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<document id="43CBF4F51A286C9AD8F15FFD666DCE7B" ID-DOI="10.1080/14772019.2016.1151958" ID-ISSN="1478-0941" ID-PMC="PMC5221741" ID-PubMed="28077930" ID-Zenodo-Dep="10883098" ID-ZooBank="A30E5364-0003-4467-B902-43A41AD456CC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711483431746" checkinUser="felipe" docAuthor="Schwarzhans, Werner, Mors, Thomas, Engelbrecht, Andrea, Reguero, Marcelo &amp; Kriwet, Jurgen" docDate="2017" docId="041B87CAFFA8FFFCD676FB5FE6DDDFC0" docLanguage="en" docName="JSystPalaeontol.15.2.147-170.pdf" docOrigin="Journal of Systematic Palaeontology 15 (2)" docSource="http://dx.doi.org/10.1080/14772019.2016.1151958" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.2:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="2" docTitle="Hoplobrotula Gill 1863" docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="F822FFB2FFA4FFF0D753FFE4E35DDA07" masterDocTitle="Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)" masterLastPageNumber="170" masterPageNumber="147" pageNumber="158" updateTime="1711483851815" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="50F1FA4BD316C9CDD8D1369AF675C160">Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)</mods:title>
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<mods:namePart id="87C8523F86B043FF4D80B05B75A60CEC">Schwarzhans, Werner</mods:namePart>
<mods:affiliation id="16EEC78A74466A225EBA6C8C0CCEBE09">Ahrensburger Weg 103, D- 22359 Hamburg, Germany; &amp; Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, DK- 2100 Copenhagen, Denmark;</mods:affiliation>
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<mods:namePart id="B22CFE25B74265AECD62BB889F8D1FAD">Mors, Thomas</mods:namePart>
<mods:affiliation id="1DE50056D4023B0DD251ED92ED403BD2">Swedish Museum of Natural History, Department of Palaeobiology, P. O. Box 5007, SE- 10405 Stockholm, Sweden;</mods:affiliation>
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<mods:namePart id="7B872D34F574F6A84B389FB9EE35E37A">Engelbrecht, Andrea</mods:namePart>
<mods:affiliation id="E170454B2466E9E6170F267A57CA375A">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:namePart id="F83914ACF5FCA4BE39CE37B622AE1D51">Reguero, Marcelo</mods:namePart>
<mods:affiliation id="E7B9CBA060B7812E28746C58899E9C17">Museo de La Plata, Division ́ Paleontolog ́ ıa de Vertebrados, Paseo del Bosque s / n, B 1900 FWA La Plata, Argentina</mods:affiliation>
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<mods:namePart id="4005CA7F51072BCE8C7B90A3653C901A">Kriwet, Jurgen</mods:namePart>
<mods:affiliation id="34263D0B741E83992FC2B4AAFE78D30B">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:date id="F24FCD35D09CEA92500F9B0AF12846D2">2017</mods:date>
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<mods:number id="37575FF3B3393CE3251F57DE0E080128">2016-03-16</mods:number>
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<mods:number id="DA6722CD42CB7BA0AB52264487E599D3">15</mods:number>
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<mods:identifier id="1DA06B941D1CB1AB9595F62E67ACCE4B" type="ISSN">1478-0941</mods:identifier>
<mods:identifier id="F72C0AA613C55C562D8F2317D0B6C839" type="PMC">PMC5221741</mods:identifier>
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<treatment id="041B87CAFFA8FFFCD676FB5FE6DDDFC0" ID-DOI="http://doi.org/10.5281/zenodo.10883172" ID-Zenodo-Dep="10883172" LSID="urn:lsid:plazi:treatment:041B87CAFFA8FFFCD676FB5FE6DDDFC0" httpUri="http://treatment.plazi.org/id/041B87CAFFA8FFFCD676FB5FE6DDDFC0" lastPageNumber="158" pageId="12" pageNumber="158">
<subSubSection id="C4A86557FFA8FFFCD676FB5FE1D5DED5" box="[293,648,1211,1234]" pageId="12" pageNumber="158" type="nomenclature">
<paragraph id="8C0D36DCFFA8FFFCD676FB5FE1D5DED5" blockId="12.[246,695,1052,1266]" box="[293,648,1211,1234]" pageId="12" pageNumber="158">
<heading id="D74581B0FFA8FFFCD676FB5FE1D5DED5" box="[293,648,1211,1234]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="158" reason="8">
<emphasis id="BEC6EACEFFA8FFFCD676FB5FE170DED5" bold="true" box="[293,557,1211,1234]" pageId="12" pageNumber="158">
<taxonomicName id="4BB24D5FFFA8FFFCD676FB5FE2E4DED5" ID-CoL="4YLG" ID-ENA="990983" authorityName="Gill" authorityYear="1863" box="[293,441,1211,1234]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD676FB5FE2E4DED5" bold="true" box="[293,441,1211,1234]" italics="true" pageId="12" pageNumber="158">Hoplobrotula</emphasis>
</taxonomicName>
?
