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<document id="14394E42A7AB1CE60E070D256582D540" ID-DOI="10.11646/zootaxa.5496.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13331013" ID-ZooBank="D993FA3C-03B0-4232-B762-0F24CA6EBF08" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723805670933" checkinUser="plazi" docAuthor="Watanabe, Kyohei &amp; Riedel, Matthias" docDate="2024" docId="7723B944E9224A6FFF3CFD77FC69FC9E" docLanguage="en" docName="zootaxa.5496.1.2.pdf" docOrigin="Zootaxa 5496 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syzeuctus yamatonis Watanabe &amp; Riedel, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="70" masterDocId="8B1AC13CE9024A4CFFABFFC4FFFEFF8B" masterDocTitle="Revision of the genus Syzeuctus Förster, 1869 (Hymenoptera, Ichneumonidae, Banchinae) from Japan" masterLastPageNumber="71" masterPageNumber="35" pageNumber="67" updateTime="1723807098958" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="091667243A181B9708E33818222A7ED1">Revision of the genus Syzeuctus Förster, 1869 (Hymenoptera, Ichneumonidae, Banchinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="EBFD6597FF48D8E370F658303AB7C893" type="personal">
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<mods:namePart id="F0B6FBC4A4AA39E6DC8E68BC73C290B5">Watanabe, Kyohei</mods:namePart>
<mods:affiliation id="E5702875DD82014A362FDF9BFE54ACA7">Kanagawa Prefectural Museum of Natural History, Iriuda 499, Odawara, Kanagawa 250 - 0031, Japan.</mods:affiliation>
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<mods:namePart id="20807F44547ADB583FD2B7BBC80FBFB6">Riedel, Matthias</mods:namePart>
<mods:affiliation id="E12D50CBC51584B002A1967A6B4BDA60">Blumenlage 22 C, D- 29683, Bad Fallingbostel, Germany.</mods:affiliation>
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<mods:title id="27ACB04BFF1153C1F695CE2BE7E2C301">Zootaxa</mods:title>
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<mods:date id="B9029321BD3CFA72E9421828F4CD3B55">2024</mods:date>
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<mods:number id="830F7B11834F8FD5FB048F3BDFB1DEB4">2024-08-16</mods:number>
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<mods:number id="398F1AECD85FD39A2924EEEFEBA227C6">5496</mods:number>
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<mods:url id="3C11BD3B77613A9CE1E8EC8A6FD67DD1">http://dx.doi.org/10.11646/zootaxa.5496.1.2</mods:url>
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<treatment id="7723B944E9224A6FFF3CFD77FC69FC9E" ID-DOI="http://doi.org/10.5281/zenodo.13330982" ID-Zenodo-Dep="13330982" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E" httpUri="http://treatment.plazi.org/id/7723B944E9224A6FFF3CFD77FC69FC9E" lastPageId="35" lastPageNumber="70" pageId="32" pageNumber="67">
<subSubSection id="B7905BD9E9224A6CFF3CFD77FE08FD46" box="[151,502,691,717]" pageId="32" pageNumber="67" type="nomenclature">
<paragraph id="FF350852E9224A6CFF3CFD77FE08FD46" blockId="32.[151,578,691,753]" box="[151,502,691,717]" pageId="32" pageNumber="67">
<heading id="A47DBF3EE9224A6CFF3CFD77FE08FD46" bold="true" box="[151,502,691,717]" fontSize="11" level="1" pageId="32" pageNumber="67" reason="1">
<emphasis id="CDFED440E9224A6CFF3CFD77FE08FD46" bold="true" box="[151,502,691,717]" pageId="32" pageNumber="67">
<taxonomicName id="388A73D1E9224A6CFF3CFD77FE6FFD46" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[151,401,691,717]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="67" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9224A6CFF3CFD77FE6FFD46" bold="true" box="[151,401,691,717]" italics="true" pageId="32" pageNumber="67">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6CD693BE9224A6CFE32FD77FE08FD46" box="[409,502,691,717]" pageId="32" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF3CFD13FDBCFD7A" box="[151,578,727,753]" pageId="32" pageNumber="67" type="description">
<paragraph id="FF350852E9224A6CFF3CFD13FDBCFD7A" blockId="32.[151,578,691,753]" box="[151,578,727,753]" pageId="32" pageNumber="67">
(
<figureCitation id="67B114D7E9224A6CFF34FD13FEFEFD7A" box="[159,256,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.13331015" httpUri="https://zenodo.org/record/13331015/files/figure.png" pageId="32" pageNumber="67">Figs 1 L</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFEA7FD13FE8BFD7A" box="[268,373,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="32" pageNumber="67">2 N, R, S</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFE2AFD13FE28FD7A" box="[385,470,727,753]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." figureDoi="http://doi.org/10.5281/zenodo.13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="32" pageNumber="67">22 AF</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFE4BFD13FDC4FD7A" box="[480,570,727,753]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13331071" httpUri="https://zenodo.org/record/13331071/files/figure.png" pageId="32" pageNumber="67">23 AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF3CFCDBFB7AFBF6" pageId="32" pageNumber="67" type="materials_examined">
<paragraph id="FF350852E9224A6CFF3CFCDBFB7AFBF6" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF3CFCDBFEE1FCB3" bold="true" box="[151,287,798,825]" pageId="32" pageNumber="67">Type series</emphasis>
.
