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<document id="1A30EE69A397BEAA7F618B50943626EC" ID-DOI="10.11646/zootaxa.3995.1.16" ID-GBIF-Dataset="82bfdbc0-a51d-4076-a101-c43e886312c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="235985" ID-ZooBank="2640F866-AB2B-46F4-8498-5F7721092B71" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461277781053" checkinUser="plazi" docAuthor="Morales-Núñez, Andrés G. &amp; Heard, Richard W." docDate="2015" docId="5B2A87E8FFDF30605FD1F97DE50DF99E" docLanguage="en" docName="zt03995p188.pdf" docOrigin="Zootaxa 3995 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cacoheterotanais Morales-Núñez &amp; Heard, 2015, gen. n." docType="treatment" docVersion="10" lastPageNumber="172" masterDocId="A713FF90FFDE30635F46FFA5E105FFCE" masterDocTitle="Cacoheterotanais rogerbamberi, a new genus and species of leptocheliid Tanaidacea (Crustacea: Peracarida) from shelf-waters of the Gulf of Mexico" masterLastPageNumber="188" masterPageNumber="169" pageNumber="170" updateTime="1698613312762" updateUser="plazi">
<mods:mods id="D75DB7DEE2E16CEDE99A78F9EEE1F8F3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="720773613FF259D22BEE438DC23F5E24">
<mods:title id="F449BB6BA462C82A9BDC261B5C17F07C">Cacoheterotanais rogerbamberi, a new genus and species of leptocheliid Tanaidacea (Crustacea: Peracarida) from shelf-waters of the Gulf of Mexico</mods:title>
</mods:titleInfo>
<mods:name id="215D701AAAA5F7FBA3F751CBBDCD6FEB" type="personal">
<mods:role id="02728DBCA2DDBBBE520C10F72837835F">
<mods:roleTerm id="DD3C852F4BD0BA9C5CE6480E53C6736D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81EC037D5EC590F3800F75AF6574B0E4">Morales-Núñez, Andrés G.</mods:namePart>
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<mods:name id="2975A0536A3B3482CB0A41D7A1F531E2" type="personal">
<mods:role id="904E7D373CB97DF6E8D32202BA1BB036">
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<mods:namePart id="65BB422C2B784EA920A10C84E1CA26FE">Heard, Richard W.</mods:namePart>
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<mods:title id="F0D5A99A1A1FB8DFF98011E065263C24">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94C7140A319B11C963D211F008D1D2C5">
<mods:date id="442B4A2A8EB6C3768F7B42FE1684D29B">2015</mods:date>
<mods:detail id="8D52A037A09B8AB74359AECE06AC4D70" type="volume">
<mods:number id="33976C0A7EE47A2E515C1942CB150054">3995</mods:number>
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<mods:detail id="6FAB342D5BD7A66306A3FBD0F9DC5A1B" type="issue">
<mods:number id="7BDB35D18EE8C70E22DE459FE547588C">1</mods:number>
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<mods:classification id="B189D4BC670F47908CAF3431EE4AD4D7">journal article</mods:classification>
<mods:identifier id="ECE5424300F94A492166E28F0A5F96E1" type="DOI">10.11646/zootaxa.3995.1.16</mods:identifier>
<mods:identifier id="A220DC80C0E0474A9ED26AC512EC0186" type="GBIF-Dataset">82bfdbc0-a51d-4076-a101-c43e886312c6</mods:identifier>
<mods:identifier id="0F44B4CD4BDCFCA6F6B3D55DFBAE6563" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1034C93D8B172FC96C51AF052DE5882A" type="Zenodo-Dep">235985</mods:identifier>
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</mods:mods>
<treatment id="5B2A87E8FFDF30605FD1F97DE50DF99E" ID-DOI="http://doi.org/10.5281/zenodo.5670391" ID-GBIF-Taxon="119639905" ID-Zenodo-Dep="5670391" LSID="urn:lsid:plazi:treatment:5B2A87E8FFDF30605FD1F97DE50DF99E" httpUri="http://treatment.plazi.org/id/5B2A87E8FFDF30605FD1F97DE50DF99E" lastPageId="3" lastPageNumber="172" pageId="1" pageNumber="170">
<subSubSection id="9B996575FFDF30625FD1F97DE5ACF8F9" pageId="1" pageNumber="170" type="nomenclature">
<paragraph id="D33C36FEFFDF30625FD1F97DE0C1F93C" blockId="1.[151,452,1752,1778]" box="[151,452,1752,1778]" pageId="1" pageNumber="170">
<heading id="88748192FFDF30625FD1F97DE0C1F93C" bold="true" box="[151,452,1752,1778]" fontSize="11" level="1" pageId="1" pageNumber="170" reason="1">
<emphasis id="E1F7EAECFFDF30625FD1F97DE0C1F93C" bold="true" box="[151,452,1752,1778]" pageId="1" pageNumber="170">
<taxonomicName id="14834D7DFFDF30625FD1F97DE06FF93C" ID-CoL="84MS6" box="[151,362,1752,1778]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E1F7EAECFFDF30625FD1F97DE06FF93C" bold="true" box="[151,362,1752,1778]" italics="true" pageId="1" pageNumber="170">Cacoheterotanais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAC45797FFDF30625E37F97DE0C1F93C" box="[369,452,1752,1778]" pageId="1" pageNumber="170" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D33C36FEFFDF30625FD1F8BBE5ACF8F9" blockId="1.[151,1436,1822,2027]" box="[151,1193,1822,1847]" pageId="1" pageNumber="170">
<emphasis id="E1F7EAECFFDF30625FD1F8BBE01EF8F9" bold="true" box="[151,283,1822,1847]" pageId="1" pageNumber="170">Synonymy.</emphasis>
<emphasis id="E1F7EAECFFDF30625E67F8BAE0FFF8F9" box="[289,506,1823,1847]" italics="true" pageId="1" pageNumber="170">
“
<taxonomicName id="14834D7DFFDF30625E6FF8BAE0FFF8F9" box="[297,506,1823,1847]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">Pseudonototanais</taxonomicName>
</emphasis>
sp. B”:
<bibRefCitation id="B7124B0FFFDF30625D10F8BBE22BF8F9" box="[598,814,1822,1847]" pageId="1" pageNumber="170" refString="Heard, R. W., Hansknecht, T. &amp; Larsen K. (2004) An illustrated identification guide to Florida Tanaidacea (Crustacea: Peracarida) occurring in depths of less than 200 m. Florida Department of Environmental Protection. Division of Water Resource Management, Tallahassee, Florida, 163 pp. Available from: http: // publicfiles. dep. state. fl. us / dear / labs / biology / biokeys / tanaidacea. pdf (Accessed 3 Aug. 2015)" type="book">
<collectingCountry id="AB94766EFFDF30625D10F8BBE39EF8F9" box="[598,667,1822,1847]" name="Heard Island and McDonald Islands" pageId="1" pageNumber="170">Heard</collectingCountry>
<emphasis id="E1F7EAECFFDF30625DE7F885E3D4F8F9" box="[673,721,1823,1847]" italics="true" pageId="1" pageNumber="170">et al</emphasis>
. (2004)
</bibRefCitation>
; Morales-Núñez &amp;
<collectingCountry id="AB94766EFFDF30625B4DF8BBE555F8F9" box="[1035,1104,1822,1847]" name="Heard Island and McDonald Islands" pageId="1" pageNumber="170">Heard</collectingCountry>
(2013).