<emphasis id="BEC6EACEFFA8FFFCD69EFB5FE170DED5" bold="true" box="[461,557,1211,1234]" italics="true" pageId="12" pageNumber="158">antipoda</emphasis>
</emphasis>
sp. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD638FB3FE11CDEF5" box="[363,577,1243,1266]" pageId="12" pageNumber="158" type="description">
<paragraph id="8C0D36DCFFA8FFFCD638FB3FE11CDEF5" blockId="12.[246,695,1052,1266]" box="[363,577,1243,1266]" pageId="12" pageNumber="158">
(
<figureCitation id="14892A59FFA8FFFCD627FB3FE2ACDEF5" box="[372,497,1243,1266]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="12" pageNumber="158">Figs 5A—C</figureCitation>
,
<figureCitation id="14892A59FFA8FFFCD6AFFB3FE167DEF5" box="[508,570,1243,1266]" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="12" pageNumber="158">6O, P</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD7CCFAC5E194DFCA" pageId="12" pageNumber="158" type="materials_examined">
<paragraph id="8C0D36DCFFA8FFFCD7CCFAC5E13DDF5F" blockId="12.[159,782,1313,1368]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD7CCFAC5E257DF3F" bold="true" box="[159,266,1313,1336]" pageId="12" pageNumber="158">Material.</emphasis>
<typeStatus id="5309887EFFA8FFFCD64BFAC5E2DDDF3F" box="[280,384,1313,1336]" pageId="12" pageNumber="158" type="holotype">Holotype</typeStatus>
: NRM-PZ P.15984 (
<figureCitation id="14892A59FFA8FFFCD533FAC5E184DF3F" box="[608,729,1313,1336]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="12" pageNumber="158">Figs 5A, B</figureCitation>
,
<figureCitation id="14892A59FFA8FFFCD5B5FAC5E3ECDF5F" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="12" pageNumber="158">6O, P</figureCitation>
).
<typeStatus id="5309887EFFA8FFFCD797FAA5E27BDF5F" box="[196,294,1345,1368]" pageId="12" pageNumber="158" type="paratype">Paratype</typeStatus>
: NRM-PZ P.15985 (
<figureCitation id="14892A59FFA8FFFCD552FAA5E109DF5F" box="[513,596,1345,1368]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="12" pageNumber="158">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8C0D36DCFFA8FFFCD7CCFA93E194DFCA" blockId="12.[159,781,1399,1485]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD7CCFA93E271DF89" bold="true" box="[159,300,1399,1422]" pageId="12" pageNumber="158">Occurrence.</emphasis>
Telm 5 unit;
<taxonomicName id="4BB24D5FFFA8FFFCD688FA93E17FDF89" authorityName="Scopoli" authorityYear="1777" box="[475,546,1399,1422]" class="Gastropoda" family="Naticidae" genus="Natica" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="158" phylum="Mollusca" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD688FA93E17FDF89" box="[475,546,1399,1422]" italics="true" pageId="12" pageNumber="158">Natica</emphasis>
</taxonomicName>
horizon,
<taxonomicName id="4BB24D5FFFA8FFFCD5CAFA93E1A4DF89" box="[665,761,1399,1422]" class="Bivalvia" family="Cucullaeidae" genus="Cucullea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Arcida" pageId="12" pageNumber="158" phylum="Mollusca" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD5CAFA93E1A4DF89" box="[665,761,1399,1422]" italics="true" pageId="12" pageNumber="158">Cucullea</emphasis>
</taxonomicName>
I member, La Meseta Formation, late Ypresian, early Eocene. Site IAA 1/90, Seymour Island,
<collectingCountry id="F4A5764CFFA8FFFCD501FA52E198DFCA" box="[594,709,1462,1485]" name="Antarctica" pageId="12" pageNumber="158">Antarctica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD7CCFA08E151DC44" pageId="12" pageNumber="158" type="etymology">
<paragraph id="8C0D36DCFFA8FFFCD7CCFA08E151DC44" blockId="12.[159,781,1516,1603]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD7CCFA08E27CDC04" bold="true" box="[159,289,1516,1539]" pageId="12" pageNumber="158">Etymology.</emphasis>
Named for its occurrence on the opposite side of the globe when compared to other Eocene species of the genus, known from Europe.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD7CCF986E2A4DCBE" pageId="12" pageNumber="158" type="diagnosis">
<paragraph id="8C0D36DCFFA8FFFCD7CCF986E2A4DCBE" blockId="12.[159,781,1634,1721]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD7CCF986E249DC7E" bold="true" box="[159,276,1634,1657]" pageId="12" pageNumber="158">Diagnosis.</emphasis>
OL:OH = 1.75—1.95. Dorsal rim with low, broad predorsal lobe.