<materialsCitation id="4FE2020FE9224A6CFE84FCDBFEBAFCD6" collectingDate="2009-05-27" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" county="Honshu" location="Enkaizan" municipality="Yokohama City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
<emphasis id="CDFED440E9224A6CFE84FCDBFE62FCB2" bold="true" box="[303,412,799,825]" pageId="32" pageNumber="67">
<typeStatus id="2031B6F0E9224A6CFE84FCDBFE62FCB2" box="[303,412,799,825]" pageId="32" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
: F,
<emphasis id="CDFED440E9224A6CFE60FCDAFDDBFCB3" bold="true" box="[459,549,798,824]" pageId="32" pageNumber="67">
<collectingCountry id="879D48C2E9224A6CFE60FCDAFDDBFCB3" box="[459,549,798,824]" name="Japan" pageId="32" pageNumber="67">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E9224A6CFD9DFCDBFD6DFCB2" bold="true" box="[566,659,799,825]" pageId="32" pageNumber="67">
<collectingCounty id="165470DEE9224A6CFD9DFCDBFD6DFCB2" box="[566,659,799,825]" pageId="32" pageNumber="67">Honshu</collectingCounty>
</emphasis>
,
<collectingRegion id="3D4EC6B0E9224A6CFD0FFCDBFCA3FCB2" box="[676,861,799,825]" country="Japan" name="Kanagawa" pageId="32" pageNumber="67">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFCC2FCDBFBE2FCB2" box="[873,1052,799,825]" pageId="32" pageNumber="67">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFB81FCDBFB6BFCB2" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB81FCDBFB6BFCB2" box="[1066,1173,799,825]" country="Japan" county="Honshu" municipality="Yokohama City" name="Enkaizan" pageId="32" pageNumber="67" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E9224A6CFB08FCDBFAD5FCB2" box="[1187,1323,799,825]" pageId="32" pageNumber="67" value="2009-05-27">
<collectingDate id="9B70D77AE9224A6CFB08FCDBFAD5FCB2" box="[1187,1323,799,825]" pageId="32" pageNumber="67" value="2009-05-27">27. V. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFA92FCDBFA62FCB2" box="[1337,1436,799,825]" pageId="32" pageNumber="67">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFF7BFC87FEC2FCD6" box="[208,316,835,861]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE9224A6CFEE4FC87FE70FC0A" collectingDate="2008-08-18" collectingDateMax="2008-08-25" collectingDateMin="2008-08-18" collectionCode="KPMNH" collectorName="K. Konishi" country="Japan" location="Hitsujigaoka" municipality="Sapporo City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<emphasis id="CDFED440E9224A6CFEE4FC87FE39FCD6" bold="true" box="[335,455,835,861]" pageId="32" pageNumber="67">
<typeStatus id="2031B6F0E9224A6CFEE4FC87FE39FCD6" box="[335,455,835,861]" pageId="32" pageNumber="67" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="CDFED440E9224A6CFE7CFC86FDCFFCD7" bold="true" box="[471,561,834,860]" pageId="32" pageNumber="67">
<collectingCountry id="879D48C2E9224A6CFE7CFC86FDCFFCD7" box="[471,561,834,860]" name="Japan" pageId="32" pageNumber="67">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E9224A6CFDE1FC86FD3EFCD7" bold="true" box="[586,704,834,860]" pageId="32" pageNumber="67">
<collectingRegion id="3D4EC6B0E9224A6CFDE1FC86FD3EFCD7" box="[586,704,834,860]" country="Japan" name="Hokkaido" pageId="32" pageNumber="67">Hokkaido</collectingRegion>
</emphasis>
] 1 F,
<collectingMunicipality id="1F519228E9224A6CFCAFFC87FC65FCD6" box="[772,923,835,861]" pageId="32" pageNumber="67">Sapporo City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC0CFC87FBC7FCD6" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC0CFC87FBC7FCD6" box="[935,1081,835,861]" country="Japan" municipality="Sapporo City" name="Hitsujigaoka" pageId="32" pageNumber="67" stateProvince="Hokkaido">Hitsujigaoka</location>
,
<date id="8B342E92E9224A6CFBEEFC86FAEAFCD6" box="[1093,1300,834,861]" pageId="32" pageNumber="67" value="2008-08-18" valueMax="2008-08-25" valueMin="2008-08-18">
<collectingDate id="9B70D77AE9224A6CFBEEFC86FAEAFCD6" box="[1093,1300,834,861]" pageId="32" pageNumber="67" value="2008-08-18" valueMax="2008-08-25" valueMin="2008-08-18">1825. VIII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFA8BFC87FA62FCD6" box="[1312,1436,835,861]" pageId="32" pageNumber="67">K. Konishi</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFEB1FCA3FE78FC0A" box="[282,390,871,897]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFE36FCA3FB95FC0A" box="[413,1131,870,897]" collectingDate="2010-07-23" collectingDateMax="2010-07-25" collectingDateMin="2010-07-23" collectionCode="EUM" collectorName="K. Takashino" country="Japan" location="Hitsujigaoka" municipality="Sapporo City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 F, ditto,
