410 lines
50 KiB
XML
410 lines
50 KiB
XML
<document id="16D8F5AF41A77FD95E40F00CEAF9D363" ID-DOI="10.5281/zenodo.273232" ID-GBIF-Dataset="1d0652a6-d4b0-48a0-83af-d6283dc9bbaf" ID-ISSN="11755326" ID-Zenodo-Dep="273232" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459720815360" checkinUser="plazi" docAuthor="Dimitrov, Dimitar & Ribera, Carles" docDate="2005" docId="081B87BF756EFFE1FEC0FEF4FE7DFC37" docLanguage="en" docName="zt00982.pdf" docOrigin="Zootaxa 982" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ossinissa Dimitrov & Ribera, 2005, new genus" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="F422FFC7756CFFE6FFC8FFE8FFA6FFE9" masterDocTitle="Description of Ossinissa, a new pholcid genus from the Canary Islands (Araneae: Pholcidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1698209309543" updateUser="plazi">
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<mods:titleInfo id="FD602B1D069C88F881B48D293A4A7F39">
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<mods:title id="844DDD2E454C497394B4904EF978267E">Description of Ossinissa, a new pholcid genus from the Canary Islands (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="212D2A494CDA4B0A8D3B1999862698BA">Dimitrov, Dimitar</mods:namePart>
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<mods:namePart id="38E8E2CEA2F8D808FB26491C790DCFC1">Ribera, Carles</mods:namePart>
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<mods:typeOfResource id="548C27B42DD46E73A76A8357DC339B02">text</mods:typeOfResource>
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<mods:title id="1031A9584B71441793CF319B02B16C34">Zootaxa</mods:title>
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<mods:part id="BC601CA41D7C798C801B670EFD21B807">
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<mods:date id="0E477CFB38B42BDEEFAB57ADAA9FAB3E">2005</mods:date>
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<mods:identifier id="2FE8294F41FA60FFAA42A6B6F71A80F7" type="DOI">10.5281/zenodo.273232</mods:identifier>
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<treatment id="081B87BF756EFFE1FEC0FEF4FE7DFC37" ID-DOI="http://doi.org/10.5281/zenodo.6265562" ID-GBIF-Taxon="119335580" ID-Zenodo-Dep="6265562" LSID="urn:lsid:plazi:treatment:081B87BF756EFFE1FEC0FEF4FE7DFC37" httpUri="http://treatment.plazi.org/id/081B87BF756EFFE1FEC0FEF4FE7DFC37" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
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<subSubSection id="C8A86522756EFFE4FEC0FEF4FDA4FEDC" box="[264,514,283,310]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="800D36A9756EFFE4FEC0FEF4FDA4FEDC" blockId="2.[264,514,283,310]" box="[264,514,283,310]" pageId="2" pageNumber="3">
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<heading id="DB4581C5756EFFE4FEC0FEF4FDA4FEDC" bold="true" box="[264,514,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="B2C6EABB756EFFE4FEC0FEF4FDA4FEDC" bold="true" box="[264,514,283,310]" pageId="2" pageNumber="3">
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<taxonomicName id="47B24D2A756EFFE4FEC0FEF4FEDCFEDF" ID-CoL="63GRC" box="[264,378,284,310]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B2C6EABB756EFFE4FEC0FEF4FEDCFEDF" bold="true" box="[264,378,284,310]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A9F557C0756EFFE4FE4AFEF3FDA4FEDC" box="[386,514,283,309]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A86522756EFFE4FEC0FE90FC14FE7B" box="[264,946,376,402]" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="800D36A9756EFFE4FEC0FE90FC14FE7B" blockId="2.[264,1323,376,1885]" box="[264,946,376,402]" pageId="2" pageNumber="3">
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||
<emphasis id="B2C6EABB756EFFE4FEC0FE90FE68FE7B" bold="true" box="[264,462,376,402]" pageId="2" pageNumber="3">
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<typeStatus id="5F09880B756EFFE4FEC0FE90FEE3FE7B" box="[264,325,376,402]" pageId="2" pageNumber="3">Type</typeStatus>
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species. —
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</emphasis>
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<taxonomicName id="47B24D2A756EFFE4FE1EFE90FC14FE7B" authority="Wunderlich 1992" authorityName="Wunderlich" authorityYear="1992" box="[470,946,376,402]" class="Arachnida" family="Pholcidae" genus="Spermophorides" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="justoi">
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<emphasis id="B2C6EABB756EFFE4FE1EFE90FD79FE78" box="[470,735,376,401]" italics="true" pageId="2" pageNumber="3">Spermophorides justoi</emphasis>
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<bibRefCitation id="E4234B58756EFFE4FD2EFE90FC14FE7B" author="Wunderlich" box="[742,946,376,402]" pageId="2" pageNumber="11" refString="Wunderlich, J. (1992) Die Spinnenfauna der Makaronesischen Inseln. Beitrage zur Araneologie, 1, 619 pp." type="book" year="1992">Wunderlich 1992</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A86522756EFFE4FEF0FE48FD9DFDE4" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="800D36A9756EFFE4FEF0FE48FD9DFDE4" blockId="2.[264,1323,376,1885]" pageId="2" pageNumber="3">
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<emphasis id="B2C6EABB756EFFE4FEF0FE48FE4DFE53" bold="true" box="[312,491,416,442]" pageId="2" pageNumber="3">Etymology. —</emphasis>
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The generic name honors
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<taxonomicName id="47B24D2A756EFFE4FCE5FE48FC39FE53" box="[813,927,416,442]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Ossinissa</taxonomicName>