<collectionCode id="EAA3AE19FFA8FFFCD6DEF966E299DC9E" box="[397,452,1666,1689]" pageId="12" pageNumber="158">OCL</collectionCode>
:
<collectionCode id="EAA3AE19FFA8FFFCD69BF966E2A3DC9E" box="[456,510,1666,1689]" pageId="12" pageNumber="158">CCL</collectionCode>
= 2.5—2.75. Sulcus and particularly ostium very narrow.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD7CCF93CE78BDBF4" pageId="12" pageNumber="158" type="description">
<paragraph id="8C0D36DCFFA8FFFCD7CCF93CE1BADDA9" blockId="12.[159,781,1752,1966]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD7CCF93CE277DCE8" bold="true" box="[159,298,1752,1775]" pageId="12" pageNumber="158">Description.</emphasis>
Relatively small, elongate and thin otoliths up to
<quantity id="4B4A9B39FFA8FFFCD7B3F913E27DDD09" box="[224,288,1783,1806]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="158" unit="mm" value="4.0">4 mm</quantity>
in length (
<typeStatus id="5309887EFFA8FFFCD6CBF913E2A9DD09" box="[408,500,1783,1806]" pageId="12" pageNumber="158" type="holotype">holotype</typeStatus>
<quantity id="4B4A9B39FFA8FFFCD6AEF913E10DDD09" box="[509,592,1783,1806]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="158" unit="mm" value="3.9">3.9 mm</quantity>
long). OH:OT = 2.6. Dorsal rim rather regularly and gently curved with broad, not much expanded predorsal lobe. Ventral rim regularly curved, deepest slightly in front of its middle. Anterior tip tapering, inferior, below level of ostium; posterior tip narrow, tapering, at level of cauda. All rims smooth.
</paragraph>
<paragraph id="8C0D36DCFFA8FFFCD434FF59E78BDBF4" blockId="12.[845,1467,189,499]" pageId="12" pageNumber="158">Inner face moderately convex, smooth, with very narrow, shallow, slightly supramedian sulcus. Ostium very narrow, about 2.5 to 3 times the length of cauda, reaching close to anterior rim of otolith; cauda short, slightly deepened, with rounded tip, terminating at some distance from posterior rim of otolith. Ventral margin of sulcus distinctly indented at ostial—caudal joint. Dorsal depression indistinct, small; ventral furrow faint, fading towards posterior, close to ventral rim of otolith. Outer face slightly convex, less than inner face, smooth.</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFA8FFFCD41EFDF7E6DDDFC0" pageId="12" pageNumber="158" type="discussion">
<paragraph id="8C0D36DCFFA8FFFCD41EFDF7E70DDEC0" blockId="12.[845,1467,531,1479]" pageId="12" pageNumber="158">
<emphasis id="BEC6EACEFFA8FFFCD41EFDF7E0E6D82D" bold="true" box="[845,955,531,554]" pageId="12" pageNumber="158">Remarks.</emphasis>
<taxonomicName id="4BB24D5FFFA8FFFCD499FDF7E706D82D" authorityName="Gill" authorityYear="1863" box="[970,1115,531,554]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD499FDF7E706D82D" box="[970,1115,531,554]" italics="true" pageId="12" pageNumber="158">Hoplobrotula</emphasis>
</taxonomicName>
?
