treatments-xml/data/77/23/B9/7723B944E91C4A6CFF3CFB43FD1BFDEA.xml

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<mods:title id="8E0D64737F3A3FD89C5DCFAF266183C7">Revision of the genus Syzeuctus Förster, 1869 (Hymenoptera, Ichneumonidae, Banchinae) from Japan</mods:title>
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<mods:date id="D483085A0A45AF11B1B4660F8D988A83">2024</mods:date>
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<paragraph id="FF350852E91C4A52FF3CFB43FDB7FB2B" blockId="30.[151,585,1158,1221]" box="[151,585,1158,1185]" pageId="30" pageNumber="65">
<heading id="A47DBF3EE91C4A52FF3CFB43FDB7FB2B" bold="true" box="[151,585,1158,1185]" fontSize="11" level="1" pageId="30" pageNumber="65" reason="1">
<taxonomicName id="388A73D1E91C4A52FF3CFB43FDB7FB2B" ID-CoL="545JL" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[151,585,1158,1185]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="65" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91C4A52FF3CFB43FDB7FB2B" bold="true" box="[151,585,1158,1185]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF3CFB43FE60FB2A" bold="true" box="[151,414,1158,1185]" italics="true" pageId="30" pageNumber="65">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91C4A52FE0DFB42FDB7FB2B" author="Uchida, T." box="[422,585,1158,1184]" pageId="30" pageNumber="65" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</emphasis>
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<subSubSection id="B7905BD9E91C4A52FF3CFB6FFE56FB4E" box="[151,424,1195,1221]" pageId="30" pageNumber="65" type="description">
<paragraph id="FF350852E91C4A52FF3CFB6FFE56FB4E" blockId="30.[151,585,1158,1221]" box="[151,424,1195,1221]" pageId="30" pageNumber="65">
(
<figureCitation id="67B114D7E91C4A52FF34FB6FFEFAFB4E" box="[159,260,1195,1221]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.13331015" httpUri="https://zenodo.org/record/13331015/files/figure.png" pageId="30" pageNumber="65">Figs 1 K</figureCitation>
,
<figureCitation id="67B114D7E91C4A52FEA4FB6FFEC1FB4E" box="[271,319,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="30" pageNumber="65">2 M</figureCitation>
,
<figureCitation id="67B114D7E91C4A52FEE0FB6FFE5EFB4E" box="[331,416,1195,1221]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." figureDoi="http://doi.org/10.5281/zenodo.13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="30" pageNumber="65">21 AF</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="DF9A2F7CE91C4A52FF3CFB2BFDD0FA8C" box="[151,558,1263,1287]" pageId="30" pageNumber="65">
<treatmentCitation id="7E2B2E43E91C4A52FF3CFB2BFDD4FA8C" author="Uchida, T." box="[151,554,1263,1287]" page="95" pageId="30" pageNumber="65" year="1928">
<taxonomicName id="388A73D1E91C4A52FF3CFB2BFDD4FA8C" ID-CoL="545JL" authority="Uchida, 1928: 95" authorityName="Uchida" authorityPageNumber="95" authorityYear="1928" box="[151,554,1263,1287]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="65" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91C4A52FF3CFB2BFE8CFA8C" box="[151,370,1263,1287]" italics="true" pageId="30" pageNumber="65">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91C4A52FED3FB2BFDD4FA8C" author="Uchida, T." box="[376,554,1263,1287]" pageId="30" pageNumber="65" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928: 95</bibRefCitation>
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.
