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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.857.29011" ID-GBIF-Dataset="2b50a048-8eb3-4ef4-b2ad-fb21cc83c29c" ID-PMC="PMC6606672" ID-Pensoft-Pub="1313-2970-857-85" ID-PubMed="31303806" ID-ZBK="8B50D262C5074697A93466F977031F1D" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-857-85" ModsDocOrigin="ZooKeys 857" ModsDocTitle="The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species" checkinTime="1561532585439" checkinUser="pensoft" docAuthor="Liu, Zhen &amp; Jiang, Shi-hong" docDate="2019" docId="5E03B6B0147AD02D8E0AB8298C4F56DA" docLanguage="en" docName="ZooKeys 857: 85-104" docOrigin="ZooKeys 857" docSource="http://dx.doi.org/10.3897/zookeys.857.29011" docTitle="Scutellathous Kishii 1955" docType="treatment" docVersion="5" lastPageNumber="85" masterDocId="43751E4C815BFFE5FFE8FF9AFFE2FFF2" masterDocTitle="The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species" masterLastPageNumber="104" masterPageNumber="85" pageNumber="85" updateTime="1668167437205" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Zhen</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Shi-hong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>857</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>85</mods:start>
<mods:end>104</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.857.29011</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.857.29011</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-857-85</mods:identifier>
<mods:identifier type="ZBK">8B50D262C5074697A93466F977031F1D</mods:identifier>
<mods:identifier type="ZooBank">8B50D262C5074697A93466F977031F1D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158387731" LSID="urn:lsid:plazi:treatment:5E03B6B0147AD02D8E0AB8298C4F56DA" httpUri="http://treatment.plazi.org/id/5E03B6B0147AD02D8E0AB8298C4F56DA" lastPageNumber="85" pageId="0" pageNumber="85">
<subSubSection pageId="0" pageNumber="85" type="nomenclature">
<paragraph pageId="0" pageNumber="85">
<taxonomicName authority="Kishii, 1955" authorityName="Kishii" authorityYear="1955" class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous Kishii, 1955</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="reference_group">
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
Kishii, 1955: 79 (
<typeStatus>type</typeStatus>
species:
<taxonomicName class="Insecta" family="Elateridae" genus="Athous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athous comes" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="comes">Athous comes</taxonomicName>
Lewis, 1894: 200 (Sapporo,
<collectingCountry name="Japan">Japan</collectingCountry>
; by original designation);
<bibRefCitation author="Ohira, H" journalOrPublisher="Nature and Insects" pageId="0" pageNumber="85" pagination="19 - 24" title="Elateridae in Japan (V)." volume="5" year="1970">
<normalizedToken originalValue="Ôhira">Ohira</normalizedToken>
1970
</bibRefCitation>
: 22;
<bibRefCitation author="Gurjeva, EL" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="85" pagination="96 - 113" title="Stroenie grudnogo otdela zhukov-shchelkunov (Coleoptera, Elateridae) I znachenie ego priznakov dlya sistemy semeistva [Thoracic structure of click beetles (Coleoptera, Elateridae) and the significance of the structural characters for the system of the family]." volume="53" year="1974">Gurjeva 1974</bibRefCitation>
: 108;
<bibRefCitation author="Kishii, T" journalOrPublisher="Kishii, Kyoto" pageId="0" pageNumber="85" title="A taxonomic study of the Japanese Elateridae (Coleoptera), with the keys to the subfamilies, tribes and genera. T." year="1987">Kishii 1987</bibRefCitation>
: 91;
<bibRefCitation author="Park, JS" journalOrPublisher="Report of the Korean Association for Conservation of Nature" pageId="0" pageNumber="85" pagination="158 - 213" title="Faunistic Study on the Insect from Hamyang-gun and Paemsagol area of Mt Chiri." volume="31" year="1993">Park et al. 1993</bibRefCitation>
: 179;
<bibRefCitation author="Suzuki, W" journalOrPublisher="Miscellaneous Reports of the Hiwa Museum for Natural History" pageId="0" pageNumber="85" pagination="1 - 348" title="Catalogue of the Family Elateridae (Coleoptera) of Taiwan." volume="38" year="1999">Suzuki 1999</bibRefCitation>
: 113;
<bibRefCitation author="Kishii, T" journalOrPublisher="Special publication of the Japan Coleopterological Society" pageId="0" pageNumber="85" pagination="205 - 214" title="Revision of the genus Scutellathous from Japan, with descriptions of two new species and taxonomic notes on its relatives (Elateridae, Coleoptera) &quot; Some new forms of Elateridae in Japan (XXXI) &quot;." volume="1" year="2001">Kishii 2001</bibRefCitation>
: 206;
<bibRefCitation author="Cate, PC" editor="Loebl, I" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="85" title="Vol. 4. Apollo Books, Stenstrup" url="https://doi.org/10.1163/9789004309142_003" year="2007">Cate et al. 2007</bibRefCitation>
: 172;
<bibRefCitation author="Han, T" journalOrPublisher="International Journal of Industrial Entomology" pageId="0" pageNumber="85" pagination="72 - 77" title="A taxonomic note on the genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae) in Korea." url="10.7852/ijie.2016.33.2.72" volume="33" year="2016">Han et al. 2016</bibRefCitation>
: 72.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="diagnosis">
<paragraph pageId="0" pageNumber="85">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="85">
Body length:
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11-16 mm</quantity>
; frons triangularly depressed behind supra antennal carina, which is strongly thickened and overhanging labrum and nasale, carina elevated above part of frons immediately posterior to it in dorsal view; supra-orbital groove broadly excavated; antennae serrate from 3rd antennomere; pronotum mostly longer than wide, pronotal disc with weak median depression; hind angles of pronotum unicarinate; sublateral incisions at posterior margin of pronotum small, or tooth-like; prosternal sutures not grooved anteriorly; apical end of tarsomeres 2 and 3 lobed beneath; aedeagus with simple and narrow parameres apices, without apico-lateral expansion at apex, penis gradually narrowed and acute at apex (after