<emphasis id="BEC6EACEFFA8FFFCD33CFDF7E793D82D" box="[1135,1230,531,554]" italics="true" pageId="12" pageNumber="158">antipoda</emphasis>
is a typical representative of the many small ophidiid otoliths, which are so common in the Palaeogene. It clearly differs from the species described from the Eocene of Europe such as
<taxonomicName id="4BB24D5FFFA8FFFCD23CFD97E7D1D8AE" authority="(Sulc, 1932)" baseAuthorityName="Sulc" baseAuthorityYear="1932" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="biscaica">
<emphasis id="BEC6EACEFFA8FFFCD23CFD97E0A1D8AE" italics="true" pageId="12" pageNumber="158">Hoplobrotula biscaica</emphasis>
(Sulc, 1932)
</taxonomicName>
,
<taxonomicName id="4BB24D5FFFA8FFFCD3CBFD76E6E9D8AE" authority="Nolf, 1980" authorityName="Nolf" authorityYear="1980" box="[1176,1460,658,681]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="greenwoodi">
<emphasis id="BEC6EACEFFA8FFFCD3CBFD76E66AD8AE" box="[1176,1335,658,681]" italics="true" pageId="12" pageNumber="158">H. greenwoodi</emphasis>
<bibRefCitation id="E8234B2DFFA8FFFCD26CFD76E6E9D8AE" author="Nolf, D." box="[1343,1460,658,681]" pageId="12" pageNumber="158" pagination="71 - 195" refId="ref16789" refString="Nolf, D. 1980. Etude monographique des otolithes des Ophidiiformes actuels et r´evision des esp`eces fossils (Pisces, Teleostei). Mededelingen van de Werkgroup voor Tertiaire en Kwartaire Geologie, 17, 71 - 195." type="journal article" year="1980">Nolf, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4BB24D5FFFA8FFFCD41EFD56E760D8CE" authority="Nolf, 1980" authorityName="Nolf" authorityYear="1980" box="[845,1085,690,713]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="robusta">
<emphasis id="BEC6EACEFFA8FFFCD41EFD56E09DD8CE" box="[845,960,690,713]" italics="true" pageId="12" pageNumber="158">H. robusta</emphasis>
<bibRefCitation id="E8234B2DFFA8FFFCD49BFD56E760D8CE" author="Nolf, D." box="[968,1085,690,713]" pageId="12" pageNumber="158" pagination="71 - 195" refId="ref16789" refString="Nolf, D. 1980. Etude monographique des otolithes des Ophidiiformes actuels et r´evision des esp`eces fossils (Pisces, Teleostei). Mededelingen van de Werkgroup voor Tertiaire en Kwartaire Geologie, 17, 71 - 195." type="journal article" year="1980">Nolf, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4BB24D5FFFA8FFFCD31FFD56E0D0D8EE" authority="(Dante &amp; Frizzell, 1965)" baseAuthorityName="Dante &amp; Frizzell" baseAuthorityYear="1965" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="melrosensis">
<emphasis id="BEC6EACEFFA8FFFCD31FFD56E7B6D8CE" box="[1100,1259,690,713]" italics="true" pageId="12" pageNumber="158">H. melrosensis</emphasis>
(Dante &amp; Frizzell, 1965)
</taxonomicName>
,
<taxonomicName id="4BB24D5FFFA8FFFCD4C9FD36E797D8EE" authority="(Schubert, 1916)" baseAuthorityName="Schubert" baseAuthorityYear="1916" box="[922,1226,722,745]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="waltoni">
<emphasis id="BEC6EACEFFA8FFFCD4C9FD36E751D8EE" box="[922,1036,722,745]" italics="true" pageId="12" pageNumber="158">H. waltoni</emphasis>
(
<bibRefCitation id="E8234B2DFFA8FFFCD34FFD36E79DD8EE" author="Schubert, R." box="[1052,1216,722,745]" pageId="12" pageNumber="158" pagination="277 - 288" refId="ref17635" refString="Schubert, R. 1916. Obereoz ¨ ane Otolithen vom Barton Cliff bei Christchurch (Hampshire). Jahrbuch der Kaiserlich- Koniglichen ¨ Geologischen Gesellschaft, 65, 277 - 288." type="journal article" year="1916">Schubert, 1916</bibRefCitation>
)
</taxonomicName>
, as well as
<taxonomicName id="4BB24D5FFFA8FFFCD21FFD36E721D90E" authority="Konig, 1825" authorityName="Konig" authorityYear="1825" family="Ophidiidae" genus="Ampheristus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="toliapicus">