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</paragraph>
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<emphasis id="CDFED440E91C4A52FF3CFAF1FE79FAC4" bold="true" box="[151,391,1333,1359]" pageId="30" pageNumber="65">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE91C4A52FE3CFAF1FA7EFAC4" box="[407,1408,1333,1360]" collectingDate="1966-07-15" collectingDateMax="1966-07-18" collectingDateMin="1966-07-15" collectionCode="SEHU" collectorName="K. Kusigemati" country="Japan" location="Mt. Daisetsu" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
<emphasis id="CDFED440E91C4A52FE3CFAF1FE0FFAC4" bold="true" box="[407,497,1333,1359]" pageId="30" pageNumber="65">
<collectingCountry id="879D48C2E91C4A52FE3CFAF1FE0FFAC4" box="[407,497,1333,1359]" name="Japan" pageId="30" pageNumber="65">JAPAN</collectingCountry>
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: [
<emphasis id="CDFED440E91C4A52FDA0FAF1FD7FFAC4" bold="true" box="[523,641,1333,1359]" pageId="30" pageNumber="65">
<collectingRegion id="3D4EC6B0E91C4A52FDA0FAF1FD7FFAC4" box="[523,641,1333,1359]" country="Japan" name="Hokkaido" pageId="30" pageNumber="65">Hokkaido</collectingRegion>
</emphasis>
] 1 F,
<location id="FA555E89E91C4A52FD62FAF2FCA3FAC4" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FD62FAF2FCA3FAC4" box="[713,861,1333,1360]" country="Japan" name="Mt. Daisetsu" pageId="30" pageNumber="65" stateProvince="Hokkaido">Mt. Daisetsu</location>
,
<date id="8B342E92E91C4A52FCC0FAF1FBCDFAC4" box="[875,1075,1333,1360]" pageId="30" pageNumber="65" value="1966-07-15" valueMax="1966-07-18" valueMin="1966-07-15">
<collectingDate id="9B70D77AE91C4A52FCC0FAF1FBCDFAC4" box="[875,1075,1333,1360]" pageId="30" pageNumber="65" value="1966-07-15" valueMax="1966-07-18" valueMin="1966-07-15">1518. VII. 1966</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FBEBFAF2FB1BFAC4" box="[1088,1253,1333,1360]" pageId="30" pageNumber="65">K. Kusigemati</collectorName>
leg. (
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)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FA25FAF1FC23FAF8" collectingDate="2012-07-16" collectionCode="KPMNH" collectorName="M. Ito" country="Japan" county="Horokanai Town" location="Uryu" municipality="Moshiri" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
1 F,
<collectingCounty id="165470DEE91C4A52FF1FFA9DFE8BFAFF" box="[180,373,1369,1396]" pageId="30" pageNumber="65">Horokanai Town</collectingCounty>
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<location id="FA555E89E91C4A52FE40FA9EFDD9FAFF" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FE40FA9EFDD9FAFF" box="[491,551,1370,1396]" country="Japan" county="Horokanai Town" municipality="Moshiri" name="Uryu" pageId="30" pageNumber="65" stateProvince="Hokkaido">Uryu</location>
,
<date id="8B342E92E91C4A52FD9FFA9DFD2FFAF8" box="[564,721,1369,1396]" pageId="30" pageNumber="65" value="2012-07-16">
<collectingDate id="9B70D77AE91C4A52FD9FFA9DFD2FFAF8" box="[564,721,1369,1396]" pageId="30" pageNumber="65" value="2012-07-16">16. VII. 2012</collectingDate>
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,
<collectorName id="527F6D84E91C4A52FD75FA9EFCD8FAFF" box="[734,806,1370,1396]" pageId="30" pageNumber="65">M. Ito</collectorName>
leg. (
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;
<materialsCitation id="4FE2020FE91C4A52FC42FA9DFDFBFA1C" collectingDate="2007-06-22" collectingDateMax="2007-07-10" collectingDateMin="2007-06-22" collectionCode="ZSM" collectorName="A. Ueda" country="Japan" location="Kuriyama Town" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
1 F,
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,
<date id="8B342E92E91C4A52FB4EFA9DFF2CFA1C" pageId="30" pageNumber="65" value="2007-06-22" valueMax="2007-07-10" valueMin="2007-06-22">
<collectingDate id="9B70D77AE91C4A52FB4EFA9DFF2CFA1C" pageId="30" pageNumber="65" value="2007-06-22" valueMax="2007-07-10" valueMin="2007-06-22">22. VI.10. VII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FF77FABAFEC4FA1C" box="[220,314,1405,1432]" pageId="30" pageNumber="65">A. Ueda</collectorName>