<date id="8B342E92E9224A6CFDB7FCA2FD16FC0A" box="[540,744,870,897]" pageId="32" pageNumber="67" value="2010-07-23" valueMax="2010-07-25" valueMin="2010-07-23">
<collectingDate id="9B70D77AE9224A6CFDB7FCA2FD16FC0A" box="[540,744,870,897]" pageId="32" pageNumber="67" value="2010-07-23" valueMax="2010-07-25" valueMin="2010-07-23">2325. VII. 2010</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFD5CFCA3FC6EFC0A" box="[759,912,871,897]" pageId="32" pageNumber="67">K. Takashino</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFBB5FCA3FB9CFC0A" box="[1054,1122,871,897]" pageId="32" pageNumber="67">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFBD1FCA3FB5EFC2E" collectingDate="2012-08-21" collectionCode="KPMNH" collectorName="Y. Matsubara &amp; K. Fukuda" country="Japan" location="Mamurogawa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Yamagata" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFB2FFCA3FB1FFC0A" bold="true" box="[1156,1249,871,897]" pageId="32" pageNumber="67">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9224A6CFA85FCA3FF2EFC2E" country="Japan" name="Yamagata" pageId="32" pageNumber="67">Yamagata Pref.</collectingRegion>
,
<location id="FA555E89E9224A6CFF70FC4FFE3EFC2E" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFF70FC4FFE3EFC2E" box="[219,448,907,933]" country="Japan" name="Mamurogawa Town" pageId="32" pageNumber="67" stateProvince="Yamagata">Mamurogawa Town</location>
,
<date id="8B342E92E9224A6CFE60FC4FFD91FC2E" box="[459,623,907,933]" pageId="32" pageNumber="67" value="2012-08-21">
<collectingDate id="9B70D77AE9224A6CFE60FC4FFD91FC2E" box="[459,623,907,933]" pageId="32" pageNumber="67" value="2012-08-21">21. VIII. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFDD2FC4FFCF0FC2E" box="[633,782,907,933]" pageId="32" pageNumber="67">Y. Matsubara</collectorName>
&amp;
<collectorName id="527F6D84E9224A6CFC98FC4FFC54FC2E" box="[819,938,907,933]" pageId="32" pageNumber="67">K. Fukuda</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFB87FC4FFB66FC2E" box="[1068,1176,907,933]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFB07FC4FFD40FC42" collectingDate="2009-06-19" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Mamurogawa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 M, same locality of
<typeStatus id="2031B6F0E9224A6CFF3CFC6BFF02FC42" box="[151,252,943,969]" pageId="32" pageNumber="67" type="holotype">holotype</typeStatus>
,
<date id="8B342E92E9224A6CFEA3FC6BFE65FC42" box="[264,411,943,969]" pageId="32" pageNumber="67" value="2009-06-19">
<collectingDate id="9B70D77AE9224A6CFEA3FC6BFE65FC42" box="[264,411,943,969]" pageId="32" pageNumber="67" value="2009-06-19">19. VI. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFE0CFC6BFDF7FC42" box="[423,521,943,969]" pageId="32" pageNumber="67">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFDE1FC6BFD48FC42" box="[586,694,943,969]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFD60FC6BFD8CFC66" collectingDate="2013-07-16" collectionCode="KPMNH" collectorName="S. Shimizu" country="Japan" county="Myokou" location="Suginosawa" municipality="City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Niigata" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFD56FC6BFC6FFC42" box="[765,913,943,969]" country="Japan" name="Niigata" pageId="32" pageNumber="67">Niigata Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9224A6CFC36FC6BFC01FC42" box="[925,1023,943,969]" pageId="32" pageNumber="67">Myokou</collectingCounty>
,
<collectingMunicipality id="1F519228E9224A6CFBA0FC6BFBC2FC42" box="[1035,1084,943,969]" pageId="32" pageNumber="67">City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFBE2FC6BFB2DFC42" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFBE2FC6BFB2DFC42" box="[1097,1235,943,969]" country="Japan" county="Myokou" municipality="City" name="Suginosawa" pageId="32" pageNumber="67" stateProvince="Niigata">Suginosawa</location>
,
<location id="FA555E89E9224A6CFB74FC6BFA66FC42" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB74FC6BFA66FC42" box="[1247,1432,943,969]" country="Japan" county="Myokou" municipality="City" name="Mt. Sasagamine" pageId="32" pageNumber="67" stateProvince="Niigata">Mt. Sasagamine</location>