<emphasis id="BEC6EACEFFA8FFFCD21FFD36E0BFD90E" italics="true" pageId="12" pageNumber="158">Ampheristus toliapicus</emphasis>
Konig, 1825
</taxonomicName>
and
<taxonomicName id="4BB24D5FFFA8FFFCD3EDFD16E097D92E" authority="(Stinton &amp; Nolf, 1970)" baseAuthorityName="Stinton &amp; Nolf" baseAuthorityYear="1970" family="Ophidiidae" genus="Ampheristus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="lerichei">
<emphasis id="BEC6EACEFFA8FFFCD3EDFD16E668D90E" box="[1214,1333,754,777]" italics="true" pageId="12" pageNumber="158">A. lerichei</emphasis>
(Stinton &amp; Nolf, 1970)
</taxonomicName>
(see
<bibRefCitation id="E8234B2DFFA8FFFCD356FCF6E72ED92E" author="Nolf, D." box="[1029,1139,786,809]" pageId="12" pageNumber="158" pagination="71 - 195" refId="ref16789" refString="Nolf, D. 1980. Etude monographique des otolithes des Ophidiiformes actuels et r´evision des esp`eces fossils (Pisces, Teleostei). Mededelingen van de Werkgroup voor Tertiaire en Kwartaire Geologie, 17, 71 - 195." type="journal article" year="1980">Nolf 1980</bibRefCitation>
,
<bibRefCitation id="E8234B2DFFA8FFFCD3D1FCF6E7EAD92E" author="Nolf, D." box="[1154,1207,786,809]" pageId="12" pageNumber="158" refId="ref16894" refString="Nolf, D. 2013. The diversity of fish otoliths, past and present. Royal Belgian Institute of Natural Sciences, Brussels, 222 pp., 359 pls." type="book" year="2013">2013</bibRefCitation>
for figures) by the very narrow and long ostium and the low predorsal lobe. Otoliths of
<taxonomicName id="4BB24D5FFFA8FFFCD4FFFCB6E6E6D96E" authority="(Schubert, 1916)" baseAuthorityName="Schubert" baseAuthorityYear="1916" box="[940,1467,850,873]" family="Ophidiidae" genus="Hoplobrotuloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="bartonensis">
<emphasis id="BEC6EACEFFA8FFFCD4FFFCB6E7AFD96E" box="[940,1266,850,873]" italics="true" pageId="12" pageNumber="158">Hoplobrotuloides bartonensis</emphasis>
(
<bibRefCitation id="E8234B2DFFA8FFFCD254FCB6E6ECD96E" author="Schubert, R." box="[1287,1457,850,873]" pageId="12" pageNumber="158" pagination="277 - 288" refId="ref17635" refString="Schubert, R. 1916. Obereoz ¨ ane Otolithen vom Barton Cliff bei Christchurch (Hampshire). Jahrbuch der Kaiserlich- Koniglichen ¨ Geologischen Gesellschaft, 65, 277 - 288." type="journal article" year="1916">Schubert, 1916</bibRefCitation>
)
</taxonomicName>
are similarly elongate, but differ in the absence of a predorsal lobe, the flat ventral rim and the broadly expanded posterior tip (see
<bibRefCitation id="E8234B2DFFA8FFFCD355FC55E72FD9CF" author="Nolf, D." box="[1030,1138,945,968]" pageId="12" pageNumber="158" pagination="71 - 195" refId="ref16789" refString="Nolf, D. 1980. Etude monographique des otolithes des Ophidiiformes actuels et r´evision des esp`eces fossils (Pisces, Teleostei). Mededelingen van de Werkgroup voor Tertiaire en Kwartaire Geologie, 17, 71 - 195." type="journal article" year="1980">Nolf 1980</bibRefCitation>
;
<bibRefCitation id="E8234B2DFFA8FFFCD3D3FC55E615D9CF" author="Schwarzhans, W." box="[1152,1352,945,968]" pageId="12" pageNumber="158" pagination="63 - 122" refId="ref17755" refString="Schwarzhans, W. 1981. Vergleichende morphologische Untersuchungen an rezenten und fossilen Otolithen der Ordnung Ophidiiformes. Berliner geowissenschaftliche Abhandlungen, (A), 32, 63 - 122." type="journal article" year="1981">Schwarzhans 1981</bibRefCitation>
). It is possible that
<emphasis id="BEC6EACEFFA8FFFCD496FC35E0BDD9EF" box="[965,992,977,1000]" italics="true" pageId="12" pageNumber="158">H.</emphasis>
?