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, the last king of El Hierro Island before its conquest by the Norman Jean de Bethencourt at the beginning of the 15th century. Gender is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A86522756EFFE4FEF0FDF2FE61FAEC" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="800D36A9756EFFE4FEF0FDF2FDDBFBD4" blockId="2.[264,1323,376,1885]" pageId="2" pageNumber="3">
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<emphasis id="B2C6EABB756EFFE4FEF0FDF2FE7BFDDD" bold="true" box="[312,477,538,564]" pageId="2" pageNumber="3">Diagnosis. —</emphasis>
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The opisthosoma is cylindrical in males and oval in females (
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<figureCitation id="18892A2C756EFFE4FB0CFDF3FABBFDDC" box="[1220,1309,539,565]" captionStart="FIGURES 25 – 29" captionStartId="5.[264,382,760,784]" captionTargetBox="[264,1323,289,732]" captionTargetId="figure@5.[264,1323,289,733]" captionTargetPageId="5" captionText="FIGURES 25 – 29. Ossinissa justoi new combination, female. 25. Opisthosoma, lateral; 26. Anterior lateral spinnerets, ventral; 27. Ocular area, dorsal; 28. Spinnerets, ventral; 29. Posterior median spinnerets, ventral. Scale bars: 25: 500 µm; 26: 20 µm; 27: 100 µm; 29: 10 µm; 28: 50 µm." httpUri="https://zenodo.org/record/273234/files/figure.png" pageId="2" pageNumber="3">Fig. 25</figureCitation>
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). Prosoma and habitus as in Figs. 21–22. Ocular area elevated and placed close to the center of the prosoma. Distinguished from closely related genera (
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<emphasis id="B2C6EABB756EFFE4FC09FD83FEC5FD45" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="47B24D2A756EFFE4FC09FD83FB84FD6D" box="[961,1058,619,644]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Pholcus</taxonomicName>
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,
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<taxonomicName id="47B24D2A756EFFE4FBE6FD83FB75FD6D" box="[1070,1235,619,644]" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Micropholcus</taxonomicName>
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,
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<taxonomicName id="47B24D2A756EFFE4FB16FD83FEC5FD45" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Leptopholcus</taxonomicName>
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</emphasis>
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and
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<taxonomicName id="47B24D2A756EFFE4FE57FD7BFD98FD45" box="[415,574,659,684]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B2C6EABB756EFFE4FE57FD7BFD98FD45" box="[415,574,659,684]" italics="true" pageId="2" pageNumber="3">Spermophora</emphasis>
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</taxonomicName>
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) by the shape of the prosoma (Fig. 21) and the morphology of the procursus with characteristic dorsoretrolateral apophysis and concave shape. A useful character for identification purposes is the shape of the tarsal organ (
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<figureCitation id="18892A2C756EFFE4FBE6FD0BFB0BFD14" box="[1070,1197,739,765]" captionStart="FIGURES 1 – 13" captionStartId="3.[264,383,1707,1731]" captionTargetBox="[264,1322,393,1680]" captionTargetId="figure@3.[264,1323,393,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 13. Ossinissa justoi new combination, male. 1. Uncus, retrolateral; 2. Apex of the procursus, retrolateral; 3. Chelicerae, frontal; 4. Procursus, retrolateral; 5. Uncus, dorsoretrolateral; 6. Tarsal organ, lateral; 7. Palp, dorsoretrolateral; 8. Palp with procursus, dorsal; 9. Distal apophyses of the chelicerae, frontolateral; 10. Ocular area, frontal; 11. Procursus, ventral; 12. Male palp, prolateral; 13. Tarsal organ, frontolateral. Scale bars: 1 – 5, 10 – 11: 100 µm; 7 – 8, 12: 200 µm; 4: 50 µm; 13: 40 µm; 6: 20 µm; 9: 10 µm." httpUri="https://zenodo.org/record/273233/files/figure.png" pageId="2" pageNumber="3">Figs. 6, 13</figureCitation>
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), which in
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<taxonomicName id="47B24D2A756EFFE4FEC0FCE3FEDEFCCD" box="[264,376,779,804]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B2C6EABB756EFFE4FEC0FCE3FEDEFCCD" box="[264,376,779,804]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A9F557C0756EFFE4FEB7FCE3FE5DFCCC" box="[383,507,779,805]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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is flat (sensu
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<bibRefCitation id="E4234B58756EFFE4FD56FCE3FC8AFCCC" author="Huber" box="[670,812,779,805]" pageId="2" pageNumber="11" refString="Huber, B. A. (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History, 254, 1 - 348." type="journal article" year="2000">Huber 2000</bibRefCitation>
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,
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<bibRefCitation id="E4234B58756EFFE4FCFFFCE3FCD0FCCC" author="Huber" box="[823,886,779,805]" pageId="2" pageNumber="11" refString="Huber, B. A. (2003) High species diversity in one of the dominant groups of spiders in East African montane forests (Araneae: Pholcidae: Buitinga n. gen., Spermophora Hentz). Zoological Journal of the Linnean Society, 137, 555 - 619." type="journal article" year="2003">2003</bibRefCitation>
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), while in all closely related genera it is capsulated. Although the small size and the globular shape of the female opisthosoma may lead to some confusion with
