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<document id="CA981FCCB926C075959A2AD10644885C" ID-DOI="10.11646/zootaxa.3946.4.3" ID-GBIF-Dataset="46db289d-0964-4a71-8b5c-ab81e23e961f" ID-ISSN="1175-5326" ID-Zenodo-Dep="238608" ID-ZooBank="7A5F142A-EEDE-453A-9CB5-241917A83921" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038442617" checkinUser="plazi" docAuthor="Rifkind, Jacques" docDate="2015" docId="0151C53EFFB4FFBE9DDCF899477789F2" docLanguage="en" docName="zt03946p552.pdf" docOrigin="Zootaxa 3946 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymatodera doda Rifkind, n. sp." docType="treatment" docVersion="8" lastPageNumber="527" masterDocId="FD68BD46FFB3FFB69D4BFF9B462E8F6A" masterDocTitle="New species of Cymatodera Gray (Coleoptera: Cleridae: Tillinae) from México and Central America, with notes on others" masterLastPageNumber="552" masterPageNumber="519" pageNumber="526" updateTime="1698574954642" updateUser="plazi">
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<mods:title id="C00D7E098CB33F31FB24B8D08421AD7C">New species of Cymatodera Gray (Coleoptera: Cleridae: Tillinae) from México and Central America, with notes on others</mods:title>
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<mods:namePart id="1A3196CA5BB69C4F9ADB65F959C56A9B">Rifkind, Jacques</mods:namePart>
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<mods:date id="735E9F9BE668C56890BECB94C679A360">2015</mods:date>
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<treatment id="0151C53EFFB4FFBE9DDCF899477789F2" ID-DOI="http://doi.org/10.5281/zenodo.6110737" ID-GBIF-Taxon="119609532" ID-Zenodo-Dep="6110737" LSID="urn:lsid:plazi:treatment:0151C53EFFB4FFBE9DDCF899477789F2" httpUri="http://treatment.plazi.org/id/0151C53EFFB4FFBE9DDCF899477789F2" lastPageId="8" lastPageNumber="527" pageId="7" pageNumber="526">
<subSubSection id="C1E227A3FFB4FFB19DDCF89947038854" pageId="7" pageNumber="526" type="nomenclature">
<paragraph id="89477428FFB4FFB19DDCF899440A8877" blockId="7.[151,548,1794,1854]" box="[151,548,1794,1821]" pageId="7" pageNumber="526">
<heading id="D20FC344FFB4FFB19DDCF899440A8877" bold="true" box="[151,548,1794,1821]" fontSize="11" level="1" pageId="7" pageNumber="526" reason="1">
<emphasis id="BB8CA83AFFB4FFB19DDCF899440A8877" bold="true" box="[151,548,1794,1821]" pageId="7" pageNumber="526">
<taxonomicName id="4EF80FABFFB4FFB19DDCF89947FB8877" authority="Rifkind" authorityName="Rifkind" box="[151,469,1794,1821]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="526" phylum="Arthropoda" rank="species" species="doda" status="sp. nov.">
<emphasis id="BB8CA83AFFB4FFB19DDCF89947428876" bold="true" box="[151,364,1794,1820]" italics="true" pageId="7" pageNumber="526">Cymatodera doda</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="A0BF1541FFB4FFB19CAAF898440A8877" box="[481,548,1795,1821]" pageId="7" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="89477428FFB4FFB19DDCF8BE47038854" blockId="7.[151,548,1794,1854]" box="[151,301,1829,1854]" pageId="7" pageNumber="526">
(
<figureCitation id="11C368ADFFB4FFB19DD4F8BE470B8854" box="[159,293,1829,1854]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="7" pageNumber="526">Figs. 1518</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFB4FFBE9DDCF8F647378EC6" lastPageId="8" lastPageNumber="527" pageId="7" pageNumber="526" type="materials_examined">
<paragraph id="89477428FFB4FFBE9DDCF8F647378EC6" blockId="7.[151,1436,1901,2034]" lastBlockId="8.[151,1437,151,1688]" lastPageId="8" lastPageNumber="527" pageId="7" pageNumber="526">
<emphasis id="BB8CA83AFFB4FFB19DDCF8F6477588EC" bold="true" box="[151,347,1901,1926]" pageId="7" pageNumber="526">
<typeStatus id="5643CA8AFFB4FFB19DDCF8F646FC88EC" box="[151,210,1901,1926]" pageId="7" pageNumber="526">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="5643CA8AFFB4FFB19C2FF8F647E288EC" box="[356,460,1901,1926]" pageId="7" pageNumber="526" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F1EF34B8FFB4FFB19F56F8F6445B88EC" box="[541,629,1901,1926]" name="Mexico" pageId="7" pageNumber="526">México</collectingCountry>
, Jalisco, Est. Biol. Chamela,
<date id="FD4652E8FFB4FFB19E8CF8F6420188EC" box="[967,1071,1901,1926]" pageId="7" pageNumber="526" value="1916-07-08">VII-8/16</date>
-[19]85, at light, J. Chemsak, H. Katsura, A. &amp; E. Michelbacher, collectors.
<typeStatus id="5643CA8AFFB4FFB19FDDF80A44D088C0" box="[662,766,1937,1962]" pageId="7" pageNumber="526" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="EFE9ECEDFFB4FFB19EE9F80945DD88C0" LSID="urn:lsid:biocol.org:col:1013" box="[930,1011,1938,1962]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="7" pageNumber="526">EMEC</collectionCode>
.
<typeStatus id="5643CA8AFFB4FFB19948F809425B88C0" box="[1027,1141,1938,1962]" pageId="7" pageNumber="526" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F1EF34B8FFB4FFB199C8F80942DA88C0" box="[1155,1268,1938,1962]" name="Mexico" pageId="7" pageNumber="526">MEXICO</collectingCountry>
, JALISCO: 1, Chamela, Vic. Est.
<collectionCode id="EFE9ECEDFFB4FFB19C25F82D47E888A4" box="[366,454,1974,1998]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="526">UNAM</collectionCode>
,
<date id="FD4652E8FFB4FFB19C9AF82D445688A4" box="[465,632,1974,1998]" pageId="7" pageNumber="526" value="1993-07-09" valueMax="1993-07-19" valueMin="1993-07-09">9-19-VII-1993</date>
, J. Huether; 1, 8.6 km. N Chamela, mv + bl,
<date id="FD4652E8FFB4FFB19927F82D429088A4" box="[1132,1214,1973,1998]" pageId="7" pageNumber="526" value="1918-07">18 July</date>
, 1987, R. Turnbow; OAXACA: 1, 2.1 mi. NW Totolapan,
<date id="FD4652E8FFB4FFB19F0DF84144D88898" box="[582,758,2010,2034]" pageId="7" pageNumber="526" value="1980-08-07">August 7, 1980</date>
, Schaffner and Friedlander;
<quantity id="4E00D9CDFFB4FFB1997DF84142B68898" box="[1078,1176,2009,2034]" metricMagnitude="3" metricUnit="m" metricValue="1.97949312" pageId="7" pageNumber="526" unit="mi" value="1.23">1, 23 mi</quantity>
. south Matias Romero,
<date id="FD4652E8FFBBFFBE9DDCFF0C47238FDA" box="[151,269,151,176]" pageId="8" pageNumber="527" value="1962-04-06">IV-6-1962</date>
, F. D. Parker, L. A. Stange, collectors;
<collectingCountry id="F1EF34B8FFBBFFBE9F8AFF0345408FDA" box="[705,878,152,176]" name="Nicaragua" pageId="8" pageNumber="527">NICARAGUA</collectingCountry>
: 1,
<collectingCountry id="F1EF34B8FFBBFFBE9ED8FF0C45DB8FDA" box="[915,1013,151,176]" name="Grenada" pageId="8" pageNumber="527">Granada</collectingCountry>
, Res. Silv. Domitila,
<date id="FD4652E8FFBBFFBE99A9FF0C433E8FDA" box="[1250,1296,151,176]" pageId="8" pageNumber="527">vi-2</date>
/5-2013, day collecting,
<geoCoordinate id="ECCC12EFFFBBFFBE9C5DFF2747AE8FBE" box="[278,384,188,212]" direction="north" orientation="latitude" pageId="8" pageNumber="527" precision="555" value="11.71">11.71° N</geoCoordinate>
,
<geoCoordinate id="ECCC12EFFFBBFFBE9CC5FF2747D38FBE" box="[398,509,188,213]" direction="west" orientation="longitude" pageId="8" pageNumber="527" precision="555" value="-85.95">85.95° W</geoCoordinate>
, el. abt.
<quantity id="4E00D9CDFFBBFFBE9F28FF2744F78FBE" box="[611,729,188,213]" metricMagnitude="1" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="8" pageNumber="527" unit="m" value="80.0" valueMax="100.0" valueMin="60.0">60100 m</quantity>
, B. Raber, D. Heffern &amp; E. van den Berghe; 1, same data as previous except collected at “merc. vapor lite”;
<collectingCountry id="F1EF34B8FFBBFFBE9FF7FF7B454B8F92" box="[700,869,224,248]" name="Costa Rica" pageId="8" pageNumber="527">COSTA RICA</collectingCountry>
, GUANACASTE PROVINCE: 2, La Pacifica, nr Cañas,
<date id="FD4652E8FFBBFFBE9DA2FE9F474C8E76" box="[233,354,260,284]" pageId="8" pageNumber="527" value="1989-06-04">VI-4-1989</date>
, F. Hovore, coll.; 1, Hacienda La Pacifica,
<date id="FD4652E8FFBBFFBE9E74FE9F45A48E76" box="[831,906,260,284]" pageId="8" pageNumber="527">9 June</date>
, 1973, 75 meters, Ginter Ekis; 1, “La Pacifica,” nr Cañas,
<date id="FD4652E8FFBBFFBE9C43FEB347AB8E2A" box="[264,389,296,320]" pageId="8" pageNumber="527" value="1920-05-21">20/21 May</date>
, 1985, F. T. Hovore, coll.; 1, La Pacifica, nr. Cañas,
<date id="FD4652E8FFBBFFBE9E9FFEBC42168E2A" box="[980,1080,295,320]" pageId="8" pageNumber="527" value="1922-05-26">V-22-26</date>
-[19]84, E. Riley, D. Rider &amp; D. LeDoux; 1, Est. Murciélago,
<quantity id="4E00D9CDFFBBFFBE9C97FED7443B8E0F" box="[476,533,332,357]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="527" unit="km" value="8.0">8 km</quantity>
Suroeste de Cuajiniquil,
<quantity id="4E00D9CDFFBBFFBE9E64FED745598E0E" box="[815,887,332,356]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="527" unit="m" value="100.0">100 m</quantity>
,
<date id="FD4652E8FFBBFFBE9EC9FED7421D8E0E" box="[898,1075,332,357]" pageId="8" pageNumber="527" value="1993-05-03" valueMax="1993-05-08" valueMin="1993-05-03">
3 a 8
<bibRefCitation id="ED6909D9FFBBFFBE9EF5FED6421D8E0E" author="May" box="[958,1075,332,357]" pageId="8" pageNumber="527" refString="Ramamoorthy, T. P., Bye, R., Lot, A. &amp; Fa, J. E. (Eds.) (1993) Biological diversity of Mexico: origins and distribution. Oxford University Press, New York, pp. 1 - 812." type="book chapter" year="1993">May 1993</bibRefCitation>
</date>
, F. A. Quesada [
<collectionCode id="EFE9ECEDFFBBFFBE99A3FED743BC8E0E" box="[1256,1426,332,356]" pageId="8" pageNumber="527">INBIO CODE</collectionCode>
:] LN320300, 347200,
<collectionCode id="EFE9ECEDFFBBFFBE9CD9FEEB47CE8EE2" box="[402,480,368,392]" pageId="8" pageNumber="527">INBIO</collectionCode>
CR1001, 194408.
<typeStatus id="5643CA8AFFBBFFBE9FF1FEEB45098EE2" box="[698,807,368,392]" pageId="8" pageNumber="527" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="EFE9ECEDFFBBFFBE9EA6FEEB421A8EE2" box="[1005,1076,368,392]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="527">CNIN</collectionCode>
,
<collectionCode id="EFE9ECEDFFBBFFBE9909FEEB42B48EED" box="[1090,1178,368,391]" httpUri="http://grbio.org/cool/brtj-zfcn" name="W.F. Barr Entomological Collection" pageId="8" pageNumber="527">WFBM</collectionCode>
,
<collectionCode id="EFE9ECEDFFBBFFBE99ECFEEB42DE8EE2" box="[1191,1264,368,392]" pageId="8" pageNumber="527">DJHC</collectionCode>
,
<collectionCode id="EFE9ECEDFFBBFFBE99B6FEEB43648EE2" box="[1277,1354,368,392]" pageId="8" pageNumber="527">EGRC</collectionCode>
,
<collectionCode id="EFE9ECEDFFBBFFBE9813FEEB43B28EE2" box="[1368,1436,368,392]" pageId="8" pageNumber="527">JNRC</collectionCode>
and
<collectionCode id="EFE9ECEDFFBBFFBE9D8DFE0F473B8EC6" box="[198,277,404,428]" pageId="8" pageNumber="527">RHTC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFBBFFBE9D8CFE2C43298D56" pageId="8" pageNumber="527" type="diagnosis">
<paragraph id="89477428FFBBFFBE9D8CFE2C43298D56" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CFE2C476F8EBA" bold="true" box="[199,321,439,464]" pageId="8" pageNumber="527">Diagnosis.</emphasis>
Males are unique among known congeners by virtue of their nipple-like metathoracic projections and the unusual shape of the sixth abdominal ventrite (
<figureCitation id="11C368ADFFBBFFBE9E58FE4745B18E9F" box="[787,927,476,501]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="8" pageNumber="527">Figs. 1617</figureCitation>
). The nearly cristate condition of the elytral integument between the rows of punctures will serve to distinguish females of
<taxonomicName id="4EF80FABFFBBFFBE9968FD9B42538D72" box="[1059,1149,512,536]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="527" phylum="Arthropoda" rank="species" species="doda">
<emphasis id="BB8CA83AFFBBFFBE9968FD9B42538D72" box="[1059,1149,512,536]" italics="true" pageId="8" pageNumber="527">C. doda</emphasis>
</taxonomicName>
from other small, robust
<taxonomicName id="4EF80FABFFBBFFBE9DDCFDBF470C8D56" box="[151,290,548,572]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="BB8CA83AFFBBFFBE9DDCFDBF470C8D56" box="[151,290,548,572]" italics="true" pageId="8" pageNumber="527">Cymatodera</emphasis>
</taxonomicName>
species possessing similar subserrate antennae (e. g. paler specimens of
<taxonomicName id="4EF80FABFFBBFFBE9904FDBF42D58D56" authority="Barr" authorityName="Barr" box="[1103,1275,548,572]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="527" phylum="Arthropoda" rank="species" species="sobara">
<emphasis id="BB8CA83AFFBBFFBE9904FDBF42908D56" box="[1103,1214,548,572]" italics="true" pageId="8" pageNumber="527">C. sobara</emphasis>
Barr
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFBBFFBE9D8CFDDC44A88AAA" pageId="8" pageNumber="527" type="description">
<paragraph id="89477428FFBBFFBE9D8CFDDC47868A66" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CFDDC477F8D0A" bold="true" box="[199,337,583,608]" pageId="8" pageNumber="527">Description</emphasis>
(
<typeStatus id="5643CA8AFFBBFFBE9C2AFDDC47FE8D0A" box="[353,464,583,608]" pageId="8" pageNumber="527" type="holotype">Holotype</typeStatus>
). Length: 5.20 mm. Form: small, stout, body rather deep in dorsoventral cross section (
<figureCitation id="11C368ADFFBBFFBE9DD5FDF746C58DEF" box="[158,235,620,645]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="8" pageNumber="527">Fig 15</figureCitation>
). Color: reddish testaceous; head and pronotum reddish brown, sterna a little darker. Head: surface closely, rather shallowly punctate, moderately set with short, fine, mostly reclinate, pale setae; eyes large, coarsely faceted; antennae elongate (reaching beyond elytral humeri when laid alongside), antennomeres rather robust: antennomeres 210 subserrate; 2 much shorter than 3; 3 slightly shorter than 4; 11 longer than 10, ovate-elongate and gradually tapered apically. Pronotum: constricted before middle and more acutely inflected posterior to median lobe; surface rather rough, closely punctate and arenose; vestiture as on head but with an intermixture of a few longer, erect, more robust, brownish setae. Scutellum: densely setose. Elytra: subrectangulate, ratio of length to maximum width 39:20, rather broadly flattened above, slightly depressed at middle of disk, deep in cross section with a precipitous apical slope; surface cribratepunctate; punctures large, closely arranged in longitudinal series (punctures somewhat smaller posteriorly, but attaining apices); area between punctures narrow, subcristate; vestiture sparse, composed of erect and suberect, rather stout, testaceous setae of moderate length. Metaventrite: uniquely shaped; surface shining, sparsely setose, set with rather large punctures laterally; posterior deeply sulcate at middle, each side produced posteriorly into an elongate, slightly divergent and slightly downturned nipplelike tubercle (
<figureCitation id="11C368ADFFBBFFBE9C49FBA447798B32" box="[258,343,1087,1112]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="8" pageNumber="527">Fig. 16</figureCitation>
). Abdomen: ventrite 5 deeply, subtriangularly emarginate posteriorly; ventrite 6 (
<figureCitation id="11C368ADFFBBFFBE99BEFBA443668B32" box="[1269,1352,1087,1112]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="8" pageNumber="527">Fig. 17</figureCitation>
) raised and flattened ventrally, campanulate (narrower anteriorly), with a darkly sclerotized “false” posterior margin deeply Ushaped, and paler true margin slightly, arcuately emarginate posteriorly; tergite 6 acutely deflected ventrally, oblique laterally, broadly arcuately emarginate posteriorly. Aedeagus with lateral lobes rather robust, subacute posteriorly; phallus lanceolate apically. Legs: each metatibia with a distinct sinuate carina internally at proximate end (
<figureCitation id="11C368ADFFBBFFBE9C0CFB6F47B58A66" box="[327,411,1268,1293]" captionStart="FIGURES 15 18. 15" captionStartId="9.[151,264,1562,1585]" captionTargetBox="[169,1415,189,1529]" captionTargetId="figure@9.[152,1435,175,1541]" captionTargetPageId="9" captionText="FIGURES 15 18. 15. Habitus of Cymatodera doda Rifkind, n. sp. (holotype male). 16. Cymatodera doda (holotype male): detail of metaventrite, showing nipplelike projections. 17. Pygidium (ventral view) of male Cymatodera doda. 18. Cymatodera doda: detail of male metatibial carina." httpUri="https://zenodo.org/record/238612/files/figure.png" pageId="8" pageNumber="527">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="89477428FFBBFFBE9D8CFA8C44A88AAA" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CFA8C47158A5A" bold="true" box="[199,315,1303,1328]" pageId="8" pageNumber="527">Variation.</emphasis>
Some specimens exhibit a slight darkening of the elytral dorsal integument ante- and postmedially; two specimens from Jalisco have the faint indication of a pale median elytra fascia. Females have abdominal ventrite 5 with sides oblique, hind margin rather broadly, shallowly emarginate posteriorly; ventrite 6 small, arcuate posteriorly, more or less conjointly rounded with tergite 6. Females lack tubercles on the metaventrite and the enlarged metatibial carinae of the males.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFBBFFBE9D8CFA57423D8962" pageId="8" pageNumber="527" type="etymology">
<paragraph id="89477428FFBBFFBE9D8CFA57423D8962" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CFA5747608A8F" bold="true" box="[199,334,1484,1509]" pageId="8" pageNumber="527">Etymology.</emphasis>
The new species is named for Carol Ann
<taxonomicName id="4EF80FABFFBBFFBE9E54FA5745708A8F" box="[799,862,1484,1509]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="528" phylum="Arthropoda" rank="species" species="doda">Doda</taxonomicName>
, Americas pioneering ecdysiast and 1960s cultural icon. The patronymic is treated as a noun in apposition for the sake of euphony.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFBBFFBE9D8CF98F4534893A" pageId="8" pageNumber="527" type="distribution">
<paragraph id="89477428FFBBFFBE9D8CF98F4534893A" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CF98F474E8947" bold="true" box="[199,352,1556,1581]" pageId="8" pageNumber="527">Distribution.</emphasis>
Known from the Pacific versant of
<collectingCountry id="F1EF34B8FFBBFFBE9E4CF98F45728947" box="[775,860,1556,1581]" name="Mexico" pageId="8" pageNumber="527">México</collectingCountry>
from Jalisco to the Isthmus of Tehuantepec, then south to western
<collectingCountry id="F1EF34B8FFBBFFBE9C1EF9AC47E6893A" box="[341,456,1591,1616]" name="Nicaragua" pageId="8" pageNumber="527">Nicaragua</collectingCountry>
and northwestern
<collectingCountry id="F1EF34B8FFBBFFBE9FD3F9A34538893A" box="[664,790,1591,1616]" name="Costa Rica" pageId="8" pageNumber="527">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFBBFFBE9D8CF9C7477789F2" pageId="8" pageNumber="527" type="biology_ecology">
<paragraph id="89477428FFBBFFBE9D8CF9C7477789F2" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="527">
<emphasis id="BB8CA83AFFBBFFBE9D8CF9C74709891F" bold="true" box="[199,295,1628,1653]" pageId="8" pageNumber="527">Biology.</emphasis>
This species appears to be associated with tropical deciduous forest throughout its range. It has been collected at light.
</paragraph>
</subSubSection>
</treatment>
</document>