</paragraph>
</subSubSection>
<subSubSection id="9B996575FFDF30605F81F8E7E422FED3" lastPageId="3" lastPageNumber="172" pageId="1" pageNumber="170" type="diagnosis">
<paragraph id="D33C36FEFFDF30615F81F8E7E1DDFD1B" blockId="1.[151,1436,1822,2027]" lastBlockId="2.[119,1877,590,775]" lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170">
<emphasis id="E1F7EAECFFDF30625F81F8E7E044F895" bold="true" box="[199,321,1858,1883]" pageId="1" pageNumber="170">Diagnosis.</emphasis>
<emphasis id="E1F7EAECFFDF30625E0CF8E6E086F895" box="[330,387,1859,1883]" italics="true" pageId="1" pageNumber="170">Male</emphasis>
:
<emphasis id="E1F7EAECFFDF30625ED3F8E6E302F895" box="[405,519,1859,1883]" italics="true" pageId="1" pageNumber="170">Antennule</emphasis>
with peduncle having three articles; flagellum with three articles, article-1 with proximal and distal cluster of aesthetascs; article-2 with distal cluster aesthetascs; article-3 minute, terminating with four simple setae of varying length.
<emphasis id="E1F7EAECFFDF30625D2FF82EE3C3F86C" box="[617,710,1931,1954]" italics="true" pageId="1" pageNumber="170">Antenna</emphasis>
with one mid-dorsal and two distal (one dorsal and one ventral) setiform spines on the second article.
<emphasis id="E1F7EAECFFDF30625D10F80AE280F809" box="[598,901,1967,1991]" italics="true" pageId="1" pageNumber="170">Buccal mass (mouthparts)</emphasis>
obsolete.
<emphasis id="E1F7EAECFFDF30625CB9F80AE57DF809" box="[1023,1144,1967,1991]" italics="true" pageId="1" pageNumber="170">Maxilliped</emphasis>
reduced, appearing nonfunctional.
<emphasis id="E1F7EAECFFDF30625E53F876E07FF825" box="[277,378,2003,2027]" italics="true" pageId="1" pageNumber="170">Cheliped</emphasis>
forcipate with carpus lateral face having length greater than depth; propodus with length about
<emphasis id="E1F7EAECFFDC30615F31FD3BE1DDFD1B" bold="true" pageId="2" pageNumber="171">
Character
<emphasis id="E1F7EAECFFDC30615E25FD3BE2DBFD7C" bold="true" box="[355,990,670,690]" italics="true" pageId="2" pageNumber="171">
<taxonomicName id="14834D7DFFDC30615E25FD3BE0CDFD7C" box="[355,456,670,690]" class="Malacostraca" family="Leptocheliidae" genus="Cocotanais" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Cocotanais</taxonomicName>
<taxonomicName id="14834D7DFFDC30615D1AFD3BE2DBFD7C" authority="Makassaritanai" authorityName="Makassaritanai" box="[604,990,670,690]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Heterotanais Makassaritanai</taxonomicName>
</emphasis>
s
<emphasis id="E1F7EAECFFDC30615BC8FD3BE616FD7C" bold="true" box="[1166,1811,670,690]" italics="true" pageId="2" pageNumber="171">
<taxonomicName id="14834D7DFFDC30615BC8FD3BE5C8FD7C" box="[1166,1229,670,690]" class="Malacostraca" family="Leptocheliidae" genus="Ogleus" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Ogleus</taxonomicName>
<taxonomicName id="14834D7DFFDC30615A3AFD3BE71EFD7C" box="[1404,1563,670,690]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Pseudonototanais</taxonomicName>
<taxonomicName id="14834D7DFFDC30615930FD3BE616FD7C" box="[1654,1811,670,690]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus" status="gen. nov.">Cacoheterotanais</taxonomicName>
</emphasis>
<taxonomicNameLabel id="FAC45797FFDC3061585FFD3BE650FD7F" box="[1817,1877,670,689]" pageId="2" pageNumber="171" rank="genus">gen. n.</taxonomicNameLabel>
Antennule
</emphasis>
</paragraph>
<caption id="87FC6676FFDC30615F31FDEAE1BAFD59" ID-Table-UUID="87FC6676FFDC30615F31FDEAE1BAFD59" httpUri="http://table.plazi.org/id/87FC6676FFDC30615F31FDEAE1BAFD59" pageId="2" pageNumber="171" targetBox="[119,1887,832,1621]" targetIsTable="true" targetPageId="2">
<paragraph id="D33C36FEFFDC30615F31FDEAE1BAFD59" blockId="2.[119,1877,590,775]" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615F31FDEAE5E4FDAD" bold="true" box="[119,1249,590,611]" pageId="2" pageNumber="171">
TABLE 1. Some characters for separating the extant genera within the
<taxonomicName id="14834D7DFFDC30615C74FDEBE2DFFDAC" box="[818,986,590,610]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDC30615C74FDEBE2DFFDAC" bold="true" box="[818,986,590,610]" italics="true" pageId="2" pageNumber="171">Pseudonototanais</emphasis>
</taxonomicName>
complex and
<taxonomicName id="14834D7DFFDC30615B27FDEBE5DEFDAC" box="[1121,1243,590,610]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDC30615B27FDEBE5DEFDAC" bold="true" box="[1121,1243,590,610]" italics="true" pageId="2" pageNumber="171">Heterotanais</emphasis>
</taxonomicName>
.
</emphasis>
Sieg (1973) indicates both simple setae and aesthetascs present on terminal article-2. Confirmation of the presence or absence of a minute third terminal article bearing onlysimple setae is needed.
</paragraph>
</caption>
<paragraph id="D33C36FEFFDC30615FFFFD7FE00DFD20" blockId="2.[119,1877,590,775]" box="[185,264,730,750]" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615FFFFD7FE00DFD20" box="[185,264,730,750]" italics="true" pageId="2" pageNumber="171">peduncle</emphasis>
</paragraph>
<paragraph id="D33C36FEFFDC30605E64FD51E422FED3" blockId="2.[119,1877,590,775]" lastBlockId="3.[151,1437,151,1616]" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
3 articles 3 articles 3 articles 3 articles 2 articles 3 articles twice depth, ventro-distal margin with pair of subacute processes (one disto-laterally and other disto-medially) to seat long, slightly curved dactylus when closed; fixed finger of propodus directed downward, nearly at right angle to posterior (ventral) margin of propodus, with low tooth or apophysis sub-distally on incisive margin.
<emphasis id="E1F7EAECFFDD30605A76FF45E499FF36" box="[1328,1436,224,248]" italics="true" pageId="3" pageNumber="172">Uropodal</emphasis>
endopod with three subequal articles; exopod uni-articulate, small not reaching past endopodal article-1.
</paragraph>
</subSubSection>
<paragraph id="D33C36FEFFDC30615FF0FCE5E7E6F99B" pageId="2" pageNumber="171">
<table id="A183C45EFFDCCF9C5F31FCE5E65AF99B" box="[119,1887,832,1621]" gridcols="7" gridrows="6" pageId="2" pageNumber="171">
<tr id="6DB334BCFFDCCF9C5F31FCE5E65AFC06" box="[119,1887,832,968]" gridrow="0" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31FCE5E00DFC06" box="[119,264,832,968]" gridcol="0" gridrow="0" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615FF0FCE5E00DFC9A" box="[182,264,832,852]" italics="true" pageId="2" pageNumber="171">flagellum</emphasis>
</th>
<th id="2E625DC0FFDCCF9C5E64FCE5E30DFC06" box="[290,520,832,968]" gridcol="1" gridrow="0" pageId="2" pageNumber="171">5 articles; aesthetascs present on articles 1 to 4; article-1 elongate with proximal and distal cluster of aesthetascs</th>
<th id="2E625DC0FFDCCF9C5D65FCE5E20CFC06" box="[547,777,832,968]" gridcol="2" gridrow="0" pageId="2" pageNumber="171">2 articles*; aesthetascs present on articles1 to 2; article-1 elongate with only distal cluster.</th>
<th id="2E625DC0FFDCCF9C5C62FCE5E517FC06" box="[804,1042,832,968]" gridcol="3" gridrow="0" pageId="2" pageNumber="171">4 to 5 articles; aesthetascs present on articles 1 to 4; article-1 elongate with proximal and distal cluster.</th>
<th id="2E625DC0FFDCCF9C5B68FCE5E42DFC06" box="[1070,1320,832,968]" gridcol="4" gridrow="0" pageId="2" pageNumber="171">6 articles; aesthetascs present on articles 1 to 5; article-1 elongate with only distal cluster.</th>
<th id="2E625DC0FFDCCF9C5A01FCE5E74AFC06" box="[1351,1615,832,968]" gridcol="5" gridrow="0" pageId="2" pageNumber="171">3 articles; aesthetascs present on articles 1 to 3; article-1 elongate with proximal and distal cluster.</th>
<th id="2E625DC0FFDCCF9C592CFCE5E65AFC06" box="[1642,1887,832,968]" gridcol="6" gridrow="0" pageId="2" pageNumber="171">3 articles; aesthetascs present on articles 1 to 2; article-1 elongate with proximal and distal cluster.</th>
</tr>
<tr id="6DB334BCFFDCCF9C5F31FC4FE65AFB80" box="[119,1887,1002,1102]" gridrow="1" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31FC4FE00DFB80" box="[119,264,1002,1102]" gridcol="0" gridrow="1" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615F31FC4FE1C2FC33" bold="true" box="[119,199,1002,1021]" pageId="2" pageNumber="171">Antenna</emphasis>
<emphasis id="E1F7EAECFFDC30615FFBFBA7E00DFBD8" box="[189,264,1026,1046]" italics="true" pageId="2" pageNumber="171">article-2</emphasis>
</th>
<td id="2E625DC0FFDCCF9C5E64FC4FE30DFB80" box="[290,520,1002,1102]" gridcol="1" gridrow="1" pageId="2" pageNumber="171">with 2 distal spines (dorsally and ventrally)</td>
<td id="2E625DC0FFDCCF9C5D65FC4FE20CFB80" box="[547,777,1002,1102]" gridcol="2" gridrow="1" pageId="2" pageNumber="171">with 2 distal spines (dorsally and ventrally)</td>
<td id="2E625DC0FFDCCF9C5C62FC4FE517FB80" box="[804,1042,1002,1102]" gridcol="3" gridrow="1" pageId="2" pageNumber="171">with 2 distal spines (dorsally and ventrally)</td>
<td id="2E625DC0FFDCCF9C5B68FC4FE42DFB80" box="[1070,1320,1002,1102]" gridcol="4" gridrow="1" pageId="2" pageNumber="171">with 2 distal spines (dorsally and ventrally)</td>
<td id="2E625DC0FFDCCF9C5A01FC4FE74AFB80" box="[1351,1615,1002,1102]" gridcol="5" gridrow="1" pageId="2" pageNumber="171">with 3 spines (2 distal and one mid-dorsal)</td>
<td id="2E625DC0FFDCCF9C592CFC4FE65AFB80" box="[1642,1887,1002,1102]" gridcol="6" gridrow="1" pageId="2" pageNumber="171">with 3 spines (2 distal and one mid-dorsal)</td>
</tr>
<tr id="6DB334BCFFDCCF9C5F31FBD5E65AFB3F" box="[119,1887,1136,1265]" gridrow="2" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31FBD5E00DFB3F" box="[119,264,1136,1265]" gridcol="0" gridrow="2" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615F31FBD5E1CFFB4D" bold="true" box="[119,202,1136,1155]" pageId="2" pageNumber="171">Cheliped</emphasis>
<emphasis id="E1F7EAECFFDC30615F95FB00E00DFB79" box="[211,264,1189,1207]" italics="true" pageId="2" pageNumber="171">merus</emphasis>
</th>
<td id="2E625DC0FFDCCF9C5E64FBD5E30DFB3F" box="[290,520,1136,1265]" gridcol="1" gridrow="2" pageId="2" pageNumber="171">Not extending posterodistally under carpus to margin of propodus.</td>
<td id="2E625DC0FFDCCF9C5D65FBD5E20CFB3F" box="[547,777,1136,1265]" gridcol="2" gridrow="2" pageId="2" pageNumber="171">Not extending posterodistally under carpus to margin of propodus.</td>
<td id="2E625DC0FFDCCF9C5C62FBD5E517FB3F" box="[804,1042,1136,1265]" gridcol="3" gridrow="2" pageId="2" pageNumber="171">Not extending posterodistally under carpus to margin of propodus.</td>
<td id="2E625DC0FFDCCF9C5B68FBD5E42DFB3F" box="[1070,1320,1136,1265]" gridcol="4" gridrow="2" pageId="2" pageNumber="171">Extending posterodistally under carpus to or very near margin of propodus.</td>
<td id="2E625DC0FFDCCF9C5A01FBD5E74AFB3F" box="[1351,1615,1136,1265]" gridcol="5" gridrow="2" pageId="2" pageNumber="171">Extending posterodistally under carpus to or very near margin of propodus</td>
<td id="2E625DC0FFDCCF9C592CFBD5E65AFB3F" box="[1642,1887,1136,1265]" gridcol="6" gridrow="2" pageId="2" pageNumber="171">Not extending posterodistally under carpus to margin of propodus.</td>
</tr>
<tr id="6DB334BCFFDCCF9C5F31FAB7E65AFA5A" box="[119,1887,1298,1428]" gridrow="3" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31FAB7E00DFA5A" box="[119,264,1298,1428]" gridcol="0" gridrow="3" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615F8BFAB6E00DFAEB" box="[205,264,1299,1317]" italics="true" pageId="2" pageNumber="171">carpus</emphasis>
</th>
<td id="2E625DC0FFDCCF9C5E64FAB7E30DFA5A" box="[290,520,1298,1428]" gridcol="1" gridrow="3" pageId="2" pageNumber="171">width of lateral face&gt; two thirds of length, with carpal flanges</td>
<td id="2E625DC0FFDCCF9C5D65FAB7E20CFA5A" box="[547,777,1298,1428]" gridcol="2" gridrow="3" pageId="2" pageNumber="171">width of lateral face&gt; two thirds of length, with carpal flanges</td>
<td id="2E625DC0FFDCCF9C5C62FAB7E517FA5A" box="[804,1042,1298,1428]" gridcol="3" gridrow="3" pageId="2" pageNumber="171">width of lateral face &lt;two thirds of length, without carpal flanges</td>
<td id="2E625DC0FFDCCF9C5B68FAB7E42DFA5A" box="[1070,1320,1298,1428]" gridcol="4" gridrow="3" pageId="2" pageNumber="171">width of lateral face&gt; two thirds of length, with carpal flanges</td>
<td id="2E625DC0FFDCCF9C5A01FAB7E74AFA5A" box="[1351,1615,1298,1428]" gridcol="5" gridrow="3" pageId="2" pageNumber="171">width of lateral face&gt; two thirds of length, with carpal flanges</td>
<td id="2E625DC0FFDCCF9C592CFAB7E65AFA5A" box="[1642,1887,1298,1428]" gridcol="6" gridrow="3" pageId="2" pageNumber="171">width of lateral face&gt; two thirds of length, with carpal flanges</td>
</tr>
<tr id="6DB334BCFFDCCF9C5F31FA10E65AF9CD" box="[119,1887,1461,1539]" gridrow="4" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31FA10E00DF9CD" box="[119,264,1461,1539]" gridcol="0" gridrow="4" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615FF5FA10E00DFA28" box="[179,264,1461,1510]" italics="true" pageId="2" pageNumber="171">Propodus (chela)</emphasis>
</th>
<td id="2E625DC0FFDCCF9C5E64FA10E30DF9CD" box="[290,520,1461,1539]" gridcol="1" gridrow="4" pageId="2" pageNumber="171">without wide ovoid gap between dactylus and tip of fixed finger</td>
<td id="2E625DC0FFDCCF9C5D65FA10E20CF9CD" box="[547,777,1461,1539]" gridcol="2" gridrow="4" pageId="2" pageNumber="171">without wide ovoid gap</td>
<td id="2E625DC0FFDCCF9C5C62FA10E517F9CD" box="[804,1042,1461,1539]" gridcol="3" gridrow="4" pageId="2" pageNumber="171">with wide ovoid gap</td>
<td id="2E625DC0FFDCCF9C5B68FA10E42DF9CD" box="[1070,1320,1461,1539]" gridcol="4" gridrow="4" pageId="2" pageNumber="171">without wide ovoid gap</td>
<td id="2E625DC0FFDCCF9C5A01FA10E74AF9CD" box="[1351,1615,1461,1539]" gridcol="5" gridrow="4" pageId="2" pageNumber="171">without wide ovoid gap</td>
<td id="2E625DC0FFDCCF9C592CFA10E65AF9CD" box="[1642,1887,1461,1539]" gridcol="6" gridrow="4" pageId="2" pageNumber="171">without wide ovoid gap</td>
</tr>
<tr id="6DB334BCFFDCCF9C5F31F986E65AF99B" box="[119,1887,1571,1621]" gridrow="5" pageId="2" pageNumber="171">
<th id="2E625DC0FFDCCF9C5F31F986E00DF99B" box="[119,264,1571,1621]" gridcol="0" gridrow="5" pageId="2" pageNumber="171">
<emphasis id="E1F7EAECFFDC30615FE2F981E00DF9F9" box="[164,264,1572,1592]" italics="true" pageId="2" pageNumber="171">fixed finger</emphasis>
</th>
<td id="2E625DC0FFDCCF9C5E64F986E30DF99B" box="[290,520,1571,1621]" gridcol="1" gridrow="5" pageId="2" pageNumber="171">styliform</td>
<td id="2E625DC0FFDCCF9C5D65F986E20CF99B" box="[547,777,1571,1621]" gridcol="2" gridrow="5" pageId="2" pageNumber="171">club shaped</td>
<td id="2E625DC0FFDCCF9C5C62F986E517F99B" box="[804,1042,1571,1621]" gridcol="3" gridrow="5" pageId="2" pageNumber="171">subdistal apophysis on grasping edge</td>
<td id="2E625DC0FFDCCF9C5B68F986E42DF99B" box="[1070,1320,1571,1621]" gridcol="4" gridrow="5" pageId="2" pageNumber="171">subdistal, dentate, apophysis on grasping edge</td>
<td id="2E625DC0FFDCCF9C5A01F986E74AF99B" box="[1351,1615,1571,1621]" gridcol="5" gridrow="5" pageId="2" pageNumber="171">styliform</td>
<td id="2E625DC0FFDCCF9C592CF986E65AF99B" box="[1642,1887,1571,1621]" gridcol="6" gridrow="5" pageId="2" pageNumber="171">subdistalapophysison grasping edge</td>
</tr>
</table>
</paragraph>
<subSubSection id="9B996575FFDD30605F81FE8DE0A9FEAB" pageId="3" pageNumber="172" type="description">
<paragraph id="D33C36FEFFDD30605F81FE8DE0A9FEAB" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FE8DE0A3FE8E" box="[199,422,296,320]" italics="true" pageId="3" pageNumber="172">Female: Maxilliped</emphasis>
with inner plate having three pairs of long simple setae distally.
<emphasis id="E1F7EAECFFDD30605BC2FE8DE5F5FE8E" box="[1156,1264,296,320]" italics="true" pageId="3" pageNumber="172">Uropodal</emphasis>
endopod, as in male, with three articles.
</paragraph>
</subSubSection>
<subSubSection id="9B996575FFDD30605F81FECAE212FE46" box="[199,791,367,392]" pageId="3" pageNumber="172" type="materials_examined">
<paragraph id="D33C36FEFFDD30605F81FECAE212FE46" blockId="3.[151,1437,151,1616]" box="[199,791,367,392]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FECAE067FE46" bold="true" box="[199,354,367,392]" pageId="3" pageNumber="172">
<typeStatus id="0C38885CFFDD30605F81FECAE007FE46" box="[199,258,367,392]" pageId="3" pageNumber="172">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="14834D7DFFDD30605E2FFED5E3D4FE46" box="[361,721,368,392]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="rogerbamberi" status="sp. nov.">
<emphasis id="E1F7EAECFFDD30605E2FFED5E3D4FE46" box="[361,721,368,392]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais rogerbamberi</emphasis>
</taxonomicName>
<emphasis id="E1F7EAECFFDD30605D91FECAE212FE46" bold="true" box="[727,791,367,392]" pageId="3" pageNumber="172">
<taxonomicNameLabel id="FAC45797FFDD30605D91FECAE212FE46" box="[727,791,367,392]" pageId="3" pageNumber="172" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9B996575FFDD30605F81FE31E43FFE3A" pageId="3" pageNumber="172" type="etymology">
<paragraph id="D33C36FEFFDD30605F81FE31E53EFE62" blockId="3.[151,1437,151,1616]" box="[199,1083,404,429]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FE31E043FE63" bold="true" box="[199,326,404,429]" pageId="3" pageNumber="172">Etymology</emphasis>
. From the Greek
<emphasis id="E1F7EAECFFDD30605D4AFE31E357FE65" box="[524,594,404,427]" italics="true" pageId="3" pageNumber="172">Cacos</emphasis>
= bad, plus the generic name
<emphasis id="E1F7EAECFFDD30605CE4FE30E53EFE62" box="[930,1083,405,428]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605CE4FE30E532FE62" box="[930,1079,405,428]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Heterotanais</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="D33C36FEFFDD30605F81FE12E0ACFE1E" blockId="3.[151,1437,151,1616]" box="[199,425,439,464]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FE12E023FE1E" bold="true" box="[199,294,439,464]" pageId="3" pageNumber="172">Gender.</emphasis>
Masculine.
</paragraph>
<paragraph id="D33C36FEFFDD30605F81FE79E43FFE3A" blockId="3.[151,1437,151,1616]" box="[199,1338,476,501]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FE79E06CFE3B" bold="true" box="[199,361,476,501]" pageId="3" pageNumber="172">Known range</emphasis>
. North
<collectingCountry id="AB94766EFFDD30605E86FE79E326FE3B" box="[448,547,476,501]" name="United States of America" pageId="3" pageNumber="172">America</collectingCountry>
, Gulf of
<collectingCountry id="AB94766EFFDD30605DC1FE79E3DAFE3B" box="[647,735,476,501]" name="Mexico" pageId="3" pageNumber="172">Mexico</collectingCountry>
, Eastern Continental shelf off the Florida West Coast.
</paragraph>
</subSubSection>
<subSubSection id="9B996575FFDD30605F81FE5AE50DF99E" pageId="3" pageNumber="172" type="discussion">
<paragraph id="D33C36FEFFDD30605F81FE5AE2E6FD4B" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="172">
<emphasis id="E1F7EAECFFDD30605F81FE5AE03EFDD6" bold="true" box="[199,315,511,536]" pageId="3" pageNumber="172">Remarks.</emphasis>
Systematically, the genera within the
<taxonomicName id="14834D7DFFDD30605DA5FDA5E2A9FDD6" box="[739,940,512,536]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605DA5FDA5E2A9FDD6" box="[739,940,512,536]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
complex (i.e.
<taxonomicName id="14834D7DFFDD30605B0AFDA5E5CEFDD9" box="[1100,1227,512,535]" class="Malacostraca" family="Leptocheliidae" genus="Cocotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605B0AFDA5E5CEFDD9" box="[1100,1227,512,535]" italics="true" pageId="3" pageNumber="172">Cocotanais</emphasis>
</taxonomicName>
;
<taxonomicName id="14834D7DFFDD30605B9FFDA5E491FDD6" box="[1241,1428,512,536]" class="Malacostraca" family="Leptocheliidae" genus="Makassaritanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605B9FFDA5E491FDD6" box="[1241,1428,512,536]" italics="true" pageId="3" pageNumber="172">Makassaritanais</emphasis>
</taxonomicName>
;
<taxonomicName id="14834D7DFFDD30605FD1FD81E099FDF3" authority="Morales-Nunez" authorityName="Morales-Nunez" box="[151,412,548,573]" class="Malacostraca" family="Leptocheliidae" genus="Ogleus" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605FD1FD81E1E3FDF2" box="[151,230,548,572]" italics="true" pageId="3" pageNumber="172">Ogleus</emphasis>
Morales-Núñez
</taxonomicName>
&amp;
<collectingCountry id="AB94766EFFDD30605E85FD81E30FFDF3" box="[451,522,548,573]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="172">Heard</collectingCountry>
, 2013;
<taxonomicName id="14834D7DFFDD30605D19FD81E22DFDF2" box="[607,808,548,572]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605D19FD81E22DFDF2" box="[607,808,548,572]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
and
<taxonomicName id="14834D7DFFDD30605C26FD80E2F4FDF2" box="[864,1009,549,572]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605C26FD80E2F4FDF2" box="[864,1009,549,572]" italics="true" pageId="3" pageNumber="172">Heterotanais</emphasis>
</taxonomicName>
) appear to represent a highly derived group displaying a mixture of characters (see
<tableCitation id="9E010345FFDD30605DAAFDE2E24CFDAE" box="[748,841,583,608]" captionStart="TABLE 1" captionStartId="2.[119,196,591,611]" captionTargetBox="[119,1887,832,1621]" captionTargetPageId="2" captionText="TABLE 1. Some characters for separating the extant genera within the Pseudonototanais complex and Heterotanais. Sieg (1973) indicates both simple setae and aesthetascs present on terminal article- 2. Confirmation of the presence or absence of a minute third terminal article bearing onlysimple setae is needed." httpUri="http://table.plazi.org/id/87FC6676FFDC30615F31FDEAE1BAFD59" pageId="3" pageNumber="172" tableUuid="87FC6676FFDC30615F31FDEAE1BAFD59">Table 1</tableCitation>
) confounded by the probability of homoplasy.
<taxonomicName id="14834D7DFFDD30605FD1FDC9E058FD4A" box="[151,349,620,644]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605FD1FDC9E058FD4A" box="[151,349,620,644]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
shares a unique combination of some of these characters.
</paragraph>
<paragraph id="D33C36FEFFDD30605F81FD2AE381FB26" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="172">
The males of
<taxonomicName id="14834D7DFFDD30605E22FD35E32FFD66" box="[356,554,656,680]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605E22FD35E32FFD66" box="[356,554,656,680]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
and
<taxonomicName id="14834D7DFFDD30605D22FD35E228FD66" box="[612,813,656,680]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605D22FD35E228FD66" box="[612,813,656,680]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
have an antennule with three flagella articles, uropods with three endopodal articles, and article-2 of the antenna with three setiform spines, which appear to distinguish them the other male members of the genus-group. The male of the
<taxonomicName id="14834D7DFFDD30605C9AFD7DE5A7FD3E" box="[988,1186,728,752]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605C9AFD7DE5A7FD3E" box="[988,1186,728,752]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="14834D7DFFDD30605FD1FD59E065FCDA" box="[151,352,764,788]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605FD1FD59E065FCDA" box="[151,352,764,788]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
by having three instead of two apparent antennular peduncle articles (
<figureCitation id="4BB82A7BFFDD30605B33FD59E5CCFCDA" box="[1141,1225,764,789]" captionStart="FIGURE 10" captionStartId="15.[151,250,1877,1899]" captionTargetBox="[183,1410,402,1752]" captionTargetId="figure@15.[168,1428,375,1861]" captionTargetPageId="15" captionText="FIGURE 10. Male antennules. A, Pseudonototanais werthi; B, Ogleus pilarae; C, Heterotanais oerstedii; D, Makassaritanais angustus; E, Cocotanais puravida; F, M. bamberi; G, Cacoheterotanais rogerbamberi gen. et sp. n. Male antennas. H, M. bamberi; I, Cacoheterotanais rogerbamberi gen. et sp. n. Figures modified from: A, Shiino 1978; B, Morales-Núñez &amp; Heard 2013; C, Krøyer 1842; D, F, and H, Guţu 2012; E, Esquete et al. 2013; G and I, Morales-Núñez &amp; Heard (this study). Not to scale." httpUri="https://zenodo.org/record/235991/files/figure.png" pageId="3" pageNumber="172">Figs 10</figureCitation>
G, A, respectively) and a chela with a distinctly different fixed finger. The male of
<emphasis id="E1F7EAECFFDD30605CDDFC85E56DFCF6" box="[923,1128,800,824]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605CDDFC85E561FCF6" box="[923,1124,800,824]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Cacoheterotanais</taxonomicName>
,
</emphasis>
like those of
<emphasis id="E1F7EAECFFDD30605A53FC85E05DFC92" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605A53FC85E492FCF9" box="[1301,1431,800,823]" class="Malacostraca" family="Leptocheliidae" genus="Cocotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Cocotanais</taxonomicName>
,
<taxonomicName id="14834D7DFFDD30605FD1FCE1E051FC92" box="[151,340,836,860]" class="Malacostraca" family="Leptocheliidae" genus="Makassaritanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Makassaritanais</taxonomicName>
,
</emphasis>
and
<emphasis id="E1F7EAECFFDD30605EDFFCE1E36DFC92" box="[409,616,836,860]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605EDFFCE1E360FC92" box="[409,613,836,860]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Pseudonototanais</taxonomicName>
,
</emphasis>
has an antennule with the first apparent flagella article bearing both proximal and distal clusters of aesthetascs (
<figureCitation id="4BB82A7BFFDD30605DD6FCC2E3E2FC4E" box="[656,743,871,896]" captionStart="FIGURE 10" captionStartId="15.[151,250,1877,1899]" captionTargetBox="[183,1410,402,1752]" captionTargetId="figure@15.[168,1428,375,1861]" captionTargetPageId="15" captionText="FIGURE 10. Male antennules. A, Pseudonototanais werthi; B, Ogleus pilarae; C, Heterotanais oerstedii; D, Makassaritanais angustus; E, Cocotanais puravida; F, M. bamberi; G, Cacoheterotanais rogerbamberi gen. et sp. n. Male antennas. H, M. bamberi; I, Cacoheterotanais rogerbamberi gen. et sp. n. Figures modified from: A, Shiino 1978; B, Morales-Núñez &amp; Heard 2013; C, Krøyer 1842; D, F, and H, Guţu 2012; E, Esquete et al. 2013; G and I, Morales-Núñez &amp; Heard (this study). Not to scale." httpUri="https://zenodo.org/record/235991/files/figure.png" pageId="3" pageNumber="172">Figs 10</figureCitation>
G, E, D–F, A, respectively). In contrast, only a single distal cluster of aesthetascs is present on first flagella article-1for the males within the genera
<taxonomicName id="14834D7DFFDD30605BC4FC28E416FC6A" box="[1154,1299,909,932]" class="Malacostraca" family="Leptocheliidae" genus="Heterotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605BC4FC28E416FC6A" box="[1154,1299,909,932]" italics="true" pageId="3" pageNumber="172">Heterotanais</emphasis>
</taxonomicName>
and
<taxonomicName id="14834D7DFFDD30605A0AFC29E49EFC6A" box="[1356,1435,908,932]" class="Malacostraca" family="Leptocheliidae" genus="Ogleus" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605A0AFC29E49EFC6A" box="[1356,1435,908,932]" italics="true" pageId="3" pageNumber="172">Ogleus</emphasis>
</taxonomicName>
(
<figureCitation id="4BB82A7BFFDD30605FD9FC0AE1FDFC06" box="[159,248,943,968]" captionStart="FIGURE 10" captionStartId="15.[151,250,1877,1899]" captionTargetBox="[183,1410,402,1752]" captionTargetId="figure@15.[168,1428,375,1861]" captionTargetPageId="15" captionText="FIGURE 10. Male antennules. A, Pseudonototanais werthi; B, Ogleus pilarae; C, Heterotanais oerstedii; D, Makassaritanais angustus; E, Cocotanais puravida; F, M. bamberi; G, Cacoheterotanais rogerbamberi gen. et sp. n. Male antennas. H, M. bamberi; I, Cacoheterotanais rogerbamberi gen. et sp. n. Figures modified from: A, Shiino 1978; B, Morales-Núñez &amp; Heard 2013; C, Krøyer 1842; D, F, and H, Guţu 2012; E, Esquete et al. 2013; G and I, Morales-Núñez &amp; Heard (this study). Not to scale." httpUri="https://zenodo.org/record/235991/files/figure.png" pageId="3" pageNumber="172">Figs 10</figureCitation>
C, B, respectively). The chela for the male of
<taxonomicName id="14834D7DFFDD30605C51FC15E2D8FC06" box="[791,989,944,968]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605C51FC15E2D8FC06" box="[791,989,944,968]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
appears to be transitional between
<emphasis id="E1F7EAECFFDD30605A3BFC14E1FEFC22" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605A3BFC14E1F2FC22" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="bamberi">M. bamberi</taxonomicName>
,
</emphasis>
and those of the genera
<taxonomicName id="14834D7DFFDD30605D61FC71E3A2FC25" box="[551,679,980,1003]" class="Malacostraca" family="Leptocheliidae" genus="Cocotanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605D61FC71E3A2FC25" box="[551,679,980,1003]" italics="true" pageId="3" pageNumber="172">Cocotanais</emphasis>
</taxonomicName>
and
<emphasis id="E1F7EAECFFDD30605DA3FC71E2B3FC22" box="[741,950,980,1004]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605DA3FC71E2B4FC22" box="[741,945,980,1004]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Pseudonototanais</taxonomicName>
.
</emphasis>
The latter two genera lack the sub-distal apophysis or low blunt tooth on the subdistal grasping margin of the fixed finger (
<figureCitation id="4BB82A7BFFDD30605B01FC52E599FBDE" box="[1095,1180,1015,1040]" captionStart="FIGURE 11" captionStartId="16.[151,250,1685,1707]" captionTargetBox="[193,1421,216,1575]" captionTargetId="figure@16.[151,1436,193,1663]" captionTargetPageId="16" captionText="FIGURE 11. Chelipeds. A, Pseudonototanais werthi; B, Ogleus pilarae; C, Heterotanais oerstedii; D, Makassaritanais angustus; E, Cocotanais puravida; F, M. bamberi; G, Cacoheterotanais rogerbamberi gen. et sp. n. Uropods. H, M. bamberi; I," httpUri="https://zenodo.org/record/235992/files/figure.png" pageId="3" pageNumber="172">Figs 11</figureCitation>
E, C, A, respectively). This structure is present in
<emphasis id="E1F7EAECFFDD30605E82FBB9E3E4FBFA" box="[452,737,1052,1076]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605E82FBB9E384FBFA" box="[452,641,1052,1076]" class="Malacostraca" family="Leptocheliidae" genus="Makassaritanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Makassaritanais</taxonomicName>
,
<taxonomicName id="14834D7DFFDD30605DCAFBB9E3DBFBFA" box="[652,734,1052,1076]" class="Malacostraca" family="Leptocheliidae" genus="Ogleus" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">Ogleus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="14834D7DFFDD30605C51FBB9E2D8FBFA" box="[791,989,1052,1076]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605C51FBB9E2D8FBFA" box="[791,989,1052,1076]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
(
<figureCitation id="4BB82A7BFFDD30605CADFBB9E567FBFA" box="[1003,1122,1052,1077]" captionStart="FIGURE 11" captionStartId="16.[151,250,1685,1707]" captionTargetBox="[193,1421,216,1575]" captionTargetId="figure@16.[151,1436,193,1663]" captionTargetPageId="16" captionText="FIGURE 11. Chelipeds. A, Pseudonototanais werthi; B, Ogleus pilarae; C, Heterotanais oerstedii; D, Makassaritanais angustus; E, Cocotanais puravida; F, M. bamberi; G, Cacoheterotanais rogerbamberi gen. et sp. n. Uropods. H, M. bamberi; I," httpUri="https://zenodo.org/record/235992/files/figure.png" pageId="3" pageNumber="172">Figures 11</figureCitation>
D, F, B, G, respectively) and maybe homologous with the “teeth” occurring on the fixed finger of males within genus
<emphasis id="E1F7EAECFFDD30605BD3FBE5E499FB96" box="[1173,1436,1088,1112]" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605BD3FBE5E465FB96" box="[1173,1376,1088,1112]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="sensu">
Leptochelia
<taxonomicNameLabel id="FAC45797FFDD30605A64FBE4E465FB96" box="[1314,1376,1089,1112]" pageId="3" pageNumber="172" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato.
</emphasis>
<tableCitation id="9E010345FFDD30605FD1FBC1E1F2FBB2" box="[151,247,1124,1149]" captionStart="TABLE 1" captionStartId="2.[119,196,591,611]" captionTargetBox="[119,1887,832,1621]" captionTargetPageId="2" captionText="TABLE 1. Some characters for separating the extant genera within the Pseudonototanais complex and Heterotanais. Sieg (1973) indicates both simple setae and aesthetascs present on terminal article- 2. Confirmation of the presence or absence of a minute third terminal article bearing onlysimple setae is needed." httpUri="http://table.plazi.org/id/87FC6676FFDC30615F31FDEAE1BAFD59" pageId="3" pageNumber="172" tableUuid="87FC6676FFDC30615F31FDEAE1BAFD59">Table 1</tableCitation>
presents a synopsis of these and additional characters for separating the genera within the
<taxonomicName id="14834D7DFFDD30605FD1FB2DE065FB6E" box="[151,352,1160,1184]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605FD1FB2DE065FB6E" box="[151,352,1160,1184]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
complex. The female of
<taxonomicName id="14834D7DFFDD30605D3BFB2DE246FB6E" box="[637,835,1160,1184]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605D3BFB2DE246FB6E" box="[637,835,1160,1184]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
differs from those of other genera of the complex by having its uropodal endopod with three articles of similar length; those for the other genera have from 4–5 articles or incipient articles often of variable length.
</paragraph>
<paragraph id="D33C36FEFFDD30605F81FB51E50DF99E" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="172">
<taxonomicName id="14834D7DFFDD30605F81FB51E088FAC2" box="[199,397,1268,1292]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605F81FB51E088FAC2" box="[199,397,1268,1292]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
and the other members of the
<taxonomicName id="14834D7DFFDD30605DABFB51E2B3FAC2" box="[749,950,1268,1292]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605DABFB51E2B3FAC2" box="[749,950,1268,1292]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
complex appear to have more affinities of the
<taxonomicName id="14834D7DFFDD30605F87FAB2E062FAFE" box="[193,359,1303,1328]" class="Malacostraca" family="Leptocheliidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="subFamily" subFamily="Leptocheliinae">Leptocheliinae</taxonomicName>
than the other currently recognized subfamilies as defined by
<bibRefCitation id="B7124B0FFFDD30605B6DFAB2E5DFFAFE" author="Bamber" box="[1067,1242,1303,1328]" pageId="3" pageNumber="172" refString="Bamber, R. N. (2013) A re-assessment of Konarus Bamber, 2006 and sympatric leptocheliids from Australasia, and of Pseudoleptochelia Lang, 1973 (Crustacea: Peracarida: Tanaidacea). Zootaxa, 3694 (1), 1 - 39. http: // dx. doi. org / 10.11646 / zootaxa. 3694.1.1" type="journal article" year="2013">Bamber (2013)</bibRefCitation>
. There, however, still appears to be confusion in the systematics of this genus-group (see Morales &amp;
<collectingCountry id="AB94766EFFDD30605B22FA99E5ACFA9B" box="[1124,1193,1340,1365]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="172">Heard</collectingCountry>
2013). Based on the variable morphological features exhibited within it, the generic status of many of the species is uncertain; however, we reluctantly decided to establish the genus
<taxonomicName id="14834D7DFFDD30605DDFFA21E25AFA52" box="[665,863,1412,1436]" class="Malacostraca" family="Leptocheliidae" genus="Cacoheterotanais" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605DDFFA21E25AFA52" box="[665,863,1412,1436]" italics="true" pageId="3" pageNumber="172">Cacoheterotanais</emphasis>
</taxonomicName>
to accommodate the new species from the Gulf of
<collectingCountry id="AB94766EFFDD30605FD1FA02E1F5FA0E" box="[151,240,1447,1472]" name="Mexico" pageId="3" pageNumber="172">Mexico</collectingCountry>
. The designation of a new genus within this group of six mostly monotypic genera is based on sound morphological differences, but at this time we are unable to determine whether or not they are systematically or phylogenetically definitive. Nevertheless, we follow Morales-Núñez &amp;
<collectingCountry id="AB94766EFFDD30605CADFA4AE535F9C6" box="[1003,1072,1519,1544]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="172">Heard</collectingCountry>
(2013) in believing that until molecular studies can be conducted, the status of genera within the “
<taxonomicName id="14834D7DFFDD30605CD5F9B1E559F9E2" box="[915,1116,1556,1580]" class="Malacostraca" family="Leptocheliidae" genus="Pseudonototanais" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="E1F7EAECFFDD30605CD5F9B1E559F9E2" box="[915,1116,1556,1580]" italics="true" pageId="3" pageNumber="172">Pseudonototanais</emphasis>
</taxonomicName>
complex” (as well as for the
<taxonomicName id="14834D7DFFDD30605FD1F992E038F99E" box="[151,317,1591,1616]" class="Malacostraca" family="Leptocheliidae" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="172" phylum="Arthropoda" rank="family">Leptocheliidae</taxonomicName>
as a whole), cannot be resolved with a high degree of certainty.
</paragraph>
</subSubSection>
</treatment>
</document>
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