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<taxonomicName id="47B24D2A756EFFE4FD53FCB3FC9DFC9D" box="[667,827,859,884]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B2C6EABB756EFFE4FD53FCB3FC9DFC9D" box="[667,827,859,884]" italics="true" pageId="2" pageNumber="3">Spermophora</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="47B24D2A756EFFE4FCACFCB3FB82FC9D" box="[868,1060,859,884]" class="Arachnida" family="Pholcidae" genus="Spermophorides" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B2C6EABB756EFFE4FCACFCB3FB82FC9D" box="[868,1060,859,884]" italics="true" pageId="2" pageNumber="3">Spermophorides</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="47B24D2A756EFFE4FBFCFCB3FB02FC9D" box="[1076,1188,859,884]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B2C6EABB756EFFE4FBFCFCB3FB02FC9D" box="[1076,1188,859,884]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A9F557C0756EFFE4FB65FCB3FA8DFC9C" box="[1197,1323,859,885]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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can be easily distinguished by the presence of a welldeveloped uncus (
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<figureCitation id="18892A2C756EFFE4FB9AFC6BFB65FC74" box="[1106,1219,899,925]" captionStart="FIGURES 1 – 13" captionStartId="3.[264,383,1707,1731]" captionTargetBox="[264,1322,393,1680]" captionTargetId="figure@3.[264,1323,393,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 13. Ossinissa justoi new combination, male. 1. Uncus, retrolateral; 2. Apex of the procursus, retrolateral; 3. Chelicerae, frontal; 4. Procursus, retrolateral; 5. Uncus, dorsoretrolateral; 6. Tarsal organ, lateral; 7. Palp, dorsoretrolateral; 8. Palp with procursus, dorsal; 9. Distal apophyses of the chelicerae, frontolateral; 10. Ocular area, frontal; 11. Procursus, ventral; 12. Male palp, prolateral; 13. Tarsal organ, frontolateral. Scale bars: 1 – 5, 10 – 11: 100 µm; 7 – 8, 12: 200 µm; 4: 50 µm; 13: 40 µm; 6: 20 µm; 9: 10 µm." httpUri="https://zenodo.org/record/273233/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 5</figureCitation>
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, 14, 20), the structure of the procursus (Fig. 24) and the presence of eight eyes. Additionally, the absence of the “bump” of the epigynum in
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<taxonomicName id="47B24D2A756EFFE4FCECFC3BFC32FC05" box="[804,916,979,1004]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B2C6EABB756EFFE4FCECFC3BFC32FC05" box="[804,916,979,1004]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A9F557C0756EFFE4FC69FC3BFB87FC04" box="[929,1057,979,1005]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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clearly separates this genus from
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<taxonomicName id="47B24D2A756EFFE4FE5CFC13FDF2FBFD" box="[404,596,1019,1044]" class="Arachnida" family="Pholcidae" genus="Spermophorides" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B2C6EABB756EFFE4FE5CFC13FDF2FBFD" box="[404,596,1019,1044]" italics="true" pageId="2" pageNumber="3">Spermophorides</emphasis>
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</taxonomicName>
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, and the lack of pockets in the vulva is a useful trait when differentiating from
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<taxonomicName id="47B24D2A756EFFE4FE1FFBCBFDD1FBD5" box="[471,631,1059,1084]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B2C6EABB756EFFE4FE1FFBCBFDD1FBD5" box="[471,631,1059,1084]" italics="true" pageId="2" pageNumber="3">Spermophora</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="800D36A9756EFFE4FEF0FBA3FE61FAEC" blockId="2.[264,1323,376,1885]" pageId="2" pageNumber="3">
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The most important characters that distinguish
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<taxonomicName id="47B24D2A756EFFE4FCADFBA3FC73FB8D" box="[869,981,1099,1124]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B2C6EABB756EFFE4FCADFBA3FC73FB8D" box="[869,981,1099,1124]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A9F557C0756EFFE4FC16FBA3FBFAFB8C" box="[990,1116,1099,1125]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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from smallsized
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<taxonomicName id="47B24D2A756EFFE4FEC0FB9BFEC0FB65" box="[264,358,1139,1164]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B2C6EABB756EFFE4FEC0FB9BFEC0FB65" box="[264,358,1139,1164]" italics="true" pageId="2" pageNumber="3">Pholcus</emphasis>
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</taxonomicName>
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species are: the cavity of the procursus; the dorsoretrolateral apophysis of the procursus, and the shape of the tarsal organ. The absence of a heavily sclerotized structure around the pore plates in the female genitalia of
|
||
<taxonomicName id="47B24D2A756EFFE4FC8EFB2BFC10FB35" box="[838,950,1219,1244]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756EFFE4FC8EFB2BFC10FB35" box="[838,950,1219,1244]" italics="true" pageId="2" pageNumber="3">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756EFFE4FC76FB2BFB9DFB34" box="[958,1083,1219,1245]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
|
||
is also a good diagnostic character.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8A86522756EFFE1FEF0FAFAFC51FCFF" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="800D36A9756EFFE5FEF0FAFAFDD1FEB7" blockId="2.[264,1323,376,1885]" lastBlockId="3.[264,1322,284,350]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="B2C6EABB756EFFE4FEF0FAFAFE69FAC5" bold="true" box="[312,463,1298,1324]" pageId="2" pageNumber="3">Description.</emphasis>
