583,608]" pageId="38" pageNumber="139">Xiaheishan</collectingMunicipality>
,
<quantity id="4C909B12AE7BB616DC844508FE0F0010" box="[397,468,583,608]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="139" unit="km" value="35.0">35 km</quantity>
SE
<location id="8EB7602CAE7BB616DF0A4508FD590010" LSID="urn:lsid:plazi:treatment:03C187E1AE78B616DD9E4603FED90164:8EB7602CAE7BB616DF0A4508FD590010" box="[515,642,583,608]" country="China" latitude="24.837778" longLatPrecision="21" longitude="98.76195" municipality="Xiaheishan" name="Tengchong" pageId="38" pageNumber="139" stateProvince="Yunnan">Tengchong</location>
,
<geoCoordinate id="EE5C5030AE7BB616DF844508FCCF0010" box="[653,788,583,608]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="15" value="24.837778">24°50'16&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DE164508FC790010" box="[799,930,583,608]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="15" value="98.76195">98°45'43&quot;E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616DEA74507FBDF002F" box="[942,1028,584,608]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="38" pageNumber="139" unit="m" value="2110.0">
<quantity id="4C909B12AE7BB616DEA74507FBDF002F" box="[942,1028,584,608]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="38" pageNumber="139" unit="m" value="2110.0">2110 m</quantity>
</elevation>
, deciduous forest, on
<collectingDate id="EF92E9DFAE7BB616D9F34507FAB70010" box="[1274,1388,584,608]" pageId="38" pageNumber="139" value="2007-05-30">
<date id="FFD61037AE7BB616D9F34507FAB70010" box="[1274,1388,584,608]" pageId="38" pageNumber="139" value="2007-05-30">30/v/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616D8754507FF2800F5" pageId="38" pageNumber="139">M. Schülke</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616DC004523FEBB00F4" box="[265,352,620,644]" pageId="38" pageNumber="139">ZMHB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE7BB616DC7A4523FA5A00D8" ID-GBIF-Occurrence="1658411539" collectingDate="2007-05-31" collectionCode="ZMHB" collectorName="M. Schulke" country="China" county="Baoshan Prefecture" elevation="2200" latitude="24.825556" location="Tengchong" longLatPrecision="21" longitude="98.76833" municipality="Gaoligong Shan" pageId="38" pageNumber="139" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B91CEAE5AE7BB616DC7A4523FD9200F5" bold="true" box="[371,585,620,645]" pageId="38" pageNumber="139">
<specimenCount id="9D6EFD7EAE7BB616DC7A4523FE0C00F5" box="[371,471,620,645]" pageId="38" pageNumber="139" type="female">1 female</specimenCount>
<typeStatus id="54D38855AE7BB616DCD74523FD9200F5" box="[478,585,620,645]" pageId="38" pageNumber="139" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingCountry id="F37F7667AE7BB616DF5E4523FD6F00F4" box="[599,692,620,644]" name="China" pageId="38" pageNumber="139">CHINA</collectingCountry>
:
<collectingRegion id="49ACF815AE7BB616DFB74522FCC800F4" box="[702,787,621,644]" country="China" name="Yunnan" pageId="38" pageNumber="139">Yunnan</collectingRegion>
[CH07-13],
<collectingCounty id="62B64E7BAE7BB616DEAA4523FB5B00F5" box="[931,1152,620,645]" pageId="38" pageNumber="139">Baoshan Prefecture</collectingCounty>
,
<collectingMunicipality id="6BB3AC8DAE7BB616D9834523FA9900F5" box="[1162,1346,620,645]" pageId="38" pageNumber="139">Gaoligong Shan</collectingMunicipality>
, E pass,
<quantity id="4C909B12AE7BB616DD9E45C0FF0400D8" box="[151,223,655,680]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="38" pageNumber="139" unit="km" value="36.0">36 km</quantity>
SE
<location id="8EB7602CAE7BB616DC1945C0FE5400D8" LSID="urn:lsid:plazi:treatment:03C187E1AE78B616DD9E4603FED90164:8EB7602CAE7BB616DC1945C0FE5400D8" box="[272,399,655,680]" country="China" county="Baoshan Prefecture" latitude="24.825556" longLatPrecision="21" longitude="98.76833" municipality="Gaoligong Shan" name="Tengchong" pageId="38" pageNumber="139" stateProvince="Yunnan">Tengchong</location>
,
<geoCoordinate id="EE5C5030AE7BB616DC9545DFFDF900D8" box="[412,546,656,680]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="15" value="24.825556">24°49'32&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DF2745C0FD7800D8" box="[558,675,655,680]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="15" value="98.76833">98°46'6&quot;E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616DFB845DFFCD200D7" box="[689,777,656,680]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="38" pageNumber="139" unit="m" value="2200.0">