,
<date id="8B342E92E9224A6CFF3CFC17FECDFC66" box="[151,307,979,1005]" pageId="32" pageNumber="67" value="2013-07-16">
<collectingDate id="9B70D77AE9224A6CFF3CFC17FECDFC66" box="[151,307,979,1005]" pageId="32" pageNumber="67" value="2013-07-16">16. VII. 2013</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFEEBFC17FE42FC66" box="[320,444,979,1005]" pageId="32" pageNumber="67">S. Shimizu</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFE55FC17FD94FC66" box="[510,618,979,1005]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFDD5FC17FC1FFC66" box="[638,993,979,1005]" collectingDate="2013-08-07" collectionCode="ZSM" collectorName="S. Shimizu" country="Japan" county="Myokou" location="Suginosawa" municipality="City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 F, ditto,
<date id="8B342E92E9224A6CFD5FFC17FC77FC66" box="[756,905,979,1005]" pageId="32" pageNumber="67" value="2013-08-07">
<collectingDate id="9B70D77AE9224A6CFD5FFC17FC77FC66" box="[756,905,979,1005]" pageId="32" pageNumber="67" value="2013-08-07">7. VIII. 2013</collectingDate>
</date>
(
<collectionCode id="999B9097E9224A6CFC32FC17FC26FC66" box="[921,984,979,1005]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="32" pageNumber="67" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFC45FC17FD01FB9A" collectingDate="2017-05-31" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Kouyoudai" municipality="Narusawa Vil." pageId="32" pageNumber="67" specimenCount="1" stateProvince="Yamanashi" typeStatus="paratype">
1 M,
<collectingRegion id="3D4EC6B0E9224A6CFB87FC17FB16FC66" box="[1068,1256,979,1005]" country="Japan" name="Yamanashi" pageId="32" pageNumber="67">Yamanashi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFB5FFC17FA66FC66" box="[1268,1432,979,1005]" pageId="32" pageNumber="67">Narusawa Vil.</collectingMunicipality>
,
<location id="FA555E89E9224A6CFF3CFC33FEE8FB9A" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFF3CFC33FEE8FB9A" box="[151,278,1015,1041]" country="Japan" municipality="Narusawa Vil." name="Kouyoudai" pageId="32" pageNumber="67" stateProvince="Yamanashi">Kouyoudai</location>
,
<date id="8B342E92E9224A6CFE88FC33FE54FB9A" box="[291,426,1015,1041]" pageId="32" pageNumber="67" value="2017-05-31">
<collectingDate id="9B70D77AE9224A6CFE88FC33FE54FB9A" box="[291,426,1015,1041]" pageId="32" pageNumber="67" value="2017-05-31">31. V. 2017</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFE1CFC33FDB7FB9A" box="[439,585,1015,1041]" pageId="32" pageNumber="67">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFD20FC33FD09FB9A" box="[651,759,1015,1041]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFCA0FC33FDFCFBBE" collectingDate="1982-07-29" collectionCode="KPMNH" collectorName="T. Murota" country="Japan" county="Oono City" location="Ropponnoki" municipality="Koike" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Fukui" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFC94FC33FC3DFB9A" box="[831,963,1015,1041]" country="Japan" name="Fukui" pageId="32" pageNumber="67">Fukui Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9224A6CFC64FC33FBB9FB9A" box="[975,1095,1015,1041]" pageId="32" pageNumber="67">Oono City</collectingCounty>
,
<collectingMunicipality id="1F519228E9224A6CFBFFFC33FB65FB9A" box="[1108,1179,1015,1041]" pageId="32" pageNumber="67">Koike</collectingMunicipality>
,
<location id="FA555E89E9224A6CFB02FC33FAC8FB9A" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB02FC33FAC8FB9A" box="[1193,1334,1015,1041]" country="Japan" county="Oono City" municipality="Koike" name="Ropponnoki" pageId="32" pageNumber="67" stateProvince="Fukui">Ropponnoki</location>
,
<date id="8B342E92E9224A6CFAE8FC33FF2CFBBE" pageId="32" pageNumber="67" value="1982-07-29">
<collectingDate id="9B70D77AE9224A6CFAE8FC33FF2CFBBE" pageId="32" pageNumber="67" value="1982-07-29">29. VII. 1982</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFF76FBDFFEB3FBBE" box="[221,333,1051,1077]" pageId="32" pageNumber="67">T. Murota</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFE25FBDFFE04FBBE" box="[398,506,1051,1077]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFDA5FBDFFE72FBD2" collectingDate="2011-08-06" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Mt. Hyonosen" municipality="Wakasa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Tottori" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFD94FBDFFD32FBBE" box="[575,716,1051,1077]" country="Japan" name="Tottori" pageId="32" pageNumber="67">Tottori Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFD7CFBDFFC89FBBE" box="[727,887,1051,1077]" pageId="32" pageNumber="67">Wakasa Town</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC28FBDFFBD8FBBE" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC28FBDFFBD8FBBE" box="[899,1062,1051,1077]" country="Japan" municipality="Wakasa Town" name="Mt. Hyonosen" pageId="32" pageNumber="67" stateProvince="Tottori">Mt. Hyonosen</location>