|
||
— Ochreyellowish pholcids. Total body length in males slightly smaller than in females. Prosoma shape as in Fig. 21. The ocular area moderately elevated and situated roughly in the center of the prosoma. Eight eyes. In the
|
||
<typeStatus id="5F09880B756EFFE4FBD6FA8BFBF7FA94" box="[1054,1105,1379,1405]" pageId="2" pageNumber="3">type</typeStatus>
|
||
species, eyes are reduced, especially the AME. The arrangements of the eyes as in most eighteyed pholcids; ALE, PME and PLE grouped in two triads placed on the lateral margins of the ocular area. Lateral triads are placed on two short cylindrical outgrowths. AME situated between the triads on the frontal side of the ocular area. Ocular area surrounded by a slight depression of the prosoma. Male chelicerae (
|
||
<figureCitation id="18892A2C756EFFE4FD07F9C3FC85F9AC" box="[719,803,1579,1605]" captionStart="FIGURES 1 – 13" captionStartId="3.[264,383,1707,1731]" captionTargetBox="[264,1322,393,1680]" captionTargetId="figure@3.[264,1323,393,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 13. Ossinissa justoi new combination, male. 1. Uncus, retrolateral; 2. Apex of the procursus, retrolateral; 3. Chelicerae, frontal; 4. Procursus, retrolateral; 5. Uncus, dorsoretrolateral; 6. Tarsal organ, lateral; 7. Palp, dorsoretrolateral; 8. Palp with procursus, dorsal; 9. Distal apophyses of the chelicerae, frontolateral; 10. Ocular area, frontal; 11. Procursus, ventral; 12. Male palp, prolateral; 13. Tarsal organ, frontolateral. Scale bars: 1 – 5, 10 – 11: 100 µm; 7 – 8, 12: 200 µm; 4: 50 µm; 13: 40 µm; 6: 20 µm; 9: 10 µm." httpUri="https://zenodo.org/record/273233/files/figure.png" pageId="2" pageNumber="3">Figs. 3</figureCitation>
|
||
, 17) with distal apophyses (
|
||
<figureCitation id="18892A2C756EFFE4FBB8F9C3FB11F9AC" box="[1136,1207,1579,1605]" captionStart="FIGURES 1 – 13" captionStartId="3.[264,383,1707,1731]" captionTargetBox="[264,1322,393,1680]" captionTargetId="figure@3.[264,1323,393,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 13. Ossinissa justoi new combination, male. 1. Uncus, retrolateral; 2. Apex of the procursus, retrolateral; 3. Chelicerae, frontal; 4. Procursus, retrolateral; 5. Uncus, dorsoretrolateral; 6. Tarsal organ, lateral; 7. Palp, dorsoretrolateral; 8. Palp with procursus, dorsal; 9. Distal apophyses of the chelicerae, frontolateral; 10. Ocular area, frontal; 11. Procursus, ventral; 12. Male palp, prolateral; 13. Tarsal organ, frontolateral. Scale bars: 1 – 5, 10 – 11: 100 µm; 7 – 8, 12: 200 µm; 4: 50 µm; 13: 40 µm; 6: 20 µm; 9: 10 µm." httpUri="https://zenodo.org/record/273233/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
|
||
) carrying modified hairs (sensu
|
||
<bibRefCitation id="E4234B58756EFFE4FDDAF9BBFD06F984" author="Huber" box="[530,672,1619,1645]" pageId="2" pageNumber="11" refString="Huber, B. A. (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History, 254, 1 - 348." type="journal article" year="2000">Huber 2000</bibRefCitation>
|
||
) and proximolateral apophyses. In females, chelicerae bear no apophyses. Male palpal trochanter with retrolateral process, femur cylindrical with ventral bulge. Procursus robust and narrower at the base (Fig. 24), with a cavity open toward the bulb. Distally, procursus with well developed dorsoretrolateral apophysis. Genital bulb with three projections: uncus (Fig. 20), appendix and embolus. Embolus with membranous structure. Leg formula 1423. Legs and body covered with short hairs. Opisthosoma almost cylindrical in males and globular in females. Spinnerets (
|
||
<figureCitation id="18892A2C756EFFE4FB5EF8ABFA84F8B4" box="[1174,1314,1859,1885]" captionStart="FIGURES 25 – 29" captionStartId="5.[264,382,760,784]" captionTargetBox="[264,1323,289,732]" captionTargetId="figure@5.[264,1323,289,733]" captionTargetPageId="5" captionText="FIGURES 25 – 29. Ossinissa justoi new combination, female. 25. Opisthosoma, lateral; 26. Anterior lateral spinnerets, ventral; 27. Ocular area, dorsal; 28. Spinnerets, ventral; 29. Posterior median spinnerets, ventral. Scale bars: 25: 500 µm; 26: 20 µm; 27: 100 µm; 29: 10 µm; 28: 50 µm." httpUri="https://zenodo.org/record/273234/files/figure.png" pageId="2" pageNumber="3">Figs. 26–29</figureCitation>
|
||
) situated ventrally close to the end of the opisthosoma. Epigynum elevated. Vulva with heavily sclerotized valve ridge.
|
||
</paragraph>
|
||
<caption id="D4CD6621756FFFE5FEC0F943FD2AF899" httpUri="https://zenodo.org/record/273233/files/figure.png" pageId="3" pageNumber="4" targetBox="[264,1322,393,1680]" targetPageId="3">
|
||
<paragraph id="800D36A9756FFFE5FEC0F943FD2AF899" blockId="3.[264,1323,1707,1904]" pageId="3" pageNumber="4">
|
||
<emphasis id="B2C6EABB756FFFE5FEC0F943FE62F92A" bold="true" box="[264,452,1707,1731]" pageId="3" pageNumber="4">FIGURES 1–13.</emphasis>
|
||
<taxonomicName id="47B24D2A756FFFE5FE06F943FDDEF92B" box="[462,632,1707,1730]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="justoi" status="comb. nov.">
|
||
<emphasis id="B2C6EABB756FFFE5FE06F943FDDEF92B" box="[462,632,1707,1730]" italics="true" pageId="3" pageNumber="4">Ossinissa justoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756FFFE5FD4AF943FCE6F92B" box="[642,832,1707,1730]" pageId="3" pageNumber="4">new combination</taxonomicNameLabel>
|
||