leg., MsT (
<collectionCode id="999B9097E91C4A52FE16FAB9FE02FA1C" box="[445,508,1405,1431]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="30" pageNumber="65" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FDBAFABAFF08FA30" collectingDate="1926-07-30" collectionCode="SEHU" collectorName="S. Hirayama" country="Japan" location="Mt. Takaosan" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Tokyo" typeStatus="lectotype">
[
<emphasis id="CDFED440E91C4A52FDB1FAB9FD89FA1C" bold="true" box="[538,631,1405,1431]" pageId="30" pageNumber="65">Honshu</emphasis>
] 1 F (
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),
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<location id="FA555E89E91C4A52FC39FABAFBD3FA1C" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FC39FABAFBD3FA1C" box="[914,1069,1405,1432]" country="Japan" name="Mt. Takaosan" pageId="30" pageNumber="65" stateProvince="Tokyo">Mt. Takaosan</location>
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<date id="8B342E92E91C4A52FB92FAB9FB2DFA1C" box="[1081,1235,1405,1432]" pageId="30" pageNumber="65" value="1926-07-30">
<collectingDate id="9B70D77AE91C4A52FB92FAB9FB2DFA1C" box="[1081,1235,1405,1432]" pageId="30" pageNumber="65" value="1926-07-30">30. VII. 1926</collectingDate>
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,
<collectorName id="527F6D84E91C4A52FB74FAB9FA95FA1C" box="[1247,1387,1405,1431]" pageId="30" pageNumber="65">S. Hirayama</collectorName>
leg. (
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)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FEA9FA65FB30FA30" box="[258,1230,1441,1468]" collectingDate="2004-05-26" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Enkaizan" municipality="Yokohama City" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Kanagawa" typeStatus="lectotype">
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FE9FFA66FE15FA30" box="[308,491,1441,1468]" country="Japan" name="Kanagawa" pageId="30" pageNumber="65">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FE5EFA65FD58FA30" box="[501,678,1441,1467]" pageId="30" pageNumber="65">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E91C4A52FD19FA65FCE3FA30" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FD19FA65FCE3FA30" box="[690,797,1441,1467]" country="Japan" municipality="Yokohama City" name="Enkaizan" pageId="30" pageNumber="65" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E91C4A52FC82FA65FC53FA30" box="[809,941,1441,1468]" pageId="30" pageNumber="65" value="2004-05-26">
<collectingDate id="9B70D77AE91C4A52FC82FA65FC53FA30" box="[809,941,1441,1468]" pageId="30" pageNumber="65" value="2004-05-26">26. V. 2004</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FC12FA66FBE4FA30" box="[953,1050,1441,1468]" pageId="30" pageNumber="65">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FBF1FA65FB38FA30" box="[1114,1222,1441,1467]" pageId="30" pageNumber="65">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FB71FA65FC7BFA54" collectingDate="1951-05-22" collectionCode="MNHAH" collectorName="A. Nagatomi" country="Japan" location="Sakazukiyama" municipality="Tamba" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hyogo" typeStatus="lectotype">
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FAA7FA66FA66FA30" box="[1292,1432,1441,1468]" country="Japan" name="Hyogo" pageId="30" pageNumber="65">Hyogo Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FF3CFA01FF18FA54" box="[151,230,1477,1503]" pageId="30" pageNumber="65">Tamba</collectingMunicipality>
,
<location id="FA555E89E91C4A52FF58FA01FE67FA54" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FF58FA01FE67FA54" box="[243,409,1477,1503]" country="Japan" municipality="Tamba" name="Sakazukiyama" pageId="30" pageNumber="65" stateProvince="Hyogo">Sakazukiyama</location>
,
<date id="8B342E92E91C4A52FE0DFA01FDD5FA6B" box="[422,555,1477,1504]" pageId="30" pageNumber="65" value="1951-05-22">
<collectingDate id="9B70D77AE91C4A52FE0DFA01FDD5FA6B" box="[422,555,1477,1504]" pageId="30" pageNumber="65" value="1951-05-22">22. V. 1951</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FD9CFA02FD37FA54" box="[567,713,1477,1504]" pageId="30" pageNumber="65">A. Nagatomi</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FCA7FA01FC83FA54" box="[780,893,1477,1503]" pageId="30" pageNumber="65">MNHAH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FC3AFA02FC2CF988" collectingDate="2014-05-21" collectingDateMax="2014-05-25" collectingDateMin="2014-05-21" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Matsuyamasougou-park" municipality="Matsuyama City" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Ehime" typeStatus="lectotype">
[
<emphasis id="CDFED440E91C4A52FC30FA01FBF6FA54" bold="true" box="[923,1032,1477,1503]" pageId="30" pageNumber="65">Shikoku]</emphasis>
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FBEFFA01FB31FA54" box="[1092,1231,1477,1503]" country="Japan" name="Ehime" pageId="30" pageNumber="65">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FB70FA02FA66FA54" box="[1243,1432,1477,1504]" pageId="30" pageNumber="65">Matsuyama City</collectingMunicipality>
,
<location id="FA555E89E91C4A52FF3CFA2DFE57F988" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FF3CFA2DFE57F988" box="[151,425,1513,1539]" country="Japan" municipality="Matsuyama City" name="Matsuyamasougou-park" pageId="30" pageNumber="65" stateProvince="Ehime">Matsuyamasougou-park</location>
,
<date id="8B342E92E91C4A52FE1FFA2DFD9FF988" box="[436,609,1513,1540]" pageId="30" pageNumber="65" value="2014-05-21" valueMax="2014-05-25" valueMin="2014-05-21">
<collectingDate id="9B70D77AE91C4A52FE1FFA2DFD9FF988" box="[436,609,1513,1540]" pageId="30" pageNumber="65" value="2014-05-21" valueMax="2014-05-25" valueMin="2014-05-21">2125. V. 2014</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FDC7FA2EFCFAF98F" box="[620,772,1514,1540]" pageId="30" pageNumber="65">E. Yamamoto</collectorName>
leg., MsT (
<collectionCode id="999B9097E91C4A52FC2EFA2DFC37F988" box="[901,969,1513,1539]" pageId="30" pageNumber="65">EUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A6CFF6CF9C9FDD9FE26" lastPageId="32" lastPageNumber="67" pageId="30" pageNumber="65" type="description">
<paragraph id="FF350852E91C4A52FF6CF9C9FCABF9C7" blockId="30.[151,1437,1333,2008]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CF9C9FEAFF9AC" bold="true" box="[199,337,1549,1575]" pageId="30" pageNumber="65">Description</emphasis>
.
<emphasis id="CDFED440E91C4A52FEF5F9C9FE4BF9AC" bold="true" box="[350,437,1549,1575]" pageId="30" pageNumber="65">Female</emphasis>
(n = 6). Body polished and covered with punctures and silvery setae; length 9.0
<quantity id="3872A5B7E91C4A52FA9FF9C9FA67F9A3" box="[1332,1433,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="30" pageNumber="65" unit="mm" value="10.2">10.2 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E91C4A52FF6CF991FDF3F8E0" blockId="30.[151,1437,1333,2008]" pageId="30" pageNumber="65">
Head 0.6 × as long as wide. Combined length of clypeus and face 1.051.1 × facial width. Clypeus smooth except for dorsal margin punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.4 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.651.9 × as long as OD. OOL 1.41.5 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E91C4A52FC22F8CDFC05F8A8" box="[905,1019,1801,1827]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." figureDoi="http://doi.org/10.5281/zenodo.13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="30" pageNumber="65">Fig. 21 D</figureCitation>
). Antenna with 3842 flagellomeres. FL I 3.84.4 × as long as maximum depth in lateral view and 1.551.75 × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E91C4A53FF6CF8B2FE45FE02" blockId="30.[151,1437,1333,2008]" lastBlockId="31.[151,1437,151,393]" lastPageId="31" lastPageNumber="66" pageId="30" pageNumber="65">