,
<date id="8B342E92E9224A6CFB98FBDFFB36FBBE" box="[1075,1224,1051,1077]" pageId="32" pageNumber="67" value="2011-08-06">
<collectingDate id="9B70D77AE9224A6CFB98FBDFFB36FBBE" box="[1075,1224,1051,1077]" pageId="32" pageNumber="67" value="2011-08-06">6. VIII. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFB7FFBDFFA9AFBBE" box="[1236,1380,1051,1077]" pageId="32" pageNumber="67">K. Watanabe</collectorName>
leg.,
<collectingMethod id="26CB7045E9224A6CFF3CFBFBFEF6FBD2" box="[151,264,1087,1113]" pageId="32" pageNumber="67">Light trap</collectingMethod>
(
<collectionCode id="999B9097E9224A6CFEB3FBFBFE7AFBD2" box="[280,388,1087,1113]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFE33FBFBFA66FBD2" box="[408,1432,1086,1113]" collectingDate="1955-07-29" collectionCode="EUM" collectorName="S. Ueda" country="Japan" location="Jyojyusha" municipality="Mt. Ishizuchi" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFE09FBFAFDF8FBD3" bold="true" box="[418,518,1086,1112]" pageId="32" pageNumber="67">Shikoku</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9224A6CFDE1FBFBFD2BFBD2" box="[586,725,1087,1113]" country="Japan" name="Ehime" pageId="32" pageNumber="67">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFD4AFBFBFC85FBD2" box="[737,891,1087,1113]" pageId="32" pageNumber="67">Mt. Ishizuchi</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC2CFBFBFC04FBD2" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC2CFBFBFC04FBD2" box="[903,1018,1087,1113]" country="Japan" municipality="Mt. Ishizuchi" name="Jyojyusha" pageId="32" pageNumber="67" stateProvince="Ehime">Jyojyusha</location>
,
<date id="8B342E92E9224A6CFBACFBFBFB5DFBD2" box="[1031,1187,1086,1113]" pageId="32" pageNumber="67" value="1955-07-29">
<collectingDate id="9B70D77AE9224A6CFBACFBFBFB5DFBD2" box="[1031,1187,1086,1113]" pageId="32" pageNumber="67" value="1955-07-29">29. VII. 1955</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFB04FBFBFAF7FBD2" box="[1199,1289,1087,1113]" pageId="32" pageNumber="67">S. Ueda</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFAE7FBFBFA6EFBD2" box="[1356,1424,1087,1113]" pageId="32" pageNumber="67">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFF3CFBA7FB7EFBF6" box="[151,1152,1123,1149]" collectingDate="1961-07-11" collectionCode="MNHAH" collectorName="A. Nakanishi" country="Japan" location="Cape Sata" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFF0AFBA7FF00FBF6" bold="true" box="[161,254,1123,1149]" pageId="32" pageNumber="67">Kyushu</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9224A6CFEE1FBA7FDF4FBF6" box="[330,522,1123,1149]" country="Japan" name="Kagoshima" pageId="32" pageNumber="67">Kagoshima Pref.</collectingRegion>
,
<location id="FA555E89E9224A6CFDBFFBA7FD76FBF6" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFDBFFBA7FD76FBF6" box="[532,648,1123,1149]" country="Japan" name="Cape Sata" pageId="32" pageNumber="67" stateProvince="Kagoshima">Cape Sata</location>
,
<date id="8B342E92E9224A6CFD38FBA7FCD4FBF6" box="[659,810,1123,1149]" pageId="32" pageNumber="67" value="1961-07-11">
<collectingDate id="9B70D77AE9224A6CFD38FBA7FCD4FBF6" box="[659,810,1123,1149]" pageId="32" pageNumber="67" value="1961-07-11">11. VII. 1961</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFC9EFBA7FC37FBF6" box="[821,969,1123,1149]" pageId="32" pageNumber="67">A. Nakanishi</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFBA3FBA7FB87FBF6" box="[1032,1145,1123,1149]" pageId="32" pageNumber="67">MNHAH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6FFF6CFB42FA75FEEE" lastPageId="35" lastPageNumber="70" pageId="32" pageNumber="67" type="description">
<paragraph id="FF350852E9224A6CFF6CFB42FC36FB4E" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFB42FEAFFB2B" bold="true" box="[199,337,1158,1184]" pageId="32" pageNumber="67">Description</emphasis>
.