, male. 1. Uncus, retrolateral; 2. Apex of the procursus, retrolateral; 3. Chelicerae, frontal; 4. Procursus, retrolateral; 5. Uncus, dorsoretrolateral; 6. Tarsal organ, lateral; 7. Palp, dorsoretrolateral; 8. Palp with procursus, dorsal; 9. Distal apophyses of the chelicerae, frontolateral; 10. Ocular area, frontal; 11. Procursus, ventral; 12. Male palp, prolateral; 13. Tarsal organ, frontolateral. Scale bars: 1–5, 10–11: 100 µm; 7–8, 12: 200 µm; 4: 50 µm; 13: 40 µm; 6: 20 µm; 9: 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="800D36A97568FFE2FEC0F954FE3DF8B7" blockId="4.[264,1323,1724,1886]" pageId="4" pageNumber="5">
|
||
<emphasis id="B2C6EABB7568FFE2FEC0F954FE68F93D" bold="true" box="[264,462,1724,1748]" pageId="4" pageNumber="5">FIGURES 14–24.</emphasis>
|
||
<taxonomicName id="47B24D2A7568FFE2FE1DF955FDDAF93D" box="[469,636,1725,1748]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="justoi" status="comb. nov.">
|
||
<emphasis id="B2C6EABB7568FFE2FE1DF955FDDAF93D" box="[469,636,1725,1748]" italics="true" pageId="4" pageNumber="5">Ossinissa justoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C07568FFE2FD4BF954FC98F93A" box="[643,830,1724,1747]" pageId="4" pageNumber="5">new combination</taxonomicNameLabel>
|
||
, male (14–15, 17, 20, 22, 24), female (16, 18– 19, 21, 23). 14. Male palp, retrolateral; 15. Male palp, prolateral; 16. Vulva, dorsal; 17. Male chelicerae, frontal; 18. Female prosoma, lateral; 19. Epigynum, ventral; 20. Uncus, retrolateral; 21. Female body, lateral; 22. Male body, dorsal; 23. Epigynum, lateral; 24. Procursus, dorsal. Scale bars:
|
||
<quantity id="474A9B4C7568FFE2FE8AF8AFFE31F8B7" box="[322,407,1863,1886]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="D4CD66217569FFE3FEC0FD10FB6BFCB2" httpUri="https://zenodo.org/record/273234/files/figure.png" pageId="5" pageNumber="6" targetBox="[264,1323,289,732]" targetPageId="5">
|
||
<paragraph id="800D36A97569FFE3FEC0FD10FB6BFCB2" blockId="5.[264,1324,760,859]" pageId="5" pageNumber="6">
|
||
<emphasis id="B2C6EABB7569FFE3FEC0FD10FE76FCF9" bold="true" box="[264,464,760,784]" pageId="5" pageNumber="6">FIGURES 25–29.</emphasis>
|
||
<taxonomicName id="47B24D2A7569FFE3FE11FD11FD24FCF9" box="[473,642,761,784]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="justoi" status="comb. nov.">
|
||
<emphasis id="B2C6EABB7569FFE3FE11FD11FD24FCF9" box="[473,642,761,784]" italics="true" pageId="5" pageNumber="6">Ossinissa justoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C07569FFE3FD43FD11FCEEFCF9" box="[651,840,761,784]" pageId="5" pageNumber="6">new combination</taxonomicNameLabel>
|
||
, female. 25. Opisthosoma, lateral; 26. Anterior lateral spinnerets, ventral; 27. Ocular area, dorsal; 28. Spinnerets, ventral; 29. Posterior median spinnerets, ventral. Scale bars: 25: 500 µm; 26: 20 µm; 27: 100µm; 29: 10 µm; 28: 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="800D36A97569FFE3FEF0FC79FB1CFB75" blockId="5.[264,1324,913,1420]" pageId="5" pageNumber="6">
|
||
<emphasis id="B2C6EABB7569FFE3FEF0FC79FDEAFC42" bold="true" box="[312,588,913,939]" pageId="5" pageNumber="6">Generic relationships.</emphasis>
|
||
— This genus shares several synapomorphies with the members of the genus
|
||
<taxonomicName id="47B24D2A7569FFE3FE1DFC52FD95FC3A" box="[469,563,954,979]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB7569FFE3FE1DFC52FD95FC3A" box="[469,563,954,979]" italics="true" pageId="5" pageNumber="6">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
. The most important of these is the presence of a welldeveloped uncus. Similarities can be observed in the general structure of the male palp (the shape of femur, tibia and trochanter). The distribution of the eyes is also similar to that in
|
||
<taxonomicName id="47B24D2A7569FFE3FB0DFBE2FA85FBCA" box="[1221,1315,1034,1059]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB7569FFE3FB0DFBE2FA85FBCA" box="[1221,1315,1034,1059]" italics="true" pageId="5" pageNumber="6">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
. However, despite these similarities, the structure of the procursus in
|
||
<taxonomicName id="47B24D2A7569FFE3FBFCFBDAFB02FBA2" box="[1076,1188,1074,1099]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB7569FFE3FBFCFBDAFB02FBA2" box="[1076,1188,1074,1099]" italics="true" pageId="5" pageNumber="6">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C07569FFE3FB65FBDAFA8DFBA5" box="[1197,1323,1074,1100]" pageId="5" pageNumber="6" rank="genus">new genus</taxonomicNameLabel>
|
||
differs greatly; the tarsal organ is flat while that in
|
||
<taxonomicName id="47B24D2A7569FFE3FC91FBB2FC1EFB9A" box="[857,952,1114,1139]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB7569FFE3FC91FBB2FC1EFB9A" box="[857,952,1114,1139]" italics="true" pageId="5" pageNumber="6">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
is capsulated; the chelicerae do not have frontal prominences and the morphology of the female vulva is distinct.
|
||
</paragraph>
|
||
<paragraph id="800D36A97569FFE3FEF0FB42FEEBFAFD" blockId="5.[264,1324,913,1420]" pageId="5" pageNumber="6">
|
||
Although
|
||
<taxonomicName id="47B24D2A7569FFE3FE66FB42FDB5FB2A" box="[430,531,1194,1219]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="justoi">
|
||
<emphasis id="B2C6EABB7569FFE3FE66FB42FDB5FB2A" box="[430,531,1194,1219]" italics="true" pageId="5" pageNumber="6">O. justoi</emphasis>
|
||
</taxonomicName>
|
||
was placed initially in
|
||
<emphasis id="B2C6EABB7569FFE3FCEDFB42FC4BFB2A" box="[805,1005,1194,1219]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="47B24D2A7569FFE3FCEDFB42FC4EFB2A" box="[805,1000,1194,1219]" class="Arachnida" family="Pholcidae" genus="Spermophorides" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Spermophorides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
it does not share any character with the members of this genus except for the globular shape of the female opisthosoma.