Mesosoma coarsely and densely punctate, 1.4 × as long as maximum height. Pronotum with rugulose area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E91C4A52FD9AF879FD61F85C" box="[561,671,1981,2007]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." figureDoi="http://doi.org/10.5281/zenodo.13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="30" pageNumber="65">Fig. 21 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E91D4A53FC60FF53FBCDFF3A" box="[971,1075,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.13331015" httpUri="https://zenodo.org/record/13331015/files/figure.png" pageId="31" pageNumber="66">Fig. 1 K</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum present and complete. Fore wing length
<quantity id="3872A5B7E91D4A53FAE3FF7FFF38FF72" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.9" metricValueMin="6.9" pageId="31" pageNumber="66" unit="mm" value="7.9" valueMax="8.9" valueMin="6.9">6.98.9 mm</quantity>
. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole shorter than areolet (
<figureCitation id="67B114D7E91D4A53FF34FEC7FEF3FE96" box="[159,269,259,285]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." figureDoi="http://doi.org/10.5281/zenodo.13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="31" pageNumber="66">Fig. 21 F</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.15.9 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.94.1: 2.0: 1.4: 0.7: 0.8. Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E91D4A53FEE2FEABFE51FE02" box="[329,431,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (OS)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); EG: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—AE, GN: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13331017" httpUri="https://zenodo.org/record/13331017/files/figure.png" pageId="31" pageNumber="66">Fig. 2 M</figureCitation>
).
</paragraph>
<caption id="ABF558DAE91D4A53FF3CF933FAFEF8B8" ID-DOI="http://doi.org/10.5281/zenodo.13331067" ID-Zenodo-Dep="13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="31" pageNumber="66" startId="31.[151,250,1783,1807]" targetBox="[151,1436,417,1759]" targetPageId="31" targetType="figure">
<paragraph id="FF350852E91D4A53FF3CF933FAFEF8B8" blockId="31.[151,1437,1783,1843]" pageId="31" pageNumber="66">
<emphasis id="CDFED440E91D4A53FF3CF933FEDEF884" bold="true" box="[151,288,1783,1807]" pageId="31" pageNumber="66">FIGURE 21.</emphasis>
<taxonomicName id="388A73D1E91D4A53FE8CF933FD6DF884" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[295,659,1783,1807]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="66" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91D4A53FE8CF933FDFCF884" box="[295,514,1783,1807]" italics="true" pageId="31" pageNumber="66">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91D4A53FDA3F933FD6DF884" author="Uchida, T." box="[520,659,1783,1807]" pageId="31" pageNumber="66" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings.
</paragraph>
</caption>
<paragraph id="FF350852E91D4A53FF6CF8A6FBE1F86C" blockId="31.[151,1436,1889,2024]" pageId="31" pageNumber="66">Metasoma. T I 1.91.95 × as long as maximum width, finely punctate except for smooth areas on median longitudinal area, area along posterior margin and lateral sides. T II 1.251.4 × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 3.353.6 × as long as hind tibia, ca. 1.4 × as long as fore wing, and as long as or slightly longer than body length.</paragraph>
<paragraph id="FF350852E9224A6CFF6CFF53FCABFE02" blockId="32.[151,1437,151,609]" pageId="32" pageNumber="67">
Colouration (
<figureCitation id="67B114D7E9224A6CFEF4FF53FE08FF3A" box="[351,502,151,177]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." figureDoi="http://doi.org/10.5281/zenodo.13331067" httpUri="https://zenodo.org/record/13331067/files/figure.png" pageId="32" pageNumber="67">Figs 21 AF</figureCitation>
). Body (excluding wings and legs) black. Clypeus yellow except for dorsal part, mandible except for base and teeth, ventral part of pedicel, and tegula yellow. Ventral part of scape and basal segments of flagellum tinged with brown to reddish-brown. Fore and mid legs yellow except for largely black coxae and partly darkened mid trochanter. Hind leg black to blackish-brown except for trochantellus, base and apex of femur, and basal part of tibia yellow to yellowish-brown. Membranous parts of metasomal sternites largely dark yellowish-brown. Ovipositor reddish-brown. Wings hyaline, without distinct apical clouded area of fore wing. Veins and pterostigma brown except for yellowish-brown wing base.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFE57FDD9FE26" blockId="32.[151,1437,151,609]" box="[199,551,403,429]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE57FEFAFE26" bold="true" box="[199,260,403,429]" pageId="32" pageNumber="67">Male</emphasis>
. Unknown (see remarks).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF6CFE72FE36FE7E" pageId="32" pageNumber="67" type="distribution">
<paragraph id="FF350852E9224A6CFF6CFE72FA95FE5A" blockId="32.[151,1437,151,609]" box="[199,1387,438,465]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE72FEA7FE5B" bold="true" box="[199,345,438,464]" pageId="32" pageNumber="67">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9224A6CFECCFE73FE59FE5A" box="[359,423,439,465]" name="Japan" pageId="32" pageNumber="67">Japan</collectingCountry>
(
<collectingRegion id="3D4EC6B0E9224A6CFE1DFE73FDD7FE5A" box="[438,553,439,465]" country="Japan" name="Hokkaido" pageId="32" pageNumber="67">Hokkaido</collectingRegion>
, Honshu, and Shikoku),
<collectingCountry id="879D48C2E9224A6CFC96FE73FC7EFE5A" box="[829,896,439,465]" name="China" pageId="32" pageNumber="67">China</collectingCountry>
and
<collectingCountry id="879D48C2E9224A6CFC1DFE73FC00FE5A" box="[950,1022,439,465]" name="South Korea" pageId="32" pageNumber="67">Korea</collectingCountry>
. New to Hokkaido and Shikoku.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFE1EFE36FE7E" blockId="32.[151,1437,151,609]" box="[199,456,474,501]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE1EFEBDFE7F" bold="true" box="[199,323,474,500]" pageId="32" pageNumber="67">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF6CFE3BFD1BFDEA" pageId="32" pageNumber="67" type="discussion">
<paragraph id="FF350852E9224A6CFF6CFE3BFD1BFDEA" blockId="32.[151,1437,151,609]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE3BFECAFD92" bold="true" box="[199,308,511,537]" pageId="32" pageNumber="67">Remarks</emphasis>
.
<bibRefCitation id="9B1B75A3E9224A6CFEEEFE3BFE2FFD92" author="Iwata, K." box="[325,465,510,537]" pageId="32" pageNumber="67" pagination="63 - 74" refId="ref23060" refString="Iwata, K. (1958) Ovarian eggs of 233 species of the Japanese Ichneumonidae (Hymenoptera). Acta Hymenopterologica, 1 (1), 63 - 74." type="journal article" year="1958">Iwata (1958</bibRefCitation>
,
<bibRefCitation id="9B1B75A3E9224A6CFE74FE3BFDE6FD92" author="Iwata, K." box="[479,536,511,537]" pageId="32" pageNumber="67" pagination="115 - 169" refId="ref23092" refString="Iwata, K. (1960) The comparative anatomy of the ovary in Hymenoptera. Part V. Ichneumonidae. Acta Hymenopterologica, 1 (2), 115 - 169." type="journal article" year="1960">1960</bibRefCitation>
) described the ovarian eggs and ovary.
<bibRefCitation id="9B1B75A3E9224A6CFC4DFE3BFB45FD92" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[998,1211,511,537]" pageId="32" pageNumber="67" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E9224A6CFB87FE3BFB99FD92" box="[1068,1127,511,537]" italics="true" pageId="32" pageNumber="67">et al.</emphasis>
(2020)
</bibRefCitation>
described the male of this species, but it may be misidentified because of the clouded apex of fore wing in his male in contrast to the entirely hyaline wing in the female of
<taxonomicName id="388A73D1E9224A6CFDEFFD83FD20FDEA" authorityName="Uchida" authorityYear="1928" box="[580,734,583,609]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="67" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E9224A6CFDEFFD83FD20FDEA" box="[580,734,583,609]" italics="true" pageId="32" pageNumber="67">S. takaozanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>