<quantity id="4C909B12AE7BB616DFB845DFFCD200D7" box="[689,777,656,680]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="38" pageNumber="139" unit="m" value="2200.0">2200 m</quantity>
</elevation>
, deciduous forest, on
<collectingDate id="EF92E9DFAE7BB616D90A45DFFBAE00D8" box="[1027,1141,656,680]" pageId="38" pageNumber="139" value="2007-05-31">
<date id="FFD61037AE7BB616D90A45DFFBAE00D8" box="[1027,1141,656,680]" pageId="38" pageNumber="139" value="2007-05-31">31/v/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616D98F45DFFAD000D8" box="[1158,1291,655,680]" pageId="38" pageNumber="139">M. Schülke</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616D82B45DFFAA200D8" box="[1314,1401,656,680]" pageId="38" pageNumber="139">ZMHB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE7BB616D88745C0FF250164" ID-GBIF-Occurrence="1658411540" collectingDate="2007-06-03" collectionCode="ZMHB" collectorName="M. Schulke" country="China" elevation="2280" latitude="24.491945" location="Luxi" longLatPrecision="21" longitude="98.88278" municipality="Dehong Dai Autonomous Prefecture" pageId="38" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B91CEAE5AE7BB616D88745C0FE9900BD" bold="true" pageId="38" pageNumber="139">
<specimenCount id="9D6EFD7EAE7BB616D88745C0FF0A00BD" pageId="38" pageNumber="139" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE7BB616DDDE45FBFE9900BD" box="[215,322,692,717]" pageId="38" pageNumber="139" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingCountry id="F37F7667AE7BB616DC5845FBFE7500BC" box="[337,430,692,716]" name="China" pageId="38" pageNumber="139">CHINA</collectingCountry>
:
<collectingRegion id="49ACF815AE7BB616DCB145FAFDD500BC" box="[440,526,693,716]" country="China" name="Yunnan" pageId="38" pageNumber="139">Yunnan</collectingRegion>
[CH07-19],
<collectingMunicipality id="6BB3AC8DAE7BB616DF9445FBFBE100BD" box="[669,1082,692,717]" pageId="38" pageNumber="139">Dehong Dai Autonomous Prefecture</collectingMunicipality>
, mountain range
<quantity id="4C909B12AE7BB616D9F245FBFA9A00BD" box="[1275,1345,692,717]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="38" pageNumber="139" unit="km" value="31.0">31 km</quantity>
E
<location id="8EB7602CAE7BB616D86945FBFA4300BD" LSID="urn:lsid:plazi:treatment:03C187E1AE78B616DD9E4603FED90164:8EB7602CAE7BB616D86945FBFA4300BD" box="[1376,1432,692,717]" country="China" latitude="24.491945" longLatPrecision="21" longitude="98.88278" municipality="Dehong Dai Autonomous Prefecture" name="Luxi" pageId="38" pageNumber="139" stateProvince="Yunnan">Luxi</location>
,
<geoCoordinate id="EE5C5030AE7BB616DD9E4597FEC60080" box="[151,285,728,752]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="15" value="24.491945">24°29'31&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DC244598FE6A0080" box="[301,433,727,752]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="15" value="98.88278">98°52'58&quot;E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616DCC84597FDC6009F" box="[449,541,728,752]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="38" pageNumber="139" unit="m" value="2280.0">
<quantity id="4C909B12AE7BB616DCC84597FDC6009F" box="[449,541,728,752]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="38" pageNumber="139" unit="m" value="2280.0">2280 m</quantity>
</elevation>
, secondary pine forest with old deciduous trees, on
<collectingDate id="EF92E9DFAE7BB616D9854598FB230080" box="[1164,1272,727,752]" pageId="38" pageNumber="139" value="2007-06-03">
<date id="FFD61037AE7BB616D9854598FB230080" box="[1164,1272,727,752]" pageId="38" pageNumber="139" value="2007-06-03">3/vi/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616D8054597FA4E0080" box="[1292,1429,727,752]" pageId="38" pageNumber="139">M. Schülke</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616DD9645B3FF2D0164" box="[159,246,764,788]" pageId="38" pageNumber="139">ZMHB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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