|
||
</paragraph>
|
||
<paragraph id="800D36A97569FFE3FEF0FACAFE5FFA65" blockId="5.[264,1324,913,1420]" pageId="5" pageNumber="6">
|
||
Relationships were examined by means of a cladistic analysis. The data matrix used for the analysis is listed in Appendix 1. The matrix was based on
|
||
<bibRefCitation id="E4234B587569FFE3FBFCFAA2FB65FA8D" author="Huber" box="[1076,1219,1354,1380]" pageId="5" pageNumber="11" refString="Huber, B. A. (2003) High species diversity in one of the dominant groups of spiders in East African montane forests (Araneae: Pholcidae: Buitinga n. gen., Spermophora Hentz). Zoological Journal of the Linnean Society, 137, 555 - 619." type="journal article" year="2003">Huber 2003</bibRefCitation>
|
||
but was modified as follows:
|
||
</paragraph>
|
||
<paragraph id="800D36A97569FFE3FEC0FA2AFD40F9BD" blockId="5.[264,1324,1474,1780]" pageId="5" pageNumber="6">
|
||
1 Several taxa were deleted, mainly New World genera and ninetines, which would not have contributed to the determination of
|
||
<taxonomicName id="47B24D2A7569FFE3FCDEFA02FC20F9EA" box="[790,902,1514,1539]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB7569FFE3FCDEFA02FC20F9EA" box="[790,902,1514,1539]" italics="true" pageId="5" pageNumber="6">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C07569FFE3FC46FA02FBACF9ED" box="[910,1034,1514,1540]" pageId="5" pageNumber="6" rank="genus">new genus</taxonomicNameLabel>
|
||
and its close relatives. 2
|
||
<taxonomicName id="47B24D2A7569FFE3FEF0F9FAFE3BF9C2" box="[312,413,1554,1579]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="justoi">
|
||
<emphasis id="B2C6EABB7569FFE3FEF0F9FAFE3BF9C2" box="[312,413,1554,1579]" italics="true" pageId="5" pageNumber="6">O. justoi</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<taxonomicName id="47B24D2A7569FFE3FDD3F9FAFDDFF9C2" box="[539,633,1554,1579]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB7569FFE3FDD3F9FAFDDFF9C2" box="[539,633,1554,1579]" italics="true" pageId="5" pageNumber="6">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
species from the Canary Islands were added (
|
||
<emphasis id="B2C6EABB7569FFE3FB5FF9FAFDBFF9BA" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="47B24D2A7569FFE3FB5FF9FAFE30F9BA" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fuerteventurensis">P. fuerteventurensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="47B24D2A7569FFE3FE6AF9D2FDBFF9BA" box="[418,537,1594,1619]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ornatus">P. ornatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="47B24D2A7569FFE3FD99F9D2FD72F9BA" box="[593,724,1594,1619]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gomerae">
|
||
<emphasis id="B2C6EABB7569FFE3FD99F9D2FD72F9BA" box="[593,724,1594,1619]" italics="true" pageId="5" pageNumber="6">P. gomerae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="800D36A97569FFE3FEC0F98AFC45F91D" blockId="5.[264,1324,1474,1780]" pageId="5" pageNumber="6">3 One character was added: 60 – shape of the opisthosoma; 0 – cylindrical, 1 – elevated (the opisthosoma does not present regular height). When the opisthosoma was elevated in one of the two sexes, the species was coded as 1. (Note that the total number of characters is 61 but they are numbered starting from 0).</paragraph>
|
||
<paragraph id="800D36A97569FFE3FEF0F8C2FC39F885" blockId="5.[264,1323,1834,1900]" pageId="5" pageNumber="6">Using a previously published matrix from which several nonrelated taxa were deleted resulted in a high number of noninformative characters.</paragraph>
|
||
<caption id="D4CD6621756AFFE0FEC0FA8FFBCCF9C5" httpUri="https://zenodo.org/record/273235/files/figure.png" pageId="6" pageNumber="7" targetBox="[264,1323,326,1359]" targetPageId="6">
|
||
<paragraph id="800D36A9756AFFE0FEC0FA8FFBCCF9C5" blockId="6.[264,1323,1383,1580]" pageId="6" pageNumber="7">
|
||
<emphasis id="B2C6EABB756AFFE0FEC0FA8FFE75FA96" bold="true" box="[264,467,1383,1407]" pageId="6" pageNumber="7">FIGURES 30–39.</emphasis>
|
||
<taxonomicName id="47B24D2A756AFFE0FE15FA80FD2FFA96" box="[477,649,1384,1407]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="justoi" status="comb. nov.">
|
||
<emphasis id="B2C6EABB756AFFE0FE15FA80FD2FFA96" box="[477,649,1384,1407]" italics="true" pageId="6" pageNumber="7">Ossinissa justoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756AFFE0FD5CFA8FFCF5FA97" box="[660,851,1383,1406]" pageId="6" pageNumber="7">new combination</taxonomicNameLabel>
|
||
, male. Bulbal projections (arrow indicates embolus), dorsoprolateral. Figures 31, 33, 38.
|
||
<taxonomicName id="47B24D2A756AFFE0FCCEFA62FC13FA48" box="[774,949,1418,1441]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B2C6EABB756AFFE0FCCEFA62FC13FA48" box="[774,949,1418,1441]" italics="true" pageId="6" pageNumber="7">Pholcus ornatus</emphasis>
|
||
</taxonomicName>
|
||
, male: 31. Spinnerets, ventral; 33. Gonopore, ventral; 38. Male palp, prolateral. Figures 32, 35–36.