<emphasis id="CDFED440E9224A6CFEF5FB43FE4BFB2A" bold="true" box="[350,437,1159,1185]" pageId="32" pageNumber="67">Female</emphasis>
(n = 8). Body polished and covered with punctures and silvery setae; length 9.010.8 (HT: 10.8) mm. Setae on head and mesosoma not exceptionally long and erect
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFB0BFA7BFA6E" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
Head 0.6 × as long as wide. Combined length of clypeus and face 0.951.1 (HT: 1.0) × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.50.6 (HT: 0.5) × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.61.8 (HT: 1.7) × as long as OD. OOL 1.21.4 (HT: 1.2) × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9224A6CFF34FA63FEECFA4A" box="[159,274,1447,1473]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." figureDoi="http://doi.org/10.5281/zenodo.13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="32" pageNumber="67">Fig. 22 D</figureCitation>
). Antenna with 4244 (HT: 44) flagellomeres. FL I 4.14.7 (HT: 4.7) × as long as maximum depth in lateral view and 1.51.85 (HT: 1.75) × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFA2BFAD9F8C6" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
Mesosoma coarsely and densely punctate, 1.41.45 (HT: 1.4) × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate ventrally, sparsely punctate dorsally; speculum with a large smooth area (
<figureCitation id="67B114D7E9224A6CFB31F9F3FAF6F9DA" box="[1178,1288,1591,1617]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." figureDoi="http://doi.org/10.5281/zenodo.13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="32" pageNumber="67">Fig. 22 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally (
<figureCitation id="67B114D7E9224A6CFD09F99FFCF9F9FE" box="[674,775,1627,1653]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.13331015" httpUri="https://zenodo.org/record/13331015/files/figure.png" pageId="32" pageNumber="67">Fig. 1 L</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum largely absent medially, obsolete laterally. Fore wing length 7.68.2 (HT: 8.2) mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E9224A6CFEBDF903FE79F96A" box="[278,391,1735,1761]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." figureDoi="http://doi.org/10.5281/zenodo.13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="32" pageNumber="67">Fig. 22 B</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.25.7 (HT: 5.25) × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.04.1 (HT: 4.1): 2.0: 1.3: 0.60.7 (HT: 0.7): 0.70.8 (HT: 0.8). Hind tarsal claws simple except for minute subbasal teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9224A6CFB12F8F7FAE4F8C6" box="[1209,1306,1843,1869]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="32" pageNumber="67">Fig. 2 N</figureCitation>
).
</paragraph>
<paragraph id="FF350852E9224A6CFF6CF893FDDBF856" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">Metasoma. T I 1.92.4 (HT: 2.2) × as long as maximum width, finely punctate laterally, smooth medially. T II 1.351.5 (HT: 1.5) × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine and dense punctures. Ovipositor sheath 2.12.5 (HT: 2.3) × as long as hind tibia, ca. 0.80.85 (HT: 0.8) × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE9234A6DFF3CF9A8FA6AF923" ID-DOI="http://doi.org/10.5281/zenodo.13331069" ID-Zenodo-Dep="13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="33" pageNumber="68" startId="33.[151,248,1644,1668]" targetBox="[151,1436,181,1619]" targetPageId="33" targetType="figure">
<paragraph id="FF350852E9234A6DFF3CF9A8FA6AF923" blockId="33.[151,1437,1644,1704]" pageId="33" pageNumber="68">
<emphasis id="CDFED440E9234A6DFF3CF9A8FEE0F90F" bold="true" box="[151,286,1644,1668]" pageId="33" pageNumber="68">FIGURE 22.</emphasis>
<taxonomicName id="388A73D1E9234A6DFE8EF9A9FE0DF90F" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[293,499,1645,1668]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="68" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9234A6DFE8EF9A9FE0DF90F" box="[293,499,1645,1668]" italics="true" pageId="33" pageNumber="68">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9234A6DFE51F9A8FDB4F90F" bold="true" box="[506,586,1644,1668]" pageId="33" pageNumber="68">
<taxonomicNameLabel id="D6CD693BE9234A6DFE51F9A8FDB4F90F" box="[506,586,1644,1668]" pageId="33" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus.
</paragraph>
</caption>
<paragraph id="FF350852E9234A6EFF6CF912FA9DFE02" blockId="33.[151,1437,1750,2029]" lastBlockId="34.[151,1437,151,393]" lastPageId="34" lastPageNumber="69" pageId="33" pageNumber="68">
Colouration (
<figureCitation id="67B114D7E9234A6DFEF7F912FE10F97A" box="[348,494,1750,1777]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." figureDoi="http://doi.org/10.5281/zenodo.13331069" httpUri="https://zenodo.org/record/13331069/files/figure.png" pageId="33" pageNumber="68">Figs 22 AF</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, malar space, a pair of small yellow spots below antennal sockets, orbits on face, frons, vertex and gena, and palpi yellow. Antenna blackish-brown to reddish-brown. Scape and pedicel usually partly tinged with yellow ventrally. Two spots of collar, a L-shape marking of dorsal margin of pronotum, a pair of antero-lateral markings and a median marking of mesoscutum, subtegular ridge, antero-dorsal and postero-ventral spots of mesopleuron, scutellum except for a median black spot, postscutellum, posterior spot of metapleuron, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow. Scutellum sometimes entirely yellow. Metasomal tergites black except for a pair of basal spots of T I and posterior margins of T I to T III yellow to ivory. Posterior margins of T IV sometimes tinged with yellow. Membranous parts of metasomal sternites largely yellow. Subgenital plate blackish-brown, usually partly tinged with yellowish-brown. Ovipositor reddish-brown. Wings hyaline.Apical clouded area of fore wing slightly to weakly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow; mid coxa partly tinged with blackish-brown basally. Hind coxa black except for a dorsal spot and apex yellow to yellowish-brown. Hind trochanter black to blackish-brown. Hind trochantellus reddish-yellow. Hind femur reddish-brown except for apex tinged with yellow. Hind tibia and tibial spurs yellowish-brown to whitish-brown except for apical part of tibia blackish-brown. Hind tarsus brown, partly tinged with yellow.
</paragraph>
<caption id="ABF558DAE9204A6EFF3CF862FE73F868" ID-DOI="http://doi.org/10.5281/zenodo.13331071" ID-Zenodo-Dep="13331071" httpUri="https://zenodo.org/record/13331071/files/figure.png" pageId="34" pageNumber="69" startId="34.[151,250,1958,1982]" targetBox="[151,1436,417,1935]" targetPageId="34" targetType="figure">
<paragraph id="FF350852E9204A6EFF3CF862FE73F868" blockId="34.[151,1437,1958,2019]" pageId="34" pageNumber="69">
<emphasis id="CDFED440E9204A6EFF3CF862FEDFF835" bold="true" box="[151,289,1958,1982]" pageId="34" pageNumber="69">FIGURE 23.</emphasis>
<taxonomicName id="388A73D1E9204A6EFE83F863FE07F835" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[296,505,1959,1982]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="69" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9204A6EFE83F863FE07F835" box="[296,505,1959,1982]" italics="true" pageId="34" pageNumber="69">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9204A6EFDAAF863FDACF834" bold="true" box="[513,594,1959,1983]" pageId="34" pageNumber="69">
<taxonomicNameLabel id="D6CD693BE9204A6EFDAAF863FDACF834" box="[513,594,1959,1983]" pageId="34" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view.
</paragraph>
</caption>
<paragraph id="FF350852E9214A6FFF6CFF53FA75FEEE" blockId="35.[151,1437,150,789]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFF53FEFAFF3A" bold="true" box="[199,260,151,177]" pageId="35" pageNumber="70">Male</emphasis>
(n = 3). Similar to female (
<figureCitation id="67B114D7E9214A6FFD9CFF53FD32FF3A" box="[567,716,151,177]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13331071" httpUri="https://zenodo.org/record/13331071/files/figure.png" pageId="35" pageNumber="70">Figs 23 AD</figureCitation>
). A male with body length
<quantity id="3872A5B7E9214A6FFC51FF52FBADFF3A" box="[1018,1107,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="70" unit="mm" value="8.5">8.5 mm</quantity>
. Malar space 0.9 × as long as basal mandibular width. Tergal spiracles enlarged and with circular felt-like area, its length less than 0.2× as long as T I (
<figureCitation id="67B114D7E9214A6FFF4FFF1BFE9DFF72" box="[228,355,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="35" pageNumber="70">Figs 2 R, S</figureCitation>
,
<figureCitation id="67B114D7E9214A6FFEDBFF1BFE55FF72" box="[368,427,223,249]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13331071" httpUri="https://zenodo.org/record/13331071/files/figure.png" pageId="35" pageNumber="70">23 D</figureCitation>
). Basal part of T I punctate laterally (
<figureCitation id="67B114D7E9214A6FFCE7FF1BFC59FF72" box="[844,935,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="35" pageNumber="70">Fig. 2 S</figureCitation>
). Hind femur 5.36.3 × as long as maximum depth in lateral view. T I 2.42.8 × as long as maximum width. T II 1.72.05 × as long as maximum width. Yellow area of body larger than female. Face yellow except for a small median longitudinal spot. Two yellow spots of mesopleuron united into a single C-shaped marking. Posterior segments of metasoma tinged with reddish-brown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFEAAFC62FE02" box="[199,924,366,393]" pageId="35" pageNumber="70" type="distribution">
<paragraph id="FF350852E9214A6FFF6CFEAAFC62FE02" blockId="35.[151,1437,150,789]" box="[199,924,366,393]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFEAAFEA7FE03" bold="true" box="[199,345,366,392]" pageId="35" pageNumber="70">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9214A6FFECCFEABFE59FE02" box="[359,423,367,393]" name="Japan" pageId="35" pageNumber="70">Japan</collectingCountry>
(
<collectingRegion id="3D4EC6B0E9214A6FFE1DFEABFDD7FE02" box="[438,553,367,393]" country="Japan" name="Hokkaido" pageId="35" pageNumber="70">Hokkaido</collectingRegion>
, Honshu, Shikoku, and Kyushu).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFE57FE36FE5A" pageId="35" pageNumber="70" type="etymology">
<paragraph id="FF350852E9214A6FFF6CFE57FC35FE26" blockId="35.[151,1437,150,789]" box="[199,971,403,429]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE57FEB6FE26" bold="true" box="[199,328,403,429]" pageId="35" pageNumber="70">Etymology</emphasis>
. The specific name is from old name of
<collectingCountry id="879D48C2E9214A6FFCA5FE57FCB0FE26" box="[782,846,403,429]" name="Japan" pageId="35" pageNumber="70">Japan</collectingCountry>
“Yamato”.
</paragraph>
<paragraph id="FF350852E9214A6FFF6CFE72FE36FE5A" blockId="35.[151,1437,150,789]" box="[199,456,438,465]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE72FEBDFE5B" bold="true" box="[199,323,438,464]" pageId="35" pageNumber="70">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFE1FFC69FC9E" pageId="35" pageNumber="70" type="discussion">
<paragraph id="FF350852E9214A6FFF6CFE1FFC69FC9E" blockId="35.[151,1437,150,789]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE1FFECAFE7E" bold="true" box="[199,308,475,501]" pageId="35" pageNumber="70">Remarks</emphasis>
. This species is closely related to
<taxonomicName id="388A73D1E9214A6FFD69FE1FFCAEFE7E" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[706,848,475,501]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="CDFED440E9214A6FFD69FE1FFCAEFE7E" box="[706,848,475,501]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9214A6FFCF2FE1FFC4BFE7E" bold="true" box="[857,949,475,501]" pageId="35" pageNumber="70">
<taxonomicNameLabel id="D6CD693BE9214A6FFCF2FE1FFC4BFE7E" box="[857,949,475,501]" pageId="35" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Both species were collected same locality (
<collectingRegion id="3D4EC6B0E9214A6FFF35FE3BFEB1FD92" box="[158,335,511,537]" country="Japan" name="Yamagata" pageId="35" pageNumber="70">Yamagata Pref.</collectingRegion>
, Mamurogawa Vil.) on same day. This species can be distinguished by the punctures on upper part of mesopleuron which are irregular and sparser than lower part (same density in
<taxonomicName id="388A73D1E9214A6FFBBDFDE7FB5FFDB6" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[1046,1185,547,573]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis">
<emphasis id="CDFED440E9214A6FFBBDFDE7FB5FFDB6" box="[1046,1185,547,573]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
) and the first to fourth hind tarsomeres usually more or less darkened (ivory in
<taxonomicName id="388A73D1E9214A6FFCB7FD83FC57FDEA" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[796,937,583,609]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis">
<emphasis id="CDFED440E9214A6FFCB7FD83FC57FDEA" box="[796,937,583,609]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
). In the key provided by
<bibRefCitation id="9B1B75A3E9214A6FFB65FD83FA93FDEA" author="Riedel, M." box="[1230,1389,583,609]" pageId="35" pageNumber="70" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel (2021)</bibRefCitation>
, the females and males runs to couplet 47 (female of
<taxonomicName id="388A73D1E9214A6FFD14FDAFFCCCFD0E" authorityName="Seyrig" authorityYear="1928" box="[703,818,619,645]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFD14FDAFFCCCFD0E" box="[703,818,619,645]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
) and 7 (male of
<taxonomicName id="388A73D1E9214A6FFC5BFDAFFB9AFD0E" authorityName="Seyrig" authorityYear="1928" box="[1008,1124,619,645]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC5BFDAFFB9AFD0E" box="[1008,1124,619,645]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), but differs by the antenna with 4244 flagellomeres (3840 flagellomeres in female of
<taxonomicName id="388A73D1E9214A6FFC95FD4BFC4FFD22" authorityName="Seyrig" authorityYear="1928" box="[830,945,655,681]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC95FD4BFC4FFD22" box="[830,945,655,681]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the malar space 0.50.6 × as long as basal mandibular width (
<quantity id="3872A5B7E9214A6FFEC5FD77FE4FFD46" box="[366,433,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.7779999999999998" pageId="35" pageNumber="70" unit="in" value="0.7">0.7 in</quantity>
female of
<taxonomicName id="388A73D1E9214A6FFD87FD77FD5EFD46" authorityName="Seyrig" authorityYear="1928" box="[556,672,691,717]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFD87FD77FD5EFD46" box="[556,672,691,717]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the median part of face largely black (largely yellow in female of
<taxonomicName id="388A73D1E9214A6FFF3CFD13FEF4FD7A" authorityName="Seyrig" authorityYear="1928" box="[151,266,727,753]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFF3CFD13FEF4FD7A" box="[151,266,727,753]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the yellow markings of frons and vertex narrower than
<taxonomicName id="388A73D1E9214A6FFC23FD13FC05FD7A" authorityName="Seyrig" authorityYear="1928" box="[904,1019,727,753]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC23FD13FC05FD7A" box="[904,1019,727,753]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
, and the hind coxa black with yellow apical marking(s) (yellow with black spot in the male of
<taxonomicName id="388A73D1E9214A6FFCB8FD3FFC78FC9E" authorityName="Seyrig" authorityYear="1928" box="[787,902,763,789]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFCB8FD3FFC78FC9E" box="[787,902,763,789]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>