<bibRefCitation author="Han, T" journalOrPublisher="International Journal of Industrial Entomology" pageId="0" pageNumber="85" pagination="72 - 77" title="A taxonomic note on the genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae) in Korea." url="10.7852/ijie.2016.33.2.72" volume="33" year="2016">Han et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="distribution">
<paragraph pageId="0" pageNumber="85">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="85">
<collectingCountry name="China">China</collectingCountry>
,
<collectingCountry name="Japan">Japan</collectingCountry>
, and
<collectingCountry name="South Korea">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="remarks">
<paragraph pageId="0" pageNumber="85">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="85">
Based on a study of the descriptions and photos of
<taxonomicName class="Insecta" family="Elateridae" genus="Athous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Athous</taxonomicName>
(
<taxonomicName family="Elateridae" lsidName="" pageId="0" pageNumber="85" rank="family">Elateridae</taxonomicName>
,
<taxonomicName lsidName="" pageId="0" pageNumber="85" rank="subfamily" subfamily="Dendrometrinae">Dendrometrinae</taxonomicName>
) species from North America, we found the
<taxonomicName class="Insecta" family="Elateridae" genus="Athous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athous cucullatus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="cucullatus">Athous cucullatus</taxonomicName>
(Say, 1825) species-group shares many characters with
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
(Becker, 1979). These include head flattened with triangular depression; frontal carina prominent, well elevated above labrum; eyes large; punctures on pronotum umbillicate; tarsomeres 2- and 3-lobed; and male genitalia with parameres lacking subapical lateral tooth and dorsal carina. However, lobes on tarsomeres 2 and 3 are smaller in
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
, the carinae on the pronotal hind angles are present and sharp in all
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
(absent or present in
<taxonomicName lsidName="A. cucullatus" pageId="0" pageNumber="85" rank="species" species="cucullatus">A. cucullatus</taxonomicName>
group), and base of pronotum with sublateral incisions near hind angles in all
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
(some without incisions in
<taxonomicName lsidName="A. cucullatus" pageId="0" pageNumber="85" rank="species" species="cucullatus">A. cucullatus</taxonomicName>
group). Further comparison of members of the North American
<taxonomicName class="Insecta" family="Elateridae" genus="Athous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athous cucullatus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="cucullatus">Athous cucullatus</taxonomicName>
species group to
<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
is needed.
</paragraph>
<subSection pageId="0" pageNumber="85" type="key to the male species of scutellathous kishii from china">
<paragraph pageId="0" pageNumber="85">
Key to the male species of
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
Kishii from
<collectingCountry name="China">China</collectingCountry>
</paragraph>
<paragraph pageId="0" pageNumber="85">
<table pageId="0" pageNumber="85">
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">7b2e9a</td>
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">4</td>
</tr>
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
8
<normalizedToken originalValue="ad">a-d</normalizedToken>
7f
</td>
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName lsidName="S. nanlingensis" pageId="0" pageNumber="85" rank="species" species="nanlingensis">S. nanlingensis</taxonomicName>
sp.
<date>
nov.
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
4
<normalizedToken originalValue="ac">a-c</normalizedToken>
11e2f11b
</td>
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">3</td>
</tr>
</date>
</td>
</tr>
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">9a10c</td>
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName lsidName="S. quadrata" pageId="0" pageNumber="85" rank="species" species="quadrata">S. quadrata</taxonomicName>
sp.
<date>
nov.
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">1a2e</td>
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName lsidName="S. habenularis" pageId="0" pageNumber="85" rank="species" species="habenularis">S. habenularis</taxonomicName>
sp. nov.
</td>
</tr>
</date>
</td>
</tr>
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName lsidName="S. spinosus" pageId="0" pageNumber="85" rank="species" species="spinosus">S. spinosus</taxonomicName>
Platia &amp; Schimmel, 2007
</td>
</tr>
<tr pageId="0" pageNumber="85">
<td colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName lsidName="S. yamashitai" pageId="0" pageNumber="85" rank="species" species="yamashitai">S. yamashitai</taxonomicName>
Arimoto, 1992
</td>
</tr>
</table>
</paragraph>
</subSection>
<subSection pageId="0" pageNumber="85" type="checklist of scutellathous from china">
<paragraph pageId="0" pageNumber="85">
Checklist of
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
from
<collectingCountry name="China">China</collectingCountry>
</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
sp. nov. [
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
)]
</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous nanlingensis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="nanlingensis">Scutellathous nanlingensis</taxonomicName>
sp. nov. [
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Guangdong">Guangdong</collectingRegion>
)]
</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous quadrata" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="quadrata">Scutellathous quadrata</taxonomicName>
sp. nov. [
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Zhejiang">Zhejiang</collectingRegion>
)]
</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous spinosus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="spinosus">Scutellathous spinosus</taxonomicName>
Platia &amp; Schimmel, 2007 [
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Taiwan)">Taiwan)</collectingRegion>
]
</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous yamashitai" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="yamashitai">Scutellathous yamashitai</taxonomicName>
Arimoto, 1992 [
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Taiwan)">Taiwan)</collectingRegion>
]
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>