|
||
<taxonomicName id="47B24D2A756AFFE0FC03FA45FB79FA2D" box="[971,1247,1453,1476]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fuerteventurensis">
|
||
<emphasis id="B2C6EABB756AFFE0FC03FA45FB79FA2D" box="[971,1247,1453,1476]" italics="true" pageId="6" pageNumber="7">Pholcus fuerteventurensis</emphasis>
|
||
</taxonomicName>
|
||
, male: 32. Spinnerets, ventral; 35. Gonopore, ventral; 36. Male palp, prolateral. Figures 34, 37, 39.
|
||
<taxonomicName id="47B24D2A756AFFE0FB27FA38FE29F9E0" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gomerae">
|
||
<emphasis id="B2C6EABB756AFFE0FB27FA38FE29F9E0" italics="true" pageId="6" pageNumber="7">Pholcus gomerae</emphasis>
|
||
</taxonomicName>
|
||
, male: 34. Gonopore, ventral; 37. Spinnerets, ventral; 39. Male palp, ventroprolateral. Scale bars: 30, 33–34: 100µm; 31–32, 36–37, 39: 200µm; 38: 500µm; 35: 50µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="800D36A9756AFFE1FEF0F988FE53FE6F" blockId="6.[264,1323,1632,1898]" lastBlockId="7.[264,1323,284,990]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
Running NONA with hold 10000 and mult*1000 gave as a result 81 most parsimonious trees with 94 steps. The strict consensus of these trees is given in Appendix II. In addition, the stability of the clades was estimated by calculating the bootstrap and jackknife support using NONA. In all trees generated by the analysis,
|
||
<taxonomicName id="47B24D2A756AFFE0FC00F930FB9EF918" box="[968,1080,1752,1777]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756AFFE0FC00F930FB9EF918" box="[968,1080,1752,1777]" italics="true" pageId="6" pageNumber="7">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756AFFE0FB88F930FB1AF91B" box="[1088,1212,1752,1778]" pageId="6" pageNumber="7" rank="genus">new genus</taxonomicNameLabel>
|
||
is placed in a clade formed by
|
||
<taxonomicName id="47B24D2A756AFFE0FE34F8E8FD38F8F0" box="[508,670,1792,1817]" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FE34F8E8FD38F8F0" box="[508,670,1792,1817]" italics="true" pageId="6" pageNumber="7">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="47B24D2A756AFFE0FD64F8E8FCADF8F0" box="[684,779,1792,1817]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FD64F8E8FCADF8F0" box="[684,779,1792,1817]" italics="true" pageId="6" pageNumber="7">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47B24D2A756AFFE0FC8AF8E8FC46F8F0" box="[834,992,1792,1817]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FC8AF8E8FC46F8F0" box="[834,992,1792,1817]" italics="true" pageId="6" pageNumber="7">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
. In this group
|
||
<taxonomicName id="47B24D2A756AFFE0FB41F8E8FA8CF8F0" box="[1161,1322,1792,1817]" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FB41F8E8FA8CF8F0" box="[1161,1322,1792,1817]" italics="true" pageId="6" pageNumber="7">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
occupies in most of the cases a basal position, and
|
||
<taxonomicName id="47B24D2A756AFFE0FCA4F8C0FC7BF8A8" box="[876,989,1832,1857]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756AFFE0FCA4F8C0FC7BF8A8" box="[876,989,1832,1857]" italics="true" pageId="6" pageNumber="7">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756AFFE0FC2DF8C0FBC2F8AB" box="[997,1124,1832,1858]" pageId="6" pageNumber="7" rank="genus">new genus</taxonomicNameLabel>
|
||
is recognized as the closest relative of
|
||
<taxonomicName id="47B24D2A756AFFE0FDC5F8B8FDCDF880" box="[525,619,1872,1897]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FDC5F8B8FDCDF880" box="[525,619,1872,1897]" italics="true" pageId="6" pageNumber="7">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
. In all cladograms
|
||
<taxonomicName id="47B24D2A756AFFE0FC99F8B8FC16F880" box="[849,944,1872,1897]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756AFFE0FC99F8B8FC16F880" box="[849,944,1872,1897]" italics="true" pageId="6" pageNumber="7">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
appears to be paraphyletic with respect to
|
||
<taxonomicName id="47B24D2A756BFFE1FE4BFEF4FD87FEDC" box="[387,545,284,309]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FE4BFEF4FD87FEDC" box="[387,545,284,309]" italics="true" pageId="7" pageNumber="8">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
and this is supported by both bootstrap and jackknife. This could be one more evidence for the possible synonymy of
|
||
<taxonomicName id="47B24D2A756BFFE1FCB0FEACFC70FEB4" box="[888,982,324,349]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FCB0FEACFC70FEB4" box="[888,982,324,349]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47B24D2A756BFFE1FBD9FEACFB09FEB4" box="[1041,1199,324,349]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FBD9FEACFB09FEB4" box="[1041,1199,324,349]" italics="true" pageId="7" pageNumber="8">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E4234B58756BFFE1FB08FEACFEE0FE6F" author="Brignoli" pageId="7" pageNumber="10" refString="Brignoli, P. M. (1980) Sur le genre Leptopholcus Simon, 1893 (Araneae, Pholcidae). Revue de Zoologie Africaine, 94 (3), 649 - 655." type="journal article" year="1980">Brignoli, 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E4234B58756BFFE1FE9AFE84FE4EFE6F" author="Huber" box="[338,488,364,390]" pageId="7" pageNumber="11" refString="Huber, B. A. (2001) The pholcids of Australia (Araneae; Pholcidae): Taxonomy, biogeography, and relationships. Bulletin of the American Museum of Natural History, 260, 1 - 144." type="journal article" year="2001">Huber, 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="800D36A9756BFFE1FEF0FE7CFDB6FE17" blockId="7.[264,1323,284,990]" pageId="7" pageNumber="8">The application of implied weighting with PeeWee (algorithm that resolves character conflict in favor of characters that present less homoplasy) with K from 1–6 produced the same result as NONA.</paragraph>