<emphasis id="BEC6EACEFFA8FFFCD4A9FC35E704D9EF" box="[1018,1113,977,1000]" italics="true" pageId="12" pageNumber="158">antipoda</emphasis>
represents yet another extinct genus related to the extant genus
<taxonomicName id="4BB24D5FFFA8FFFCD39FFC15E600DE0F" authorityName="Gill" authorityYear="1863" box="[1228,1373,1009,1032]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD39FFC15E600DE0F" box="[1228,1373,1009,1032]" italics="true" pageId="12" pageNumber="158">Hoplobrotula</emphasis>
</taxonomicName>
and the fossil
<taxonomicName id="4BB24D5FFFA8FFFCD4C1FBF5E74ADE2F" authorityName="König" authorityYear="1825" box="[914,1047,1041,1064]" family="Ophidiidae" genus="Ampheristus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="genus">
<emphasis id="BEC6EACEFFA8FFFCD4C1FBF5E74ADE2F" box="[914,1047,1041,1064]" italics="true" pageId="12" pageNumber="158">Ampheristus</emphasis>
</taxonomicName>
, but a thorough review of the group would be required first. The only other species from the Eocene of the Southern Hemisphere is
<taxonomicName id="4BB24D5FFFA8FFFCD3BFFBB5E7D4DE8F" authority="Schwarzhans, 1980" authorityName="Schwarzhans" authorityYear="1980" family="Ophidiidae" genus="Ampheristus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="158" phylum="Chordata" rank="species" species="sinuocaudatus">
<emphasis id="BEC6EACEFFA8FFFCD3BFFBB5E0F0DE8F" italics="true" pageId="12" pageNumber="158">Ampheristus sinuocaudatus</emphasis>
<bibRefCitation id="E8234B2DFFA8FFFCD4E4FB95E7D4DE8F" author="Schwarzhans, W." box="[951,1161,1137,1160]" pageId="12" pageNumber="158" pagination="1 - 269" refId="ref17706" refString="Schwarzhans, W. 1980. Die terti ¨ are Teleosteer-Fauna Neuseelands, rekonstruiert anhand von Otolithen. Berliner geowissenschaftliche Abhandlungen (A), 26, 1 - 211. [English translation 1984 in New Zealand Geological Survey Report, 113, 1 - 269.]" type="journal article" year="1980">Schwarzhans, 1980</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F4A5764CFFA8FFFCD383FB95E638DE8F" box="[1232,1381,1137,1160]" name="New Zealand" pageId="12" pageNumber="158">New Zealand</collectingCountry>
, which, however, shows a cauda nearly of the length of the ostium and distinctly deepened.
</paragraph>
<paragraph id="8C0D36DCFFA8FFFCD434FB34E6DDDFC0" blockId="12.[845,1467,531,1479]" pageId="12" pageNumber="158">
Otolith data suggest that ophidiiforms were one of the dominant teleost groups in the warm shallow seas, for instance in Europe during the Palaeogene. Temperate seas, such as probably existed in the middle Paleocene of
<collectingCountry id="F4A5764CFFA8FFFCD2D5FAD4E0DEDF60" name="Denmark" pageId="12" pageNumber="158">Denmark</collectingCountry>
or the Eocene of
<collectingCountry id="F4A5764CFFA8FFFCD31AFAB4E785DF60" box="[1097,1240,1360,1383]" name="New Zealand" pageId="12" pageNumber="158">New Zealand</collectingCountry>
and South Australia, were comparatively sparse in ophidiiform otoliths and have also yielded far fewer species. This observation is again corroborated by the finds in the Eocene of
<collectingCountry id="F4A5764CFFA8FFFCD25EFA54E621DFC0" box="[1293,1404,1456,1479]" name="Antarctica" pageId="12" pageNumber="158">Antarctica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>