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<document id="002F9BBDFA18F92DE7157F23810755F4" ID-DOI="10.11646/zootaxa.3652.2.2" ID-GBIF-Dataset="0d490ce9-0872-4a4a-8721-ca1a10de1879" ID-ISSN="1175-5326" ID-Zenodo-Dep="219927" ID-ZooBank="B721A1E5-707A-476F-A3E7-E1B0D9559706" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460609319723" checkinUser="plazi" docAuthor="Chipman, Ariel D., Dor, Neta &amp; Bonato, Lucio" docDate="2013" docId="03B987F9143CFFE0CDB7D40BF757B7BE" docLanguage="en" docName="zt03652p248.pdf" docOrigin="Zootaxa 3652 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clinopodes escherichii Verhoeff 1896" docType="treatment" docVersion="9" lastPageNumber="236" masterDocId="FF80FF811438FFE4CD20D717F30BB12D" masterDocTitle="Diversity and biogeography of Israeli geophilomorph centipedes (Chilopoda: Geophilomorpha)" masterLastPageNumber="248" masterPageNumber="232" pageNumber="236" updateTime="1698328166891" updateUser="plazi">
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<mods:title id="A482EB65665600390A811ED8434F4217">Diversity and biogeography of Israeli geophilomorph centipedes (Chilopoda: Geophilomorpha)</mods:title>
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<mods:namePart id="63237FC7E31D09E57DE3F6DBE0F29FE3">Chipman, Ariel D.</mods:namePart>
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<mods:namePart id="A7477A24B557C7C75F9CBFF4DF82B4F7">Dor, Neta</mods:namePart>
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<mods:namePart id="8B176BBAC8028C7AFFD03B7A7FD726CD">Bonato, Lucio</mods:namePart>
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<mods:date id="9906B887A8285A7E8724AB4FE7B8A2A7">2013</mods:date>
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<treatment id="03B987F9143CFFE0CDB7D40BF757B7BE" ID-DOI="http://doi.org/10.5281/zenodo.5678033" ID-GBIF-Taxon="119568158" ID-Zenodo-Dep="5678033" LSID="urn:lsid:plazi:treatment:03B987F9143CFFE0CDB7D40BF757B7BE" httpUri="http://treatment.plazi.org/id/03B987F9143CFFE0CDB7D40BF757B7BE" lastPageNumber="236" pageId="4" pageNumber="236">
<subSubSection id="C30A6564143CFFE0CDB7D40BF757B7BE" pageId="4" pageNumber="236" type="nomenclature">
<paragraph id="8BAF36EF143CFFE0CDB7D40BF172B218" blockId="4.[151,633,795,822]" box="[151,633,795,822]" pageId="4" pageNumber="236">
<heading id="D0E78183143CFFE0CDB7D40BF172B218" bold="true" box="[151,633,795,822]" fontSize="11" level="1" pageId="4" pageNumber="236" reason="1">
<taxonomicName id="4C104D6C143CFFE0CDB7D40BF172B218" authority="Verhoeff, 1896" authorityName="Verhoeff" authorityYear="1896" box="[151,633,795,822]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0CDB7D40BF172B218" bold="true" box="[151,633,795,822]" pageId="4" pageNumber="236">
<emphasis id="B964EAFD143CFFE0CDB7D40BF2A2B21B" bold="true" box="[151,425,796,822]" italics="true" pageId="4" pageNumber="236">Clinopodes escherichii</emphasis>
(Verhoeff, 1896)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF36EF143CFFE0CDB7D475F787B2B3" blockId="4.[151,1436,866,1683]" pageId="4" pageNumber="236">
Published records: “Chuldah bei Jaffa” [Hulda, near Jaffa] (Verhoeff 1925, sub
<taxonomicName id="4C104D6C143CFFE0C91CD474F680B256" box="[1084,1419,867,891]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="flavidus" subSpecies="noduliger">
<emphasis id="B964EAFD143CFFE0C91CD474F680B256" box="[1084,1419,867,891]" italics="true" pageId="4" pageNumber="236">Geophilus flavidus noduliger</emphasis>
</taxonomicName>
); “Ein Hemed (Aqua Bella)”, “Ein Yael”, ”Nes Harim” (Zapparoli 1995, sub
<taxonomicName id="4C104D6C143CFFE0C923D490F777B2B3" box="[1027,1148,902,926]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="flavidus">
<emphasis id="B964EAFD143CFFE0C923D490F777B2B3" box="[1027,1148,902,926]" italics="true" pageId="4" pageNumber="236">C. flavidus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BAF36EF143CFFE0CDE9D4BCF79AB2CB" blockId="4.[151,1436,866,1683]" pageId="4" pageNumber="236">
New records: Hermon, Emek Bolan (
<specimenCount id="9D16FD66143CFFE0CF55D4BDF00AB2EF" box="[629,769,938,962]" pageId="4" pageNumber="236" type="generic">6 specimens</specimenCount>
2012); Hermon, Emek HaMan (
<specimenCount id="9D16FD66143CFFE0C951D4BDF7E4B2EF" box="[1137,1263,938,962]" pageId="4" pageNumber="236" type="generic">1 specimen</specimenCount>
2012); Khirbet Saida (
<specimenCount id="9D16FD66143CFFE0CDC7D4D9F26FB2CB" box="[231,356,974,998]" pageId="4" pageNumber="236" type="generic">1 specimen</specimenCount>
1964); Mt. Qadarim (
<specimenCount id="9D16FD66143CFFE0CF7ED4D9F1D0B2CB" box="[606,731,974,998]" pageId="4" pageNumber="236" type="generic">1 specimen</specimenCount>
2011); Nahal Oren (
<specimenCount id="9D16FD66143CFFE0CEE5D4D9F749B2CB" box="[965,1090,974,998]" pageId="4" pageNumber="236" type="generic">1 specimen</specimenCount>
2011).
</paragraph>
<paragraph id="8BAF36EF143CFFE0CDE9D4E5F3C2B55B" blockId="4.[151,1436,866,1683]" pageId="4" pageNumber="236">
Distribution in
<collectingCountry id="F307767F143CFFE0CC5ED4E5F2CBB527" box="[382,448,1010,1034]" name="Israel" pageId="4" pageNumber="236">Israel</collectingCountry>
: most specimens are from cool high altitude Mediterranean sites such as the upper Galilee, Hermon and Jerusalem mountains, with average temperatures in the range 1622˚C and annual precipitation between
<quantity id="4CE89B0A143CFFE0CCAED32DF13FB57F" box="[398,564,1082,1106]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="12.5" metricValueMin="4.5" pageId="4" pageNumber="236" unit="mm" value="850.0" valueMax="1250.0" valueMin="450.0">4501250 mm</quantity>
. The only exception is a published record from Hulda (Verhoeff 1925) (
<figureCitation id="132B2A6A143CFFE0C871D32DF3AEB55B" captionStart="FIGURE 3" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[380,1139,222,1903]" captionTargetId="figure@9.[363,1178,193,1938]" captionTargetPageId="9" captionText="FIGURE 3. Sampling locations and presumptive distribution sites for members of the Geophilidae and Mecistocephalidae. A) Clinopodes escherichii. B) Geophilus cf. carpophagus. C) Pachymerium ferrugineum. D) Stenotaenia naxia. E) Stenotaenia palaestina. F) Mecistocephalus evansi." httpUri="https://zenodo.org/record/219930/files/figure.png" pageId="4" pageNumber="236">Figure 3</figureCitation>
A).
</paragraph>
<paragraph id="8BAF36EF143CFFE0CDE9D395F186B593" blockId="4.[151,1436,866,1683]" pageId="4" pageNumber="236">
Global distribution: the species is widespread from the Carpathians and the Balkan Peninsula, through regions around the Black Sea, to
<collectingCountry id="F307767F143CFFE0CC90D3B1F11BB593" box="[432,528,1190,1214]" name="Turkey" pageId="4" pageNumber="236">Anatolia</collectingCountry>
and
<collectingCountry id="F307767F143CFFE0CF67D3B1F182B593" box="[583,649,1190,1214]" name="Israel" pageId="4" pageNumber="236">Israel</collectingCountry>
.
</paragraph>
<paragraph id="8BAF36EF143CFFE0CDE9D3DDF757B7BE" blockId="4.[151,1436,866,1683]" pageId="4" pageNumber="236">
Taxonomic and nomenclatural notes. According to the preliminary reassessment of the entire genus
<taxonomicName id="4C104D6C143CFFE0CDB7D3F9F2A3B42B" authority="Koch, 1847" authorityName="Koch" authorityYear="1847" box="[151,424,1262,1286]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAFD143CFFE0CDB7D3F9F21CB42B" box="[151,279,1262,1286]" italics="true" pageId="4" pageNumber="236">Clinopodes</emphasis>
Koch, 1847
</taxonomicName>
by Bonato
<emphasis id="B964EAFD143CFFE0CF16D3E7F178B42B" box="[566,627,1262,1287]" italics="true" pageId="4" pageNumber="236">et al.</emphasis>
(2011), the specimens examined by us agree in morphology with
<taxonomicName id="4C104D6C143CFFE0C8A2D3F8F219B406" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0C8A2D3F8F219B406" italics="true" pageId="4" pageNumber="236">C. escherichii</emphasis>
</taxonomicName>
and not with
<taxonomicName id="4C104D6C143CFFE0CC8ED204F09AB407" authority="Bonato, Iorio &amp; Minelli, 2011" authorityName="Bonato, Iorio &amp; Minelli" authorityYear="2011" box="[430,913,1298,1323]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="verhoeffi">
<emphasis id="B964EAFD143CFFE0CC8ED204F138B406" box="[430,563,1299,1323]" italics="true" pageId="4" pageNumber="236">C. verhoeffi</emphasis>
Bonato, Iorio &amp; Minelli, 2011
</taxonomicName>
, which was described from
<collectingCountry id="F307767F143CFFE0C9ECD205F601B407" box="[1228,1290,1298,1322]" name="Israel" pageId="4" pageNumber="236">Israel</collectingCountry>
(see below). The two species differ mainly in the extent of the sternal pore-fields on the posterior leg-bearing segments and the pattern of the coxal pores.
<taxonomicName id="4C104D6C143CFFE0CCE4D24CF0B6B45F" authority="Verhoeff, 1925" authorityName="Verhoeff" authorityYear="1925" box="[452,957,1370,1395]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="flavidus" subSpecies="noduliger">
<emphasis id="B964EAFD143CFFE0CCE4D24CF006B45E" box="[452,781,1371,1395]" italics="true" pageId="4" pageNumber="236">Geophilus flavidus noduliger</emphasis>
Verhoeff, 1925
</taxonomicName>
is recognized here as a junior synonym of
<taxonomicName id="4C104D6C143CFFE0CDB7D269F140B4BB" authority="Verhoeff 1896" authorityName="Verhoeff" authorityYear="1896" box="[151,587,1406,1430]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0CDB7D269F297B4BB" box="[151,412,1406,1430]" italics="true" pageId="4" pageNumber="236">Clinopodes escherichii</emphasis>
Verhoeff 1896
</taxonomicName>
(
<emphasis id="B964EAFD143CFFE0CF7FD268F1C9B4BA" bold="true" box="[607,706,1407,1431]" pageId="4" pageNumber="236">new syn</emphasis>
.). After examining the
<typeStatus id="54AB884D143CFFE0CEF8D269F731B4BB" box="[984,1082,1406,1430]" pageId="4" pageNumber="236" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C104D6C143CFFE0C948D268F650B4BB" box="[1128,1371,1406,1430]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="flavidus" subSpecies="noduliger">
<emphasis id="B964EAFD143CFFE0C948D268F650B4BB" box="[1128,1371,1406,1430]" italics="true" pageId="4" pageNumber="236">G. flavidus noduliger</emphasis>
</taxonomicName>
from Hulda (preserved in the Zoologische Staatsammlung München) and comparing it with specimens of
<taxonomicName id="4C104D6C143CFFE0C820D2B4F697B496" box="[1280,1436,1443,1467]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0C820D2B4F697B496" box="[1280,1436,1443,1467]" italics="true" pageId="4" pageNumber="236">C. escherichii</emphasis>
</taxonomicName>
from
<collectingCountry id="F307767F143CFFE0CDF4D2D1F219B4F3" box="[212,274,1478,1502]" name="Israel" pageId="4" pageNumber="236">Israel</collectingCountry>
and
<collectingCountry id="F307767F143CFFE0CC6AD2D1F291B4F3" box="[330,410,1478,1502]" name="Turkey" pageId="4" pageNumber="236">Turkey</collectingCountry>
(Bonato
<emphasis id="B964EAFD143CFFE0CF23D2DFF137B4F3" box="[515,572,1478,1503]" italics="true" pageId="4" pageNumber="236">et al.</emphasis>
2011), we found that
<taxonomicName id="4C104D6C143CFFE0CE14D2D0F72BB4F3" box="[820,1056,1478,1502]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="flavidus" subSpecies="noduliger">
<emphasis id="B964EAFD143CFFE0CE14D2D0F72BB4F3" box="[820,1056,1478,1502]" italics="true" pageId="4" pageNumber="236">G. flavidus noduliger</emphasis>
</taxonomicName>
matches fully with
<taxonomicName id="4C104D6C143CFFE0C820D2D0F697B4F3" box="[1280,1436,1478,1502]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0C820D2D0F697B4F3" box="[1280,1436,1478,1502]" italics="true" pageId="4" pageNumber="236">C. escherichii</emphasis>
</taxonomicName>
even in the characters that are considered of diagnostic value between species in the genus
<taxonomicName id="4C104D6C143CFFE0C981D2FCF62BB72E" box="[1185,1312,1515,1539]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAFD143CFFE0C981D2FCF62BB72E" box="[1185,1312,1515,1539]" italics="true" pageId="4" pageNumber="236">Clinopodes</emphasis>
</taxonomicName>
(including the relative size of the marginal denticles on the forcipular coxosternite, the extent of chitin-lines, the number of legs, the relative extent of the pore-fields on the most posterior trunk segments, the arrangement of the coxal organs). Published records of
<taxonomicName id="4C104D6C143CFFE0CCC7D140F16BB743" box="[487,608,1622,1646]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="flavidus">
<emphasis id="B964EAFD143CFFE0CCC7D140F16BB743" box="[487,608,1622,1646]" italics="true" pageId="4" pageNumber="236">C. flavidus</emphasis>
</taxonomicName>
from
<collectingCountry id="F307767F143CFFE0CF86D141F1EFB743" box="[678,740,1622,1646]" name="Israel" pageId="4" pageNumber="236">Israel</collectingCountry>
(Zapparoli 1995) are reinterpreted as most probably referring to
<taxonomicName id="4C104D6C143CFFE0CD94D16CF244B7BE" box="[180,335,1659,1683]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="escherichii">
<emphasis id="B964EAFD143CFFE0CD94D16CF244B7BE" box="[180,335,1659,1683]" italics="true" pageId="4" pageNumber="236">C. escherichii</emphasis>
</taxonomicName>
instead of
<taxonomicName id="4C104D6C143CFFE0CCEBD16CF144B7BE" box="[459,591,1659,1683]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="236" phylum="Arthropoda" rank="species" species="verhoeffi">
<emphasis id="B964EAFD143CFFE0CCEBD16CF144B7BE" box="[459,591,1659,1683]" italics="true" pageId="4" pageNumber="236">C. verhoeffi</emphasis>
</taxonomicName>
, on the basis of the number of trunk segments.
</paragraph>
</subSubSection>
</treatment>
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