|
||
<paragraph id="800D36A9756BFFE1FEF0FDE4FD9BFD2F" blockId="7.[264,1323,284,990]" pageId="7" pageNumber="8">
|
||
The placement of
|
||
<taxonomicName id="47B24D2A756BFFE1FDC7FDE4FDD9FDCC" box="[527,639,524,549]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756BFFE1FDC7FDE4FDD9FDCC" box="[527,639,524,549]" italics="true" pageId="7" pageNumber="8">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756BFFE1FD4FFDE4FCA2FDCF" box="[647,772,524,550]" pageId="7" pageNumber="8" rank="genus">new genus</taxonomicNameLabel>
|
||
outside
|
||
<taxonomicName id="47B24D2A756BFFE1FCA2FDE4FC6EFDCC" box="[874,968,524,549]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FCA2FDE4FC6EFDCC" box="[874,968,524,549]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
in all of the analyses, and the fact that the clade
|
||
<taxonomicName id="47B24D2A756BFFE1FE2CFDDCFDE4FDA4" box="[484,578,564,589]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FE2CFDDCFDE4FDA4" box="[484,578,564,589]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="47B24D2A756BFFE1FDACFDDCFCA4FDA4" box="[612,770,564,589]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FDACFDDCFCA4FDA4" box="[612,770,564,589]" italics="true" pageId="7" pageNumber="8">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
appears in all cladograms and is supported by bootstrap and jackknife, upholds the existence of
|
||
<taxonomicName id="47B24D2A756BFFE1FC9EFDB4FC61FD9C" box="[854,967,604,629]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756BFFE1FC9EFDB4FC61FD9C" box="[854,967,604,629]" italics="true" pageId="7" pageNumber="8">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756BFFE1FC07FDB4FBEBFD9F" box="[975,1101,604,630]" pageId="7" pageNumber="8" rank="genus">new genus</taxonomicNameLabel>
|
||
as separate genus. Although these results suggest that
|
||
<taxonomicName id="47B24D2A756BFFE1FD75FD6CFCFDFD74" box="[701,859,644,669]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FD75FD6CFCFDFD74" box="[701,859,644,669]" italics="true" pageId="7" pageNumber="8">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<taxonomicName id="47B24D2A756BFFE1FBF2FD6CFB3EFD74" box="[1082,1176,644,669]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FBF2FD6CFB3EFD74" box="[1082,1176,644,669]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
, this would need further investigation.
|
||
</paragraph>
|
||
<paragraph id="800D36A9756BFFE1FEF0FD3CFC51FCFF" blockId="7.[264,1323,284,990]" pageId="7" pageNumber="8">
|
||
In summary, we conclude that
|
||
<taxonomicName id="47B24D2A756BFFE1FD6DFD3CFCB3FD04" box="[677,789,724,749]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B2C6EABB756BFFE1FD6DFD3CFCB3FD04" box="[677,789,724,749]" italics="true" pageId="7" pageNumber="8">Ossinissa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756BFFE1FCD6FD3CFC3DFD07" box="[798,923,724,750]" pageId="7" pageNumber="8" rank="genus">new genus</taxonomicNameLabel>
|
||
is closely related with the genera in the
|
||
<taxonomicName id="47B24D2A756BFFE1FE99FD14FE16FCFC" box="[337,432,764,789]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FE99FD14FE16FCFC" box="[337,432,764,789]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
group:
|
||
<taxonomicName id="47B24D2A756BFFE1FDC3FD14FDCFFCFC" box="[523,617,764,789]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FDC3FD14FDCFFCFC" box="[523,617,764,789]" italics="true" pageId="7" pageNumber="8">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="47B24D2A756BFFE1FDB0FD14FCBCFCFC" box="[632,794,764,789]" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FDB0FD14FCBCFCFC" box="[632,794,764,789]" italics="true" pageId="7" pageNumber="8">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47B24D2A756BFFE1FC9BFD14FC57FCFC" box="[851,1009,764,789]" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2C6EABB756BFFE1FC9BFD14FC57FCFC" box="[851,1009,764,789]" italics="true" pageId="7" pageNumber="8">Leptopholcus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8A86522756BFFE1FEF0FCCCFE7DFC37" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="800D36A9756BFFE1FEF0FCCCFE4CFC67" blockId="7.[264,1323,284,990]" pageId="7" pageNumber="8">
|
||
<emphasis id="B2C6EABB756BFFE1FEF0FCCCFE7EFCD7" bold="true" box="[312,472,804,830]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
– The distribution of this genus is restricted. The Cueva de Don Justo and the Cueva de los Pocitos on El Hierro Island are the only two locations where it has ever been reported.
|
||
</paragraph>
|
||
<paragraph id="800D36A9756BFFE1FEF0FC74FE7DFC37" blockId="7.[264,1323,284,990]" pageId="7" pageNumber="8">
|
||
<emphasis id="B2C6EABB756BFFE1FEF0FC74FD80FC5F" bold="true" box="[312,550,924,950]" pageId="7" pageNumber="8">Included species. –</emphasis>
|
||
Currently, this genus includes only one species,
|
||
<taxonomicName id="47B24D2A756BFFE1FBB8FC74FA8CFC5C" box="[1136,1322,924,949]" class="Arachnida" family="Pholcidae" genus="Ossinissa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="justoi" status="comb. nov.">
|
||
<emphasis id="B2C6EABB756BFFE1FBB8FC74FA8CFC5C" box="[1136,1322,924,949]" italics="true" pageId="7" pageNumber="8">Ossinissa justoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A9F557C0756BFFE1FEC0FC2CFE71FC37" box="[264,471,964,990]" pageId="7" pageNumber="8">new combination</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |